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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZBK="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo &amp; Balke, Michael" docDate="2018" docId="0FA25962B8B22EAF53ED8A1E28528DA9" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina menyamya Shaverdo &amp; Balke, sp. n." docType="treatment" docUuid="B6435BF0-3067-44BD-946E-8772C350CAD1" docUuidSource="ZooBank" docVersion="7" lastPageNumber="27" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="27" updateTime="1668166558239" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sagata, Katayo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>803</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>7</mods:start>
<mods:end>70</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
<mods:identifier type="ZBK">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
<mods:identifier type="ZooBank">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154125611" LSID="urn:lsid:zoobank.org:act:B6435BF0-3067-44BD-946E-8772C350CAD1" httpUri="http://treatment.plazi.org/id/0FA25962B8B22EAF53ED8A1E28528DA9" lastPageNumber="27" pageId="20" pageNumber="27">
<subSubSection pageId="20" pageNumber="27" type="nomenclature">
<paragraph pageId="20" pageNumber="27">
13.
<taxonomicName LSID="http://zoobank.org/B6435BF0-3067-44BD-946E-8772C350CAD1" authority="Shaverdo &amp; Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina menyamya" order="Coleoptera" pageId="20" pageNumber="27" phylum="Arthropoda" rank="species" species="menyamya">Exocelina menyamya Shaverdo &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="20" pageNumber="27">sp. n.</taxonomicNameLabel>
Figs 20, 44
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="reference_group">
<paragraph pageId="20" pageNumber="27">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="20" pageNumber="27" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
undescribed sp. MB1377:
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
: supplementary figs 1-4, table 2;
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
: supplementary figs S1-S2, table S3.
</paragraph>
<paragraph pageId="20" pageNumber="27">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina menyamya" order="Coleoptera" pageId="20" pageNumber="27" phylum="Arthropoda" rank="species" species="menyamya">Exocelina menyamya</taxonomicName>
_New_Guinea_MB1377:
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
: supplementary information S5-S6.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="type locality">
<paragraph pageId="20" pageNumber="27">Type locality.</paragraph>
<paragraph pageId="20" pageNumber="27">
Papua New Guinea: Morobe Province, Menyamya, Mount Inji,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-7.2376666">07°14.26'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="146.02333">146°01.40'E</geoCoordinate>
, 1500 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="type material">
<paragraph pageId="20" pageNumber="27">Type material.</paragraph>
<paragraph pageId="20" pageNumber="27">Holotype: male &quot;Papua New Guinea: Morobe, Menyamya, Mt Inji, deep well, 1500m, 14.xi.2006, 07.14.264S 146.01.400E, Balke &amp; Kinibel (PNG 98)&quot;, &quot;M.Balke 1377&quot; [green] (ZSM).</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="description">
<paragraph pageId="20" pageNumber="27">Description.</paragraph>
<paragraph pageId="20" pageNumber="27">Body size and form: Beetle medium-sized: TL-H 4.25 mm, TL 4.8 mm, MW 2.2 mm, with oblong-oval habitus.</paragraph>
<paragraph pageId="20" pageNumber="27">Coloration: Brown, with reddish pronotum. Head brown, with slightly darker areas posterior to eyes. Pronotum broadly reddish laterally and dark brown medially from anterior to posterior margins. Elytra uniformly brown, in the middle with traces of narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish brown, especially metathoracic legs (Fig. 20).</paragraph>
<paragraph pageId="20" pageNumber="27">Surface sculpture: Matt dorsally. Head with rather dense, coarse punctation (spaces between punctures 1-2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum and elytra with distinct punctation, sparser and finer than on head. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation.</paragraph>
<paragraph pageId="20" pageNumber="27">Structures: Pronotum with distinct lateral bead. Its lateral sides with longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively short, broad, slightly convex and smooth in the middle, with distinct lateral bead and few lateral setae, lateral sides slightly flattened. Abdominal ventrite 6 slightly truncate.</paragraph>
<paragraph pageId="20" pageNumber="27">
Male: Antennae simple (Fig. 20). Protarsomere 4 with large, thick, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of more than 50 and posterior row of 13 relatively long, thin setae (Fig. 44D). Median lobe in lateral view narrowed to apex, with thin, slightly curved upwards apex; in ventral view, narrowed before apex, with apex rounded, of shape of a baby pacifier, with distinct ventral impression. Paramere slightly concave on dorsal side and with long, dense, thin setae situated along dorsal margin, subdistal setae denser, proximal setae much sparser, setae in middle shorter, thinner (Fig. 44
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Abdominal ventrite 6 with 19-22 lateral striae on each side.
</paragraph>
<paragraph pageId="20" pageNumber="27">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="affinities">
<paragraph pageId="20" pageNumber="27">Affinities.</paragraph>
<paragraph pageId="20" pageNumber="27">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina menyamya" order="Coleoptera" pageId="20" pageNumber="27" phylum="Arthropoda" rank="species" species="menyamya">Exocelina menyamya</taxonomicName>
sp. n. is similar to
<taxonomicName lsidName="E. casuarina" pageId="20" pageNumber="27" rank="species" species="casuarina">E. casuarina</taxonomicName>
and
<taxonomicName lsidName="E. fume" pageId="20" pageNumber="27" rank="species" species="fume">E. fume</taxonomicName>
in body size, shape, and coloration but can be distinguished from them by the completely different shape of the median lobe and presence of the pronotal bead.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="distribution">
<paragraph pageId="20" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="20" pageNumber="27">Papua New Guinea: Morobe Province. The species is known only from the type locality (Fig. 50).</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="27" type="etymology">
<paragraph pageId="20" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="20" pageNumber="27">The species is named after Menyamya Village. The name is a noun in the nominative singular standing in apposition.</paragraph>
</subSubSection>
</treatment>
</document>