345 lines
57 KiB
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345 lines
57 KiB
XML
<document ID-DOI="10.3853/j.2201-4349.74.2022.1797" ID-ISSN="2201-4349" ID-Zenodo-Dep="7177884" approvalRequired="12" approvalRequired_for_textStreams="12" checkinTime="1665305558210" checkinUser="felipe" docAuthor="Sinclair, Bradley J. & Saigusa, Toyohei" docDate="2022" docId="021487E73D2CFF9F0C64FC7E0CEFF850" docLanguage="en" docName="RecAustMus.74.3.75–98.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (3)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1797" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.1:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="1" docTitle="Gondwanodromia Sinclair & Saigusa 2022, gen. nov." docType="treatment" docVersion="2" lastPageNumber="77" masterDocId="FE2DFF9F3D2DFF9D0D63FFAE0803FFB9" masterDocTitle="A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae)" masterLastPageNumber="98" masterPageNumber="75" pageNumber="76" updateTime="1665324056033" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sinclair, Bradley J.</mods:namePart>
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<mods:affiliation>Canadian National Collection of Insects & Canadian Food Inspection Agency, OPL-Entomology, K. W. Neatby Building, C. E. F., 960 Carling Avenue, Ottawa, ON, K 1 A 0 C 6 Canada</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Saigusa, Toyohei</mods:namePart>
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<mods:affiliation>7 - 1 - 402 Baikoen 2 - chome, Chuo-ku, Fukuoka-shi 810 - 0035, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="series">
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<mods:title>Rec. Aust. Mus.</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2022-04-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>74</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>98</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3853/j.2201-4349.74.2022.1797</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.2201-4349.74.2022.1797</mods:identifier>
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<mods:identifier type="ISSN">2201-4349</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7177884</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:021487E73D2CFF9F0C64FC7E0CEFF850" httpUri="http://treatment.plazi.org/id/021487E73D2CFF9F0C64FC7E0CEFF850" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">
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<subSubSection pageId="1" pageNumber="76" type="nomenclature">
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<paragraph blockId="1.[263,606,976,1003]" box="[263,606,976,1003]" pageId="1" pageNumber="76">
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<heading bold="true" box="[263,606,976,1003]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="76" reason="2">
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<emphasis bold="true" box="[263,606,976,1003]" pageId="1" pageNumber="76">
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<taxonomicName authority="Sinclair & Saigusa, 2022" authorityName="Sinclair & Saigusa" authorityYear="2022" box="[263,485,976,1003]" class="Insecta" family="Empididae" genus="Gondwanodromia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[263,485,976,1003]" italics="true" pageId="1" pageNumber="76">Gondwanodromia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[494,606,977,1003]" pageId="1" pageNumber="76" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="1.[135,734,1027,1045]" box="[135,734,1027,1045]" pageId="1" pageNumber="76">
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<uri box="[135,734,1027,1045]" pageId="1" pageNumber="76">
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urn:lsid:zoobank.org:act:
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<uuid box="[332,734,1027,1045]" pageId="1" pageNumber="76">CBBBF605-C250-40CC-A747-E523086022D4</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="76" type="description">
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<paragraph blockId="1.[151,615,1067,1143]" pageId="1" pageNumber="76">
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“new genus 3”
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<bibRefCitation author="Sinclair, B. J. & J. M. Cumming" box="[298,611,1067,1089]" pageId="1" pageNumber="76" pagination="161 - 186" refId="ref16127" refString="Sinclair, B. J., and J. M. Cumming. 2000. Revision of the genus Apterodromia (Diptera: Empidoidea), with a redefinition of the tribe Ocydromiini. Records of the Australian Museum 52 (2): 161 - 186. https: // doi. org / 10.3853 / j. 0067 - 1975.52.2000.1313" type="journal article" year="2000">Sinclair & Cumming, 2000: 185</bibRefCitation>
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. “Undescribed genus”
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<bibRefCitation author="Saigusa, T." box="[362,547,1094,1116]" pageId="1" pageNumber="76" pagination="211" refId="ref15999" refString="Saigusa, T. 2006. Torrenticolous Empididae from northern Argentine Patagonia. In Abstracts Volume. 6 th International Congress of Dipterology, ed. M. Suwa, p. 211. Fukuoka, Japan." type="book chapter" year="2006">Saigusa, 2006: 211</bibRefCitation>
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.
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</paragraph>
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<paragraph blockId="1.[151,615,1067,1143]" box="[151,560,1121,1143]" pageId="1" pageNumber="76">
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“Undescribed genus A”
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<bibRefCitation author="Sinclair, B. J." box="[384,556,1121,1143]" pageId="1" pageNumber="76" pagination="30 - 36" refId="ref16071" refString="Sinclair, B. J. 2011. Revision of the New Zealand genus Adipsomyia (Diptera: Empidoidea: Brachystomatidae: Trichopezinae), with key to local Empidoid family and selected genus groups. The New Zealand Entomologist 34: 30 - 36. https: // doi. org / 10.1080 / 00779962.2011.9722205" type="journal article" year="2011">Sinclair, 2011: 31</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="76" type="type_taxon">
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<paragraph blockId="1.[121,629,1164,1188]" box="[121,629,1164,1188]" pageId="1" pageNumber="76">
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<emphasis bold="true" box="[121,262,1164,1188]" pageId="1" pageNumber="76">
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<typeStatus box="[121,177,1164,1188]" pageId="1" pageNumber="76">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName authority="Sinclair & Saigusa, 2022" authorityName="Sinclair & Saigusa" authorityYear="2022" box="[276,540,1164,1188]" class="Insecta" family="Empididae" genus="Gondwanodromia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="mikae" status="sp. nov.">
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<emphasis box="[276,540,1164,1188]" italics="true" pageId="1" pageNumber="76">Gondwanodromia mikae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[546,629,1165,1188]" pageId="1" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="1.[121,749,1209,1468]" pageId="1" pageNumber="76">
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<emphasis bold="true" box="[121,302,1209,1234]" pageId="1" pageNumber="76">Taxonomic note</emphasis>
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. This new genus was originally identified as “new genus 3” from
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<collectingCountry box="[390,489,1239,1263]" name="Australia" pageId="1" pageNumber="76">Australia</collectingCountry>
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(
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<bibRefCitation author="Sinclair, B. J. & J. M. Cumming" pageId="1" pageNumber="76" pagination="161 - 186" refId="ref16127" refString="Sinclair, B. J., and J. M. Cumming. 2000. Revision of the genus Apterodromia (Diptera: Empidoidea), with a redefinition of the tribe Ocydromiini. Records of the Australian Museum 52 (2): 161 - 186. https: // doi. org / 10.3853 / j. 0067 - 1975.52.2000.1313" type="journal article" year="2000">Sinclair & Cumming, 2000</bibRefCitation>
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).
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<bibRefCitation author="Saigusa, T." box="[195,353,1268,1292]" pageId="1" pageNumber="76" pagination="211" refId="ref15999" refString="Saigusa, T. 2006. Torrenticolous Empididae from northern Argentine Patagonia. In Abstracts Volume. 6 th International Congress of Dipterology, ed. M. Suwa, p. 211. Fukuoka, Japan." type="book chapter" year="2006">Saigusa (2006)</bibRefCitation>
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listed some morphological characters of “undescribed genus” based on an undescribed species from northern Patagonia, which is now the
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<typeStatus box="[588,634,1327,1351]" pageId="1" pageNumber="76">type</typeStatus>
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species of
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<taxonomicName authorityName="Sinclair & Saigusa" authorityYear="2022" box="[121,313,1356,1380]" class="Insecta" family="Empididae" genus="Gondwanodromia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="genus">
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<emphasis box="[121,313,1356,1380]" italics="true" pageId="1" pageNumber="76">Gondwanodromia</emphasis>
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</taxonomicName>
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. Later,
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<bibRefCitation author="Sinclair, B. J." box="[391,549,1356,1380]" pageId="1" pageNumber="76" pagination="30 - 36" refId="ref16071" refString="Sinclair, B. J. 2011. Revision of the New Zealand genus Adipsomyia (Diptera: Empidoidea: Brachystomatidae: Trichopezinae), with key to local Empidoid family and selected genus groups. The New Zealand Entomologist 34: 30 - 36. https: // doi. org / 10.1080 / 00779962.2011.9722205" type="journal article" year="2011">Sinclair (2011)</bibRefCitation>
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identified a similar new genus from
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<collectingCountry box="[299,445,1386,1410]" name="New Zealand" pageId="1" pageNumber="76">New Zealand</collectingCountry>
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, referred to as “Undescribed genus A”. These generic concepts have been re-evaluated and they are now considered congeneric.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="76" type="diagnosis">
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<paragraph blockId="1.[121,749,1489,1924]" pageId="1" pageNumber="76">
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<emphasis bold="true" box="[121,238,1489,1513]" pageId="1" pageNumber="76">Diagnosis</emphasis>
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. The genus is readily distinguished by the following combination of characters: eyes with or without ommatrichia; both sexes dichoptic with broad frons lacking setae; postpedicel subtriangular, gradually tapered with apical stylus or postpedicel strap-like, elongate, without apical stylus, bearing simply peg-like apical sensillum; palpus drooping, narrow and elongate (
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<figureCitation box="[543,613,1666,1690]" captionStart="Figures 1–4" captionStartId="2.[166,243,1439,1461]" captionTargetBox="[166,1466,134,1428]" captionTargetId="figure-135@2.[166,1466,134,1428]" captionTargetPageId="2" captionText="Figures 1–4. Photographs of living Gondwanodromia and habitats:(1)Arroyo Torrontegui,Argentina; (2)Arroyo Torrontegui,Argentina, with collector Mika Sugimoto; (3) female, Altos del Lircay, Chile (photograph © Steve Marshall); (4) Dead Horse Gap trail, Kosciusko National Park, Australia; November 1994." figureDoi="http://doi.org/10.5281/zenodo.7177886" httpUri="https://zenodo.org/record/7177886/files/figure.png" pageId="1" pageNumber="76">Figs 3</figureCitation>
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,
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<figureCitation box="[626,656,1666,1690]" captionStart="Figures 30–33" captionStartId="12.[167,244,1403,1425]" captionTargetBox="[169,1459,133,1388]" captionTargetId="figure-10@12.[167,1467,121,1396]" captionTargetPageId="12" captionText="Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177902" httpUri="https://zenodo.org/record/7177902/files/figure.png" pageId="1" pageNumber="76">30</figureCitation>
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); female labrum more robust than male;
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<emphasis box="[448,492,1696,1722]" italics="true" pageId="1" pageNumber="76">
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R
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<subScript attach="left" box="[464,492,1708,1722]" fontSize="6" pageId="1" pageNumber="76">4+5</subScript>
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</emphasis>
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branched, veins without setae; male terminalia symmetrical, unrotated, held upright in part by enlarged sternite 8, erect, arched anteriorly, epandrium and hypandrium not fused, epandrium without dorsal bridge, phallus tubular, narrow; female tergite 7 without posterior fringe, female tergite 8 with or without short internal medial apodeme and female syntergite 9+10 (acanthophorites) with numerous short thickened setae.
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76" type="description">
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||
<paragraph blockId="1.[121,749,1945,2028]" lastBlockId="1.[793,1422,151,762]" pageId="1" pageNumber="76">
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<emphasis bold="true" box="[121,256,1945,1969]" pageId="1" pageNumber="76">Description</emphasis>
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.
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<emphasis box="[274,330,1946,1970]" italics="true" pageId="1" pageNumber="76">Male</emphasis>
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.
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<emphasis bold="true" box="[348,410,1946,1970]" pageId="1" pageNumber="76">Head</emphasis>
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. Dichoptic, eyes usually bare (
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<figureCitation box="[129,207,1975,1999]" captionStart="Figures 30–33" captionStartId="12.[167,244,1403,1425]" captionTargetBox="[169,1459,133,1388]" captionTargetId="figure-10@12.[167,1467,121,1396]" captionTargetPageId="12" captionText="Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177902" httpUri="https://zenodo.org/record/7177902/files/figure.png" pageId="1" pageNumber="76">Fig. 30</figureCitation>
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); frons very broad and bare; face broad, shorter than membranous clypeus; ocellar triangle prominent with pair of ocellar setae inserted anterior or anteromedial to lateral ocelli and 1–2 pairs of shorter setae; upper postocular seta arched over eye; 2–3 pairs of vertical setae. Antenna dark, scape subequal to or longer than pedicel, with short dorsal setae; pedicel with circlet of stout setae; postpedicel subtriangular, gradually tapered, subequal or slightly longer than apical stylus (
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<figureCitation box="[868,948,327,352]" captionStart="Figures 13–16" captionStartId="8.[167,243,1448,1470]" captionTargetBox="[166,1466,151,1435]" captionTargetId="figure-9@8.[166,1466,151,1435]" captionTargetPageId="8" captionText="Figures 13–16.Argentinian and New Zealand species of Gondwanodromia, lateral view:(13) G.femorata sp.nov., male habitus, holotype, scale bar = 1.0 mm; (14) G. femorata sp. nov., male terminalia, scale bar = 0.25 mm; (15) G. femorata sp. nov., female postabdomen, scale bar = 0.25 mm; (16) G. mikae sp. nov., male habitus, holotype, scale bar = 0.75 mm. Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177892" httpUri="https://zenodo.org/record/7177892/files/figure.png" pageId="1" pageNumber="76">Figs 13</figureCitation>
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,
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<figureCitation box="[959,988,328,352]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">37</figureCitation>
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,
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<figureCitation box="[997,1065,327,352]" captionStart="Figures 43–47" captionStartId="18.[166,243,1827,1849]" captionTargetBox="[166,1459,151,1808]" captionTargetId="figure-9@18.[166,1466,151,1808]" captionTargetPageId="18" captionText="Figures 43–47. New Zealand species of Gondwanodromia: (43) G. tongariro sp. nov, holotype, male terminalia, scale bar = 0.25 mm; (44) G. femorata sp. nov., female midleg, scale bar = 0.5 mm (courtesy Canterbury Museum); (45) G. femorata sp. nov., female abdomen, scale bar = 0.5 mm (courtesy Canterbury Museum); (46) G. wardi sp. nov., holotype (abdomen dissected), lateral view, scale bar = 0.5 mm (courtesy Canterbury Museum); (47) G. wardi sp. nov., holotype (abdomen dissected), dorsal view, scale bar = 0.5 mm (courtesy Canterbury Museum). Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177910" httpUri="https://zenodo.org/record/7177910/files/figure.png" pageId="1" pageNumber="76">44, 46</figureCitation>
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), or postpedicel strap-like, greatly lengthened, pubescent, without apical stylus (
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<figureCitation box="[1262,1341,357,381]" captionStart="Figures 13–16" captionStartId="8.[167,243,1448,1470]" captionTargetBox="[166,1466,151,1435]" captionTargetId="figure-9@8.[166,1466,151,1435]" captionTargetPageId="8" captionText="Figures 13–16.Argentinian and New Zealand species of Gondwanodromia, lateral view:(13) G.femorata sp.nov., male habitus, holotype, scale bar = 1.0 mm; (14) G. femorata sp. nov., male terminalia, scale bar = 0.25 mm; (15) G. femorata sp. nov., female postabdomen, scale bar = 0.25 mm; (16) G. mikae sp. nov., male habitus, holotype, scale bar = 0.75 mm. Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177892" httpUri="https://zenodo.org/record/7177892/files/figure.png" pageId="1" pageNumber="76">Figs 16</figureCitation>
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,
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<figureCitation box="[1351,1379,357,381]" captionStart="Figures 19–23" captionStartId="10.[167,244,1062,1084]" captionTargetBox="[167,1466,151,1052]" captionTargetId="figure-10@10.[166,1466,151,1052]" captionTargetPageId="10" captionText="Figures 19–23. Gondwanodromia mikae sp. nov.: (19) labellum, lateral view, scale bar = 0.05 mm; (20) wing, scale bar = 1.0 mm; (21) male antenna, lateral view, scale bar = 0.1 mm; (22) tarsomere 5, dorsal view, scale bar = 0.05 mm; (23) tarsomere 5, lateral view, scale bar = 0.05 mm. Abbreviations: pped – postpedicel; psdtrch – pseudotrachea." figureDoi="http://doi.org/10.5281/zenodo.7177896" httpUri="https://zenodo.org/record/7177896/files/figure.png" pageId="1" pageNumber="76">21</figureCitation>
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,
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<figureCitation box="[1389,1418,357,381]" captionStart="Figures 30–33" captionStartId="12.[167,244,1403,1425]" captionTargetBox="[169,1459,133,1388]" captionTargetId="figure-10@12.[167,1467,121,1396]" captionTargetPageId="12" captionText="Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177902" httpUri="https://zenodo.org/record/7177902/files/figure.png" pageId="1" pageNumber="76">30</figureCitation>
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,
|
||
<figureCitation box="[794,824,386,410]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">39</figureCitation>
|
||
); apex with short, peg-like, or longer sensillum. Proboscis elongate, longer than eye height (
|
||
<figureCitation box="[1145,1223,415,440]" captionStart="Figures 30–33" captionStartId="12.[167,244,1403,1425]" captionTargetBox="[169,1459,133,1388]" captionTargetId="figure-10@12.[167,1467,121,1396]" captionTargetPageId="12" captionText="Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177902" httpUri="https://zenodo.org/record/7177902/files/figure.png" pageId="1" pageNumber="76">Fig. 30</figureCitation>
|
||
). Palpus drooping, arched and elongate along edge of proboscis (
|
||
<figureCitation box="[1295,1365,445,469]" captionStart="Figures 1–4" captionStartId="2.[166,243,1439,1461]" captionTargetBox="[166,1466,134,1428]" captionTargetId="figure-135@2.[166,1466,134,1428]" captionTargetPageId="2" captionText="Figures 1–4. Photographs of living Gondwanodromia and habitats:(1)Arroyo Torrontegui,Argentina; (2)Arroyo Torrontegui,Argentina, with collector Mika Sugimoto; (3) female, Altos del Lircay, Chile (photograph © Steve Marshall); (4) Dead Horse Gap trail, Kosciusko National Park, Australia; November 1994." figureDoi="http://doi.org/10.5281/zenodo.7177886" httpUri="https://zenodo.org/record/7177886/files/figure.png" pageId="1" pageNumber="76">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation box="[1379,1409,445,469]" captionStart="Figures 30–33" captionStartId="12.[167,244,1403,1425]" captionTargetBox="[169,1459,133,1388]" captionTargetId="figure-10@12.[167,1467,121,1396]" captionTargetPageId="12" captionText="Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177902" httpUri="https://zenodo.org/record/7177902/files/figure.png" pageId="1" pageNumber="76">30</figureCitation>
|
||
), one-half to three-quarters length of labrum; bearing fine setae longer than width of palpus. Lacinia slightly shorter than palpus, stylus-like; stipes long and narrow. Labrum stout, straight; epipharyngeal blades robust, articulated; epipharyngeal carina absent. Hypopharynx stout, straight, subequal in length to labrum. Labellum well developed, fleshy, with numerous short setae; prementum with several setae; 6–8 pseudotracheae well developed (
|
||
<figureCitation box="[1244,1321,679,704]" captionStart="Figures 19–23" captionStartId="10.[167,244,1062,1084]" captionTargetBox="[167,1466,151,1052]" captionTargetId="figure-10@10.[166,1466,151,1052]" captionTargetPageId="10" captionText="Figures 19–23. Gondwanodromia mikae sp. nov.: (19) labellum, lateral view, scale bar = 0.05 mm; (20) wing, scale bar = 1.0 mm; (21) male antenna, lateral view, scale bar = 0.1 mm; (22) tarsomere 5, dorsal view, scale bar = 0.05 mm; (23) tarsomere 5, lateral view, scale bar = 0.05 mm. Abbreviations: pped – postpedicel; psdtrch – pseudotrachea." figureDoi="http://doi.org/10.5281/zenodo.7177896" httpUri="https://zenodo.org/record/7177896/files/figure.png" pageId="1" pageNumber="76">Fig. 19</figureCitation>
|
||
). Clypeal ridge long and slender, as long as cibarium, articulated together with labrum at labrofulcral articulation point.
|
||
</paragraph>
|
||
<paragraph blockId="1.[794,1422,783,1541]" pageId="1" pageNumber="76">
|
||
<emphasis bold="true" box="[794,880,784,808]" pageId="1" pageNumber="76">Thorax</emphasis>
|
||
. Prosternum fused to proepisterna; proepisternum with scattered setae. Scutal setae long and conspicuous. Acrostichal setae uniserial or biserial, with pair of upright anterior setae; dorsocentral setae uniserial, anterior pair divergent from row and increasing in length posteriorly; 2–3 long notopleural setae; 1–2 pairs of scutellar setae on each side. Laterotergite with short setae; dorsal mesepimeron pocket present.
|
||
<emphasis box="[1146,1203,990,1013]" italics="true" pageId="1" pageNumber="76">Wing</emphasis>
|
||
(
|
||
<figureCitation box="[1222,1313,989,1013]" captionStart="Figures 13–16" captionStartId="8.[167,243,1448,1470]" captionTargetBox="[166,1466,151,1435]" captionTargetId="figure-9@8.[166,1466,151,1435]" captionTargetPageId="8" captionText="Figures 13–16.Argentinian and New Zealand species of Gondwanodromia, lateral view:(13) G.femorata sp.nov., male habitus, holotype, scale bar = 1.0 mm; (14) G. femorata sp. nov., male terminalia, scale bar = 0.25 mm; (15) G. femorata sp. nov., female postabdomen, scale bar = 0.25 mm; (16) G. mikae sp. nov., male habitus, holotype, scale bar = 0.75 mm. Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177892" httpUri="https://zenodo.org/record/7177892/files/figure.png" pageId="1" pageNumber="76">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation box="[1330,1360,989,1013]" captionStart="Figures 19–23" captionStartId="10.[167,244,1062,1084]" captionTargetBox="[167,1466,151,1052]" captionTargetId="figure-10@10.[166,1466,151,1052]" captionTargetPageId="10" captionText="Figures 19–23. Gondwanodromia mikae sp. nov.: (19) labellum, lateral view, scale bar = 0.05 mm; (20) wing, scale bar = 1.0 mm; (21) male antenna, lateral view, scale bar = 0.1 mm; (22) tarsomere 5, dorsal view, scale bar = 0.05 mm; (23) tarsomere 5, lateral view, scale bar = 0.05 mm. Abbreviations: pped – postpedicel; psdtrch – pseudotrachea." figureDoi="http://doi.org/10.5281/zenodo.7177896" httpUri="https://zenodo.org/record/7177896/files/figure.png" pageId="1" pageNumber="76">20</figureCitation>
|
||
,
|
||
<figureCitation box="[1376,1407,989,1013]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">38</figureCitation>
|
||
). Relatively slender, infuscate; anal lobe weakly developed, without alular incision; alula not developed; basal costal seta present; pterostigma faint, elongate-oval overlapping apex of
|
||
<emphasis box="[892,916,1107,1133]" italics="true" pageId="1" pageNumber="76">
|
||
R
|
||
<subScript attach="left" box="[908,916,1119,1133]" fontSize="6" pageId="1" pageNumber="76">1</subScript>
|
||
</emphasis>
|
||
.
|
||
<collectingRegion box="[935,998,1106,1130]" country="New Zealand" name="West Coast" pageId="1" pageNumber="76">Costa</collectingRegion>
|
||
circumambient with biserial row of slender marginal setae, sometimes with erect dorsal setae;
|
||
<emphasis box="[794,820,1165,1188]" italics="true" pageId="1" pageNumber="76">Sc</emphasis>
|
||
incomplete, ending short of costa; all veins bare;
|
||
<emphasis box="[1377,1421,1166,1192]" italics="true" pageId="1" pageNumber="76">
|
||
R
|
||
<subScript attach="left" box="[1393,1421,1178,1192]" fontSize="6" pageId="1" pageNumber="76">4+5</subScript>
|
||
</emphasis>
|
||
branched;
|
||
<emphasis box="[913,938,1195,1221]" italics="true" pageId="1" pageNumber="76">
|
||
R
|
||
<subScript attach="left" box="[930,938,1207,1221]" fontSize="6" pageId="1" pageNumber="76">4</subScript>
|
||
</emphasis>
|
||
slightly sinuous, ending slightly closer to
|
||
<emphasis box="[794,839,1224,1250]" italics="true" pageId="1" pageNumber="76">
|
||
R
|
||
<subScript attach="left" box="[810,839,1236,1250]" fontSize="6" pageId="1" pageNumber="76">2+3</subScript>
|
||
</emphasis>
|
||
than to
|
||
<emphasis box="[928,972,1224,1250]" italics="true" pageId="1" pageNumber="76">
|
||
R
|
||
<subScript attach="left" box="[944,972,1236,1250]" fontSize="6" pageId="1" pageNumber="76">4+5</subScript>
|
||
</emphasis>
|
||
; cell
|
||
<emphasis box="[1034,1067,1223,1247]" italics="true" pageId="1" pageNumber="76">dm</emphasis>
|
||
emitting 3 veins, reaching wing margin; cell
|
||
<emphasis box="[934,973,1254,1277]" italics="true" pageId="1" pageNumber="76">cua</emphasis>
|
||
rounded with
|
||
<emphasis box="[1136,1184,1253,1276]" italics="true" pageId="1" pageNumber="76">CuA</emphasis>
|
||
recurrent;
|
||
<emphasis box="[1307,1422,1253,1276]" italics="true" pageId="1" pageNumber="76">CuA+CuP</emphasis>
|
||
reduced to short streak. Halter with 2–3 setae below knob, occasionally shaft rather elongate.
|
||
<emphasis box="[1187,1239,1312,1335]" italics="true" pageId="1" pageNumber="76">Legs</emphasis>
|
||
. Fore tibia with anteroventral apical comb; hind tibia with posteroventral apical comb. Mid tibia often slightly to strongly swollen with modified setae (
|
||
<figureCitation box="[1051,1140,1399,1424]" captionStart="Figures 13–16" captionStartId="8.[167,243,1448,1470]" captionTargetBox="[166,1466,151,1435]" captionTargetId="figure-9@8.[166,1466,151,1435]" captionTargetPageId="8" captionText="Figures 13–16.Argentinian and New Zealand species of Gondwanodromia, lateral view:(13) G.femorata sp.nov., male habitus, holotype, scale bar = 1.0 mm; (14) G. femorata sp. nov., male terminalia, scale bar = 0.25 mm; (15) G. femorata sp. nov., female postabdomen, scale bar = 0.25 mm; (16) G. mikae sp. nov., male habitus, holotype, scale bar = 0.75 mm. Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177892" httpUri="https://zenodo.org/record/7177892/files/figure.png" pageId="1" pageNumber="76">Fig. 13</figureCitation>
|
||
). Tarsomere 5 partially flattened, truncate apically, without apical projection (
|
||
<figureCitation box="[802,927,1458,1482]" captionStart="Figures 19–23" captionStartId="10.[167,244,1062,1084]" captionTargetBox="[167,1466,151,1052]" captionTargetId="figure-10@10.[166,1466,151,1052]" captionTargetPageId="10" captionText="Figures 19–23. Gondwanodromia mikae sp. nov.: (19) labellum, lateral view, scale bar = 0.05 mm; (20) wing, scale bar = 1.0 mm; (21) male antenna, lateral view, scale bar = 0.1 mm; (22) tarsomere 5, dorsal view, scale bar = 0.05 mm; (23) tarsomere 5, lateral view, scale bar = 0.05 mm. Abbreviations: pped – postpedicel; psdtrch – pseudotrachea." figureDoi="http://doi.org/10.5281/zenodo.7177896" httpUri="https://zenodo.org/record/7177896/files/figure.png" pageId="1" pageNumber="76">Figs 22, 23</figureCitation>
|
||
); claw moderately long; pulvillus pale, highly pubescent, shorter than claw; empodium slender, with distinct ventral pubescence (
|
||
<figureCitation box="[1109,1239,1517,1541]" captionStart="Figures 19–23" captionStartId="10.[167,244,1062,1084]" captionTargetBox="[167,1466,151,1052]" captionTargetId="figure-10@10.[166,1466,151,1052]" captionTargetPageId="10" captionText="Figures 19–23. Gondwanodromia mikae sp. nov.: (19) labellum, lateral view, scale bar = 0.05 mm; (20) wing, scale bar = 1.0 mm; (21) male antenna, lateral view, scale bar = 0.1 mm; (22) tarsomere 5, dorsal view, scale bar = 0.05 mm; (23) tarsomere 5, lateral view, scale bar = 0.05 mm. Abbreviations: pped – postpedicel; psdtrch – pseudotrachea." figureDoi="http://doi.org/10.5281/zenodo.7177896" httpUri="https://zenodo.org/record/7177896/files/figure.png" pageId="1" pageNumber="76">Figs 22, 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="1.[794,1422,1562,2027]" pageId="1" pageNumber="76">
|
||
<emphasis bold="true" box="[794,905,1562,1586]" pageId="1" pageNumber="76">Abdomen</emphasis>
|
||
. Male tergite 6 often with longest setae; male sternite 7 often prolonged posterolaterally (
|
||
<figureCitation box="[1287,1362,1591,1615]" captionStart="Figures 5–9" captionStartId="4.[166,241,1899,1921]" captionTargetBox="[178,1456,151,1888]" captionTargetId="figure-9@4.[166,1466,151,1888]" captionTargetPageId="4" captionText="Figures 5–9. Australian and New Zealand species of Gondwanodromia, lateral view: (5) G. bulbosa sp. nov., male habitus, holotype, scale bar = 0.75 mm; (6) G. bulbosa sp. nov., male terminalia, scale bar = 0.25 mm; (7) G. colomatta sp. nov., male habitus, holotype, scale bar = 0.75 mm; (8) G. elongata sp. nov., male habitus, holotype (prior to abdomen dissection), scale bar = 0.5 mm; (9) G. elongata sp. nov., female habitus,scale bar=0.75 mm.Abbreviations:asur – anteriorsurstylus;cerc –cercus;epand –epandrium;psur – posteriorsurstylus;st– sternite." figureDoi="http://doi.org/10.5281/zenodo.7177888" httpUri="https://zenodo.org/record/7177888/files/figure.png" pageId="1" pageNumber="76">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation box="[1378,1409,1592,1616]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">40</figureCitation>
|
||
). Sternite 8 enlarged, broad, providing support to upright terminalia (
|
||
<figureCitation box="[915,982,1650,1674]" captionStart="Figures 5–9" captionStartId="4.[166,241,1899,1921]" captionTargetBox="[178,1456,151,1888]" captionTargetId="figure-9@4.[166,1466,151,1888]" captionTargetPageId="4" captionText="Figures 5–9. Australian and New Zealand species of Gondwanodromia, lateral view: (5) G. bulbosa sp. nov., male habitus, holotype, scale bar = 0.75 mm; (6) G. bulbosa sp. nov., male terminalia, scale bar = 0.25 mm; (7) G. colomatta sp. nov., male habitus, holotype, scale bar = 0.75 mm; (8) G. elongata sp. nov., male habitus, holotype (prior to abdomen dissection), scale bar = 0.5 mm; (9) G. elongata sp. nov., female habitus,scale bar=0.75 mm.Abbreviations:asur – anteriorsurstylus;cerc –cercus;epand –epandrium;psur – posteriorsurstylus;st– sternite." figureDoi="http://doi.org/10.5281/zenodo.7177888" httpUri="https://zenodo.org/record/7177888/files/figure.png" pageId="1" pageNumber="76">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation box="[992,1021,1650,1674]" captionStart="Figures 17–18" captionStartId="9.[121,198,1082,1104]" captionTargetBox="[121,1421,123,1071]" captionTargetId="figure-9@9.[121,1421,123,1071]" captionTargetPageId="9" captionText="Figures 17–18. Male terminalia of New Zealand andAustralian species of Gondwanodromia, lateral view: (17) G. elongata sp. nov.; (18) G. lutea sp. nov. Scale bar = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177894" httpUri="https://zenodo.org/record/7177894/files/figure.png" pageId="1" pageNumber="76">18</figureCitation>
|
||
,
|
||
<figureCitation box="[1031,1062,1650,1674]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">40</figureCitation>
|
||
).
|
||
<emphasis box="[1078,1194,1650,1674]" italics="true" pageId="1" pageNumber="76">Terminalia</emphasis>
|
||
(
|
||
<figureCitation box="[1207,1299,1650,1674]" captionStart="Figures 5–9" captionStartId="4.[166,241,1899,1921]" captionTargetBox="[178,1456,151,1888]" captionTargetId="figure-9@4.[166,1466,151,1888]" captionTargetPageId="4" captionText="Figures 5–9. Australian and New Zealand species of Gondwanodromia, lateral view: (5) G. bulbosa sp. nov., male habitus, holotype, scale bar = 0.75 mm; (6) G. bulbosa sp. nov., male terminalia, scale bar = 0.25 mm; (7) G. colomatta sp. nov., male habitus, holotype, scale bar = 0.75 mm; (8) G. elongata sp. nov., male habitus, holotype (prior to abdomen dissection), scale bar = 0.5 mm; (9) G. elongata sp. nov., female habitus,scale bar=0.75 mm.Abbreviations:asur – anteriorsurstylus;cerc –cercus;epand –epandrium;psur – posteriorsurstylus;st– sternite." figureDoi="http://doi.org/10.5281/zenodo.7177888" httpUri="https://zenodo.org/record/7177888/files/figure.png" pageId="1" pageNumber="76">Figs 5, 7</figureCitation>
|
||
,
|
||
<figureCitation box="[1309,1338,1650,1674]" captionStart="Figures 17–18" captionStartId="9.[121,198,1082,1104]" captionTargetBox="[121,1421,123,1071]" captionTargetId="figure-9@9.[121,1421,123,1071]" captionTargetPageId="9" captionText="Figures 17–18. Male terminalia of New Zealand andAustralian species of Gondwanodromia, lateral view: (17) G. elongata sp. nov.; (18) G. lutea sp. nov. Scale bar = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177894" httpUri="https://zenodo.org/record/7177894/files/figure.png" pageId="1" pageNumber="76">18</figureCitation>
|
||
,
|
||
<figureCitation box="[1348,1417,1650,1674]" captionStart="Figures 24–29" captionStartId="11.[121,197,1891,1913]" captionTargetBox="[133,1413,174,1862]" captionTargetId="figure-9@11.[121,1421,151,1862]" captionTargetPageId="11" captionText="Figures 24–29. Gondwanodromia mikae sp. nov.: (24) epandrium, dorsal view; (25) epandrium, dorsal view; (26) hypandrium and phallus, lateral view; (27) hypandrium and phallus, posterior view; (28) female terminalia, lateral view, scale bar = 0.05 mm; (29) female abdomen, lateral view. Scale bars: Figs 24–28 = 0.05 mm; Fig. 29 = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; spmth – spermatheca; sur – surstylus; tg – tergite." figureDoi="http://doi.org/10.5281/zenodo.7177898" httpUri="https://zenodo.org/record/7177898/files/figure.png" pageId="1" pageNumber="76">24–27</figureCitation>
|
||
,
|
||
<figureCitation box="[794,866,1680,1704]" captionStart="Figures 30–33" captionStartId="12.[167,244,1403,1425]" captionTargetBox="[169,1459,133,1388]" captionTargetId="figure-10@12.[167,1467,121,1396]" captionTargetPageId="12" captionText="Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177902" httpUri="https://zenodo.org/record/7177902/files/figure.png" pageId="1" pageNumber="76">31–33</figureCitation>
|
||
,
|
||
<figureCitation box="[882,912,1680,1704]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">40</figureCitation>
|
||
,
|
||
<figureCitation box="[929,961,1680,1704]" captionStart="Figures 43–47" captionStartId="18.[166,243,1827,1849]" captionTargetBox="[166,1459,151,1808]" captionTargetId="figure-9@18.[166,1466,151,1808]" captionTargetPageId="18" captionText="Figures 43–47. New Zealand species of Gondwanodromia: (43) G. tongariro sp. nov, holotype, male terminalia, scale bar = 0.25 mm; (44) G. femorata sp. nov., female midleg, scale bar = 0.5 mm (courtesy Canterbury Museum); (45) G. femorata sp. nov., female abdomen, scale bar = 0.5 mm (courtesy Canterbury Museum); (46) G. wardi sp. nov., holotype (abdomen dissected), lateral view, scale bar = 0.5 mm (courtesy Canterbury Museum); (47) G. wardi sp. nov., holotype (abdomen dissected), dorsal view, scale bar = 0.5 mm (courtesy Canterbury Museum). Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177910" httpUri="https://zenodo.org/record/7177910/files/figure.png" pageId="1" pageNumber="76">43</figureCitation>
|
||
). Held upright, directed anterodorsally, symmetrical, unrotated. Epandrium and hypandrium not fused. Epandrium rounded or divided medially, without narrow dorsal bridge. Surstylus weakly differentiated or distinct, sometimes divided into anterior and posterior surstylar processes. Postgonites not differentiated. Phallus narrow, arched, tubular; ejaculatory apodeme narrow to broad, articulated at base of phallus.
|
||
</paragraph>
|
||
<paragraph blockId="1.[794,1422,1562,2027]" lastBlockId="2.[166,795,1545,1892]" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">
|
||
<emphasis box="[824,903,1914,1938]" italics="true" pageId="1" pageNumber="76">Female</emphasis>
|
||
. Similar to male except as follows: postpedicel shorter than male; labrum more robust than male; midleg unmodified.Abdomen truncate, caudal segments withdrawn into segment 7 (
|
||
<figureCitation box="[976,1062,2002,2026]" captionStart="Figures 13–16" captionStartId="8.[167,243,1448,1470]" captionTargetBox="[166,1466,151,1435]" captionTargetId="figure-9@8.[166,1466,151,1435]" captionTargetPageId="8" captionText="Figures 13–16.Argentinian and New Zealand species of Gondwanodromia, lateral view:(13) G.femorata sp.nov., male habitus, holotype, scale bar = 1.0 mm; (14) G. femorata sp. nov., male terminalia, scale bar = 0.25 mm; (15) G. femorata sp. nov., female postabdomen, scale bar = 0.25 mm; (16) G. mikae sp. nov., male habitus, holotype, scale bar = 0.75 mm. Abbreviations: epand lb – epandrial lobe; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177892" httpUri="https://zenodo.org/record/7177892/files/figure.png" pageId="1" pageNumber="76">Figs 15</figureCitation>
|
||
,
|
||
<figureCitation box="[1077,1107,2002,2026]" captionStart="Figures 37–40" captionStartId="15.[121,198,1826,1848]" captionTargetBox="[187,1401,151,1808]" captionTargetId="figure-10@15.[121,1421,151,1808]" captionTargetPageId="15" captionText="Figures 37–40.Australian and New Zealand species of Gondwanodromia, lateral view:(37) G. mutabilis (Collin), male habitus (abdomen dissected), scale bar = 1.0 mm; (38) G. tasmanica sp. nov., female habitus, scale bar = 0.75 mm; (39) G. thredbo sp. nov., male habitus, holotype, scale bar = 0.75 mm; (40) G. thredbo sp. nov., male terminalia, scale bar = 0.25 mm.Abbreviations: a sur – anterior surstylus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177906" httpUri="https://zenodo.org/record/7177906/files/figure.png" pageId="1" pageNumber="76">38</figureCitation>
|
||
); posterior margin of tergite 7 without row of fine setae.
|
||
<emphasis box="[480,595,1545,1569]" italics="true" pageId="2" pageNumber="77">Terminalia</emphasis>
|
||
(
|
||
<figureCitation box="[612,695,1545,1569]" captionStart="Figures 10–12" captionStartId="6.[166,243,1032,1054]" captionTargetBox="[179,1466,139,1019]" captionTargetId="figure-9@6.[167,1467,134,1028]" captionTargetPageId="6" captionText="Figures 10–12. Terminalia of Gondwanodromia colomatta sp. nov., lateral view: (10) male; (11) female; (12) spermatheca. Scale bar = 0.1 mm. Abbreviations: a sur – anterior surstylus; cerc – cercus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177890" httpUri="https://zenodo.org/record/7177890/files/figure.png" pageId="2" pageNumber="77">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation box="[709,738,1545,1569]" captionStart="Figures 24–29" captionStartId="11.[121,197,1891,1913]" captionTargetBox="[133,1413,174,1862]" captionTargetId="figure-9@11.[121,1421,151,1862]" captionTargetPageId="11" captionText="Figures 24–29. Gondwanodromia mikae sp. nov.: (24) epandrium, dorsal view; (25) epandrium, dorsal view; (26) hypandrium and phallus, lateral view; (27) hypandrium and phallus, posterior view; (28) female terminalia, lateral view, scale bar = 0.05 mm; (29) female abdomen, lateral view. Scale bars: Figs 24–28 = 0.05 mm; Fig. 29 = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; spmth – spermatheca; sur – surstylus; tg – tergite." figureDoi="http://doi.org/10.5281/zenodo.7177898" httpUri="https://zenodo.org/record/7177898/files/figure.png" pageId="2" pageNumber="77">28</figureCitation>
|
||
,
|
||
<figureCitation box="[752,782,1545,1569]" captionStart="Figures 41–42" captionStartId="16.[166,243,1009,1031]" captionTargetBox="[175,1456,138,989]" captionTargetId="figure-450@16.[167,1467,130,995]" captionTargetPageId="16" captionText="Figures 41–42. Gondwanodromia tasmanica sp. nov., lateral view: (41) male terminalia; (42) female terminalia. Scale bar = 0.1 mm. Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177908" httpUri="https://zenodo.org/record/7177908/files/figure.png" pageId="2" pageNumber="77">42</figureCitation>
|
||
). Sclerites of segment 8 articulated anterolaterally; anterodorsal margin of tergite 8 with or without short, narrow apodeme, extending into segment 7; posterior margin of tergite 8 with deep, median membranous cleft, medially with transverse row of short setae. Sternite 8 without posterior marginal setae. Syntergite 9+10 (acanthophorites) divided dorsomedially, subtriangular, with row(s) of strong, erect setae. Cercus heavily sclerotized, slightly arched posterodorsally or prolonged distally. Spermathecal receptacle elongate or spherical (
|
||
<figureCitation box="[289,381,1838,1863]" captionStart="Figures 24–29" captionStartId="11.[121,197,1891,1913]" captionTargetBox="[133,1413,174,1862]" captionTargetId="figure-9@11.[121,1421,151,1862]" captionTargetPageId="11" captionText="Figures 24–29. Gondwanodromia mikae sp. nov.: (24) epandrium, dorsal view; (25) epandrium, dorsal view; (26) hypandrium and phallus, lateral view; (27) hypandrium and phallus, posterior view; (28) female terminalia, lateral view, scale bar = 0.05 mm; (29) female abdomen, lateral view. Scale bars: Figs 24–28 = 0.05 mm; Fig. 29 = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; spmth – spermatheca; sur – surstylus; tg – tergite." figureDoi="http://doi.org/10.5281/zenodo.7177898" httpUri="https://zenodo.org/record/7177898/files/figure.png" pageId="2" pageNumber="77">Figs 29</figureCitation>
|
||
,
|
||
<figureCitation box="[397,428,1838,1862]" captionStart="Figures 34–36" captionStartId="13.[121,198,1944,1966]" captionTargetBox="[128,1417,1020,1923]" captionTargetId="figure-208@13.[121,1421,1020,1929]" captionTargetPageId="13" captionText="Figures 34–36. Gondwanodromia mutabilis (Collin), lateral view: (34) male terminalia; (35) female terminalia; (36) spermatheca. Scale bar = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; sur – surstylus." figureDoi="http://doi.org/10.5281/zenodo.7177904" httpUri="https://zenodo.org/record/7177904/files/figure.png" pageId="2" pageNumber="77">36</figureCitation>
|
||
), darkly pigmented, with long, unpigmented duct; genital fork small, U-shaped.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7177886" ID-Zenodo-Dep="7177886" httpUri="https://zenodo.org/record/7177886/files/figure.png" pageId="2" pageNumber="77" startId="2.[166,243,1439,1461]" targetBox="[166,1466,134,1428]" targetPageId="2">
|
||
<paragraph blockId="2.[166,1467,1439,1514]" pageId="2" pageNumber="77">
|
||
<emphasis bold="true" box="[166,283,1439,1461]" pageId="2" pageNumber="77">Figures 1–4</emphasis>
|
||
. Photographs of living
|
||
<taxonomicName authorityName="Sinclair & Saigusa" authorityYear="2022" box="[503,676,1439,1460]" class="Insecta" family="Empididae" genus="Gondwanodromia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[503,676,1439,1460]" italics="true" pageId="2" pageNumber="77">Gondwanodromia</emphasis>
|
||
</taxonomicName>
|
||
and habitats: (
|
||
<emphasis box="[814,826,1439,1460]" italics="true" pageId="2" pageNumber="77">1</emphasis>
|
||
) Arroyo Torrontegui,Argentina; (
|
||
<emphasis box="[1147,1159,1439,1460]" italics="true" pageId="2" pageNumber="77">2</emphasis>
|
||
) Arroyo Torrontegui, Argentina, with collector Mika Sugimoto; (
|
||
<emphasis box="[476,488,1466,1487]" italics="true" pageId="2" pageNumber="77">3</emphasis>
|
||
) female, Altos del Lircay, Chile (photograph © Steve Marshall); (
|
||
<emphasis box="[1123,1135,1466,1487]" italics="true" pageId="2" pageNumber="77">4</emphasis>
|
||
) Dead Horse Gap trail, Kosciusko National Park, Australia; November 1994.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="2" pageNumber="77" type="etymology">
|
||
<paragraph blockId="2.[166,794,1913,2025]" pageId="2" pageNumber="77">
|
||
<emphasis bold="true" box="[166,288,1913,1937]" pageId="2" pageNumber="77">Etymology</emphasis>
|
||
. The generic name is from the ancient supercontinent Gondwana + the Greek,
|
||
<emphasis box="[527,603,1942,1966]" italics="true" pageId="2" pageNumber="77">dromia</emphasis>
|
||
(runner) based on its distributional range on the southern continents of South America and Australasia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="77" type="distribution">
|
||
<paragraph blockId="2.[839,1467,1545,1628]" pageId="2" pageNumber="77">
|
||
<emphasis bold="true" box="[839,978,1545,1569]" pageId="2" pageNumber="77">Distribution</emphasis>
|
||
.
|
||
<taxonomicName authorityName="Sinclair & Saigusa" authorityYear="2022" box="[990,1183,1545,1569]" class="Insecta" family="Empididae" genus="Gondwanodromia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[990,1183,1545,1569]" italics="true" pageId="2" pageNumber="77">Gondwanodromia</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed in the Southern Hemisphere, recorded from
|
||
<collectingCountry box="[1223,1321,1574,1598]" name="Australia" pageId="2" pageNumber="77">Australia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1329,1467,1574,1598]" name="New Zealand" pageId="2" pageNumber="77">New Zealand</collectingCountry>
|
||
and southern South America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="77" type="discussion">
|
||
<paragraph blockId="2.[839,1467,1649,2025]" pageId="2" pageNumber="77">
|
||
<emphasis bold="true" box="[839,943,1649,1673]" pageId="2" pageNumber="77">Remarks</emphasis>
|
||
.
|
||
<taxonomicName authorityName="Sinclair & Saigusa" authorityYear="2022" box="[958,1151,1649,1673]" class="Insecta" family="Empididae" genus="Gondwanodromia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[958,1151,1649,1673]" italics="true" pageId="2" pageNumber="77">Gondwanodromia</emphasis>
|
||
</taxonomicName>
|
||
is assigned to the subfamily
|
||
<taxonomicName authorityName="Vaillant" authorityYear="1981" box="[839,988,1678,1702]" class="Insecta" family="Brachystomatidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="subFamily" subFamily="Trichopezinae">Trichopezinae</taxonomicName>
|
||
on the basis of the modified female terminalia, especially the presence of a small apodeme on the anterior margin of tergite 8. Genera of
|
||
<taxonomicName authorityName="Vaillant" authorityYear="1981" box="[1176,1328,1737,1761]" class="Insecta" family="Brachystomatidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="subFamily" subFamily="Trichopezinae">Trichopezinae</taxonomicName>
|
||
with similar elongate antennae (i.e., strap-like postpedicel) include
|
||
<taxonomicName authority="Melander" authorityName="Melander" authorityYear="1927" box="[840,1073,1796,1820]" class="Insecta" family="Brachystomatidae" genus="Ceratempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[840,963,1796,1819]" italics="true" pageId="2" pageNumber="77">Ceratempis</emphasis>
|
||
Melander
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Melander" authorityName="Melander" authorityYear="1927" box="[1129,1364,1796,1820]" class="Insecta" family="Brachystomatidae" genus="Niphogenia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1129,1253,1796,1820]" italics="true" pageId="2" pageNumber="77">Niphogenia</emphasis>
|
||
Melander
|
||
</taxonomicName>
|
||
from the Nearctic Region, but they differ in the form of the male terminalia and the palpus is not drooping.
|
||
</paragraph>
|
||
<paragraph blockId="2.[839,1467,1649,2025]" pageId="2" pageNumber="77">
|
||
Adult flies are found in fast flowing streams and creeks, occurring on emergent rocks and boulders (
|
||
<figureCitation box="[1292,1386,1913,1937]" captionStart="Figures 1–4" captionStartId="2.[166,243,1439,1461]" captionTargetBox="[166,1466,134,1428]" captionTargetId="figure-135@2.[166,1466,134,1428]" captionTargetPageId="2" captionText="Figures 1–4. Photographs of living Gondwanodromia and habitats:(1)Arroyo Torrontegui,Argentina; (2)Arroyo Torrontegui,Argentina, with collector Mika Sugimoto; (3) female, Altos del Lircay, Chile (photograph © Steve Marshall); (4) Dead Horse Gap trail, Kosciusko National Park, Australia; November 1994." figureDoi="http://doi.org/10.5281/zenodo.7177886" httpUri="https://zenodo.org/record/7177886/files/figure.png" pageId="2" pageNumber="77">Figs 1, 4</figureCitation>
|
||
). These habitats are primarily in mountainous and cool hilly regions. Adults can be swept from the rocks or sometimes aspirated directly from the rock surfaces (
|
||
<figureCitation box="[1181,1247,2001,2025]" captionStart="Figures 1–4" captionStartId="2.[166,243,1439,1461]" captionTargetBox="[166,1466,134,1428]" captionTargetId="figure-135@2.[166,1466,134,1428]" captionTargetPageId="2" captionText="Figures 1–4. Photographs of living Gondwanodromia and habitats:(1)Arroyo Torrontegui,Argentina; (2)Arroyo Torrontegui,Argentina, with collector Mika Sugimoto; (3) female, Altos del Lircay, Chile (photograph © Steve Marshall); (4) Dead Horse Gap trail, Kosciusko National Park, Australia; November 1994." figureDoi="http://doi.org/10.5281/zenodo.7177886" httpUri="https://zenodo.org/record/7177886/files/figure.png" pageId="2" pageNumber="77">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |