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<document id="505CA5AD39059C8FC8D3F1297B635839" ID-DOI="10.1080/00222933.2021.1914237" ID-ISSN="1464-5262" ID-Zenodo-Dep="5477393" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630929801240" checkinUser="plazi" docAuthor="El-Hawagry, Magdi S." docDate="2021" docId="625787F0C301CD21FAD0FC0255052448" docLanguage="en" docName="JNATHIST.55.9-10.597-624.pdf" docOrigin="Journal of Natural History 55 (9 - 10)" docSource="http://dx.doi.org/10.1080/00222933.2021.1914237" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Anthrax sticticus Klug 1832" docType="treatment" docVersion="4" lastPageNumber="618" masterDocId="9E6EFF88C315CD34FB50FFDB577F2074" masterDocTitle="Taxonomy of the genus Anthrax Scopoli (Diptera: Bombyliidae) in Egypt" masterLastPageNumber="624" masterPageNumber="597" pageNumber="617" updateTime="1699283167991" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7590AA3040A5725F10C700B66737E647">Taxonomy of the genus Anthrax Scopoli (Diptera: Bombyliidae) in Egypt</mods:title>
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<mods:namePart id="6FAA3ECEBBDC9848EC307764AEC34F47">El-Hawagry, Magdi S.</mods:namePart>
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<mods:title id="552F31A685AC4682CDFC1A2CD30C47F1">Journal of Natural History</mods:title>
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<mods:part id="DF6596D1ADE4A85E2AAE2FE6E45A6332">
<mods:date id="6A7A59511E89B2D4CB0805A5CD4CDE47">2021</mods:date>
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<mods:number id="0A026D75D2EEF3914FC5D317CCF676CA">2021-06-11</mods:number>
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<mods:number id="A48A278A2D0EEB122870845F1D72B171">9 - 10</mods:number>
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<subSubSection id="A2E4656DC301CD20FAD0FC0254DC2387" box="[384,931,985,1011]" pageId="20" pageNumber="617" type="nomenclature">
<paragraph id="EA4136E6C301CD20FAD0FC0254DC2387" blockId="20.[384,931,985,1011]" box="[384,931,985,1011]" pageId="20" pageNumber="617">
<heading id="B109818AC301CD20FAD0FC0254DC2387" box="[384,931,985,1011]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="617" reason="5">
<taxonomicName id="2DFE4D65C301CD20FAD0FC0255B02387" ID-CoL="5VGCS" authority="Klug, 1832" authorityName="Klug" authorityYear="1832" box="[384,719,985,1011]" class="Insecta" family="Bombyliidae" genus="Anthrax" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="617" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="D88AEAF4C301CD20FAD0FC0255332387" bold="true" box="[384,588,985,1011]" italics="true" pageId="20" pageNumber="617">Anthrax sticticus</emphasis>
<bibRefCitation id="8E6F4B17C301CD20F903FC0255B02387" author="Klug JCF" box="[595,719,985,1011]" pageId="20" pageNumber="617" refId="ref11804" refString="Klug JCF. 1832. Symbolae physicae seu icones et descriptions insectorum quae exitinere Africam borealem et Asiam Occidentalem F. G. Hemprich et C. H. Ehrenberg studio novae aut illustratae rediereunt. Berlin: Mittler; p. 185." type="journal volume" year="1832">Klug, 1832</bibRefCitation>
</taxonomicName>
(
<figureCitation id="72C52A63C301CD20F987FC02542B2387" box="[727,852,985,1011]" captionStart="Figure 12" captionStartId="21.[159,224,878,901]" captionTargetBox="[167,1148,143,846]" captionTargetId="figure-208@21.[166,1149,142,846]" captionTargetPageId="21" captionText="Figure 12. Anthrax sticticus: (a) gonocoxites, ventral view; (b) hypopygium, lateral view; (c) tip of epiphallus, dorsal view; (d) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5477417" httpUri="https://zenodo.org/record/5477417/files/figure.png" pageId="20" pageNumber="617">Figures 12</figureCitation>
,
<figureCitation id="72C52A63C301CD20F80FFC0254022387" box="[863,893,985,1011]" captionStart="Figure 16" captionStartId="25.[159,224,1268,1291]" captionTargetBox="[167,1148,143,1234]" captionTargetId="figure-140@25.[166,1149,142,1236]" captionTargetPageId="25" captionText="Figure 16. Anthrax spp., habitus: (a) A. galali; (b) A. moursyi; (c) A. pharaonis, neotype; (d) A. sticticus; (e) A. trifasciatus; (f) A. zohrayensis." figureDoi="http://doi.org/10.5281/zenodo.5477425" httpUri="https://zenodo.org/record/5477425/files/figure.png" pageId="20" pageNumber="617">16</figureCitation>
(d))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2E4656DC301CD20FBF0FBD6563C24A0" pageId="20" pageNumber="617" type="reference_group">
<paragraph id="EA4136E6C301CD20FBF0FBD6531C2452" blockId="20.[160,1155,1037,1236]" box="[160,1123,1037,1062]" pageId="20" pageNumber="617">
<taxonomicName id="2DFE4D65C301CD20FBF0FBD656B82452" ID-CoL="87SJK" authority="Klug, 1832" authorityName="Klug" authorityYear="1832" box="[160,455,1037,1062]" class="Insecta" family="Bombyliidae" genus="Anthrax" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="617" phylum="Arthropoda" rank="species" species="stictica">
<emphasis id="D88AEAF4C301CD20FBF0FBD656362452" box="[160,329,1037,1062]" italics="true" pageId="20" pageNumber="617">Anthrax stictica</emphasis>
<bibRefCitation id="8E6F4B17C301CD20FA00FBD656B82452" author="Klug JCF" box="[336,455,1037,1062]" pageId="20" pageNumber="617" refId="ref11804" refString="Klug JCF. 1832. Symbolae physicae seu icones et descriptions insectorum quae exitinere Africam borealem et Asiam Occidentalem F. G. Hemprich et C. H. Ehrenberg studio novae aut illustratae rediereunt. Berlin: Mittler; p. 185." type="journal volume" year="1832">Klug, 1832</bibRefCitation>
</taxonomicName>
: pl. 30, fig. 14.
<materialsCitation id="5A963CBBC301CD20F920FBD653202452" box="[624,1119,1037,1062]" country="Saudi Arabia" location="Saudi Arabia" pageId="20" pageNumber="617" specimenCount="1">
<typeStatus id="35458844C301CD20F920FBD655D92452" box="[624,678,1037,1062]" pageId="20" pageNumber="617">Type</typeStatus>
locality:
<collectingCountry id="92E97676C301CD20F85BFBD654E72452" box="[779,920,1037,1062]" name="Saudi Arabia" pageId="20" pageNumber="617">Saudi Arabia</collectingCountry>
[as Arabia felici]
</materialsCitation>
.
</paragraph>
<paragraph id="EA4136E6C301CD20FBF0FBEB5311243D" blockId="20.[160,1155,1037,1236]" box="[160,1134,1072,1097]" pageId="20" pageNumber="617">
<treatmentCitationGroup id="CAEE11C8C301CD20FBF0FBEB5311243D" box="[160,1134,1072,1097]" pageId="20" pageNumber="617">
<treatmentCitation id="6B5F10F7C301CD20FBF0FBEB5516243D" author="Sack P" box="[160,617,1072,1097]" page="534" pageId="20" pageNumber="617" year="1909">
<taxonomicName id="2DFE4D65C301CD20FBF0FBEB5516243D" ID-CoL="GJ5W" authority="Sack, 1909: 534" authorityName="Sack" authorityPageNumber="534" authorityYear="1909" box="[160,617,1072,1097]" class="Insecta" family="Bombyliidae" genus="Argyramoeba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="617" phylum="Arthropoda" rank="species" species="polystigma">
<emphasis id="D88AEAF4C301CD20FBF0FBEB56CC243D" box="[160,435,1072,1097]" italics="true" pageId="20" pageNumber="617">Argyramoeba polystigma</emphasis>
<bibRefCitation id="8E6F4B17C301CD20FAEAFBEB5516243D" author="Sack P" box="[442,617,1072,1097]" pageId="20" pageNumber="617" pagination="501 - 548" refId="ref12098" refString="Sack P. 1909. Die palaearktischen Spongostylinen. Abh Senckenb Naturforsch Ges. 30: 501 - 548." type="journal article" year="1909">Sack, 1909: 534</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<materialsCitation id="5A963CBBC301CD20F923FBEB5315243D" box="[627,1130,1072,1097]" country="Cyprus and Greece" location="Cyprus and Greece" pageId="20" pageNumber="617" specimenCount="1">
<typeStatus id="35458844C301CD20F923FBEB55D6243D" box="[627,681,1072,1097]" pageId="20" pageNumber="617">Type</typeStatus>
locality:
<collectingCountry id="92E97676C301CD20F85DFBEB549D243D" box="[781,994,1072,1097]" isEnumeration="true" pageId="20" pageNumber="617">Cyprus and Greece</collectingCountry>
and
<collectingCountry id="92E97676C301CD20FF4CFBEB5315243D" box="[1052,1130,1072,1097]" name="Turkey" pageId="20" pageNumber="617">Turkey</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA4136E6C301CD20FBF0FB8856CF24FA" blockId="20.[160,1155,1037,1236]" pageId="20" pageNumber="617">
<taxonomicName id="2DFE4D65C301CD20FBF0FB8855872418" ID-CoL="4ZB3T" authority="Bezzi, 1920: 105" authorityName="Bezzi" authorityPageNumber="105" authorityYear="1920" box="[160,760,1107,1132]" class="Insecta" family="Bombyliidae" genus="Spongostylum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="617" phylum="Arthropoda" rank="species" species="quinquemaculatum">
<emphasis id="D88AEAF4C301CD20FBF0FB88555F2418" box="[160,544,1107,1132]" italics="true" pageId="20" pageNumber="617">Spongostylum quinquemaculatum</emphasis>
Bezzi, 1920: 105
</taxonomicName>
.
<materialsCitation id="5A963CBBC301CD20F85BFB8856D324FA" country="Guinea-Bissau" location="Guinea-Bissau" pageId="20" pageNumber="617" specimenCount="1">
<typeStatus id="35458844C301CD20F85BFB88543E2418" box="[779,833,1107,1132]" pageId="20" pageNumber="617">Type</typeStatus>
locality:
<collectingCountry id="92E97676C301CD20F8E7FB88532B2418" box="[951,1108,1107,1132]" name="Guinea-Bissau" pageId="20" pageNumber="617">Guinea-Bissau</collectingCountry>
[as Guinea Portoghese]
</materialsCitation>
.
</paragraph>
<paragraph id="EA4136E6C301CD20FBF0FB43563C24A0" blockId="20.[160,1155,1037,1236]" pageId="20" pageNumber="617">
<treatmentCitationGroup id="CAEE11C8C301CD20FBF0FB43563C24A0" pageId="20" pageNumber="617">
<treatmentCitation id="6B5F10F7C301CD20FBF0FB4355AF24C5" author="Bezzi M" box="[160,720,1176,1201]" page="170" pageId="20" pageNumber="617" year="1924">
<taxonomicName id="2DFE4D65C301CD20FBF0FB4355AF24C5" ID-CoL="4ZB3T" authority="Bezzi, 1924: 170" authorityName="Bezzi" authorityPageNumber="170" authorityYear="1924" box="[160,720,1176,1201]" class="Insecta" family="Bombyliidae" genus="Spongostylum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="617" phylum="Arthropoda" rank="species" species="quinquemaculatum">
<emphasis id="D88AEAF4C301CD20FBF0FB43556924C5" box="[160,534,1176,1201]" italics="true" pageId="20" pageNumber="617">Spongostylum quinquemaculatum</emphasis>
<bibRefCitation id="8E6F4B17C301CD20F94CFB4355AF24C5" author="Bezzi M" box="[540,720,1176,1201]" pageId="20" pageNumber="617" refId="ref11329" refString="Bezzi M. 1924. The Bombyliidae of the Ethiopian Region. London: British Museum (Natural History); p. 390." type="journal volume" year="1924">Bezzi, 1924: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<materialsCitation id="5A963CBBC301CD20F989FB4354B324C5" box="[729,972,1176,1201]" country="Senegal" location="Senegal" pageId="20" pageNumber="617" specimenCount="1">
<typeStatus id="35458844C301CD20F989FB43547024C5" box="[729,783,1176,1201]" pageId="20" pageNumber="617">Type</typeStatus>
locality:
<collectingCountry id="92E97676C301CD20F821FB4354B324C5" box="[881,972,1176,1201]" name="Senegal" pageId="20" pageNumber="617">Senegal</collectingCountry>
</materialsCitation>
. Preoccupied by Bezzi, 1920.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2E4656DC301CD20FBF0FADD546B26B4" pageId="20" pageNumber="617" type="diagnosis">
<paragraph id="EA4136E6C301CD20FBF0FADD566B2554" blockId="20.[160,1156,1286,1728]" box="[160,276,1286,1312]" pageId="20" pageNumber="617">
<emphasis id="D88AEAF4C301CD20FBF0FADD566B2554" bold="true" box="[160,276,1286,1312]" italics="true" pageId="20" pageNumber="617">Diagnosis</emphasis>
</paragraph>
<paragraph id="EA4136E6C301CD20FBF0FAF1546B26B4" blockId="20.[160,1156,1286,1728]" pageId="20" pageNumber="617">
<emphasis id="D88AEAF4C301CD20FBF0FAF156282537" box="[160,343,1322,1347]" italics="true" pageId="20" pageNumber="617">Male and female</emphasis>
: About
<quantity id="2D069B03C301CD20FAE0FAF1568E2537" box="[432,497,1322,1347]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="617" unit="mm" value="8.0">8 mm</quantity>
in length (
<figureCitation id="72C52A63C301CD20F93DFAF155A72537" box="[621,728,1322,1347]" captionStart="Figure 16" captionStartId="25.[159,224,1268,1291]" captionTargetBox="[167,1148,143,1234]" captionTargetId="figure-140@25.[166,1149,142,1236]" captionTargetPageId="25" captionText="Figure 16. Anthrax spp., habitus: (a) A. galali; (b) A. moursyi; (c) A. pharaonis, neotype; (d) A. sticticus; (e) A. trifasciatus; (f) A. zohrayensis." figureDoi="http://doi.org/10.5281/zenodo.5477425" httpUri="https://zenodo.org/record/5477425/files/figure.png" pageId="20" pageNumber="617">Figure 16</figureCitation>
(d)). Frons with black hairs mixed with some yellow-white scaly hairs. Face, occiput and pleura with white hairs mixed with a few black hairs at face. Scutum and scutellum black-haired, with some white-brown scaly hairs. Wing pattern composed of brown spots on cross-vein r-m and origin of vein R2+3, on origin of vein R4, on cross-vein m-m, on base of cell m2, and a larger one on bases of cells cua1 and dm; costal and subcostal cells infuscated brown and this infuscation extends to bases of cells br, bm and cup; it extends also to cross-vein bm-cu through the tip of cell bm and the middle of cell br. Abdomen black, with black hairs except for white hairs on first tergite and rows of white scaly hairs on hind margins of tergites giving the abdomen a banded appearance. Gonocoxites (
<figureCitation id="72C52A63C301CD20F96EF9B955D7260F" box="[574,680,1634,1659]" captionStart="Figure 12" captionStartId="21.[159,224,878,901]" captionTargetBox="[167,1148,143,846]" captionTargetId="figure-208@21.[166,1149,142,846]" captionTargetPageId="21" captionText="Figure 12. Anthrax sticticus: (a) gonocoxites, ventral view; (b) hypopygium, lateral view; (c) tip of epiphallus, dorsal view; (d) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5477417" httpUri="https://zenodo.org/record/5477417/files/figure.png" pageId="20" pageNumber="617">Figure 12</figureCitation>
(a)) truncate, triangular; epiphallus simple; aedeagus massive (
<figureCitation id="72C52A63C301CD20FAD2F95E568D26EA" box="[386,498,1669,1694]" captionStart="Figure 12" captionStartId="21.[159,224,878,901]" captionTargetBox="[167,1148,143,846]" captionTargetId="figure-208@21.[166,1149,142,846]" captionTargetPageId="21" captionText="Figure 12. Anthrax sticticus: (a) gonocoxites, ventral view; (b) hypopygium, lateral view; (c) tip of epiphallus, dorsal view; (d) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5477417" httpUri="https://zenodo.org/record/5477417/files/figure.png" pageId="20" pageNumber="617">Figure 12</figureCitation>
(b,c)). Spermatheca (
<figureCitation id="72C52A63C301CD20F98DF95E543326EA" box="[733,844,1669,1694]" captionStart="Figure 12" captionStartId="21.[159,224,878,901]" captionTargetBox="[167,1148,143,846]" captionTargetId="figure-208@21.[166,1149,142,846]" captionTargetPageId="21" captionText="Figure 12. Anthrax sticticus: (a) gonocoxites, ventral view; (b) hypopygium, lateral view; (c) tip of epiphallus, dorsal view; (d) spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5477417" httpUri="https://zenodo.org/record/5477417/files/figure.png" pageId="20" pageNumber="617">Figure 12</figureCitation>
(d)) with a balloon-shaped, well-sclerotised bulb and a long, well-sclerotised pump.
</paragraph>
</subSubSection>
<caption id="BE81666EC300CD21FBCFFCB5554A23D6" ID-DOI="http://doi.org/10.5281/zenodo.5477417" ID-Zenodo-Dep="5477417" httpUri="https://zenodo.org/record/5477417/files/figure.png" pageId="21" pageNumber="618" startId="21.[159,224,878,901]" targetBox="[167,1148,143,846]" targetPageId="21">
<paragraph id="EA4136E6C300CD21FBCFFCB5554A23D6" blockId="21.[159,1156,878,930]" pageId="21" pageNumber="618">
<emphasis id="D88AEAF4C300CD21FBCFFCB5567523F1" bold="true" box="[159,266,878,901]" pageId="21" pageNumber="618">Figure 12.</emphasis>
<taxonomicName id="2DFE4D65C300CD21FA44FCB556CB23F1" authorityName="Klug" authorityYear="1832" box="[276,436,878,901]" class="Insecta" family="Bombyliidae" genus="Anthrax" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="618" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="D88AEAF4C300CD21FA44FCB556CB23F1" box="[276,436,878,901]" italics="true" pageId="21" pageNumber="618">Anthrax sticticus</emphasis>
</taxonomicName>
: (a) gonocoxites, ventral view; (b) hypopygium, lateral view; (c) tip of epiphallus, dorsal view; (d) spermatheca.
</paragraph>
</caption>
<subSubSection id="A2E4656DC300CD21FBF0FC0755052448" pageId="21" pageNumber="618" type="materials_examined">
<paragraph id="EA4136E6C300CD21FBF0FC0756002382" blockId="21.[160,1156,988,1084]" box="[160,383,988,1014]" pageId="21" pageNumber="618">
<emphasis id="D88AEAF4C300CD21FBF0FC0756002382" bold="true" box="[160,383,988,1014]" italics="true" pageId="21" pageNumber="618">Material examined</emphasis>
</paragraph>
<paragraph id="EA4136E6C300CD21FBF0FBDB55052448" blockId="21.[160,1156,988,1084]" pageId="21" pageNumber="618">
I male, W.
<materialsCitation id="5A963CBBC300CD21FA42FBDB55092448" collectingDate="1997-05-21" collectingDateMax="1998-07-12" collectingDateMin="1997-05-21" collectionCode="MSHC" collectorName="M. El-Hawagry" latitude="28.5469" longLatPrecision="6" longitude="33.953" pageId="21" pageNumber="618" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
El-Arbaein (
<geoCoordinate id="8FCA5021C300CD21FAC1FBDB5578246D" box="[401,519,1024,1049]" degrees="28.5469" direction="north" orientation="latitude" pageId="21" pageNumber="618" precision="5" value="28.5469">28.5469°N</geoCoordinate>
,
<geoCoordinate id="8FCA5021C300CD21F940FBDB55FD246D" box="[528,642,1024,1049]" degrees="33.9530" direction="east" orientation="longitude" pageId="21" pageNumber="618" precision="5" value="33.953">33.9530°E</geoCoordinate>
),
<date id="9E401026C300CD21F9C1FBDB545E246D" box="[657,801,1024,1049]" pageId="21" pageNumber="618" value="1997-05-21">
<collectingDate id="8E04E9CEC300CD21F9C1FBDB545E246D" box="[657,801,1024,1049]" pageId="21" pageNumber="618" value="1997-05-21">21 May 1997</collectingDate>
</date>
;
<specimenCount id="FCF8FD6FC300CD21F87AFBDB54F2246D" box="[810,909,1024,1049]" pageId="21" pageNumber="618" type="female">1 female</specimenCount>
,
<specimenCount id="FCF8FD6FC300CD21F8C6FBDB549C246D" box="[918,995,1024,1049]" pageId="21" pageNumber="618" type="male">1 male</specimenCount>
, same locality,
<date id="9E401026C300CD21FBF0FBF856522448" box="[160,301,1059,1084]" pageId="21" pageNumber="618" value="1998-07-12">
<collectingDate id="8E04E9CEC300CD21FBF0FBF856522448" box="[160,301,1059,1084]" pageId="21" pageNumber="618" value="1998-07-12">12 July 1998</collectingDate>
</date>
(
<collectorName id="470B5330C300CD21FA6BFBF856A02448" box="[315,479,1059,1084]" pageId="21" pageNumber="618">M. El-Hawagry</collectorName>
, leg.) [
<collectionCode id="8CEFAE23C300CD21F976FBF8550E2448" box="[550,625,1059,1084]" pageId="21" pageNumber="618">MSHC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>