122 lines
15 KiB
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122 lines
15 KiB
XML
<document id="0BDB9E6B7EADFC9A6A376F7DD121AB97" ID-CLB-Dataset="35529" ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZooBank="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno" docDate="2015" docId="16C88AA8E00961244257B7F30556D9F3" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Philalanka moluensis E. A. Smith 1893" docType="treatment" docVersion="11" lastPageNumber="56" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="54" updateTime="1701381844169" updateUser="ExternalLinkService">
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<mods:title id="821A33FBBF9B2F5A7A61D646508E26F2">Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
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<mods:namePart id="629149141AF145A601E3B4B201152AB5">Vermeulen, Jaap J.</mods:namePart>
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<mods:namePart id="060EF394F0FE2C3975E129A0D37B46A5">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="0F4937EA7A95C85CB614870CDBD290B9">Schilthuizen, Menno</mods:namePart>
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<mods:title id="63D17420584F430962E123B352DA47D2">ZooKeys</mods:title>
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<mods:date id="E22B28F85CD15AA9C7EB9E7B9A00873B">2015</mods:date>
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<mods:number id="860017E2EAC9E8FDA46749C6F0665B74">531</mods:number>
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<mods:classification id="F5C93ED257E7C74EB24B5C72479BFEBA">journal article</mods:classification>
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<mods:identifier id="78F71F389D40DAB6284406C4C8957D5A" type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
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<treatment id="16C88AA8E00961244257B7F30556D9F3" ID-GBIF-Taxon="144057522" LSID="urn:lsid:plazi:treatment:16C88AA8E00961244257B7F30556D9F3" httpUri="http://treatment.plazi.org/id/16C88AA8E00961244257B7F30556D9F3" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
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<subSubSection id="68AC9EEE1048691D9A4176F5E6E9B5B5" pageId="53" pageNumber="54" type="multiple">
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<paragraph id="EBD8D6945581797BB7913688468FE81A" pageId="53" pageNumber="54">Taxon classification Animalia Stylommatophora Endodontidae</paragraph>
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</subSubSection>
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<subSubSection id="2B612338479F71377EC2D4824763F8A7" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="D89544F9DCD6D5A045822AE8F22886D7" pageId="53" pageNumber="54">
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<taxonomicName id="338F3B51F2AE6BAA7F57F482FEC38155" ID-CoL="4G4VF" authority="E. A. Smith, 1893" authorityName="E. A. Smith" authorityYear="1893" class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka moluensis" order="Stylommatophora" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="moluensis">Philalanka moluensis (E.A. Smith, 1893)</taxonomicName>
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Figure 44
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</paragraph>
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</subSubSection>
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<subSubSection id="8AB8650AE83A429D29C4A4247B52FDE9" pageId="54" pageNumber="55" type="reference_group">
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<paragraph id="08D267B33A34227665A3169404EDE365" pageId="54" pageNumber="55">
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<taxonomicName id="13F42B4B2EC3DB119564F8D6E5621BAC" class="Gastropoda" family="Helicarionidae" genus="Sitala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitala moluensis" order="Stylommatophora" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="moluensis">
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<pageBreakToken id="542BA230C86B5F30C625B1F082972305" pageId="54" pageNumber="55" start="start">Sitala</pageBreakToken>
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moluensis
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</taxonomicName>
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E.A. Smith, 1893: 343.
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<taxonomicName id="9E6B36AF8913471715447AECB5E2C153" class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka moluensis" order="Stylommatophora" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="moluensis">Philalanka moluensis</taxonomicName>
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(E.A. Smith).
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<bibRefCitation id="43321F371060DDDBD5BF42F11A5EA409" author="Schilthuizen, M" journalOrPublisher="Journal of Molluscan Studies" pageId="54" pageNumber="55" pagination="255 - 258" title="Microsnails at microscales in Borneo: distributions of Prosobranchia versus Pulmonata." url="10.1093/mollus/68.3.255" volume="68" year="2002">Schilthuizen et al., 2002</bibRefCitation>
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: 356-257;
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<bibRefCitation id="E52B196F0888D39521A4E979F813D220" author="Schilthuizen, M" editor="Mohamed, M" journalOrPublisher="Universiti Malaysia Sabah, Kota Kinabalu" pageId="54" pageNumber="55" pagination="93 - 98" title="The land snails of the Tabin limestone hill." volumeTitle="Tabin Limestone Scientific Expedition 2000" year="2003">Schilthuizen and Vermeulen 2003</bibRefCitation>
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: 96;
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<bibRefCitation id="C00CDF26EC16F821D14F8F6AAF1BAE30" author="Schilthuizen, M" editor="Mohamed, M" journalOrPublisher="Universiti Malaysia Sabah, Kota Kinabalu" pageId="54" pageNumber="55" title="A checklist of the land snails of the Crocker Range." volumeTitle="Crocker Range Scientific Expedition 2002" year="2004">Schilthuizen 2004</bibRefCitation>
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: 94;
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<bibRefCitation id="6F1AF8401FFC6DB32C9652543B6C2D66" author="Clements, R" journalOrPublisher="Biological Conservation" pageId="54" pageNumber="55" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
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: 2762. Type from Malaysia, Sarawak, Mulu area).
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</paragraph>
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<paragraph id="89530A4436E897831B8F20CD2CEAA483" pageId="54" pageNumber="55">
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<taxonomicName id="C682027F2D8FDA9D985CA0DA02177FA2" class="Gastropoda" family="Helicarionidae" genus="Sitala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitala inaequisculpta" order="Stylommatophora" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="inaequisculpta">Sitala inaequisculpta</taxonomicName>
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E.A. Smith, 1895: 112. Type from Malaysia, Sarawak, Mount Rabong).
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</paragraph>
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</subSubSection>
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<subSubSection id="58A980D8C670615935F1E545C81DEF38" pageId="54" pageNumber="55" type="materials_examined">
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<paragraph id="3595159DBB9FA65EF7C0CA220B4659F0" pageId="54" pageNumber="55">Examined material from Sabah.</paragraph>
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<paragraph id="582944AD1DCD5A21C4F0807A92389D0E" pageId="54" pageNumber="55">
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Interior Province. Upper Padas valley, Matang River South of Long Pasia (leg. J.J. Vermeulen, V 9812). Crocker Range N.P., Kalang Waterfalls (leg. J.J. Vermeulen, V 1178); Crocker Range N.P., Gua Laing c. 12 km North of Keningau (leg. J.J. Vermeulen, V 1110, V 1124); West of the km 10 marker on the road Tambunan-Ranau, Mahua Waterfall (leg. J.J. Vermeulen & M. Schilthuizen, V 9728). Gunung Trusmadi slopes, Gua Loloposon (leg. J.J. Vermeulen, V 13232). Pinangah valley, Batu Urun (= Bukit Sinobang) (leg. J.J. Vermeulen, V 8001, V 1166, V 1167, BOR/MOL 782; leg. M. Schilthuizen, BOR/MOL 773); Batu Punggul (leg. J.J. Vermeulen, V 1986; leg. M. Schilthuizen, BOR/MOL 778); Batu Temurung (leg. J.J. Vermeulen, V 8047, BOR/MOL 781; leg. M. Schilthuizen, BOR/MOL 777); Bukit Tinahas, East end of Batu Punggul limestone (leg. J.J. Vermeulen & M. Schilthuizen, V 7631); Gua Pungiton (leg. J.J. Vermeulen & M. Schilthuizen, V 7558, BOR/MOL 770); Gua Sanaron (leg. J.J. Vermeulen & M. Schilthuizen, V 7668; leg. M. Schilthuizen, BOR/MOL 3619, BOR/MOL 775). Sandakan Province. Kinabatangan valley, Gomantong Hill 30 km South of Sandakan (leg. J.J. Vermeulen & H. Duistermaat, V 1617). Tawau Province. Batu Baturong c. 50 km W.S.W. of Lahad Datu (leg. J.J. Vermeulen & H. Duistermaat, V 1858); North slope (leg. J.J. Vermeulen, V 7595). Segama valley,
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<normalizedToken id="D4D2434E686B228079B45D1C89199472" originalValue="‘Kirk’s" pageId="54" pageNumber="55">'Kirk's</normalizedToken>
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<normalizedToken id="0BC596F82F47596E89406171086911A0" originalValue="Cave’" pageId="54" pageNumber="55">Cave'</normalizedToken>
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8 km North of Lahad Datu (leg. J.J. Vermeulen, V 1219), West Coast Province. Kinabalu N.P., Poring Hot Springs, along path to waterfall (leg. J.J. Vermeulen, V 13009).
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</paragraph>
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<caption id="FD6B423F0397C89B5BA031579A27B9DB" pageId="54" pageNumber="55">
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<paragraph id="9A8DA426326A7BA4826D33108E265B65" pageId="54" pageNumber="55">
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Figure 44-46. 44
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<taxonomicName id="03A1768ED679F0650FE88E52B16A76E3" class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka moluensis" order="Stylommatophora" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="moluensis">Philalanka moluensis</taxonomicName>
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(E.A. Smith, 1893). 44A Frontal view, shell 2.3 mm high 44B Umbilical view (Malaysia, Sabah, Interior Province, Sepulut valley, Gua Pungiton, V 7558) 45
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<taxonomicName id="4F5E4FE026838777173B45476D98F57A" class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka obscura" order="Stylommatophora" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="obscura">Philalanka obscura</taxonomicName>
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sp. n. 45A Frontal view, shell 2.1 mm high 45B Umbilical view (Indonesia, Kalimantan Timur, Sangkulirang Peninsula, Meweding Cave near village Tabalar Ulu (RMNH.5003939, holotype) 46
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<taxonomicName id="A4C08021A90E7A0DA7AFF0C9DCDBACF9" class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka thienemanni" order="Stylommatophora" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="thienemanni">Philalanka thienemanni</taxonomicName>
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B. Rensch, 1932 46A Frontal view, shell 2.7 mm high 46B Umbilical view (Indonesia, Java, Bremi, V 4019).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="907B9D495B1B8E420911DA07C31A2933" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
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<paragraph id="22D01BA789F30258D1BB5B78762C3DF2" pageId="54" pageNumber="55">Description.</paragraph>
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<paragraph id="9DEAF7307E6B378C959230C530703AA8" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
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Shell very small, rather thin, more or less opaque, brown, very rarely yellowish, conical with approx. flat or slightly convex sides; apex rounded. Surface about dull or with a silky luster above the periphery, glossy below. Whorls convex, approx. rounded. Protoconch whorls convex; sculpture: 9-12 inconspicuous to distinct, very fine, thin spiral threads; radial sculpture absent or some subordinate wrinkles. Teleoconch: Last whorl with a distinct spiral thread coinciding with the suture of the penultimate whorl, above this 2-4 slightly less distinct threads with a very fine (just visible at 40 times magnification) spiral striation in between, particularly towards the periphery; below the periphery up to 4 fine spiral threads close to it, with very fine
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<pageBreakToken id="598EBEE1E40AFA50223D49C76A7A46FA" pageId="55" pageNumber="56" start="start">spiral</pageBreakToken>
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striation in between. Radial sculpture: densely placed, very fine (just visible at 40 times magnification), narrow riblets that may be inconspicuous locally. Umbilicus open, narrow. Dimensions: Height up to 2.9 mm; width up to 2.8 mm; diameters of the first four whorls 0.55-0.65 mm, 0.9-1.1 mm, 1.35-1.55 mm, 1.9-2.1 mm respectively; number of whorls up to 5 7/8; height aperture up to 1.2 mm; width aperture up to 1.6 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9B791CCCDE83784BFA0C5ACBFDD1731" pageId="55" pageNumber="56" type="habitat in sabah and distribution">
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<paragraph id="4A2F406C88E8F0009F247F8EA6CD75AC" pageId="55" pageNumber="56">Habitat in Sabah and distribution.</paragraph>
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<paragraph id="2ECAA3C2C9FC6B39F23637638E2C59EC" pageId="55" pageNumber="56">Primary and secondary forest on limestone and sandstone soil, up to 1100 m alt. Sabah: West part; scattered localities along East coast. Also in Sarawak. Endemic to Borneo.</paragraph>
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</subSubSection>
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<subSubSection id="AD6647591D4895F4DCFA1444E7181A39" pageId="55" pageNumber="56" type="cross diagnosis">
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<paragraph id="7694C307BB4EBA12BA317CFFDEE584D1" pageId="55" pageNumber="56">Cross diagnosis.</paragraph>
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<paragraph id="11F8619728F9FC6C65548402CE87EDB0" pageId="55" pageNumber="56">
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Resembles
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<taxonomicName id="535DF13A47AE4DC57FE0639EA47C7DEF" class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella microconus" order="Stylommatophora" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="microconus">Kaliella microconus</taxonomicName>
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(Mousson), which also occurs in Sabah, but it has much coarser and less regularly spaced spiral threads, and more convex whorls.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |