139 lines
20 KiB
XML
139 lines
20 KiB
XML
<document id="24A81D70E4AED6F3C31D5D2014174C77" ID-CLB-Dataset="31436" ID-DOI="10.11646/zootaxa.4085.2.1" ID-GBIF-Dataset="ba498adf-4dec-4fa3-8213-9d4d8c3b9dd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052535" ID-ZooBank="E41CB99C-5177-47A7-A424-2453D27E48F0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510873310154" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2016" docId="F50D3F20FFBBD35EEBE6FBAFFA73FE1C" docLanguage="en" docName="zootaxa.4085.2.1.pdf" docOrigin="Zootaxa 4085 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prionocyphon neboissi Zwick 2016, n. sp." docType="treatment" docVersion="9" lastPageNumber="175" masterDocId="09344758FFACD346EB71FFF6FFF9FFE9" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae" masterLastPageNumber="198" masterPageNumber="151" pageNumber="174" updateTime="1698499212072" updateUser="ExternalLinkService">
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<mods:title id="76DCDC4F41415F5289C09F0707C5E8AB">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<mods:namePart id="96509755D8F37C31309E470B826450F7">Zwick, Peter</mods:namePart>
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<mods:typeOfResource id="41467759F797691E31DE302C743BE01A">text</mods:typeOfResource>
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<mods:title id="E46EB85AB779A64AFEE0B1F2F130F558">Zootaxa</mods:title>
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<mods:part id="F47F6BB078B80B413C12DBDA5C87E768">
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<mods:date id="1ADA20F56778FD020A606C5EF06C9481">2016</mods:date>
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<mods:number id="A97B1AEF53CF859CF854F6E0FED2E5DB">2016-03-02</mods:number>
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<treatment id="F50D3F20FFBBD35EEBE6FBAFFA73FE1C" ID-DOI="http://doi.org/10.5281/zenodo.6076956" ID-GBIF-Taxon="137308972" ID-Zenodo-Dep="6076956" LSID="urn:lsid:plazi:treatment:F50D3F20FFBBD35EEBE6FBAFFA73FE1C" httpUri="http://treatment.plazi.org/id/F50D3F20FFBBD35EEBE6FBAFFA73FE1C" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
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<subSubSection id="35BEDDBDFFBBD351EBE6FBAFFE19FB7D" pageId="23" pageNumber="174" type="nomenclature">
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<paragraph id="7D1B8E36FFBBD351EBE6FBAFFE05FB9A" blockId="23.[151,508,1113,1173]" box="[151,508,1113,1139]" pageId="23" pageNumber="174">
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<heading id="2653395AFFBBD351EBE6FBAFFE05FB9A" bold="true" box="[151,508,1113,1139]" fontSize="11" level="1" pageId="23" pageNumber="174" reason="1">
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<emphasis id="4FD05224FFBBD351EBE6FBAFFE05FB9A" bold="true" box="[151,508,1113,1139]" pageId="23" pageNumber="174">
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<emphasis id="4FD05224FFBBD351EBE6FBAFFE4BFB9A" bold="true" box="[151,434,1113,1139]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName id="BAA4F5B5FFBBD351EBE6FBAFFE55FB9A" authority="Zwick, 2016" authorityName="Zwick" authorityYear="2016" box="[151,428,1113,1139]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="neboissi" status="sp. nov.">Prionocyphon neboissi</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="54E3EF5FFFBBD351EAC8FBAFFE05FB9A" box="[441,508,1113,1139]" pageId="23" pageNumber="174" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7D1B8E36FFBBD351EBE6FB8AFE19FB7D" blockId="23.[151,508,1113,1173]" box="[151,480,1148,1173]" pageId="23" pageNumber="174">
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(
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<figureCitation id="E59F92B3FFBBD351EBEEFB8AFED8FB7D" box="[159,289,1148,1173]" captionStart="FIGURES 42 − 44" captionStartId="19.[151,264,1681,1703]" captionTargetBox="[161,1422,905,1653]" captionTargetId="figure@19.[151,1435,897,1660]" captionTargetPageId="19" captionText="FIGURES 42 − 44. Prionocyphon neboissi n. sp., heterogenous pilosity on abdominal sternites. Figs. 42 and 43 to the same scale." httpUri="https://zenodo.org/record/1052553/files/figure.png" pageId="23" pageNumber="174">Figs 42−44</figureCitation>
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,
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<figureCitation id="E59F92B3FFBBD351EA5CFB8AFE8EFB7C" box="[301,375,1148,1173]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">54−60</figureCitation>
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,
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<tableCitation id="3026BB8DFFBBD351EAF2FB8AFE2EFB7D" box="[387,471,1148,1173]" captionStart="TABLE 2" captionStartId="17.[151,239,152,175]" captionText="TABLE 2. Character definitions and character matrix for the European type species, Prionocyphon serricornis, and most Australian Prionocyphon species. Asterisked names: not seen, information from Watts (2010 b). Not included: P. kurandaensis Watts, P. microflavidus Watts, and P. ocellatus Watts (not seen, insufficient information), and P. insolitus Watts, P. lamingtonensis Watts and P. macroflavidus Watts which are misplaced in Prionocyphon (Watts 2014)." pageId="23" pageNumber="174">Table 2</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="35BEDDBDFFBBD351EBE6FB32FE0BF9C9" pageId="23" pageNumber="174" type="materials_examined">
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<paragraph id="7D1B8E36FFBBD351EBE6FB32FE8CFACD" blockId="23.[151,1437,1220,2037]" pageId="23" pageNumber="174">
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<emphasis id="4FD05224FFBBD351EBE6FB32FE12FB34" bold="true" box="[151,491,1220,1245]" pageId="23" pageNumber="174">Type material. Holotype 1♂</emphasis>
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: Marysville,
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<emphasis id="4FD05224FFBBD351E9F6FB32FD4FFB34" bold="true" box="[647,694,1220,1245]" pageId="23" pageNumber="174">Vic.</emphasis>
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<date id="091AA8F6FFBBD351E9CEFB32FCC2FB34" box="[703,827,1220,1245]" pageId="23" pageNumber="174" value="1957-11-11">11.11.1957</date>
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A.N. \
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<taxonomicName id="BAA4F5B5FFBBD351E8E6FB32FBF3FB34" box="[919,1034,1220,1245]" class="Insecta" family="Helodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="family">Helodidae</taxonomicName>
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det. A.Newton 1981 (T-22031; on card, in poor shape) (MV).
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<emphasis id="4FD05224FFBBD351EABFFB11FD96FAE9" bold="true" box="[462,623,1255,1280]" pageId="23" pageNumber="174">1♂ Paratype:</emphasis>
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35.16S 149.06
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<emphasis id="4FD05224FFBBD351E853FB11FCA4FAE9" bold="true" box="[802,861,1255,1280]" pageId="23" pageNumber="174">ACT</emphasis>
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nr.
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<collectingMethod id="A4E5F621FFBBD351E8FBFB11FC0AFB16" box="[906,1011,1255,1280]" pageId="23" pageNumber="174">light trap</collectingMethod>
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, Black Mtn.
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<date id="091AA8F6FFBBD351EFF7FB11FAE1FAE9" box="[1158,1304,1255,1280]" pageId="23" pageNumber="174" value="1993-07-15">15 July 1993</date>
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W.Dressler under bark (ANIC).
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</paragraph>
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<paragraph id="7D1B8E36FFBBD351EBB6FAD9FE0BF9C9" blockId="23.[151,1437,1220,2037]" pageId="23" pageNumber="174">
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<emphasis id="4FD05224FFBBD351EBB6FAD9FEDFFAA1" bold="true" box="[199,294,1327,1352]" pageId="23" pageNumber="174">Habitus</emphasis>
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. BL
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<quantity id="BA5C23D3FFBBD351EA17FAC6FE3BFAAE" box="[358,450,1328,1352]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="23" pageNumber="174" unit="mm" value="2.9">2.9 mm</quantity>
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, BL/BW ~1.6. Slender, elongate oval, sides of anterior 2/3 of elytra almost parallel. Head, pronotum and scutellum with fairly large, deep punctures which are barely one diameter apart. Punctures on elytra still larger. Entire dorsal side yellowish brown, covered with yellowish shaggy hairs. Head strongly transverse, eyes bulging. Clypeus with projecting square lobes on either side of labrum. Antennal flagellum not serrate. Second flagellomere more than twice as long as wide at apex, the others shorter, about twice as long. Mandibles unmodified, maxillae and labium not studied. Subgenal ridge with distinct buttonhole, rest of underside of head and thorax not studied.
