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<document id="D6B9EA4CFB95E90C614A54EF6EBE3E52" ID-DOI="10.37520/fi.2024.006" ID-ISSN="2533-4069" ID-Zenodo-Dep="14722393" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737336788380" checkinUser="felipe" docAuthor="Cleal, Christopher J., Strullu-Derrien, Christine &amp; Spencer, Alan R. T." docDate="2024" docId="03A187BCB033FFAA74CBFB975308F867" docLanguage="en" docName="FossilImprint.80.1.35-67.pdf" docOrigin="Fossil Imprint 80 (1)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Pecopteris aspera BRONGN. 1836" docType="treatment" docVersion="2" lastPageNumber="45" masterDocId="FF98FFC4B039FFA0771DFFD7564FFFC3" masterDocTitle="Early Coal Swamp Vegetation From The Serpukhovian Lower Clackmannan Group Of Scotland" masterLastPageNumber="67" masterPageNumber="35" pageNumber="45" updateTime="1737596041418" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="44FA65D7A262F1D7F0BBC03E3BA7F239">Early Coal Swamp Vegetation From The Serpukhovian Lower Clackmannan Group Of Scotland</mods:title>
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<mods:namePart id="D0AE61BBF14925CFA1108481925F5240">Cleal, Christopher J.</mods:namePart>
<mods:affiliation id="08F64D8773FA4FAFDCD1600EF2DE9F30">Department of Earth Sciences, University of Bristol, Bristol, BS 8 1 TQ, United Kingdom; email: christopher. cleal @ bristol. ac. uk.</mods:affiliation>
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<mods:namePart id="4E26088DA42C6B13B245E79647303812">Strullu-Derrien, Christine</mods:namePart>
<mods:affiliation id="09A1C1DEDC888F2582D0695EC4BB3BB3">Institut de Systématique, Évolution, Biodiversité (UMR 7205), Muséum National dHistoire Naturelle, CNRS, Sorbonne Université, Paris, France. &amp; Science Group, The Natural History Museum, London, United Kingdom.</mods:affiliation>
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<mods:namePart id="94F255398AB60710D494FA72337F0C29">Spencer, Alan R. T.</mods:namePart>
<mods:affiliation id="F80F9E17CC8E1BC61F7A9F9C4C45F16B">Department of Earth Sciences, The Natural History Museum, London, United Kingdom. &amp; Department of Earth Science and Engineering, Imperial College London, London, United Kingdom.</mods:affiliation>
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<mods:date id="69FB30AA96D64582BF4BB2E40DC8AFF3">2024</mods:date>
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<treatment id="03A187BCB033FFAA74CBFB975308F867" LSID="urn:lsid:plazi:treatment:03A187BCB033FFAA74CBFB975308F867" httpUri="http://treatment.plazi.org/id/03A187BCB033FFAA74CBFB975308F867" lastPageNumber="45" pageId="10" pageNumber="45">
<subSubSection id="C3126521B033FFAA74CBFB975371FB9B" box="[982,1342,1088,1112]" pageId="10" pageNumber="45" type="nomenclature">
<paragraph id="8BB736AAB033FFAA74CBFB975371FB9B" blockId="10.[982,1342,1088,1112]" box="[982,1342,1088,1112]" pageId="10" pageNumber="45">
<heading id="D0FF81C6B033FFAA74CBFB975371FB9B" box="[982,1342,1088,1112]" centered="true" fontSize="8" level="2" pageId="10" pageNumber="45" reason="2">
<taxonomicName id="4C084D29B033FFAA74CBFB975371FB9B" authority="BRONGN., 1836" authorityName="BRONGN." authorityYear="1836" box="[982,1342,1088,1112]" class="Polypodiopsida" family="Asterothecaceae" genus="Pecopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="B97CEAB8B033FFAA74CBFB975371FB9B" bold="true" box="[982,1342,1088,1112]" pageId="10" pageNumber="45">
<emphasis id="B97CEAB8B033FFAA74CBFB9752D7FB94" bold="true" box="[982,1176,1088,1112]" italics="true" pageId="10" pageNumber="45">Pecopteris aspera</emphasis>
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., 1836
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<paragraph id="8BB736AAB033FFAA735DFBB7529BFBBB" blockId="10.[1088,1236,1120,1144]" box="[1088,1236,1120,1144]" pageId="10" pageNumber="45">
<figureCitation id="13332A2FB033FFAA735DFBB7529BFBBB" box="[1088,1236,1120,1144]" captionStart="Text-fig" captionStartId="11.[125,201,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-6@11.[123,783,624,1098]" captionTargetPageId="11" captionText="Text-fig.6.a:SphenophyllumtenerrimumETTINGSH. exHELMHACKER;RobroystonColliery,2.8kmSEofBishopbriggs,Lanarkshire; ca.10m below Orchard Limestone, Upper Limestone Formation; BGS Kidst.156. b, c: Pecopteris aspera BRONGN.; Cuthill Shore (near highwater mark) between Musselburgh and Prestonpans, Haddingtonshire; between Nos 4 and 5 Limestones, Upper Limestone Formation; BGS Kidst.5415 (b), BGS Kidst.4517 (c). d: Zeilleria moravica (ETTINGSH.) BUREAU; Thorneycroft No. 1 Borehole, 0.5 km NE of Cumbernauld Railway Station, Dumbartonshire; 389 m depth, Limestone Coal Formation; BGS Kidst.2420. Scale bars = 5 mm (a, b), 10 mm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.14722403" httpUri="https://zenodo.org/record/14722403/files/figure.png" pageId="10" pageNumber="45">Text-fig. 6c, d</figureCitation>
</paragraph>
<paragraph id="8BB736AAB033FFAA7456FB595375FB60" blockId="10.[843,1482,1166,1335]" box="[843,1338,1166,1188]" pageId="10" pageNumber="45">
1836
<taxonomicName id="4C084D29B033FFAA748BFB5852D3FB67" authority="BRONGN." authorityName="BRONGN." box="[918,1180,1167,1188]" class="Polypodiopsida" family="Asterothecaceae" genus="Pecopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="B97CEAB8B033FFAA748BFB58520BFB67" box="[918,1092,1167,1188]" italics="true" pageId="10" pageNumber="45">Pecopteris aspera</emphasis>
<smallCapsWord id="8D51A076B033FFAA7357FB5852D9FB61" baselines="1183,1182" box="[1098,1174,1167,1188]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Brongn" pageId="10" pageNumber="45">BRONGN</smallCapsWord>
.
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, p. 339, pl. 120.
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<paragraph id="8BB736AAB033FFAA7456FB7B53F7FB02" blockId="10.[843,1482,1166,1335]" box="[843,1464,1195,1217]" pageId="10" pageNumber="45">
1901
<taxonomicName id="4C084D29B033FFAA748BFB7B5336FB7C" authority="STERZEL" authorityName="STERZEL" box="[918,1401,1195,1217]" class="Polypodiopsida" family="Asterothecaceae" form="sturii" genus="Pecopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="form" species="aspera" subGenus="Dactylotheca">
<emphasis id="B97CEAB8B033FFAA748BFB7B55B2FB02" box="[918,1021,1196,1217]" italics="true" pageId="10" pageNumber="45">Pecopteris</emphasis>
(
<emphasis id="B97CEAB8B033FFAA7316FB7C52C4FB03" box="[1035,1163,1195,1216]" italics="true" pageId="10" pageNumber="45">Dactylotheca</emphasis>
)
<emphasis id="B97CEAB8B033FFAA7384FB7B5295FB02" box="[1177,1242,1196,1217]" italics="true" pageId="10" pageNumber="45">aspera</emphasis>
f.
<emphasis id="B97CEAB8B033FFAA73E9FB7B5368FB02" box="[1268,1319,1196,1217]" italics="true" pageId="10" pageNumber="45">sturii</emphasis>
<smallCapsWord id="8D51A076B033FFAA7230FB7B5336FB7C" baselines="1212,1211" box="[1325,1401,1196,1217]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Sterzel" pageId="10" pageNumber="45">STERZEL</smallCapsWord>
</taxonomicName>
, p. iii.
</paragraph>
<paragraph id="8BB736AAB033FFAA7456FB1E55A5FB3F" blockId="10.[843,1482,1166,1335]" pageId="10" pageNumber="45">
1923
<taxonomicName id="4C084D29B033FFAA748BFB1E5354FB1F" authority="(STERZEL) KIDSTON" authorityName="KIDSTON" baseAuthorityName="STERZEL" box="[918,1307,1225,1246]" genus="Dactylotheca" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="sturii">
<emphasis id="B97CEAB8B033FFAA748BFB1E5218FB1D" box="[918,1111,1225,1246]" italics="true" pageId="10" pageNumber="45">Dactylotheca sturii</emphasis>
(
<smallCapsWord id="8D51A076B033FFAA7371FB1E52F7FB1E" baselines="1241,1241" box="[1132,1208,1225,1246]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Sterzel" pageId="10" pageNumber="45">STERZEL</smallCapsWord>
)
<smallCapsWord id="8D51A076B033FFAA73D0FB1E5354FB1F" baselines="1241,1240" box="[1229,1307,1225,1246]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kidston" pageId="10" pageNumber="45">KIDSTON</smallCapsWord>
</taxonomicName>
, p. 395, pl. 94, figs 46.
</paragraph>
<paragraph id="8BB736AAB033FFAA7456FAD35276FAF5" blockId="10.[843,1482,1166,1335]" pageId="10" pageNumber="45">
1939
<taxonomicName id="4C084D29B033FFAA748BFAD45340FAD4" authority="(STERZEL) KIDSTON" authorityName="KIDSTON" baseAuthorityName="STERZEL" box="[918,1295,1283,1305]" genus="Dactylotheca" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="sturii">
<emphasis id="B97CEAB8B033FFAA748BFAD4521CFADA" box="[918,1107,1283,1305]" italics="true" pageId="10" pageNumber="45">Dactylotheca sturii</emphasis>
(
<smallCapsWord id="8D51A076B033FFAA7379FAD352FFFAD4" baselines="1300,1299" box="[1124,1200,1284,1305]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Sterzel" pageId="10" pageNumber="45">STERZEL</smallCapsWord>
)
<smallCapsWord id="8D51A076B033FFAA73DCFAD35340FAD4" baselines="1300,1299" box="[1217,1295,1284,1305]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kidston" pageId="10" pageNumber="45">KIDSTON</smallCapsWord>
</taxonomicName>
; Radforth, p. 747, pl. 1, figs 1022.
</paragraph>
<paragraph id="8BB736AAB033FFAA746CFA9852A8F9A5" blockId="10.[843,1483,1358,1638]" pageId="10" pageNumber="45">
D e s c r i p t i o n. Fragments of pinnae with pinnules
<quantity id="4CF09B4FB033FFAA7456FAB955EDFA45" box="[843,930,1390,1414]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="10" pageNumber="45" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">25 mm</quantity>
long,
<quantity id="4CF09B4FB033FFAA74F9FAB95274FA45" box="[996,1083,1390,1414]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="45" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide, inserted at near to right-angles. Pinnules rounded to linguaeform, broadly attached to rachis; midvein well-marked, extending for most of pinnule length, producing simple or occasionally one-formed, widely-spaced lateral veins. A few pinnules have up to 12 sporangia arranged in two rows, one on either side of midvein. Sporangia are single, oval with obtuse apex, ca.
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long, with rudimentary apical annulus.
</paragraph>
<paragraph id="8BB736AAB033FFAA746CF9A9537AF896" blockId="10.[843,1482,1661,1877]" pageId="10" pageNumber="45">
R e m a r k s. This species was recorded by
<bibRefCitation id="EF994B5BB033FFAA7268F9AA55DDF976" author="Kidston, R." pageId="10" pageNumber="45" pagination="377 - 522" refId="ref21286" refString="Kidston, R. (1924): Fossil plants of the Carboniferous rocks of Great Britain. Part 5. - Memoirs of the Geological Survey of Great Britain, Palaeontology, 2: 377 - 522." type="journal article" year="1924">Kidston (1924)</bibRefCitation>
as
<taxonomicName id="4C084D29B033FFAA74D9F94A52B5F970" authority="STERZEL" authorityName="STERZEL" box="[964,1274,1693,1717]" genus="Dactylotheca" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="sturii">
<emphasis id="B97CEAB8B033FFAA74D9F94A52D7F976" box="[964,1176,1693,1717]" italics="true" pageId="10" pageNumber="45">Dactylotheca sturii</emphasis>
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</taxonomicName>
but there is now general consensus that this species is indistinguishable from
<taxonomicName id="4C084D29B033FFAA7456F90955E1F936" box="[843,942,1758,1781]" class="Polypodiopsida" family="Asterothecaceae" genus="Pecopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="B97CEAB8B033FFAA7456F90955E1F936" box="[843,942,1758,1781]" italics="true" pageId="10" pageNumber="45">P. aspera</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF994B5BB033FFAA74FBF90A52CCF936" author="Dalinval, A." box="[998,1155,1757,1781]" pageId="10" pageNumber="45" refId="ref17597" refString="Dalinval, A. (1960): Contribution a l'etude des Pecopteridees. Les Pecopteris du bassin houiller du Nord de la France (Etudes geologiques pour l'atlas topographie souterraine, Section 1: Flore fossile, fascicule 3). - Douriez-Bataille, Lille, 222 pp., 61 pls." type="book" year="1960">Dalinval 1960</bibRefCitation>
). A detailed description (under the name
<taxonomicName id="4C084D29B033FFAA74AEF9295243F8D6" box="[947,1036,1790,1813]" genus="Dactylotheca" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="sturii">
<emphasis id="B97CEAB8B033FFAA74AEF9295243F8D6" box="[947,1036,1790,1813]" italics="true" pageId="10" pageNumber="45">D. sturii</emphasis>
</taxonomicName>
) of the reproductive structures was given by
<bibRefCitation id="EF994B5BB033FFAA746EF8CA5268F8F6" author="Radforth, N. W." box="[883,1063,1821,1845]" pageId="10" pageNumber="45" pagination="745 - 761" refId="ref23029" refString="Radforth, N. W. (1939): Further contributions to our knowledge of the fossil Schizaeaceae; genus Senftenbergia. - Transactions of the Royal Society of Edinburgh, 59: 745 - 761. https: // doi. org / 10.1017 / S 0080456800017415" type="journal article" year="1939">Radforth (1939)</bibRefCitation>
based on specimens from the same locality as those described by
<bibRefCitation id="EF994B5BB033FFAA7391F8EA537EF896" author="Kidston, R." box="[1164,1329,1853,1877]" pageId="10" pageNumber="45" pagination="377 - 522" refId="ref21286" refString="Kidston, R. (1924): Fossil plants of the Carboniferous rocks of Great Britain. Part 5. - Memoirs of the Geological Survey of Great Britain, Palaeontology, 2: 377 - 522." type="journal article" year="1924">Kidston (1924)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB736AAB033FFAA746CF8BA5386F847" blockId="10.[843,1481,1900,1956]" box="[881,1481,1900,1925]" pageId="10" pageNumber="45">O c c u r r e n c e. Upper Limestone Formation, Cuthill</paragraph>
<paragraph id="8BB736AAB033FFAA7456F85B5308F867" blockId="10.[843,1481,1900,1956]" box="[843,1351,1932,1956]" pageId="10" pageNumber="45">Shore, between Index and Orchard Limestones.</paragraph>
</subSubSection>
</treatment>
</document>