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<mods:title id="1FA4DBF8B074AD0233676404701864A0">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<treatment id="F50D3F20FFB5D35FEBE6FE08FB9EFBBF" ID-DOI="http://doi.org/10.5281/zenodo.6076960" ID-GBIF-Taxon="137308979" ID-Zenodo-Dep="6076960" LSID="urn:lsid:plazi:treatment:F50D3F20FFB5D35FEBE6FE08FB9EFBBF" httpUri="http://treatment.plazi.org/id/F50D3F20FFB5D35FEBE6FE08FB9EFBBF" lastPageNumber="176" pageId="25" pageNumber="176">
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<paragraph id="7D1B8E36FFB5D35FEBE6FE08FDBAFDF1" blockId="25.[151,579,510,570]" box="[151,579,510,536]" pageId="25" pageNumber="176">
<heading id="2653395AFFB5D35FEBE6FE08FDBAFDF1" bold="true" box="[151,579,510,536]" fontSize="11" level="1" pageId="25" pageNumber="176" reason="1">
<taxonomicName id="BAA4F5B5FFB5D35FEBE6FE08FDBAFDF1" authority="Watts, 2010 b" authorityName="Watts" authorityYear="2010" box="[151,579,510,536]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="4FD05224FFB5D35FEBE6FE08FDBAFDF1" bold="true" box="[151,579,510,536]" pageId="25" pageNumber="176">
<emphasis id="4FD05224FFB5D35FEBE6FE08FE60FDF1" bold="true" box="[151,409,510,536]" italics="true" pageId="25" pageNumber="176">Prionocyphon storeyi</emphasis>
<bibRefCitation id="1935F3C7FFB5D35FEAD1FE08FDBAFDF1" author="Watts" box="[416,579,510,536]" pageId="25" pageNumber="176" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">Watts, 2010b</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="7D1B8E36FFB5D35FEBE6FDD7FE73FDD0" blockId="25.[151,579,510,570]" box="[151,394,545,570]" pageId="25" pageNumber="176">
(
<figureCitation id="E59F92B3FFB5D35FEBEEFDD7FED8FDD3" box="[159,289,545,570]" captionStart="FIGURES 61 64" captionStartId="26.[151,264,1889,1911]" captionTargetBox="[312,1275,329,1868]" captionTargetId="figure@26.[312,1275,329,1876]" captionTargetPageId="26" captionText="FIGURES 61 64. Prionocyphon storeyi Watts, male genitalia. 61, Mt. Tamborine, arrow points at delicate tip of stylus; 62, Mossman Gorge; 63, Mt. Spec, arrow pointing at scaly surface of retracted endophallus; 64, diagram showing the tegmen and its components in grey shade, for easier distinction from the penis and its components. Abbreviations: pa, pala; pd, parameroid; pe, paramere; st, stylus; te, tegmen; tr, trigonium. Not to scale." httpUri="https://zenodo.org/record/1052559/files/figure.png" pageId="25" pageNumber="176">Figs 6164</figureCitation>
,
<tableCitation id="3026BB8DFFB5D35FEA5CFDD7FE78FDD0" box="[301,385,545,570]" captionStart="TABLE 2" captionStartId="17.[151,239,152,175]" captionText="TABLE 2. Character definitions and character matrix for the European type species, Prionocyphon serricornis, and most Australian Prionocyphon species. Asterisked names: not seen, information from Watts (2010 b). Not included: P. kurandaensis Watts, P. microflavidus Watts, and P. ocellatus Watts (not seen, insufficient information), and P. insolitus Watts, P. lamingtonensis Watts and P. macroflavidus Watts which are misplaced in Prionocyphon (Watts 2014)." pageId="25" pageNumber="176">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFFB5D35FEBE6FD9FFB9EFBBF" pageId="25" pageNumber="176" type="materials_examined">
<paragraph id="7D1B8E36FFB5D35FEBE6FD9FFF00FD23" blockId="25.[151,1437,617,1110]" pageId="25" pageNumber="176">
<materialsCitation id="CDCC846BFFB5D35FEBE6FD9FFC26FD6B" ID-GBIF-Occurrence="1701200974" box="[151,991,617,642]" collectingDate="1967-04-24" collectorName="Material" county="Colless" location="Mossman Gorge" municipality="Qld." pageId="25" pageNumber="176" specimenCount="1" specimenCount-male="1">
<emphasis id="4FD05224FFB5D35FEBE6FD9FFE9CFD6B" bold="true" box="[151,357,617,642]" pageId="25" pageNumber="176">
<collectorName id="D051EBE0FFB5D35FEBE6FD9FFF06FD6B" box="[151,255,617,642]" pageId="25" pageNumber="176">Material</collectorName>
studied.
</emphasis>
<specimenCount id="6BA245BFFFB5D35FEA1DFD9CFE6BFD6B" box="[364,402,618,642]" pageId="25" pageNumber="176" type="male">1♂</specimenCount>
:
<location id="787BD8EDFFB5D35FEAEFFD9CFDA3FD6B" LSID="urn:lsid:plazi:treatment:F50D3F20FFB5D35FEBE6FE08FB9EFBBF:787BD8EDFFB5D35FEAEFFD9CFDA3FD6B" box="[414,602,618,642]" county="Colless" municipality="Qld." name="Mossman Gorge" pageId="25" pageNumber="176">Mossman Gorge</location>
,
<emphasis id="4FD05224FFB5D35FE917FD9FFD4CFD6B" bold="true" box="[614,693,617,642]" pageId="25" pageNumber="176">
N.
<collectingMunicipality id="9D7F144CFFB5D35FE9F1FD9FFD4CFD6B" box="[640,693,617,642]" pageId="25" pageNumber="176">Qld.</collectingMunicipality>
</emphasis>
<date id="091AA8F6FFB5D35FE9CDFD9CFCB2FD6B" box="[700,843,617,642]" pageId="25" pageNumber="176" value="1967-04-24">
<collectingDate id="195E511EFFB5D35FE9CDFD9CFCB2FD6B" box="[700,843,617,642]" pageId="25" pageNumber="176" value="1967-04-24">24 Apr 1967</collectingDate>
</date>
D.H.
<collectingCounty id="947AF6BAFFB5D35FE8F4FD9FFC26FD6B" box="[901,991,617,642]" pageId="25" pageNumber="176">Colless</collectingCounty>
</materialsCitation>
;
<materialsCitation id="CDCC846BFFB5D35FE89BFD9CFB7CFD6B" ID-GBIF-Occurrence="1701200987" box="[1002,1157,618,642]" location="Mt. Spec" pageId="25" pageNumber="176" specimenCount="1" specimenCount-male="1">
<specimenCount id="6BA245BFFFB5D35FE89BFD9CFBE9FD6B" box="[1002,1040,618,642]" pageId="25" pageNumber="176" type="male">1♂</specimenCount>
:
<location id="787BD8EDFFB5D35FEF6DFD9CFB7CFD6B" LSID="urn:lsid:plazi:treatment:F50D3F20FFB5D35FEBE6FE08FB9EFBBF:787BD8EDFFB5D35FEF6DFD9CFB7CFD6B" box="[1052,1157,618,642]" name="Mt. Spec" pageId="25" pageNumber="176">Mt. Spec</location>
</materialsCitation>
.
<materialsCitation id="CDCC846BFFB5D35FEFF8FD9CFC4CFD4F" ID-GBIF-Occurrence="1701200977" collectionCode="ANIC" collectorName="J. G. Brooks &amp; Q. A. M. Lea" location="Mt. Tambourine" pageId="25" pageNumber="176" specimenCount="1" specimenCount-male="1">
NQ. 1/GB /
<collectorName id="D051EBE0FFB5D35FEE62FD9CFA65FD6B" box="[1299,1436,617,642]" pageId="25" pageNumber="176">J. G. Brooks</collectorName>
bequest 1976 (both
<collectionCode id="1BB516F3FFB5D35FEAF2FD78FE34FD4F" box="[387,461,654,678]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="25" pageNumber="176">ANIC</collectionCode>
).
<specimenCount id="6BA245BFFFB5D35FEA92FD78FDF0FD4F" box="[483,521,654,678]" pageId="25" pageNumber="176" type="male">1♂</specimenCount>
:
<location id="787BD8EDFFB5D35FE96BFD78FD2DFD4F" LSID="urn:lsid:plazi:treatment:F50D3F20FFB5D35FEBE6FE08FB9EFBBF:787BD8EDFFB5D35FE96BFD78FD2DFD4F" box="[538,724,653,678]" name="Mt. Tambourine" pageId="25" pageNumber="176">Mt. Tambourine</location>
[sic!]
<collectorName id="D051EBE0FFB5D35FE857FD78FC4CFD4F" box="[806,949,654,678]" pageId="25" pageNumber="176">Q. A.M.Lea</collectorName>
</materialsCitation>
;
<materialsCitation id="CDCC846BFFB5D35FE8B7FD78FF00FD23" ID-GBIF-Occurrence="1701200968" collectionCode="NSW, SAMA" location="Blue Mts." municipality="Blackburn" pageId="25" pageNumber="176" specimenCount="1" specimenCount-male="1">
<specimenCount id="6BA245BFFFB5D35FE8B7FD78FC15FD4F" box="[966,1004,654,678]" pageId="25" pageNumber="176" type="male">1♂</specimenCount>
:
<location id="787BD8EDFFB5D35FE88CFD7BFB8BFD4C" LSID="urn:lsid:plazi:treatment:F50D3F20FFB5D35FEBE6FE08FB9EFBBF:787BD8EDFFB5D35FE88CFD7BFB8BFD4C" box="[1021,1138,653,678]" municipality="Blackburn" name="Blue Mts." pageId="25" pageNumber="176">Blue Mts.</location>
,
<collectingMunicipality id="9D7F144CFFB5D35FEFF3FD7BFB0EFD4F" box="[1154,1271,653,678]" pageId="25" pageNumber="176">Blackburn</collectingMunicipality>
[
<emphasis id="4FD05224FFB5D35FEE7DFD7BFAB5FD4F" bold="true" box="[1292,1356,653,678]" pageId="25" pageNumber="176">
<collectionCode id="1BB516F3FFB5D35FEE7DFD7BFAB5FD4F" box="[1292,1356,653,678]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="25" pageNumber="176">NSW</collectionCode>
</emphasis>
] (both
<collectionCode id="1BB516F3FFB5D35FEBE6FD44FF14FD23" box="[151,237,690,714]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="25" pageNumber="176">SAMA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="7D1B8E36FFB5D35FEBB6FD23FC41FC97" blockId="25.[151,1437,617,1110]" pageId="25" pageNumber="176">
Differs from its relatives by the slenderness of the genitalia. Parameroids strongly sclerotized and longer than club-shaped hyaline parameres (arrow in
<figureCitation id="E59F92B3FFB5D35FE919FD0FFD42FCFB" box="[616,699,761,786]" captionStart="FIGURES 61 64" captionStartId="26.[151,264,1889,1911]" captionTargetBox="[312,1275,329,1868]" captionTargetId="figure@26.[312,1275,329,1876]" captionTargetPageId="26" captionText="FIGURES 61 64. Prionocyphon storeyi Watts, male genitalia. 61, Mt. Tamborine, arrow points at delicate tip of stylus; 62, Mossman Gorge; 63, Mt. Spec, arrow pointing at scaly surface of retracted endophallus; 64, diagram showing the tegmen and its components in grey shade, for easier distinction from the penis and its components. Abbreviations: pa, pala; pd, parameroid; pe, paramere; st, stylus; te, tegmen; tr, trigonium. Not to scale." httpUri="https://zenodo.org/record/1052559/files/figure.png" pageId="25" pageNumber="176">Fig. 61</figureCitation>
). Slender styles with terminal fringe long and distinct. Trigonium apex hook-like, in slide mounts its anteriorly diverging contour lines are seen to connect to sides of the penis. The appearance changes with position, and the shape possibly differs slightly between specimens (
<figureCitation id="E59F92B3FFB5D35FEFA4FCB7FAA2FCB3" box="[1237,1371,833,858]" captionStart="FIGURES 61 64" captionStartId="26.[151,264,1889,1911]" captionTargetBox="[312,1275,329,1868]" captionTargetId="figure@26.[312,1275,329,1876]" captionTargetPageId="26" captionText="FIGURES 61 64. Prionocyphon storeyi Watts, male genitalia. 61, Mt. Tamborine, arrow points at delicate tip of stylus; 62, Mossman Gorge; 63, Mt. Spec, arrow pointing at scaly surface of retracted endophallus; 64, diagram showing the tegmen and its components in grey shade, for easier distinction from the penis and its components. Abbreviations: pa, pala; pd, parameroid; pe, paramere; st, stylus; te, tegmen; tr, trigonium. Not to scale." httpUri="https://zenodo.org/record/1052559/files/figure.png" pageId="25" pageNumber="176">Figs 6163</figureCitation>
). The apex was mistaken for a cover of the endophallus at rest (
<bibRefCitation id="1935F3C7FFB5D35FE868FC90FC55FC97" author="Watts" box="[793,940,869,894]" pageId="25" pageNumber="176" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">Watts 2010b</bibRefCitation>
).
</paragraph>
<paragraph id="7D1B8E36FFB5D35FEBB6FC7FFDA2FBE7" blockId="25.[151,1437,617,1110]" pageId="25" pageNumber="176">
Endophallus long and slender, front end divided and armed with small spinules, further back with long spines (
<figureCitation id="E59F92B3FFB5D35FEBEEFC5BFF0AFC2F" box="[159,243,941,966]" captionStart="FIGURES 61 64" captionStartId="26.[151,264,1889,1911]" captionTargetBox="[312,1275,329,1868]" captionTargetId="figure@26.[312,1275,329,1876]" captionTargetPageId="26" captionText="FIGURES 61 64. Prionocyphon storeyi Watts, male genitalia. 61, Mt. Tamborine, arrow points at delicate tip of stylus; 62, Mossman Gorge; 63, Mt. Spec, arrow pointing at scaly surface of retracted endophallus; 64, diagram showing the tegmen and its components in grey shade, for easier distinction from the penis and its components. Abbreviations: pa, pala; pd, parameroid; pe, paramere; st, stylus; te, tegmen; tr, trigonium. Not to scale." httpUri="https://zenodo.org/record/1052559/files/figure.png" pageId="25" pageNumber="176">Fig. 64</figureCitation>
). Caudally a single dense pack of long spines present, separated by pale ring from dense pelt of long spines which spread outward upon eversion (
<figureCitation id="E59F92B3FFB5D35FE930FC27FD6CFC03" box="[577,661,977,1002]" captionStart="FIGURES 61 64" captionStartId="26.[151,264,1889,1911]" captionTargetBox="[312,1275,329,1868]" captionTargetId="figure@26.[312,1275,329,1876]" captionTargetPageId="26" captionText="FIGURES 61 64. Prionocyphon storeyi Watts, male genitalia. 61, Mt. Tamborine, arrow points at delicate tip of stylus; 62, Mossman Gorge; 63, Mt. Spec, arrow pointing at scaly surface of retracted endophallus; 64, diagram showing the tegmen and its components in grey shade, for easier distinction from the penis and its components. Abbreviations: pa, pala; pd, parameroid; pe, paramere; st, stylus; te, tegmen; tr, trigonium. Not to scale." httpUri="https://zenodo.org/record/1052559/files/figure.png" pageId="25" pageNumber="176">Fig. 61</figureCitation>
). At rest, membranous outer surface of retracted endophallus at penis tip with scaly surface (arrow in
<figureCitation id="E59F92B3FFB5D35FEA8AFC03FDB7FBE7" box="[507,590,1013,1038]" captionStart="FIGURES 61 64" captionStartId="26.[151,264,1889,1911]" captionTargetBox="[312,1275,329,1868]" captionTargetId="figure@26.[312,1275,329,1876]" captionTargetPageId="26" captionText="FIGURES 61 64. Prionocyphon storeyi Watts, male genitalia. 61, Mt. Tamborine, arrow points at delicate tip of stylus; 62, Mossman Gorge; 63, Mt. Spec, arrow pointing at scaly surface of retracted endophallus; 64, diagram showing the tegmen and its components in grey shade, for easier distinction from the penis and its components. Abbreviations: pa, pala; pd, parameroid; pe, paramere; st, stylus; te, tegmen; tr, trigonium. Not to scale." httpUri="https://zenodo.org/record/1052559/files/figure.png" pageId="25" pageNumber="176">Fig. 63</figureCitation>
).
</paragraph>
<paragraph id="7D1B8E36FFB5D35FEBB6FBEFFB9EFBBF" blockId="25.[151,1437,617,1110]" pageId="25" pageNumber="176">
The new records from southern Queensland and New South Wales close the gap between the known occurrence in North Queensland and the record from Lord Howe Island (
<bibRefCitation id="1935F3C7FFB5D35FE8B9FBC8FBA3FBBF" author="Watts" box="[968,1114,1085,1110]" pageId="25" pageNumber="176" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">Watts 2010b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>