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<mods:title id="5DEEDC37B687D2592F744FD2DCF60A15">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<mods:namePart id="B7544E55300D00D7D0B293D1D1F22472">Zwick, Peter</mods:namePart>
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<treatment id="F50D3F20FF8DD365EBE6F8CEFC47FB49" ID-DOI="http://doi.org/10.5281/zenodo.6076985" ID-GBIF-Taxon="137308983" ID-Zenodo-Dep="6076985" LSID="urn:lsid:plazi:treatment:F50D3F20FF8DD365EBE6F8CEFC47FB49" httpUri="http://treatment.plazi.org/id/F50D3F20FF8DD365EBE6F8CEFC47FB49" lastPageId="35" lastPageNumber="186" pageId="33" pageNumber="184">
<subSubSection id="35BEDDBDFF8DD367EBE6F8CEFE71F89A" pageId="33" pageNumber="184" type="nomenclature">
<paragraph id="7D1B8E36FF8DD367EBE6F8CEFD89F8BB" blockId="33.[151,624,1848,1908]" box="[151,624,1848,1874]" pageId="33" pageNumber="184">
<heading id="2653395AFF8DD367EBE6F8CEFD89F8BB" bold="true" box="[151,624,1848,1874]" fontSize="11" level="1" pageId="33" pageNumber="184" reason="1">
<taxonomicName id="BAA4F5B5FF8DD367EBE6F8CEFD89F8BB" authority="Watts, 2010 b" authorityName="Watts" authorityYear="2010" box="[151,624,1848,1874]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="yarraensis">
<emphasis id="4FD05224FF8DD367EBE6F8CEFD89F8BB" bold="true" box="[151,624,1848,1874]" pageId="33" pageNumber="184">
<emphasis id="4FD05224FF8DD367EBE6F8CEFE3FF8BB" bold="true" box="[151,454,1848,1874]" italics="true" pageId="33" pageNumber="184">Prionocyphon yarraensis</emphasis>
<bibRefCitation id="1935F3C7FF8DD367EABFF8CEFD89F8BB" author="Watts" box="[462,624,1848,1874]" pageId="33" pageNumber="184" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">Watts, 2010b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7D1B8E36FF8DD367EBE6F8ADFE71F89A" blockId="33.[151,624,1848,1908]" box="[151,392,1883,1908]" pageId="33" pageNumber="184">
(
<figureCitation id="E59F92B3FF8DD367EBEEF8ADFED9F89D" box="[159,288,1883,1908]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="33" pageNumber="184">Figs 8186</figureCitation>
,
<tableCitation id="3026BB8DFF8DD367EA5AF8ADFE86F89A" box="[299,383,1883,1908]" captionStart="TABLE 2" captionStartId="17.[151,239,152,175]" captionText="TABLE 2. Character definitions and character matrix for the European type species, Prionocyphon serricornis, and most Australian Prionocyphon species. Asterisked names: not seen, information from Watts (2010 b). Not included: P. kurandaensis Watts, P. microflavidus Watts, and P. ocellatus Watts (not seen, insufficient information), and P. insolitus Watts, P. lamingtonensis Watts and P. macroflavidus Watts which are misplaced in Prionocyphon (Watts 2014)." pageId="33" pageNumber="184">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFF8DD365EBE6F855FD40FE1C" lastPageId="35" lastPageNumber="186" pageId="33" pageNumber="184" type="materials_examined">
<paragraph id="7D1B8E36FF8DD367EBE6F855FF0CF836" blockId="33.[151,1436,1955,2015]" pageId="33" pageNumber="184">
<emphasis id="4FD05224FF8DD367EBE6F855FE9FF855" bold="true" box="[151,358,1955,1980]" pageId="33" pageNumber="184">Material studied.</emphasis>
<specimenCount id="6BA245BFFF8DD367EA1FF855FE65F852" box="[366,412,1955,1979]" pageId="33" pageNumber="184" type="male">1 ♂</specimenCount>
:
<geoCoordinate id="1890E8F1FF8DD367EAD8F855FE00F855" box="[425,505,1955,1980]" direction="south" orientation="latitude" pageId="33" pageNumber="184" precision="555" value="-36.36">36.36S</geoCoordinate>
<geoCoordinate id="1890E8F1FF8DD367E971F855FDA7F852" box="[512,606,1955,1979]" direction="east" orientation="longitude" pageId="33" pageNumber="184" precision="555" value="149.23">149.23E</geoCoordinate>
<emphasis id="4FD05224FF8DD367E916F855FD5EF855" bold="true" box="[615,679,1955,1980]" pageId="33" pageNumber="184">
<collectionCode id="1BB516F3FF8DD367E916F855FD5EF855" box="[615,679,1955,1980]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="33" pageNumber="184">NSW</collectionCode>
</emphasis>
Brown Mountain
<date id="091AA8F6FF8DD367E808F855FBEBF852" box="[889,1042,1955,1980]" pageId="33" pageNumber="184" value="1988-12-26">26 Dec. 1988</date>
C. Reid beating bushes at lookout (
<collectionCode id="1BB516F3FF8DD367EBEEF831FF13F836" box="[159,234,1991,2015]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="33" pageNumber="184">ANIC</collectionCode>
).
</paragraph>
<caption id="29DBDEBEFF8ED364EBE6F90DFBF0F8D9" httpUri="https://zenodo.org/record/1052569/files/figure.png" pageId="34" pageNumber="185" targetBox="[158,1430,205,1753]" targetPageId="34">
<paragraph id="7D1B8E36FF8ED364EBE6F90DFBF0F8D9" blockId="34.[151,1436,1787,1840]" pageId="34" pageNumber="185">
<emphasis id="4FD05224FF8ED364EBE6F90DFEAEF8F8" bold="true" box="[151,343,1787,1809]" pageId="34" pageNumber="185">FIGURES 79, 80.</emphasis>
Erect male genitalia of
<taxonomicName id="BAA4F5B5FF8ED364E923F90DFC89F8F8" authority="Watts." authorityName="Watts." box="[594,880,1787,1809]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="warra">
<emphasis id="4FD05224FF8ED364E923F90DFCD1F8F8" box="[594,808,1787,1809]" italics="true" pageId="34" pageNumber="185">Prionocyphon warra</emphasis>
Watts.
</taxonomicName>
79, lateral view; 80, dorsal view. Abbreviations: eph, endophallus; pa, pala; pd, parameroid; pe, paramere; st, style; te, tegmen; tr, trigonium.
</paragraph>
</caption>
<paragraph id="7D1B8E36FF8ED364EBB6F8AAFE63F870" blockId="34.[151,1437,1884,2017]" pageId="34" pageNumber="185">
The present specimen is
<quantity id="BA5C23D3FF8ED364EA92F8ABFDC2F89D" box="[483,571,1885,1909]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="34" pageNumber="185" unit="mm" value="3.3">3.3 mm</quantity>
long, a little less than the
<typeStatus id="A21F3094FF8ED364E81FF8AAFC2DF89C" box="[878,980,1884,1909]" pageId="34" pageNumber="185" type="holotype">holotype</typeStatus>
. Habitus and colouration agree with the description of the
<typeStatus id="A21F3094FF8ED364EA13F877FE6FF870" box="[354,406,1921,1945]" pageId="34" pageNumber="185">type</typeStatus>
.
</paragraph>
<paragraph id="7D1B8E36FF8ED365EBB6F852FADBFF11" blockId="34.[151,1437,1884,2017]" lastBlockId="35.[151,1437,151,1184]" lastPageId="35" lastPageNumber="186" pageId="34" pageNumber="185">Elongate, sides of elytra almost parallel, body fairly flat. Yellowish brown, antenna filiform, not serrate. Flagellomeres slender, 3 times longer than wide. Yellow pilosity long. Punctures on head and pronotum very fine, those on elytra large. Clypeus with large divergent lateral lobes. End of subgenal ridge raised, separated from raised edge of oral cavity by a distinct narrow gap (buttonhole). Pilosity of abdominal sternites S35 with many large sensilla, S6 and S7 only with straight socketed setae, setation on S7 twice as dense as on other sternites.</paragraph>
<paragraph id="7D1B8E36FF8FD365EBB6FEF2FD40FE1C" blockId="35.[151,1437,151,1184]" pageId="35" pageNumber="186">
The genitalia of the present beetle broke during dissection, damaging the basal plate (bp) and destroying the original connection of the tegmen and penis to the basal plate. The penis (
<figureCitation id="E59F92B3FF8FD365E893FED1FBCCFEA9" box="[994,1077,295,320]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 84</figureCitation>
) and both halves of the tegmen (
<figureCitation id="E59F92B3FF8FD365EBEEFEBAFF16FE8C" box="[159,239,332,357]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 85</figureCitation>
shows one of them) lie flat in a slide and are seen in side view. A photograph of the
<typeStatus id="A21F3094FF8FD365EFE9FEBAFB03FE8C" box="[1176,1274,332,357]" pageId="35" pageNumber="186" type="holotype">holotype</typeStatus>
genitalia slide (courtesy C.H.S. Watts) shows the intact structures, in dorsal view, as drawn by
<bibRefCitation id="1935F3C7FF8FD365EF7EFE86FE7CFE44" author="Watts" pageId="35" pageNumber="186" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">
Watts (2010b: fig. 42; reprinted and annotated in
<figureCitation id="E59F92B3FF8FD365EA58FE62FE85FE44" box="[297,380,404,429]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 86</figureCitation>
)
</bibRefCitation>
. In dorsal view, the basal plate is in the background and not seen at all while the curved tube of the pala appears like a single median structure. My description is based on both supplementary views, corresponding details are labelled in
<figureCitation id="E59F92B3FF8FD365E945FE2AFD4CFE1C" box="[564,693,476,501]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Figs 8486</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFF8FD365EBB6FE09FC47FB49" pageId="35" pageNumber="186" type="description">
<paragraph id="7D1B8E36FF8FD365EBB6FE09FA92FD41" blockId="35.[151,1437,151,1184]" pageId="35" pageNumber="186">
<emphasis id="4FD05224FF8FD365EBB6FE09FEF2FDF1" bold="true" box="[199,267,511,536]" pageId="35" pageNumber="186">Male.</emphasis>
T8 (
<figureCitation id="E59F92B3FF8FD365EA32FE09FE6CFDF1" box="[323,405,511,536]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 81</figureCitation>
) with short straight incompletely connected apodemes, plate transverse, with rounded rear margin. A mixed band of microtrichia and small setae in front of rear margin. Setae along rear margin a bit larger than others. S8 absent. T9 with curved incompletely connected apodemes, plate membranous, bare, with no defined shape (
<figureCitation id="E59F92B3FF8FD365EB95FD9AFECCFD6D" box="[228,309,620,645]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 82</figureCitation>
). S9 with wide front margin, plate membranous except a wide U-shaped sclerite band which ends in the weak sclerotisation of the widely separate rounded caudal lobes which bear setae laterally and caudally (
<figureCitation id="E59F92B3FF8FD365EE7DFD79FAA4FD41" box="[1292,1373,655,680]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 83</figureCitation>
).
</paragraph>
<paragraph id="7D1B8E36FF8FD365EBB6FD43FAB7FC69" blockId="35.[151,1437,151,1184]" pageId="35" pageNumber="186">
Tegmen H-shaped, two strong anterior rods (te1, te2), at a considerable distance from their front ends, connected by a transverse bar (tb) whose middle articulates with the pala (pa). Basal rods continuing caudally beyond this point and ending in an upcurved sclerotized (sl) and a hyaline lobe (hy). Parameres subterminally attached to rod and partly enveloping penis (
<figureCitation id="E59F92B3FF8FD365E9E1FCE9FD1CFCD1" box="[656,741,799,824]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 86</figureCitation>
). Large elongate main lobe (ml) with rounded apex, medially bearing a low ridge with a few small denticles (dr). On the outside attaches the stylus (sty), which in full view is a large hyaline lobe with a caudal fringe of long delicate hair-like processes (which are not shown in
<figureCitation id="E59F92B3FF8FD365EF9EFC91FABBFC69" box="[1263,1346,871,896]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 86</figureCitation>
).
</paragraph>
<paragraph id="7D1B8E36FF8FD365EBB6FC7AFA9FFBDC" blockId="35.[151,1437,151,1184]" pageId="35" pageNumber="186">
Pala (pa) a U-shaped tube (
<figureCitation id="E59F92B3FF8FD365E979FC7AFD99FC4D" box="[520,608,908,933]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 84</figureCitation>
). Precise shape of basal plate (bp) and original articulation of pala and middle of transverse bar unclear. It is not visible in the
<typeStatus id="A21F3094FF8FD365E861FC59FC8BFC21" box="[784,882,943,968]" pageId="35" pageNumber="186" type="holotype">holotype</typeStatus>
slide and not shown in
<figureCitation id="E59F92B3FF8FD365EFF5FC59FB21FC21" box="[1156,1240,943,968]" captionStart="FIGURES 81 86" captionStartId="36.[151,264,1842,1864]" captionTargetBox="[270,1308,203,1808]" captionTargetId="figure@36.[260,1320,195,1823]" captionTargetPageId="36" captionText="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri="https://zenodo.org/record/1052571/files/figure.png" pageId="35" pageNumber="186">Fig. 86</figureCitation>
. Trigonium (tr) a long curved tube containing a wide sperm duct (sd) with a well defined ventrobasal opening (sd-o). Sperm duct ending at trigonium tip, between the three apical tips. Ventrally from trigonium originates a slightly wider, laterally spinulose, long process with rounded apical lobe with senory pores, probably representing fused parameroids.
</paragraph>
<paragraph id="7D1B8E36FF8FD365EBB6FBC9FC47FB49" blockId="35.[151,1437,151,1184]" pageId="35" pageNumber="186">
<emphasis id="4FD05224FF8FD365EBB6FBC9FF06FBB1" bold="true" box="[199,255,1087,1112]" pageId="35" pageNumber="186">Note</emphasis>
. The new record considerably expands the range of
<taxonomicName id="BAA4F5B5FF8FD365E81FFBB7FBFDFBBE" box="[878,1028,1088,1112]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="yarraensis">
<emphasis id="4FD05224FF8FD365E81FFBB7FBFDFBBE" box="[878,1028,1088,1112]" italics="true" pageId="35" pageNumber="186">P. yarraensis</emphasis>
</taxonomicName>
which was only known from near Warburton, VIC. The connection between the penis and tegmen
<emphasis id="4FD05224FF8FD365E809FB93FC63FB95" box="[888,922,1125,1148]" italics="true" pageId="35" pageNumber="186">via</emphasis>
a basal plate agrees with
<taxonomicName id="BAA4F5B5FF8FD365EFB7FB93FAB4FB95" authorityName="Watts" authorityYear="2010" box="[1222,1357,1124,1148]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="4FD05224FF8FD365EFB7FB93FAB4FB95" box="[1222,1357,1124,1148]" italics="true" pageId="35" pageNumber="186">P. monteithi</emphasis>
</taxonomicName>
and
<taxonomicName id="BAA4F5B5FF8FD365EEF6FB93FF22FB49" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="warra">
<emphasis id="4FD05224FF8FD365EEF6FB93FF22FB49" italics="true" pageId="35" pageNumber="186">P. warra</emphasis>
</taxonomicName>
, but
<taxonomicName id="BAA4F5B5FF8FD365EA65FB7FFE5CFB76" box="[276,421,1160,1184]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="yarraensis">
<emphasis id="4FD05224FF8FD365EA65FB7FFE5CFB76" box="[276,421,1160,1184]" italics="true" pageId="35" pageNumber="186">P. yarraensis</emphasis>
</taxonomicName>
is an exceptional beetle without close relatives.
</paragraph>
</subSubSection>
</treatment>
</document>