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</paragraph>
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</subSubSection>
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<subSubSection id="35BEDDBDFFBBD35EEBB6F9DAFDB7FE61" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="description">
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<paragraph id="7D1B8E36FFBBD351EBB6F9DAFB07F93C" blockId="23.[151,1437,1220,2037]" pageId="23" pageNumber="174">
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<emphasis id="4FD05224FFBBD351EBB6F9DAFEFDF9AC" bold="true" box="[199,260,1580,1605]" pageId="23" pageNumber="174">Male</emphasis>
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. Setation of abdominal sternites heterogenous (
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<figureCitation id="E59F92B3FFBBD351E84FF9DAFC31F9AD" box="[830,968,1580,1605]" captionStart="FIGURES 42 − 44" captionStartId="19.[151,264,1681,1703]" captionTargetBox="[161,1422,905,1653]" captionTargetId="figure@19.[151,1435,897,1660]" captionTargetPageId="19" captionText="FIGURES 42 − 44. Prionocyphon neboissi n. sp., heterogenous pilosity on abdominal sternites. Figs. 42 and 43 to the same scale." httpUri="https://zenodo.org/record/1052553/files/figure.png" pageId="23" pageNumber="174">Figs 42–44</figureCitation>
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). Apodemes of T8 not longer than the transverse plate, indistinctly connected by a transverse sclerite. Caudal portion of plate covered by irregular microtrichia, the straight rear edge with sparse fringe of long setae (
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<figureCitation id="E59F92B3FFBBD351E8E7F982FC13F964" box="[918,1002,1652,1677]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">Fig. 54</figureCitation>
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). S8 missing. Apodemes of T9 longer, thin, distinctly curved, the colourless bare plate without defined shape (
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<figureCitation id="E59F92B3FFBBD351E8C0F961FBFCF959" box="[945,1029,1687,1712]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">Fig. 55</figureCitation>
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). S9 large, the wide base with paired sclerotized stripes resembling apodemes. Caudal part with long pilosity, rear edge rounded (
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<figureCitation id="E59F92B3FFBBD351EFECF94AFB08F93C" box="[1181,1265,1724,1749]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">Fig. 56</figureCitation>
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).
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</paragraph>
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<paragraph id="7D1B8E36FFBBD351EBB6F916FC5BF88D" blockId="23.[151,1437,1220,2037]" pageId="23" pageNumber="174">
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Tegmen (
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<figureCitation id="E59F92B3FFBBD351EA43F929FE7FF911" box="[306,390,1759,1784]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">Fig. 57</figureCitation>
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) large, the sclerotized front with wide U-shaped notch. The parameres are two large plates ending in triangular pointed lobes separated by narrow gap. Numerous minute sensilla (socketed setae) on outside near widest part, the slender tips with extremely fine pilosity or microtrichia; details not visible at 630×. The long and slender styles bear distally some fine microtrichia-like structures.
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</paragraph>
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<paragraph id="7D1B8E36FFBBD35EEBB6F899FA65FEA9" blockId="23.[151,1437,1220,2037]" lastBlockId="24.[151,1436,151,501]" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
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Penis (
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<figureCitation id="E59F92B3FFBBD351EA67F899FE64F861" box="[278,413,1903,1928]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">Figs 58, 59</figureCitation>
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) slender, the rounded pala only a little wider than the tongue-shaped caudal end which probably represents fused parameroids. A little beyond midlength originates the narrow, slender trigonium which resembles a fish bone. Its minimally thickened apex bears a few asperities and extends well beyond the penis tip. An additional structure of unclear homology (
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<figureCitation id="E59F92B3FFBBD351E9D3F82AFD0CF81C" box="[674,757,2012,2037]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="23" pageNumber="174">Fig. 60</figureCitation>
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) is ventrally loosely connected to the penis, consisting of a short straight rod which divides into two long struts which first lie close together but diverge caudally. Each strut ends in an inwardly curved tube with a colourless process at tip. On the side of the struts directed towards the penis is a membranous flange ending in transparent tongue-shaped lobes extending further caudad than curved horns. In the
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<typeStatus id="A21F3094FFB4D35EEBB1FEF3FED9FEF4" box="[192,288,261,285]" pageId="24" pageNumber="175" type="paratype">paratype</typeStatus>
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a very thin clear endophallus is seen to attach to the forked region of said sclerite. The wide section of the endophallus contains a few strongly curved and sharply pointed seta-like structures (inset between
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<figureCitation id="E59F92B3FFB4D35EEE7EFED1FA69FEA9" box="[1295,1424,295,320]" captionStart="FIGURES 54 − 60" captionStartId="24.[151,264,1933,1955]" captionTargetBox="[206,1354,541,1911]" captionTargetId="figure@24.[196,1366,535,1926]" captionTargetPageId="24" captionText="FIGURES 54 − 60. Prionocyphon neboissi n. sp., male. 54, T 8; 55, T 9; 56, S 9; 57, tegmen with parameres and styles; 58, penis, the accessory sclerite lies over it in oblique position (paratype); 59, 60, penis and accessory sclerite, beneath it an enlarged detail of the endophallus attaching to it (holotype). 54 − 56, 57 − 60 to the same scales, respectively." httpUri="https://zenodo.org/record/1052557/files/figure.png" pageId="24" pageNumber="175">Figs 58, 60</figureCitation>
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).
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</paragraph>
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<paragraph id="7D1B8E36FFB4D35EEBB6FEBAFE5BFE8C" blockId="24.[151,1436,151,501]" box="[199,418,332,357]" pageId="24" pageNumber="175">
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<emphasis id="4FD05224FFB4D35EEBB6FEBAFEE7FE8C" bold="true" box="[199,286,332,357]" pageId="24" pageNumber="175">Female</emphasis>
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. Unknown.
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</paragraph>
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<paragraph id="7D1B8E36FFB4D35EEBB6FE99FDB7FE61" blockId="24.[151,1436,151,501]" box="[199,590,367,392]" pageId="24" pageNumber="175">
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||
<emphasis id="4FD05224FFB4D35EEBB6FE99FF06FE61" bold="true" box="[199,255,367,392]" pageId="24" pageNumber="175">Note</emphasis>
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||
. There are no similar species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35BEDDBDFFB4D35EEBB6FE62FA73FE1C" pageId="24" pageNumber="175" type="etymology">
|
||
<paragraph id="7D1B8E36FFB4D35EEBB6FE62FA73FE1C" blockId="24.[151,1436,151,501]" pageId="24" pageNumber="175">
|
||
<emphasis id="4FD05224FFB4D35EEBB6FE62FEBFFE44" bold="true" box="[199,326,404,429]" pageId="24" pageNumber="175">Etymology</emphasis>
|
||
. Named in memory of the collector of the
|
||
<typeStatus id="A21F3094FFB4D35EE844FE62FC63FE44" box="[821,922,404,429]" pageId="24" pageNumber="175" type="holotype">holotype</typeStatus>
|
||
, the late Arturs Neboiss with whom I was in contact for a long time, because of our shared interest in Plecoptera. He was curator of the insect collection at the then National Museum of
|
||
<collectingRegion id="BF6040D4FFB4D35EEACCFE2AFDEEFE1C" box="[445,535,476,501]" country="Australia" name="Victoria" pageId="24" pageNumber="175">Victoria</collectingRegion>
|
||
in 1972. He allowed me to work in the collection, and helped in various ways.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |