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<mods:title id="1DBC177DC48B05A821C0E42FC3CE4409">Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)</mods:title>
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<emphasis id="B951750CEF63FFE95983FC1C4F1E8EFC" bold="true" box="[144,359,974,999]" italics="true" pageId="63" pageNumber="64">Lindholmiola gyria</emphasis>
(
<bibRefCitation id="EFB4D4EFEF63FFE95865FC1C4F808EFD" author="Roth, J. R." box="[374,505,974,998]" pageId="63" pageNumber="64" refId="ref68821" refString="Roth, J. R. (1839): Molluscorum species, quas in itinere per Orientem facto comites clariss. Schubert doctores M. Erdl et J. R. Roth collegerunt. Dissertatio inauguralis. - 3 - 26, Taf. 1 - 2, Munchen." type="book" year="1839">ROTH, 1839</bibRefCitation>
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<paragraph id="8B9AA91EEF63FFE95B1BFC1C4CAE8EFD" blockId="63.[144,727,974,999]" box="[520,727,974,998]" pageId="63" pageNumber="64">
<heading id="D0D21E72EF63FFE95B1BFC1C4CAE8EFD" box="[520,727,974,998]" centered="true" fontSize="36" level="2" pageId="63" pageNumber="64" reason="4">
<figureCitation id="131EB59BEF63FFE95B1BFC1C4CFA8EFD" box="[520,643,974,998]" captionStart="Figs 33–34" captionStartId="64.[181,218,1000,1018]" captionTargetBox="[181,1065,113,970]" captionTargetId="figure-144@64.[181,1065,113,970]" captionTargetPageId="64" captionText="Figs 33–34. Lindholmiola gyria. Fig. 33: syntype Helix gyria, ZSM 1781, Turkey, &quot;Cacamo Cariae&quot;, D = 14.9 mm; Fig. 34: NMBE 515613, Turkey, Vilayet Antalya, Phaselis, ruins close to Tekirova, D = 13.68 mm. — All photos Neubert/Bochud, × 3." figureDoi="http://doi.org/10.5281/zenodo.5842368" httpUri="https://zenodo.org/record/5842368/files/figure.png" pageId="63" pageNumber="64">Figs 33–34</figureCitation>
, 
<figureCitation id="131EB59BEF63FFE95B82FC1C4CD48EFD" box="[657,685,974,998]" captionStart="Fig" captionStartId="66.[181,210,680,698]" captionTargetBox="[181,1065,113,643]" captionTargetId="figure-256@66.[181,1065,113,643]" captionTargetPageId="66" captionText="Fig. 35. Distribution of Lindholmiola gyria." figureDoi="http://doi.org/10.5281/zenodo.5842370" httpUri="https://zenodo.org/record/5842370/files/figure.png" pageId="63" pageNumber="64">35</figureCitation>
, 
<figureCitation id="131EB59BEF63FFE95BAAFC1C4CAE8EFD" box="[697,727,974,998]" captionStart="Figs 44–46" captionStartId="76.[181,218,1459,1477]" captionTargetBox="[181,1065,113,1400]" captionTargetId="figure-8@76.[181,1065,113,1400]" captionTargetPageId="76" captionText="Figs 44–46. Genital organs of Lindholmiola species. Fig. 44: Lindholmiola gyria, a: situs, b: lumina of genital organs, Turkey, Vilayet Antalya, Phaselis; Fig. 45: Lindholmiola barbata, a: situs, b: lumina of genital organs, Greece, Island Kríti, Nomos Haniá, Soúda Bay; Fig. 46: Lindholmiola regisborisi, a: situs, b: lumina of genital organs, Greece, Makedonía, 400 m S of Kehrókambos." figureDoi="http://doi.org/10.5281/zenodo.5842380" httpUri="https://zenodo.org/record/5842380/files/figure.png" pageId="63" pageNumber="64">44</figureCitation>
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1839 
<taxonomicName id="4C25D29DEF63FFE959DFFBCB4FFD892A" authority="ROTH, Molluscorum" authorityName="ROTH, Molluscorum" box="[204,388,1049,1073]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DFFBCB4F3A892A" box="[204,323,1049,1073]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
ROTH
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, Molluscorum species, quas in itinere per Orientem facto comites clariss. Schubert doctores M. Erdl et J. R. Roth collegerunt: 16, Taf. 1 
<figureCitation id="131EB59BEF63FFE959A5FBB64F5E8967" box="[182,295,1124,1148]" captionStart="Figs 16–18" captionStartId="34.[181,218,1016,1034]" captionTargetBox="[181,1065,102,993]" captionTargetId="figure-143@34.[181,1065,102,995]" captionTargetPageId="34" captionText="Figs 16–18. Lindholmiola barbata. Fig. 16: NMBE 515636, Island Kríti, Nomos Iráklio, valley E from the airport Iráklio (small rounded form), D = 9.66 mm; Fig. 17: NMBE 515635, Island Kríti, Nomos Haniá, canyon ca. 4 Km N of Thériso (large flat form), D = 13.78 mm; Fig. 18: NMBE 515581, Rethimnon, Zousidi, 09.04.1993, leg. N. Chalwatzis, D = 12.49 mm. — All photos Neubert/Bochud, × 3." figureDoi="http://doi.org/10.5281/zenodo.5842354" httpUri="https://zenodo.org/record/5842354/files/figure.png" pageId="63" pageNumber="64">Fig. 17–18</figureCitation>
.
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<paragraph id="8B9AA91EEF63FFE9599DFB5B4CEE89DD" blockId="63.[142,1033,1049,1596]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFB5B4CEE89DD" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1846 
<taxonomicName id="4C25D29DEF63FFE959DFFB5B4F3989BA" authorityName="ROTH" authorityYear="1839" box="[204,320,1161,1185]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DFFB5B4F3989BA" box="[204,320,1161,1185]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
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, – L. Pfeiffer, in: Martini &amp; Chemnitz: Systematisches Conchylien-Cabinet, 1. 12 (II): 283, Taf. 124 
<figureCitation id="131EB59BEF63FFE95B0EFB7C4CE989DD" box="[541,656,1198,1222]" captionStart-0="Figs 30–32" captionStart-1="Figs 33–34" captionStartId-0="58.[181,218,829,847]" captionStartId-1="64.[181,218,1000,1018]" captionTargetBox-0="[181,1065,113,806]" captionTargetBox-1="[181,1065,113,970]" captionTargetId-0="figure-227@58.[181,1065,113,808]" captionTargetId-1="figure-144@64.[181,1065,113,970]" captionTargetPageId-0="58" captionTargetPageId-1="64" captionText-0="Figs 30–32. Lindholmiola girva. Fig. 30: syntype Helix girva SMF 7047, &quot;Albanien&quot;, D = 9.22 mm; Fig. 31: NMBE 515656, Greece, Makedonía, N boundary of Taxiárhis (small rounded form), D = 8.66 mm; Fig. 32: NMBE 515655, Greece, Makedonía, 400 m S of Kehrókambos (broad form, wide umbilicus), D = 9.75 mm. — All photos Neubert/Bochud, × 3." captionText-1="Figs 33–34. Lindholmiola gyria. Fig. 33: syntype Helix gyria, ZSM 1781, Turkey, &quot;Cacamo Cariae&quot;, D = 14.9 mm; Fig. 34: NMBE 515613, Turkey, Vilayet Antalya, Phaselis, ruins close to Tekirova, D = 13.68 mm. — All photos Neubert/Bochud, × 3." figureDoi-0="http://doi.org/10.5281/zenodo.5842366" figureDoi-1="http://doi.org/10.5281/zenodo.5842368" httpUri-0="https://zenodo.org/record/5842366/files/figure.png" httpUri-1="https://zenodo.org/record/5842368/files/figure.png" pageId="63" pageNumber="64">Fig. 31–33</figureCitation>
.
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</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFB064DB289F7" blockId="63.[142,1033,1049,1596]" box="[142,971,1236,1260]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFB064DB289F7" box="[142,971,1236,1260]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1848 
<taxonomicName id="4C25D29DEF63FFE959DDFB064F3F89F7" authorityName="ROTH" authorityYear="1839" box="[206,326,1236,1260]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DDFB064F3F89F7" box="[206,326,1236,1260]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – L. Pfeiffer, Monographia heliceorum viventium, 1: 414.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFB2B4DB0880A" blockId="63.[142,1033,1049,1596]" box="[142,969,1273,1297]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFB2B4DB0880A" box="[142,969,1273,1297]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1853 
<taxonomicName id="4C25D29DEF63FFE959D8FB2B4F3D880A" authorityName="ROTH" authorityYear="1839" box="[203,324,1273,1297]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959D8FB2B4F3D880A" box="[203,324,1273,1297]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – L. Pfeiffer, Monographia heliceorum viventium, 3: 261.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFACC4DB5882D" blockId="63.[142,1033,1049,1596]" box="[142,972,1310,1334]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFACC4DB5882D" box="[142,972,1310,1334]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1868 
<taxonomicName id="4C25D29DEF63FFE959DEFACC4F3F882D" authorityName="ROTH" authorityYear="1839" box="[205,326,1310,1334]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DEFACC4F3F882D" box="[205,326,1310,1334]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – L. Pfeiffer, Monographia heliceorum viventium, 4: 312.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFA964FC4889A" blockId="63.[142,1033,1049,1596]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFA964FC4889A" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1859 
<taxonomicName id="4C25D29DEF63FFE959DEFA964F318847" authorityName="ROTH" authorityYear="1839" box="[205,328,1348,1372]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DEFA964F318847" box="[205,328,1348,1372]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – Mousson, Vierteljahrsschrift der Naturforschenden Gesellschaft in 
<collectingRegion id="49E167FCEF63FFE9580EFABB4F11889A" box="[285,360,1385,1409]" country="Switzerland" name="Zuerich" pageId="63" pageNumber="64">Zürich</collectingRegion>
, 4: 259.
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</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFA5C4DB288BD" blockId="63.[142,1033,1049,1596]" box="[142,971,1422,1446]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFA5C4DB288BD" box="[142,971,1422,1446]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1868 
<taxonomicName id="4C25D29DEF63FFE959DEFA5C4F3F88BD" authorityName="ROTH" authorityYear="1839" box="[205,326,1422,1446]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DEFA5C4F3F88BD" box="[205,326,1422,1446]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – L. Pfeiffer, Monographia heliceorum viventium, 5: 415.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFA664DB388D7" blockId="63.[142,1033,1049,1596]" box="[142,970,1460,1484]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFA664DB388D7" box="[142,970,1460,1484]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1876 
<taxonomicName id="4C25D29DEF63FFE959D8FA664F3D88D7" authorityName="ROTH" authorityYear="1839" box="[203,324,1460,1484]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959D8FA664F3D88D7" box="[203,324,1460,1484]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – L. Pfeiffer, Monographia heliceorum viventium, 7: 466.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFA0B4D7C88EA" blockId="63.[142,1033,1049,1596]" box="[142,773,1497,1521]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFA0B4D7C88EA" box="[142,773,1497,1521]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1887 
<taxonomicName id="4C25D29DEF63FFE959DEFA0B4F3C88EA" authorityName="ROTH" authorityYear="1839" box="[205,325,1497,1521]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF63FFE959DEFA0B4F3C88EA" box="[205,325,1497,1521]" italics="true" pageId="63" pageNumber="64">Helix gyria</emphasis>
</taxonomicName>
, – Tryon, Manual of Conchology, 3: 177.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DFA2C4D6E8B0D" blockId="63.[142,1033,1049,1596]" box="[142,791,1534,1558]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DFA2C4D6E8B0D" box="[142,791,1534,1558]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1889 
<taxonomicName id="4C25D29DEF63FFE959DEFA2C4FAF8B0D" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[205,470,1534,1558]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria" subGenus="Caracolina">
<emphasis id="B951750CEF63FFE959DEFA2C4F7F8B0D" box="[205,262,1534,1558]" italics="true" pageId="63" pageNumber="64">Helix</emphasis>
(
<emphasis id="B951750CEF63FFE95806FA2C4FF58B0D" box="[277,396,1534,1558]" italics="true" pageId="63" pageNumber="64">Caracolina</emphasis>
) 
<emphasis id="B951750CEF63FFE9588FFA2C4FAF8B0D" box="[412,470,1534,1558]" italics="true" pageId="63" pageNumber="64">gyria</emphasis>
</taxonomicName>
, – Westerlund, Fauna, 2: 20.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF63FFE9599DF9F64D868B27" blockId="63.[142,1033,1049,1596]" box="[142,1023,1572,1596]" pageId="63" pageNumber="64">
<treatmentCitation id="0A848F0FEF63FFE9599DF9F64D868B27" box="[142,1023,1572,1596]" lastPageId="63" lastPageNumber="65" pageId="63" pageNumber="64">
1894 
<taxonomicName id="4C25D29DEF63FFE959D8F9F64FAC8B27" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[203,469,1572,1596]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="gyria" subGenus="Caracollina">
<emphasis id="B951750CEF63FFE959D8F9F64F7A8B27" box="[203,259,1572,1596]" italics="true" pageId="63" pageNumber="64">Helix</emphasis>
(
<emphasis id="B951750CEF63FFE95802F9F64FF48B27" box="[273,397,1572,1596]" italics="true" pageId="63" pageNumber="64">Caracollina</emphasis>
) 
<emphasis id="B951750CEF63FFE9588FF9F64FAC8B27" box="[412,469,1572,1596]" italics="true" pageId="63" pageNumber="64">gyria</emphasis>
</taxonomicName>
, – Pilsbry, in Tryon, Manual of Conchology, 9: 228.
</treatmentCitation>
</paragraph>
<caption id="DF5AF996EF1CFF9659A6FC3A4C31892B" ID-DOI="http://doi.org/10.5281/zenodo.5842368" ID-Zenodo-Dep="5842368" httpUri="https://zenodo.org/record/5842368/files/figure.png" pageId="64" pageNumber="65" startId="64.[181,218,1000,1018]" targetBox="[181,1065,113,970]" targetPageId="64">
<paragraph id="8B9AA91EEF1CFF9659A6FC3A4C31892B" blockId="64.[181,1067,1000,1072]" pageId="64" pageNumber="65">
Figs 33–34. 
<emphasis id="B951750CEF1CFF965837FC3A4FAA8EE1" bold="true" box="[292,467,1000,1018]" italics="true" pageId="64" pageNumber="65">
<taxonomicName id="4C25D29DEF1CFF965837FC3A4FB68EE1" ID-CoL="72FTT" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[292,463,1000,1018]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="65" phylum="Mollusca" rank="species" species="gyria">Lindholmiola gyria</taxonomicName>
.
</emphasis>
Fig. 33: syntype 
<taxonomicName id="4C25D29DEF1CFF965B62FC3A4D408EE1" authority=", ZSM 1781" authorityName=", ZSM" authorityYear="1781" box="[625,825,1000,1018]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="65" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF1CFF965B62FC3A4CAD8EE1" bold="true" box="[625,724,1000,1018]" italics="true" pageId="64" pageNumber="65">Helix gyria</emphasis>
, ZSM 1781
</taxonomicName>
, Turkey, &quot;Cacamo Cariae&quot;, D = 14.9 mm; Fig. 34: NMBE 515613, Turkey, Vilayet Antalya, Phaselis, ruins close to Tekirova, D = 13.68 mm. — All photos Neubert/Bochud, × 3.
</paragraph>
</caption>
<paragraph id="8B9AA91EEF1CFF9659A7FBB64F6F89F7" blockId="64.[180,1071,1124,1372]" pageId="64" pageNumber="65">
<treatmentCitation id="0A848F0FEF1CFF9659A7FBB64F6F89F7" lastPageId="64" lastPageNumber="65" pageId="64" pageNumber="64">
1898 
<taxonomicName id="4C25D29DEF1CFF9659E5FBB64C688967" authorityName=", ZSM" authorityYear="1781" box="[246,529,1124,1148]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="65" phylum="Mollusca" rank="species" species="gyria" subGenus="Gonostoma">
<emphasis id="B951750CEF1CFF9659E5FBB64F568967" box="[246,303,1124,1148]" italics="true" pageId="64" pageNumber="65">Helix</emphasis>
(
<emphasis id="B951750CEF1CFF965852FBB64FBD8967" box="[321,452,1124,1148]" italics="true" pageId="64" pageNumber="65">Gonostoma</emphasis>
) 
<emphasis id="B951750CEF1CFF9658C4FBB64C688967" box="[471,529,1124,1148]" italics="true" pageId="64" pageNumber="65">gyria</emphasis>
</taxonomicName>
, – Kobelt, in Rossmässler, E. A.: Iconographie der Land- &amp; Süsswasser-Mollusken mit vorzüglicher Berücksichtigung der europäischen noch nicht abgebildeten Arten, (2) 8 (1/4): 53, Taf. 226 Fig. 1447.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF1CFF9659A7FB2B4C49882D" blockId="64.[180,1071,1124,1372]" pageId="64" pageNumber="65">
<treatmentCitation id="0A848F0FEF1CFF9659A7FB2B4C49882D" lastPageId="64" lastPageNumber="65" pageId="64" pageNumber="64">
1918 
<taxonomicName id="4C25D29DEF1CFF9659E6FB2B4FCF880A" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[245,438,1273,1297]" class="Gastropoda" family="Trissexodontidae" genus="Caracollina" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="65" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF1CFF9659E6FB2B4FCF880A" box="[245,438,1273,1297]" italics="true" pageId="64" pageNumber="65">Caracollina gyria</emphasis>
</taxonomicName>
, – Hesse, Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 50: 109.
</treatmentCitation>
</paragraph>
<paragraph id="8B9AA91EEF1CFF9659A7FA964DD38847" blockId="64.[180,1071,1124,1372]" box="[180,938,1348,1372]" pageId="64" pageNumber="65">
<treatmentCitation id="0A848F0FEF1CFF9659A7FA964DD38847" box="[180,938,1348,1372]" lastPageId="64" lastPageNumber="65" pageId="64" pageNumber="64">
1991 
<taxonomicName id="4C25D29DEF1CFF9659E0FA964F818847" ID-CoL="7RJ6L" authority="gyria" box="[243,504,1348,1372]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="gyria" subSpecies="gyria">
<emphasis id="B951750CEF1CFF9659E0FA964F818847" box="[243,504,1348,1372]" italics="true" pageId="64" pageNumber="65">Helicodonta gyria gyria</emphasis>
</taxonomicName>
, – Maassen, Basteria 55: 123, 
<figureCitation id="131EB59BEF1CFF965A5EFA964DDA8847" box="[845,931,1348,1372]" captionStart-0="Figs" captionStart-1="Figs 3–6" captionStartId-0="3.[144,183,1240,1258]" captionStartId-1="10.[181,218,1390,1408]" captionTargetBox-0="[156,1028,117,1185]" captionTargetBox-1="[181,1065,113,1346]" captionTargetId-0="figure-6@3.[144,1028,113,1185]" captionTargetId-1="figure-8@10.[181,1065,113,1346]" captionTargetPageId-0="3" captionTargetPageId-1="10" captionText-0="Figs. 1–2. Genital organs of Helicodontidae. Fig. 1: Lindholmiola barbata; Greece, Kríti, Nomos Haniá, Bay of Soúda, 100–150 m from the mouth of the river S of the English cemetery. Fig. 2: Helicodonta obvoluta (O. F. MüLLER, 1774); Belgium, valley of the Meuse, 500 m from Annevoie in direction to Dinant, 2a) situs of genital organs, 2b) interior structure of penis. — Abbreviations used for the description of the genital organs: A = atrium, Al = albumen gland, E = Epiphallus, G = distal glandular part of epiphallus, Gl = Glandula, Hd = hermaphroditic duct, Hg = hermaphroditic gland, P = Penis, R = retractor muscle, Sp = Spermoviduct, V = vesicle, Vag = Vagina, Vd = Vas deferens." captionText-1="Figs 3–6. Lindholmiola lens. Fig. 3: syntype Helix lentiformis, ZMZ 508656, &quot;Thessalie et Attique&quot;, D = 11.2 mm; Fig. 4: NMBE 515542, Attika, Methana, 500 m before village, 12.07.1996, leg. E. Neubert, D = 12.77 mm; Fig. 5: syntype Helix lens var. callojuncta, NHMG 372, D = 12.0 mm; Fig. 6: syntype Helix lens var. elia, SMF 101961, Kumani, Elis, coll. Jetschin ex Boettger, 1883, D = 11.94 mm. — All photos Neubert/Bochud, × 3." figureDoi-0="http://doi.org/10.5281/zenodo.5842344" figureDoi-1="http://doi.org/10.5281/zenodo.5842346" httpUri-0="https://zenodo.org/record/5842344/files/figure.png" httpUri-1="https://zenodo.org/record/5842346/files/figure.png" pageId="64" pageNumber="65">Fig. 1–3</figureCitation>
.
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33FFA95EF1CFF9659C8FA5C4D8A88D7" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="8B9AA91EEF1CFF9659C8FA5C4D8A88D7" blockId="64.[182,1071,1422,1596]" pageId="64" pageNumber="65">Diagnosis: shell strongly depressed, protoconch and first teleoconch whorls slightly immersed, whorls densely coiled, extremely long bursa copulatrix.</paragraph>
</subSubSection>
<subSubSection id="C33FFA95EF1CFF9759C8FA0B4C25894D" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
<paragraph id="8B9AA91EEF1CFF9759C8FA0B4D698CC7" blockId="64.[182,1071,1422,1596]" lastBlockId="65.[142,1033,116,1260]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
Description of shell: shell strongly depressed, protoconch and first teleoconch whorls slightly immersed, basic shell colour brown; initial part of protoconch whorls smooth, followed by 1–1¼ whorls with a pattern of a few fine radial stripes; the following teleoconch whorls with increasingly stronger radial stripes, but almost lacking on the last whorls; surface of the last whorl covered by a fine granulation; sometimes with hairs, hair warts can be found in the granulated surface sculpture; 6¼–7¼, densely coiled and well rounded whorls, last whorl slightly descending before the aperture; suture deep; umbilicus broad, almost cylindrical, reaching a diameter of 
<quantity id="4CDD04FBEF1DFF975A24FEFC4DBD8C5D" box="[823,964,302,326]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="65" pageNumber="66" unit="mm" value="3.05" valueMax="3.3" valueMin="2.8">2.8–3.3 mm</quantity>
; aperture oblique, slightly bent in lateral view, and strongly ear-like in frontal view; apertural insertion with a distance of 
<quantity id="4CDD04FBEF1DFF975B5AFEAB4CAF8C8A" box="[585,726,377,401]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.3" metricValueMin="4.2" pageId="65" pageNumber="66" unit="mm" value="4.75" valueMax="5.3" valueMin="4.2">4.2–5.3 mm</quantity>
, connected by a very thin, almost indiscernible callus; peristomial rim ± sharp, laterally with a strong swelling, basally reinforced with a small umbilical shield.
</paragraph>
<paragraph id="8B9AA91EEF1DFF9759A5FE3B4D858F1A" blockId="65.[142,1033,116,1260]" box="[182,1020,489,513]" pageId="65" pageNumber="66">Measurements: H: 5.3–6.4; D: 12.3–15; aH: 4.4–5.5 (5.6–6.6); aW: 5.8–6.9.</paragraph>
<paragraph id="8B9AA91EEF1DFF9759A5FDDC4FCC8F8D" blockId="65.[142,1033,116,1260]" pageId="65" pageNumber="66">
Details of body (after 
<specimenCount id="9D236297EF1DFF9758BEFDDC4C338F3D" box="[429,586,526,550]" pageId="65" pageNumber="66" type="generic">one specimen</specimenCount>
from Phaselis): head and dorsum dark grey, flanks and tail light grey; sole cream with a greyish rim; mantle collar translucent with grey spots of pigment; secondary ureter opens 
<quantity id="4CDD04FBEF1DFF975A7FFD8B4DB98F6A" box="[876,960,601,625]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="66" unit="mm" value="1.5">1.5 mm</quantity>
apart from the respiratory pore.
</paragraph>
<paragraph id="8B9AA91EEF1DFF9759A5FD764C25894D" blockId="65.[142,1033,116,1260]" pageId="65" pageNumber="66">
Morphology of the genital organs (
<figureCitation id="131EB59BEF1DFF975B58FD764CE48FA7" box="[587,669,676,700]" captionStart="Figs 44–46" captionStartId="76.[181,218,1459,1477]" captionTargetBox="[181,1065,113,1400]" captionTargetId="figure-8@76.[181,1065,113,1400]" captionTargetPageId="76" captionText="Figs 44–46. Genital organs of Lindholmiola species. Fig. 44: Lindholmiola gyria, a: situs, b: lumina of genital organs, Turkey, Vilayet Antalya, Phaselis; Fig. 45: Lindholmiola barbata, a: situs, b: lumina of genital organs, Greece, Island Kríti, Nomos Haniá, Soúda Bay; Fig. 46: Lindholmiola regisborisi, a: situs, b: lumina of genital organs, Greece, Makedonía, 400 m S of Kehrókambos." figureDoi="http://doi.org/10.5281/zenodo.5842380" httpUri="https://zenodo.org/record/5842380/files/figure.png" pageId="65" pageNumber="66">Fig. 44</figureCitation>
): penis relatively long, slightly enlarged in its central part, penial glandular tissue covering ca 1/4 of the total penis length, penial lumen filled by a single, broad and serrated pilaster, starting at the distal penial wall and stretching to the narrow penial pore, where it recurves and ends centrally in the penis, remaining penial lumen with short perpendicular folds; vagina relatively long, filled with a long, narrow pilaster connecting to one of the atrial pilasters, glandula with a short stalk and a strongly folded part, bursa copulatrix three times the length of the glandula, bursa copulatrix narrow, same length as glandula, pedunculus reaching ca 40% of the total length of the bursa copulatrix, vesicle of bursa copulatrix elongate, double the diameter of the pedunculus; atrial lumen with several broad and short interdigitating pilasters.
</paragraph>
</subSubSection>
<subSubSection id="C33FFA95EF1DFF9759A5FBB64DA089F7" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="8B9AA91EEF1DFF9759A5FBB64DA089F7" blockId="65.[142,1033,116,1260]" pageId="65" pageNumber="66">
Differential diagnosis: The shell of 
<taxonomicName id="4C25D29DEF1DFF975B43FBB64CDE8967" authorityName=": ZSM" authorityYear="1781" box="[592,679,1124,1148]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF1DFF975B43FBB64CDE8967" box="[592,679,1124,1148]" italics="true" pageId="65" pageNumber="66">L. gyria</emphasis>
</taxonomicName>
with its broad last whorl and the slightly concave spire resembles that of 
<taxonomicName id="4C25D29DEF1DFF975B85FB5B4DF489BA" baseAuthorityName="O. F. MuLLER" baseAuthorityYear="1774" box="[662,909,1161,1185]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="species" species="obvoluta">
<emphasis id="B951750CEF1DFF975B85FB5B4DF489BA" box="[662,909,1161,1185]" italics="true" pageId="65" pageNumber="66">Helicodonta obvoluta</emphasis>
</taxonomicName>
, and thus cannot be confused with any of the other species of the genus 
<taxonomicName id="4C25D29DEF1DFF975A7FFB7C4A7889DD" authorityName="HESSE, Zoologica" authorityYear="1931" box="[876,1025,1198,1222]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="genus">
<emphasis id="B951750CEF1DFF975A7FFB7C4A7889DD" box="[876,1025,1198,1222]" italics="true" pageId="65" pageNumber="66">Lindholmiola</emphasis>
</taxonomicName>
. The extremely long bursa copulatrix differentiates it from all its congeners.
</paragraph>
</subSubSection>
<subSubSection id="C33FFA95EF1DFF9459A6FAC94DC68E76" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8B9AA91EEF1DFF9759A6FAC94F8B8849" blockId="65.[143,1027,1307,1362]" pageId="65" pageNumber="66">
<emphasis id="B951750CEF1DFF9759A6FAC94F1B882A" bold="true" box="[181,354,1307,1329]" pageId="65" pageNumber="66">
<typeStatus id="549E17BCEF1DFF9759A6FAC94E9F882A" box="[181,230,1307,1329]" pageId="65" pageNumber="66">Type</typeStatus>
specimens:
</emphasis>
<materialsCitation id="3B4DA343EF1DFF97587BFAC94F8B8849" ID-GBIF-Occurrence="3435946268" collectionCode="ZSM" country="Turkey" county="Antalya County" latitude="36.17" location="Kekova Island" longLatPrecision="713" longitude="29.87" pageId="65" pageNumber="66" specimenCode="ZSM 1781" specimenCount="1" typeStatus="syntype">
<collectingCountry id="F332E98EEF1DFF97587BFAC94FD7882A" box="[360,430,1307,1329]" name="Turkey" pageId="65" pageNumber="66">Turkey</collectingCountry>
, 
<collectingCounty id="62FBD192EF1DFF9758A4FAC94C2C882A" box="[439,597,1307,1329]" pageId="65" pageNumber="66">Antalya County</collectingCounty>
, 
<location id="8EFAFFC5EF1DFF975B4DFAC94C97882A" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975B4DFAC94C97882A" box="[606,750,1307,1329]" country="Turkey" county="Antalya County" latitude="36.17" longLatPrecision="713" longitude="29.87" municipality="County" name="Kekova Island" pageId="65" pageNumber="66" stateProvince="Antalya">Kekova Island</location>
, &quot; 
<location id="8EFAFFC5EF1DFF975BEDFAC94DD8882A" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975BEDFAC94DD8882A" box="[766,929,1307,1329]" country="Turkey" county="Antalya County" latitude="36.17" longLatPrecision="713" longitude="29.87" name="Cacamo Cariae" pageId="65" pageNumber="66">Cacamo Cariae</location>
&quot;, 
<geoCoordinate id="EE11CFD9EF1DFF975AA2FAC94A7A882A" box="[945,1027,1307,1329]" degrees="36. 17" direction="north" orientation="latitude" pageId="65" pageNumber="66" precision="555" value="36.17">36.17°N</geoCoordinate>
<geoCoordinate id="EE11CFD9EF1DFF97599CFAEE4E9A8849" box="[143,227,1340,1362]" degrees="29. 87" direction="east" orientation="longitude" pageId="65" pageNumber="66" precision="555" value="29.87">29.87°E</geoCoordinate>
, 
<typeStatus id="549E17BCEF1DFF9759FFFAEE4F478849" box="[236,318,1340,1362]" pageId="65" pageNumber="66" type="syntype">syntype</typeStatus>
<taxonomicName id="4C25D29DEF1DFF975857FAEE4F018849" authority=": ZSM 1781" authorityName=": ZSM" authorityYear="1781" box="[324,376,1340,1362]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF1DFF975857FAEE4F018849" box="[324,376,1340,1362]" italics="true" pageId="65" pageNumber="66">gyria</emphasis>
</taxonomicName>
: 
<specimenCode id="DB830165EF1DFF975895FAEE4F958849" box="[390,492,1340,1362]" pageId="65" pageNumber="66">ZSM 1781</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9AA91EEF1DFF975983FA5C4C7888BD" blockId="65.[143,1033,1422,1608]" box="[144,513,1422,1446]" pageId="65" pageNumber="66">Additional specimens examined:</paragraph>
<paragraph id="8B9AA91EEF1DFF9459A6FA624DC68E76" blockId="65.[143,1033,1422,1608]" lastBlockId="66.[181,1071,790,877]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<materialsCitation id="3B4DA343EF1DFF9759A6FA624D6088FC" ID-GBIF-Occurrence="3435945987" collectionCode="QA" lastPageId="65" lastPageNumber="67" pageId="65" pageNumber="66" specimenCode="QA 50">
<collectingCountry id="F332E98EEF1DFF9759A6FA624E8388DD" box="[181,250,1456,1478]" name="Turkey" pageId="65" pageNumber="66">
<emphasis id="B951750CEF1DFF9759A6FA624E8388DD" bold="true" box="[181,250,1456,1478]" pageId="65" pageNumber="66">Turkey</emphasis>
</collectingCountry>
, 
<collectingRegion id="49E167FCEF1DFF975815FA634F2D88DC" box="[262,340,1457,1479]" country="Turkey" name="Antalya" pageId="65" pageNumber="66">Antalya</collectingRegion>
<collectingCounty id="62FBD192EF1DFF975849FA634FDF88DC" box="[346,422,1457,1479]" pageId="65" pageNumber="66">County</collectingCounty>
, 
<collectingMunicipality id="6BFE3364EF1DFF9758BDFA634C4588DC" box="[430,572,1457,1479]" pageId="65" pageNumber="66">Kekova Island</collectingMunicipality>
(= 
<location id="8EFAFFC5EF1DFF975B73FA634C8588DC" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975B73FA634C8588DC" box="[608,764,1457,1479]" country="Turkey" county="County" latitude="54.0" longLatPrecision="0" longitude="30.5131" municipality="Kekova Island" name="Cacamo Cariae" pageId="65" pageNumber="66" stateProvince="Antalya">Cacamo Cariae</location>
, 
<locationDeviation id="9A28B03CEF1DFF975A16FA634DE588DC" box="[773,924,1456,1479]" location="Kas" pageId="65" pageNumber="66">
<quantity id="4CDD04FBEF1DFF975A16FA634D3D88DC" box="[773,836,1457,1479]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="65" pageNumber="66" unit="km" value="20.0">20 km</quantity>
E of Kaş
</locationDeviation>
), 
<collectionCode id="ED3431DBEF1DFF975AA2FA634D9988DC" box="[945,992,1457,1479]" pageId="65" pageNumber="66">UTM</collectionCode>
<specimenCode id="DB830165EF1DFF975AF6FA634ED088FC" collectionCode="QA" pageId="65" pageNumber="66">QA 50</specimenCode>
/60, 36.17°N 29.87°E, (
<bibRefCitation id="EFB4D4EFEF1DFF97588BFA034C4888FC" author="Roth, J. R." box="[408,561,1489,1511]" pageId="65" pageNumber="66" refId="ref68821" refString="Roth, J. R. (1839): Molluscorum species, quas in itinere per Orientem facto comites clariss. Schubert doctores M. Erdl et J. R. Roth collegerunt. Dissertatio inauguralis. - 3 - 26, Taf. 1 - 2, Munchen." type="book" year="1839">Roth, 1839: 16</bibRefCitation>
, 
<location id="8EFAFFC5EF1DFF975B2DFA034CE788FC" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975B2DFA034CE788FC" box="[574,670,1489,1511]" country="Turkey" county="County" latitude="54.0" longLatPrecision="0" longitude="30.5131" municipality="Kekova Island" name="Maassen" pageId="65" pageNumber="66" stateProvince="Antalya">Maassen</location>
, 1991: 124)
</materialsCitation>
; 
<materialsCitation id="3B4DA343EF1DFF975A36FA034CC48B33" ID-GBIF-Occurrence="3435946087" collectingDate="1998-09-27" collectionCode="TF, UTM" collectorName="Welter-Schultes &amp; Welter-Schultes" lastPageId="65" lastPageNumber="67" pageId="65" pageNumber="66" specimenCode="TF 31, 36">
rocks NW of lake near 
<location id="8EFAFFC5EF1DFF975982FA234E8F8B1C" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975982FA234E8F8B1C" box="[145,246,1521,1543]" country="Turkey" county="County" latitude="54.0" longLatPrecision="0" longitude="30.5131" municipality="Kekova Island" name="Beymelek" pageId="65" pageNumber="66" stateProvince="Antalya">Beymelek</location>
(= 
<quantity id="4CDD04FBEF1DFF975809FA234F358B1C" box="[282,332,1521,1543]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="65" pageNumber="66" unit="km" value="9.0">9 km</quantity>
as the crow flies 
<collectionCode id="ED3431DBEF1DFF975B11FA234C418B1C" box="[514,568,1521,1543]" pageId="65" pageNumber="66">WSW</collectionCode>
of 
<location id="8EFAFFC5EF1DFF975B4BFA234CE58B1C" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975B4BFA234CE58B1C" box="[600,668,1521,1543]" country="Turkey" county="County" latitude="54.0" longLatPrecision="0" longitude="30.5131" municipality="Kekova Island" name="Finike" pageId="65" pageNumber="66" stateProvince="Antalya">Finike</location>
), 
<collectionCode id="ED3431DBEF1DFF975BB9FA234CA08B1C" box="[682,729,1521,1543]" pageId="65" pageNumber="66">UTM</collectionCode>
<specimenCode id="DB830165EF1DFF975BCCFA234D6F8B1C" box="[735,790,1521,1543]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="65" pageNumber="66">TF 31</specimenCode>
, 
<specimenCode id="DB830165EF1DFF975A30FA234D448B1C" box="[803,829,1521,1543]" collectionCode="UTM" country="Thailand" name="Department of Mineral Resources" pageId="65" pageNumber="66">36</specimenCode>
.2726°N 30.0456°E, leg. 
<collectorName id="26D0CCC8EF1DFF9759AFF9C04F248B33" box="[188,349,1554,1576]" pageId="65" pageNumber="66">Welter-Schultes</collectorName>
<date id="FF9B8FDEEF1DFF975870F9C04FBE8B33" box="[355,455,1554,1576]" pageId="65" pageNumber="66" value="1998-09-27">
<collectingDate id="EFDF7636EF1DFF975870F9C04FBE8B33" box="[355,455,1554,1576]" pageId="65" pageNumber="66" value="1998-09-27">27.9.1998</collectingDate>
</date>
, coll. 
<collectorName id="26D0CCC8EF1DFF975B17F9C04CD08B33" box="[516,681,1554,1576]" pageId="65" pageNumber="66">Welter-Schultes</collectorName>
/1
</materialsCitation>
; 
<materialsCitation id="3B4DA343EF1DFF975BD9F9C04D4C8B53" ID-GBIF-Occurrence="3435945873" collectionCode="UTM, SMF, TF" lastPageId="65" lastPageNumber="67" pageId="65" pageNumber="66" specimenCode="TF 43, 36, SMF 284853">
Karamanbeyli pass near 
<location id="8EFAFFC5EF1DFF975AD5F9C04A7A8B33" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1DFF975AD5F9C04A7A8B33" box="[966,1027,1554,1576]" country="Turkey" county="County" latitude="54.0" longLatPrecision="0" longitude="30.5131" municipality="Kekova Island" name="Finike" pageId="65" pageNumber="66" stateProvince="Antalya">Finike</location>
(= Phinika Pass), 
<collectionCode id="ED3431DBEF1DFF975840F9E04FFB8B53" box="[339,386,1586,1608]" pageId="65" pageNumber="66">UTM</collectionCode>
<specimenCode id="DB830165EF1DFF975898F9E04FBF8B53" box="[395,454,1586,1608]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="65" pageNumber="66">TF 43</specimenCode>
, 
<specimenCode id="DB830165EF1DFF9758C5F9E04F898B53" box="[470,496,1586,1608]" collectionCode="UTM" country="Thailand" name="Department of Mineral Resources" pageId="65" pageNumber="66">36</specimenCode>
.54°N 29.98°E, 
<specimenCode id="DB830165EF1DFF975B8BF9E04D648B53" box="[664,797,1586,1608]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="65" pageNumber="66">SMF 284853</specimenCode>
/1
</materialsCitation>
; 
<materialsCitation id="3B4DA343EF1DFF945A55F9E04A5C8E37" ID-GBIF-Occurrence="3435945336" collectionCode="TF, UTM" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" specimenCode="TF 75, 36">
forest in gorge of 
<location id="8EFAFFC5EF1EFF9459A4FCC44F4C8E37" LSID="urn:lsid:plazi:treatment:038C1808EF63FF945983FC1C4D8D89F7:8EFAFFC5EF1EFF9459A4FCC44F4C8E37" box="[183,309,790,812]" country="Turkey" county="County" latitude="54.0" longLatPrecision="0" longitude="30.5131" municipality="Kekova Island" name="Kemer creek" pageId="66" pageNumber="67" stateProvince="Antalya">Kemer creek</location>
(= 
<locationDeviation id="9A28B03CEF1EFF94584BFCC44FAB8E37" box="[344,466,790,812]" location="Kemer" pageId="66" pageNumber="67">W of Kemer</locationDeviation>
), 
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<specimenCode id="DB830165EF1EFF945B07FCC44C338E37" box="[532,586,790,812]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="66" pageNumber="67">TF 75</specimenCode>
, 
<specimenCode id="DB830165EF1EFF945B45FCC44C098E37" box="[598,624,790,812]" collectionCode="UTM" country="Thailand" name="Department of Mineral Resources" pageId="66" pageNumber="67">36</specimenCode>
.5943°N 30.5049°E, (Maassen, 1991: 124)
</materialsCitation>
; 
<materialsCitation id="3B4DA343EF1EFF9459A5FCE44FE18E76" ID-GBIF-Occurrence="3435945506" collectionCode="UTM, TF" collectorName="Maassen" lastPageId="66" lastPageNumber="67" pageId="66" pageNumber="66" specimenCode="TF 84, 36">
Phaselis, ruins near Tekirova (= 
<locationDeviation id="9A28B03CEF1EFF945B16FCE44CB98E57" box="[517,704,822,844]" location="Kemer" pageId="66" pageNumber="67">
<quantity id="4CDD04FBEF1EFF945B16FCE44C3A8E57" box="[517,579,822,844]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="66" pageNumber="67" unit="km" value="10.0">10 km</quantity>
S of Kemer
</locationDeviation>
), 
<collectionCode id="ED3431DBEF1EFF945BDCFCE44C878E57" box="[719,766,822,844]" pageId="66" pageNumber="67">UTM</collectionCode>
<specimenCode id="DB830165EF1EFF945A17FCE44D448E57" box="[772,829,822,844]" collectionCode="TF" country="Thailand" name="Department of Mineral Resources" pageId="66" pageNumber="67">TF 84</specimenCode>
, 
<specimenCode id="DB830165EF1EFF945A59FCE44D1D8E57" box="[842,868,822,844]" collectionCode="UTM" country="Thailand" name="Department of Mineral Resources" pageId="66" pageNumber="67">36</specimenCode>
.4704°N 30.5131°E, (
<collectorName id="26D0CCC8EF1EFF9459A8FC854F678E76" box="[187,286,855,877]" pageId="66" pageNumber="67">Maassen</collectorName>
, 1991: 124)
</materialsCitation>
; 
<materialsCitation id="3B4DA343EF1EFF9458B0FC854DC68E76" ID-GBIF-Occurrence="3435945495" box="[419,959,855,877]" collectingDate="1999-02-26" collectorName="Subai" lastPageId="66" lastPageNumber="67" pageId="66" pageNumber="66">
leg. 
<collectorName id="26D0CCC8EF1EFF9458DDFC854C708E76" box="[462,521,855,877]" pageId="66" pageNumber="67">Subai</collectorName>
<date id="FF9B8FDEEF1EFF945B1DFC854C0C8E76" box="[526,629,855,877]" pageId="66" pageNumber="67" value="1999-02-26">
<collectingDate id="EFDF7636EF1EFF945B1DFC854C0C8E76" box="[526,629,855,877]" pageId="66" pageNumber="67" value="1999-02-26">26.2.1999</collectingDate>
</date>
, 
<accessionNumber id="947634FDEF1EFF945B92FC854CAC8E76" box="[641,725,855,877]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S16083" pageId="66" pageNumber="67" type="EnaNcbi">S 16083</accessionNumber>
/17 + 7 (juv.) + 1 (alk.).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF5AF996EF1EFF9459A6FD7A4C5D8FA1" ID-DOI="http://doi.org/10.5281/zenodo.5842370" ID-Zenodo-Dep="5842370" box="[181,548,680,698]" httpUri="https://zenodo.org/record/5842370/files/figure.png" pageId="66" pageNumber="67" startId="66.[181,210,680,698]" targetBox="[181,1065,113,643]" targetPageId="66">
<paragraph id="8B9AA91EEF1EFF9459A6FD7A4C5D8FA1" blockId="66.[181,548,680,698]" box="[181,548,680,698]" pageId="66" pageNumber="67">
Fig. 35. Distribution of 
<taxonomicName id="4C25D29DEF1EFF945868FD7A4C668FA1" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[379,543,680,698]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="gyria">
<emphasis id="B951750CEF1EFF945868FD7A4C668FA1" bold="true" box="[379,543,680,698]" italics="true" pageId="66" pageNumber="67">Lindholmiola gyria</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C33FFA95EF1EFF9459C8FC7B4D2F8967" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8B9AA91EEF1EFF9459C8FC7B4D2F8967" blockId="66.[180,1071,937,1260]" pageId="66" pageNumber="67">
Remarks: This species is probably endangered, because even empty shells are very rare. In 
<date id="FF9B8FDEEF1EFF94586AFC1C4C598EFD" box="[377,544,974,998]" pageId="66" pageNumber="67" value="1999-02">February 1999</date>
, it took several hours until the only hitherto known living specimen could be found close to Phaselis. The anatomical investigation revealed that it belongs to 
<taxonomicName id="4C25D29DEF1EFF945B44FBCB4C94892A" authorityName="HESSE, Zoologica" authorityYear="1931" box="[599,749,1049,1073]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="genus">
<emphasis id="B951750CEF1EFF945B44FBCB4C94892A" box="[599,749,1049,1073]" italics="true" pageId="66" pageNumber="67">Lindholmiola</emphasis>
</taxonomicName>
and not to 
<taxonomicName id="4C25D29DEF1EFF945A6AFBCB4A7A892A" authorityName="A. Ferussac" authorityYear="1821" box="[889,1027,1049,1073]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="genus">
<emphasis id="B951750CEF1EFF945A6AFBCB4A7A892A" box="[889,1027,1049,1073]" italics="true" pageId="66" pageNumber="67">Helicodonta</emphasis>
</taxonomicName>
. In 1991 (p. 124, 
<figureCitation id="131EB59BEF1EFF94585DFBEC4FDE894D" box="[334,423,1086,1110]" captionStart="Figs 3–6" captionStartId="10.[181,218,1390,1408]" captionTargetBox="[181,1065,113,1346]" captionTargetId="figure-8@10.[181,1065,113,1346]" captionTargetPageId="10" captionText="Figs 3–6. Lindholmiola lens. Fig. 3: syntype Helix lentiformis, ZMZ 508656, &quot;Thessalie et Attique&quot;, D = 11.2 mm; Fig. 4: NMBE 515542, Attika, Methana, 500 m before village, 12.07.1996, leg. E. Neubert, D = 12.77 mm; Fig. 5: syntype Helix lens var. callojuncta, NHMG 372, D = 12.0 mm; Fig. 6: syntype Helix lens var. elia, SMF 101961, Kumani, Elis, coll. Jetschin ex Boettger, 1883, D = 11.94 mm. — All photos Neubert/Bochud, × 3." figureDoi="http://doi.org/10.5281/zenodo.5842346" httpUri="https://zenodo.org/record/5842346/files/figure.png" pageId="66" pageNumber="67">Fig. 4–6</figureCitation>
), Maassen described 
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<emphasis id="B951750CEF1EFF945B88FBEC4D94894D" box="[667,1005,1086,1110]" italics="true" pageId="66" pageNumber="67">Helicodonta gyria wilhelminae</emphasis>
</taxonomicName>
from 
<collectingRegion id="49E167FCEF1EFF9459A5FBB64E8F8967" box="[182,246,1124,1148]" country="Greece" name="Kriti" pageId="66" pageNumber="67">Crete</collectingRegion>
, which is considered an endemic 
<taxonomicName id="4C25D29DEF1EFF945B79FBB64D2F8967" box="[618,854,1124,1148]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B951750CEF1EFF945B79FBB64C8A8967" box="[618,755,1124,1148]" italics="true" pageId="66" pageNumber="67">Helicodonta</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33FFA95EF1EFF9459CFFB5B4D8D89F7" pageId="66" pageNumber="67" type="distribution">
<paragraph id="8B9AA91EEF1EFF9459CFFB5B4D8D89F7" blockId="66.[180,1071,937,1260]" pageId="66" pageNumber="67">
Distribution (
<figureCitation id="131EB59BEF1EFF94587CFB5B4FCC89BA" box="[367,437,1161,1185]" captionStart="Fig" captionStartId="66.[181,210,680,698]" captionTargetBox="[181,1065,113,643]" captionTargetId="figure-256@66.[181,1065,113,643]" captionTargetPageId="66" captionText="Fig. 35. Distribution of Lindholmiola gyria." figureDoi="http://doi.org/10.5281/zenodo.5842370" httpUri="https://zenodo.org/record/5842370/files/figure.png" pageId="66" pageNumber="67">Fig 35</figureCitation>
): This species is only known from a small coastal stripe in the south-western part of the vilayet 
<collectingRegion id="49E167FCEF1EFF945B6BFB7C4CAB89DD" box="[632,722,1198,1222]" country="Turkey" name="Antalya" pageId="66" pageNumber="67">Antalya</collectingRegion>
. The distribution range covers the Island Kekova (= E of Kaş) and the surroundings of Kemer to Phaselis.
</paragraph>
</subSubSection>
</treatment>
</document>