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<mods:title id="AA1E87E0C674C16942CBDFCABB18A35F">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<treatment id="F50D3F20FF8CD366EBE6FC24FF18FA9F" ID-DOI="http://doi.org/10.5281/zenodo.6076979" ID-GBIF-Taxon="137308971" ID-Zenodo-Dep="6076979" LSID="urn:lsid:plazi:treatment:F50D3F20FF8CD366EBE6FC24FF18FA9F" httpUri="http://treatment.plazi.org/id/F50D3F20FF8CD366EBE6FC24FF18FA9F" lastPageNumber="183" pageId="32" pageNumber="183">
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<paragraph id="7D1B8E36FF8CD366EBE6FC24FDE7FC05" blockId="32.[151,542,978,1004]" box="[151,542,978,1004]" pageId="32" pageNumber="183">
<heading id="2653395AFF8CD366EBE6FC24FDE7FC05" bold="true" box="[151,542,978,1004]" fontSize="11" level="1" pageId="32" pageNumber="183" reason="1">
<emphasis id="4FD05224FF8CD366EBE6FC24FDE7FC05" bold="true" box="[151,542,978,1004]" pageId="32" pageNumber="183">
The
<emphasis id="4FD05224FF8CD366EBBEFC24FE2BFC05" bold="true" box="[207,466,978,1004]" italics="true" pageId="32" pageNumber="183">
<taxonomicName id="BAA4F5B5FF8CD366EBBEFC24FE83FC05" ID-CoL="6422L" ID-ENA="538624" baseAuthorityName="Watts" baseAuthorityYear="2014" box="[207,378,978,1004]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="genus">Prionocyphon</taxonomicName>
warra-
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group
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<paragraph id="7D1B8E36FF8CD366EBE6FBEFFE5DFAC7" blockId="32.[151,1437,1049,1398]" pageId="32" pageNumber="183">
The group includes species whose genitalia (
<figureCitation id="E59F92B3FF8CD366E9C5FBEFFCC2FBDB" box="[692,827,1049,1074]" captionStart-0="FIGURES 77, 78" captionStart-1="FIGURES 79, 80" captionStart-2="FIGURES 81 86" captionStartId-0="33.[151,264,1715,1737]" captionStartId-1="34.[151,264,1787,1809]" captionStartId-2="36.[151,264,1842,1864]" captionTargetBox-0="[200,1360,556,1670]" captionTargetBox-1="[158,1430,205,1753]" captionTargetBox-2="[270,1308,203,1808]" captionTargetId-0="figure@33.[188,1372,545,1677]" captionTargetId-1="figure@34.[156,1430,195,1765]" captionTargetId-2="figure@36.[260,1320,195,1823]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="36" captionText-0="FIGURES 77, 78. Male genitalia of Prionocyphon monteithi Watts (77) and P. warra Watts (78). Not to scale. Abbreviations: eph, endophallus; pa, pala; pd, parameroid; pe, paramere; st, style; te, tegmen; tr, trigonium." captionText-1="FIGURES 79, 80. Erect male genitalia of Prionocyphon warra Watts. 79, lateral view; 80, dorsal view. Abbreviations: eph, endophallus; pa, pala; pd, parameroid; pe, paramere; st, style; te, tegmen; tr, trigonium." captionText-2="FIGURES 81 86. Male genitalia of Prionocyphon yarraensis Watts. 81, T 8; 82, T 9; 83, S 9; 84, trigonium and parameroids in side view; black arrow points at opening of sperm duct (sd-o); 85, one paramere with half of the transverse bar between the halves, the broken end placed near where the undamedged components were originally connected; wide open arrow indicates direction of view of Fig. 86; 86, dorsal view of the intact genitalia (modified from Watts 2010 b: fig. 42). Compare text, also for abbreviations. 81 83 and 84, 85 to the same scales, respectively; 86 not to scale." httpUri-0="https://zenodo.org/record/1052567/files/figure.png" httpUri-1="https://zenodo.org/record/1052569/files/figure.png" httpUri-2="https://zenodo.org/record/1052571/files/figure.png" pageId="32" pageNumber="183">Figs 7786</figureCitation>
) appear skeletonized, 'skinny' because the tegmen, parameres and penis are intimately connected into a functional unit consisting essentially of only hard sclerites. The tegmen resembles a flat bowl in
<taxonomicName id="BAA4F5B5FF8CD366E942FB94FD41FB90" authorityName="Watts" authorityYear="2010" box="[563,696,1121,1145]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="4FD05224FF8CD366E942FB94FD41FB90" box="[563,696,1121,1145]" italics="true" pageId="32" pageNumber="183">P. monteithi</emphasis>
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but is reduced to a transverse band in
<taxonomicName id="BAA4F5B5FF8CD366EF12FB94FB3BFB90" box="[1123,1218,1122,1145]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="warra">
<emphasis id="4FD05224FF8CD366EF12FB94FB3BFB90" box="[1123,1218,1122,1145]" italics="true" pageId="32" pageNumber="183">P. warra</emphasis>
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. In both it supports slender parameres with an apical hook. The paramere bases are hinged with sclerites that are part of the long pala (missing in fig. 39 of
<bibRefCitation id="1935F3C7FF8CD366EAE6FB5CFDD1FB2B" author="Watts" box="[407,552,1193,1218]" pageId="32" pageNumber="183" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">Watts 2010b</bibRefCitation>
) with a high longitudinal keel. The strong lateral edges of the pala continue posteriorly as supports of the slender parameroids which are basally fused and end in a pair of spatulate sclerites. Between the tegmen and parameroids lies the slender, apically divided cone of the penis to which the unarmed endophallus is attached.
</paragraph>
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<paragraph id="7D1B8E36FF8CD366EBB6FACFFF18FA9F" blockId="32.[151,1437,1049,1398]" pageId="32" pageNumber="183">
The above homologies are largely based on an undissected specimen of
<taxonomicName id="BAA4F5B5FF8CD366EF68FACCFB84FAB8" box="[1049,1149,1338,1361]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="warra">
<emphasis id="4FD05224FF8CD366EF68FACCFB84FAB8" box="[1049,1149,1338,1361]" italics="true" pageId="32" pageNumber="183">P. warra</emphasis>
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with erect genitalia, see below.
</paragraph>
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