275 lines
39 KiB
XML
275 lines
39 KiB
XML
<document id="A9C199A49A56E7C4B7A83F40F38B5B9C" ID-CLB-Dataset="37319" ID-DOI="10.5281/zenodo.273086" ID-GBIF-Dataset="0d2332a5-1968-49d7-a570-f31d386eccb3" ID-ISSN="1175-5326" ID-Zenodo-Dep="273086" ID-ZooBank="0FA0F51A-3868-4F13-A93D-E34CA5A689F8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484409762612" checkinUser="plazi" docAuthor="Bernhard A. Huber" docDate="2017" docId="03B66F688539073DFF6AFD53289AFF5B" docLanguage="en" docName="zootaxa.4219.1.1.pdf.imf" docOrigin="Zootaxa 4219 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calapnita lehi Huber, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="FF8F171085320730FFFDFF852B77FFA3" masterDocTitle="Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="12" updateTime="1698435649618" updateUser="ExternalLinkService">
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<mods:titleInfo id="BB77A6C8C21E65E6D8728CAD5E7EBC68">
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<mods:title id="E79C195117946DF874D5096EA23D1042">Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E0066CA02CFD4DE274EBA70AB22EC755" type="personal">
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<mods:roleTerm id="D56E1DE8AC02E649DE358A544A29F035">Author</mods:roleTerm>
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<mods:namePart id="B8C7A8B82E8553FBC981664FB2ABB6DA">Bernhard A. Huber</mods:namePart>
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<mods:typeOfResource id="AC30B8C2974D74FCDB585B08DB337F2A">text</mods:typeOfResource>
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<mods:titleInfo id="39A1EDF0CC539BA986F55F1A99A04263">
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<mods:title id="B348C39FCAB4C9C271C23E2354750492">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FD3C98CD00C5161E4C4A5EDCDA846AEC">
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<mods:date id="26A23EA3D4A496152223378FFD3FE0B6">2017</mods:date>
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<mods:detail id="41F55CF4DB15F7755C6B52DCB0D48194" type="volume">
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<mods:number id="68A265C5246C46AFC3EBB4002BCBE8ED">4219</mods:number>
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</mods:detail>
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<mods:detail id="849E3E9C81ACC30F34CC83B34762A0EC" type="issue">
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<mods:number id="3F2C1401C1565757E429348156269305">1</mods:number>
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<mods:classification id="AD9A2C08432C45A994DD95D3BCB12C73">journal article</mods:classification>
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<mods:identifier id="81EF871B1B03C5567FCB90FB0D90AA63" type="CLB-Dataset">37319</mods:identifier>
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<mods:identifier id="74C36B42FAB92E4CD9F62BDDEB5D08FC" type="DOI">10.5281/zenodo.273086</mods:identifier>
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<mods:identifier id="8CBF25140CD3264CB60E3748225E265D" type="GBIF-Dataset">0d2332a5-1968-49d7-a570-f31d386eccb3</mods:identifier>
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<mods:identifier id="AA158C10086688C54838FCEBEA2D5DA1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BCD85698AC4663248C0E6E90A5B9308B" type="Zenodo-Dep">273086</mods:identifier>
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<treatment id="03B66F688539073DFF6AFD53289AFF5B" ID-DOI="http://doi.org/10.5281/zenodo.6040198" ID-GBIF-Taxon="127419953" ID-Zenodo-Dep="6040198" LSID="urn:lsid:plazi:treatment:03B66F688539073DFF6AFD53289AFF5B" httpUri="http://treatment.plazi.org/id/03B66F688539073DFF6AFD53289AFF5B" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
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<subSubSection id="C3058DF58539073BFF6AFD532A3BFCB1" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BA0DE7E8539073BFF6AFD532A3BFCB1" blockId="11.[151,419,726,786]" pageId="11" pageNumber="12">
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<taxonomicName id="4C1FA5FD8539073BFF6AFD532A35FD53" ID-CoL="93W54" box="[151,322,726,752]" class="Arachnida" family="Pholcidae" genus="Calapnita" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lehi" status="sp. nov.">
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<emphasis id="B96B026C8539073BFF6AFD532A35FD53" bold="true" box="[151,322,726,752]" italics="true" pageId="11" pageNumber="12">Calapnita lehi</emphasis>
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</taxonomicName>
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<emphasis id="B96B026C8539073BFEB4FD522AD4FD53" bold="true" box="[329,419,727,752]" pageId="11" pageNumber="12">
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<taxonomicNameLabel id="A258BF178539073BFEB4FD522AD4FD53" box="[329,419,727,752]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<figureCitation id="1324C2FB8539073BFF6AFD7C2B8CFCB1" box="[151,251,761,786]" captionStart="FIGURES 3 – 12" captionStartId="9.[151,264,1890,1912]" captionTargetBox="[151,1436,337,1869]" captionTargetId="figure@9.[151,1436,337,1869]" captionTargetPageId="9" captionText="FIGURES 3 – 12. Live specimens, Calapnita phyllicola group. 3 – 6. C. lehi sp. nov., ♂ and ♀ with egg-sac from Lambir (3 – 4); ♂ and penultimate instar ♂ from Gunung Mulu (5 – 6). 7 – 8. C. kubah sp. nov., ♂ and ♀ with egg-sac from Kubah. 9. C. deelemanae Huber, ♀ with egg-sac from Poring. 10. C. subphyllicola Deeleman-Reinhold, ♀ with egg-sac from Malagos. 11 – 12. C. anai sp. nov., ♂ from MacRitchie, ♀ with egg-sac from Lembah Anai." httpUri="https://zenodo.org/record/273089/files/figure.png" pageId="11" pageNumber="12">Figs 3–6</figureCitation>
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,
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<figureCitation id="1324C2FB8539073BFEFBFD7F2A3BFCB1" box="[262,332,762,786]" captionStart-0="FIGURES 20 – 25" captionStart-1="FIGURES 26 – 32" captionStartId-0="12.[151,264,1735,1757]" captionStartId-1="13.[151,264,1907,1929]" captionTargetBox-0="[151,1431,193,1708]" captionTargetBox-1="[151,1436,303,1885]" captionTargetId-0="figure@12.[151,1436,193,1714]" captionTargetId-1="figure@13.[151,1436,303,1885]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." captionText-1="FIGURES 26 – 32. Calapnita lehi sp. nov., SEM micrographs (ZFMK, Ar 15965 – 66). 26 – 27. Male cheliceral apophyses. 28. Right procursus, prolateral view. 29. Tip of bulbal appendix. 30. Comb-hair on male tarsus 4. 31. Male ALS. 32. Epigynum, ventral view. Scale lines: 10 µm (26, 27, 30, 31), 50 µm (28, 29, 32)." httpUri-0="https://zenodo.org/record/273091/files/figure.png" httpUri-1="https://zenodo.org/record/273092/files/figure.png" pageId="11" pageNumber="12">20–32</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3058DF58539073BFF6AFCC42874FC49" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8BA0DE7E8539073BFF6AFCC42874FC49" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
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<emphasis id="B96B026C8539073BFF6AFCC42A66FCF9" bold="true" box="[151,273,833,858]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
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Males are easily distinguished from congeners by modified male palpal femur (distinctive ventral process;
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<figureCitation id="1324C2FB8539073BFF03FCE02A22FCDD" box="[254,341,869,894]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="11" pageNumber="12">Fig. 21</figureCitation>
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), by shape of appendix (bifid, i.e. with long subdistal branch;
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<figureCitation id="1324C2FB8539073BFBBCFCE02FECFCDD" box="[1089,1179,869,894]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="11" pageNumber="12">Figs 20</figureCitation>
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,
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<figureCitation id="1324C2FB8539073BFB51FCE32FBEFCDD" box="[1196,1225,870,894]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,264,1907,1929]" captionTargetBox="[151,1436,303,1885]" captionTargetId="figure@13.[151,1436,303,1885]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Calapnita lehi sp. nov., SEM micrographs (ZFMK, Ar 15965 – 66). 26 – 27. Male cheliceral apophyses. 28. Right procursus, prolateral view. 29. Tip of bulbal appendix. 30. Comb-hair on male tarsus 4. 31. Male ALS. 32. Epigynum, ventral view. Scale lines: 10 µm (26, 27, 30, 31), 50 µm (28, 29, 32)." httpUri="https://zenodo.org/record/273092/files/figure.png" pageId="11" pageNumber="12">29</figureCitation>
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) and by shape of procursus tip (with seemingly hinged prolatero-ventral process;
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<figureCitation id="1324C2FB8539073BFC8AFC0C288BFC01" box="[887,1020,905,930]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="11" pageNumber="12">Figs 20–22</figureCitation>
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); females are difficult to distinguish externally from other representatives of the
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<taxonomicName id="4C1FA5FD8539073BFD78FC282982FC66" box="[645,757,941,965]" class="Arachnida" family="Pholcidae" genus="Calapnita" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="phyllicola">
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<emphasis id="B96B026C8539073BFD78FC282982FC66" box="[645,757,941,965]" italics="true" pageId="11" pageNumber="12">phyllicola</emphasis>
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</taxonomicName>
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group; internally they can be distinguished by almost round pore plates and absence of membranous ‘sac’ (
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<figureCitation id="1324C2FB8539073BFD5EFC542981FC49" box="[675,758,977,1002]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="11" pageNumber="12">Fig. 25</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3058DF58539073BFF3AFC7028DFFB91" pageId="11" pageNumber="12" type="etymology">
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<paragraph id="8BA0DE7E8539073BFF3AFC7028DFFB91" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
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<emphasis id="B96B026C8539073BFF3AFC702A39FBAD" bold="true" box="[199,334,1013,1038]" pageId="11" pageNumber="12">Etymology.</emphasis>
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The species is named for Dr. Charles Leh Moi Ung from
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<collectingCountry id="F3089EEE8539073BFBCDFC702FE6FBAD" box="[1072,1169,1013,1038]" name="Malaysia" pageId="11" pageNumber="12">
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<collectingRegion id="49DB109C8539073BFBCDFC702FE6FBAD" box="[1072,1169,1013,1038]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="12">Sarawak</collectingRegion>
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</collectingCountry>
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Museum Kuching, in recognition of his help before and during the Borneo expedition 2014.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3058DF58539073BFF3AFBB82E11FA8D" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8BA0DE7E8539073BFF3AFBB82A0FFB3D" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
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<materialsCitation id="3B77D4238539073BFF3AFBB82A0FFB3D" ID-GBIF-Occurrence="1424726737" collectingDate="2014-07-22" collectionCode="ZFMK" collectorName="B. A. Huber & S. B. Huber" country="Malaysia" elevation="105" latitude="4.207" location="Ar" longLatPrecision="78" longitude="114.045" municipality="Lambir Hills National Park" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B96B026C8539073BFF3AFBB82944FBF5" bold="true" box="[199,563,1085,1110]" pageId="11" pageNumber="12">
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Material examined.
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<typeStatus id="54A460DC8539073BFE3BFBB82944FBF5" box="[454,563,1085,1110]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
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</emphasis>
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. MALAYSIA-BORNEO:
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<specimenCount id="9D1915F78539073BFC90FBBB28F3FBF6" box="[877,900,1086,1109]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
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,
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<collectionCode id="ED0E46BB8539073BFC6AFBBB2892FBF6" box="[919,997,1086,1109]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="12">ZFMK</collectionCode>
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(
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<location id="8EC088A58539073BFC01FBBB2F6CFBF5" LSID="urn:lsid:plazi:treatment:03B66F688539073DFF6AFD53289AFF5B:8EC088A58539073BFC01FBBB2F6CFBF5" box="[1020,1051,1086,1110]" country="Malaysia" latitude="4.207" longLatPrecision="78" longitude="114.045" municipality="Lambir Hills National Park" name="Ar" pageId="11" pageNumber="12">Ar</location>
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15964),
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<collectingCountry id="F3089EEE8539073BFB73FBB82F86FBF5" box="[1166,1265,1085,1110]" name="Malaysia" pageId="11" pageNumber="12">Sarawak</collectingCountry>
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,
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<collectingMunicipality id="6BC444048539073BFAF8FBB82A4EFBD9" pageId="11" pageNumber="12">Lambir Hills National Park</collectingMunicipality>
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||
(4.198–
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||
<geoCoordinate id="EE2BB8B98539073BFE63FBE72A8AFBD9" box="[414,509,1122,1146]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="55" value="4.207">4.207°N</geoCoordinate>
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||
, 114.034–
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||
<geoCoordinate id="EE2BB8B98539073BFD86FBE42985FBD9" box="[635,754,1121,1146]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="55" value="114.045">114.045°E</geoCoordinate>
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||
),
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||
<elevation id="0032394D8539073BFCF7FBE428B2FBD9" box="[778,965,1121,1146]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="11" pageNumber="12" unit="m" value="105.0" valueMax="150.0" valueMin="60.0">
|
||
<quantity id="4CE7739B8539073BFCF7FBE428F5FBDA" box="[778,898,1121,1146]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="11" pageNumber="12" unit="m" value="105.0" valueMax="150.0" valueMin="60.0">60–150 m</quantity>
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a.s.l.
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</elevation>
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, underside of leaves,
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||
<date id="FFA1F8BE8539073BFB2CFBE42E27FBD9" box="[1233,1360,1121,1146]" pageId="11" pageNumber="12" value="2014-07-22">
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||
<collectingDate id="EFE501568539073BFB2CFBE42E27FBD9" box="[1233,1360,1121,1146]" pageId="11" pageNumber="12" value="2014-07-22">22.vii.2014</collectingDate>
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||
</date>
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||
(
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||
<collectorName id="26EABBA88539073BFA9AFBE72BA8FB3D" pageId="11" pageNumber="12">B.A. Huber</collectorName>
|
||
,
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||
<collectorName id="26EABBA88539073BFF17FB032A1BFB3D" box="[234,364,1157,1182]" pageId="11" pageNumber="12">S.B. Huber</collectorName>
|
||
).
|
||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BA0DE7E8539073BFF3AFB2C2E11FA8D" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
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||
<materialsCitation id="3B77D4238539073BFF3AFB2C2B8AFB45" ID-GBIF-Occurrence="1424726733" collectingDate="2014-07-22" collectionCode="ZFMK, SMK" collectorName="B. A. Huber & S. B. Huber" country="Malaysia" elevation="105" latitude="4.207" location="Ar" longLatPrecision="78" longitude="114.045" municipality="Lambir Hills National Park" pageId="11" pageNumber="12" specimenCount="15" specimenCount-female="10" specimenCount-male="5">
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||
<emphasis id="B96B026C8539073BFF3AFB2C2A09FB61" bold="true" box="[199,382,1193,1218]" pageId="11" pageNumber="12">Other material</emphasis>
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||
. MALAYSIA-BORNEO:
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||
<specimenCount id="9D1915F78539073BFD52FB2F29A5FB61" box="[687,722,1194,1218]" pageId="11" pageNumber="12" type="male">4♂</specimenCount>
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||
<specimenCount id="9D1915F78539073BFD20FB2F2876FB61" box="[733,769,1194,1218]" pageId="11" pageNumber="12" type="female">9♀</specimenCount>
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||
,
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||
<collectionCode id="ED0E46BB8539073BFCECFB2F2828FB62" box="[785,863,1194,1217]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="12">ZFMK</collectionCode>
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(Ar 15965–66), and
|
||
<specimenCount id="9D1915F78539073BFBA5FB2F2F0CFB61" box="[1112,1147,1194,1218]" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
|
||
<specimenCount id="9D1915F78539073BFB78FB2F2FDFFB61" box="[1157,1192,1194,1218]" pageId="11" pageNumber="12" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED0E46BB8539073BFB47FB2F2F8EFB61" box="[1210,1273,1194,1218]" country="Malaysia" httpUri="http://grbio.org/cool/cg7m-xnpx" name="Sarawak Museum" pageId="11" pageNumber="12">SMK</collectionCode>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B77D4238539073BFEF1FB4B28B9FB45" ID-GBIF-Occurrence="1424726744" box="[268,974,1229,1254]" collectingDate="2014-07-22" collectionCode="ZFMK" collectorName="Bor" country="Malaysia" elevation="105" latitude="4.207" location="Ar" longLatPrecision="78" longitude="114.045" municipality="Lambir Hills National Park" pageId="11" pageNumber="12" specimenCount="3" specimenCount-female="2">
|
||
<specimenCount id="9D1915F78539073BFEF1FB4B2A58FB45" box="[268,303,1230,1254]" pageId="11" pageNumber="12" type="female">2♀</specimenCount>
|
||
<specimenCount id="9D1915F78539073BFEC6FB4B2A09FB45" box="[315,382,1229,1254]" pageId="11" pageNumber="12" type="generic">1 juv.</specimenCount>
|
||
in absolute ethanol,
|
||
<collectionCode id="ED0E46BB8539073BFD85FB4B29B1FB46" box="[632,710,1230,1253]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="12">ZFMK</collectionCode>
|
||
(
|
||
<collectorName id="26EABBA88539073BFD27FB4B2872FB45" box="[730,773,1230,1254]" pageId="11" pageNumber="12">Bor</collectorName>
|
||
199), same data
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B77D4238539073BFC20FB4B29E3FA8D" ID-GBIF-Occurrence="1424726754" collectingDate="2014-07-24" collectionCode="ZFMK" collectorName="B. A. Huber & S. B. Huber" country="Malaysia" county="Lagang Cave" elevation="60" latitude="4.051" location="Ar" longLatPrecision="78" longitude="114.822" municipality="Gunung Mulu National Park" pageId="11" pageNumber="12" specimenCount="14" specimenCount-female="5" specimenCount-male="8">
|
||
<specimenCount id="9D1915F78539073BFC20FB4B2F76FB45" box="[989,1025,1230,1254]" pageId="11" pageNumber="12" type="male">8♂</specimenCount>
|
||
<specimenCount id="9D1915F78539073BFBF1FB482F58FB45" box="[1036,1071,1229,1254]" pageId="11" pageNumber="12" type="female">5♀</specimenCount>
|
||
<specimenCount id="9D1915F78539073BFBC6FB4B2FF6FB45" box="[1083,1153,1229,1254]" pageId="11" pageNumber="12" type="generic">1 juv.</specimenCount>
|
||
,
|
||
<collectionCode id="ED0E46BB8539073BFB6DFB4B2FA9FB46" box="[1168,1246,1230,1253]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="12">ZFMK</collectionCode>
|
||
(
|
||
<location id="8EC088A58539073BFB0CFB4B2E67FB45" LSID="urn:lsid:plazi:treatment:03B66F688539073DFF6AFD53289AFF5B:8EC088A58539073BFB0CFB4B2E67FB45" box="[1265,1296,1230,1254]" country="Malaysia" county="Lagang Cave" latitude="4.051" longLatPrecision="78" longitude="114.822" municipality="Gunung Mulu National Park" name="Ar" pageId="11" pageNumber="12">Ar</location>
|
||
15967–68),
|
||
<collectingCountry id="F3089EEE8539073BFF6AFB742B8DFAA9" box="[151,250,1265,1290]" name="Malaysia" pageId="11" pageNumber="12">Sarawak</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BC444048539073BFEFBFB77293AFAA9" box="[262,589,1265,1290]" pageId="11" pageNumber="12">Gunung Mulu National Park</collectingMunicipality>
|
||
, forest near
|
||
<collectingCounty id="62C1A6F28539073BFD27FB772807FAA9" box="[730,880,1266,1290]" pageId="11" pageNumber="12">Lagang Cave</collectingCounty>
|
||
(
|
||
<geoCoordinate id="EE2BB8B98539073BFC7CFB742894FAA9" box="[897,995,1265,1290]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="55" value="4.051">4.051°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2BB8B98539073BFC0DFB772F1DFAA9" box="[1008,1130,1266,1290]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="55" value="114.822">114.822°E</geoCoordinate>
|
||
),
|
||
<elevation id="0032394D8539073BFB82FB742F80FAA9" box="[1151,1271,1265,1290]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="60.0">
|
||
<quantity id="4CE7739B8539073BFB82FB742FCEFAAA" box="[1151,1209,1265,1290]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="60.0">60 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, undersides of leaves,
|
||
<date id="FFA1F8BE8539073BFF16FA902A1DFA8D" box="[235,362,1301,1326]" pageId="11" pageNumber="12" value="2014-07-24">
|
||
<collectingDate id="EFE501568539073BFF16FA902A1DFA8D" box="[235,362,1301,1326]" pageId="11" pageNumber="12" value="2014-07-24">24.vii.2014</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26EABBA88539073BFE86FA932A88FA8D" box="[379,511,1301,1326]" pageId="11" pageNumber="12">B.A. Huber</collectorName>
|
||
,
|
||
<collectorName id="26EABBA88539073BFDF4FA9329FBFA8D" box="[521,652,1301,1326]" pageId="11" pageNumber="12">S.B. Huber</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B77D4238539073BFD62FA932E15FA8D" ID-GBIF-Occurrence="1424726757" box="[671,1378,1301,1326]" collectingDate="2014-07-24" collectionCode="ZFMK" collectorName="Bor" country="Malaysia" county="Lagang Cave" elevation="60" latitude="4.051" location="Ar" longLatPrecision="78" longitude="114.822" municipality="Gunung Mulu National Park" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCount id="9D1915F78539073BFD62FA9329B5FA8D" box="[671,706,1302,1326]" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
|
||
<specimenCount id="9D1915F78539073BFD34FA93299BFA8D" box="[713,748,1302,1326]" pageId="11" pageNumber="12" type="female">1♀</specimenCount>
|
||
7 juvs in absolute ethanol,
|
||
<collectionCode id="ED0E46BB8539073BFBE2FA932F1AFA8E" box="[1055,1133,1302,1325]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="12">ZFMK</collectionCode>
|
||
(
|
||
<collectorName id="26EABBA88539073BFB81FA932FDFFA8D" box="[1148,1192,1302,1326]" pageId="11" pageNumber="12">Bor</collectorName>
|
||
236), same data
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3058DF58539073DFF3AFABC289AFF5B" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BA0DE7E8539073BFF3AFABC296FFAF1" blockId="11.[151,1437,833,2010]" box="[199,536,1337,1362]" pageId="11" pageNumber="12">
|
||
<emphasis id="B96B026C8539073BFF3AFABC2AEBFAF1" bold="true" box="[199,412,1337,1362]" pageId="11" pageNumber="12">Description. Male</emphasis>
|
||
(
|
||
<typeStatus id="54A460DC8539073BFE56FABC2967FAF1" box="[427,528,1337,1362]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AFADB2977FA1D" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">MEASUREMENTS. Total body length 5.0, carapace width 0.9. Leg 1: 41.7 (9.6 + 0.4 + 9.8 + 19.9 + 2.0), tibia 2: 6.9, tibia 3: 3.9, tibia 4: 5.9; tibia 1 L/d: 122. Distance PME-PME 240 µm, diameter PME 120 µm, distance PME- ALE ~30 µm; no trace of AME.</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AFA4F291EF9A5" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">COLOR. Entire animal mostly whitish to pale ochre-yellow, patellae and tibia-metatarsus joints slightly darker brown, abdomen monochromous whitish.</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AF99728BAF9ED" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
|
||
BODY. Habitus as in
|
||
<figureCitation id="1324C2FB8539073BFE53F9942A86F989" box="[430,497,1553,1578]" captionStart="FIGURES 3 – 12" captionStartId="9.[151,264,1890,1912]" captionTargetBox="[151,1436,337,1869]" captionTargetId="figure@9.[151,1436,337,1869]" captionTargetPageId="9" captionText="FIGURES 3 – 12. Live specimens, Calapnita phyllicola group. 3 – 6. C. lehi sp. nov., ♂ and ♀ with egg-sac from Lambir (3 – 4); ♂ and penultimate instar ♂ from Gunung Mulu (5 – 6). 7 – 8. C. kubah sp. nov., ♂ and ♀ with egg-sac from Kubah. 9. C. deelemanae Huber, ♀ with egg-sac from Poring. 10. C. subphyllicola Deeleman-Reinhold, ♀ with egg-sac from Malagos. 11 – 12. C. anai sp. nov., ♂ from MacRitchie, ♀ with egg-sac from Lembah Anai." httpUri="https://zenodo.org/record/273089/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
|
||
; ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum as wide as long (0.56), unmodified.
|
||
</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AF9DF2F68F935" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
|
||
CHELICERAE. As in
|
||
<figureCitation id="1324C2FB8539073BFE5BF9DC2A80F9D1" box="[422,503,1625,1650]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="11" pageNumber="12">Fig. 23</figureCitation>
|
||
, with pair of simple scaly apophyses near lamellae (
|
||
<figureCitation id="1324C2FB8539073BFBC1F9DC2FCCF9D1" box="[1084,1211,1625,1650]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,264,1907,1929]" captionTargetBox="[151,1436,303,1885]" captionTargetId="figure@13.[151,1436,303,1885]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Calapnita lehi sp. nov., SEM micrographs (ZFMK, Ar 15965 – 66). 26 – 27. Male cheliceral apophyses. 28. Right procursus, prolateral view. 29. Tip of bulbal appendix. 30. Comb-hair on male tarsus 4. 31. Male ALS. 32. Epigynum, ventral view. Scale lines: 10 µm (26, 27, 30, 31), 50 µm (28, 29, 32)." httpUri="https://zenodo.org/record/273092/files/figure.png" pageId="11" pageNumber="12">Figs 26, 27</figureCitation>
|
||
) and pair of distinct but whitish proximal humps; without modified hairs; without stridulatory ridges.
|
||
</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AF9272F37F886" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
|
||
PALPS. As in
|
||
<figureCitation id="1324C2FB8539073BFE9EF9242A92F919" box="[355,485,1697,1722]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="11" pageNumber="12">Figs 20–21</figureCitation>
|
||
; coxa unmodified; trochanter with simple retrolatero-ventral apophysis; femur with distinctive ventral process proximally; tarsus with short conical whitish process, procursus with small transparent rounded process from ‘knee’, with apparently hinged distal element; bulb with long weakly sclerotized embolus, large appendix with distinctive bifid tip (
|
||
<figureCitation id="1324C2FB8539073BFD9DF88829C3F885" box="[608,692,1805,1830]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,264,1907,1929]" captionTargetBox="[151,1436,303,1885]" captionTargetId="figure@13.[151,1436,303,1885]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Calapnita lehi sp. nov., SEM micrographs (ZFMK, Ar 15965 – 66). 26 – 27. Male cheliceral apophyses. 28. Right procursus, prolateral view. 29. Tip of bulbal appendix. 30. Comb-hair on male tarsus 4. 31. Male ALS. 32. Epigynum, ventral view. Scale lines: 10 µm (26, 27, 30, 31), 50 µm (28, 29, 32)." httpUri="https://zenodo.org/record/273092/files/figure.png" pageId="11" pageNumber="12">Fig. 29</figureCitation>
|
||
), small membranous basal process.
|
||
</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AF8B72AE0F831" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments not visible in dissecting microscope.</paragraph>
|
||
<paragraph id="8BA0DE7E8539073BFF3AF81828E2F879" blockId="11.[151,1437,833,2010]" pageId="11" pageNumber="12">
|
||
<emphasis id="B96B026C8539073BFF3AF8182A73F815" bold="true" box="[199,260,1949,1974]" pageId="11" pageNumber="12">Male</emphasis>
|
||
(variation). Tibia
|
||
<quantity id="4CE7739B8539073BFE2DF81B2A8CF815" box="[464,507,1949,1974]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="12" unit="in" value="1.0">1 in</quantity>
|
||
12 other males: 8.4–10.0 (mean 9.2). Males from Gunung Mulu tend to be smaller than males from Lambir Hills (tibia 1 lengths: 8.4–9.2
|
||
<emphasis id="B96B026C8539073BFD01F847286DF87A" box="[764,794,1986,2009]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
|
||
9.4–10.0).
|
||
</paragraph>
|
||
<caption id="DF608EF6853E073CFF6AF9422F33F899" httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1431,193,1708]" targetPageId="12">
|
||
<paragraph id="8BA0DE7E853E073CFF6AF9422F33F899" blockId="12.[151,1436,1735,1850]" pageId="12" pageNumber="13">
|
||
<emphasis id="B96B026C853E073CFF6AF9422A27F97E" bold="true" box="[151,336,1735,1757]" pageId="12" pageNumber="13">FIGURES 20–25</emphasis>
|
||
.
|
||
<taxonomicName id="4C1FA5FD853E073CFEA2F9422A83F97E" box="[351,500,1735,1757]" class="Arachnida" family="Pholcidae" genus="Calapnita" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lehi" status="sp. nov.">
|
||
<emphasis id="B96B026C853E073CFEA2F9422A83F97E" box="[351,500,1735,1757]" italics="true" pageId="12" pageNumber="13">Calapnita lehi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B026C853E073CFE00F9422927F97E" bold="true" box="[509,592,1735,1757]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A258BF17853E073CFE00F9422927F97E" box="[509,592,1735,1757]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZFMK, Ar 15965–66).
|
||
<emphasis id="B96B026C853E073CFCA4F94228EEF97E" bold="true" box="[857,921,1735,1757]" pageId="12" pageNumber="13">20–21</emphasis>
|
||
. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘knee’).
|
||
<emphasis id="B96B026C853E073CFB99F9632F0AF95F" bold="true" box="[1124,1149,1766,1788]" pageId="12" pageNumber="13">22</emphasis>
|
||
. Procursus, dorsal view.
|
||
<emphasis id="B96B026C853E073CFA81F9632EE2F95F" bold="true" box="[1404,1429,1766,1788]" pageId="12" pageNumber="13">23</emphasis>
|
||
. Male chelicerae, frontal view.
|
||
<emphasis id="B96B026C853E073CFE21F880296CF8B8" bold="true" box="[476,539,1797,1819]" pageId="12" pageNumber="13">24–25</emphasis>
|
||
. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23–25), 0.5 mm (20–22).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA0DE7E853E073CFF3AF8E32E05F848" blockId="12.[151,1436,1894,2027]" pageId="12" pageNumber="13">
|
||
<emphasis id="B96B026C853E073CFF3AF8E32A69F8DC" bold="true" box="[199,286,1894,1919]" pageId="12" pageNumber="13">Female</emphasis>
|
||
. In general similar to male; eye triads slightly closer together (distance PME-PME 200 µm). Tibia
|
||
<quantity id="4CE7739B853E073CFA8CF8E22EEBF8DC" box="[1393,1436,1894,1919]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="13" unit="in" value="1.0">1 in</quantity>
|
||
<specimenCount id="9D1915F7853E073CFF6AF80E2A60F800" box="[151,279,1930,1955]" pageId="12" pageNumber="13" type="female">14 females</specimenCount>
|
||
: 7.0–8.0 (mean 7.5). Females from Gunung Mulu tend to be smaller than females from Lambir Hills (tibia 1 lengths: 7.0–7.4
|
||
<emphasis id="B96B026C853E073CFE4DF82A2AB8F865" box="[432,463,1967,1990]" italics="true" pageId="12" pageNumber="13">vs.</emphasis>
|
||
7.3–8.0). Epigynum weakly sclerotized, internal anterior arch visible through cuticle (
|
||
<figureCitation id="1324C2FB853E073CFF62F8572B83F848" box="[159,244,2002,2027]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="12" pageNumber="13">Fig. 24</figureCitation>
|
||
); with simple short posterior ‘knob’ on extensible wide scape (
|
||
<figureCitation id="1324C2FB853E073CFC49F8572F7FF848" box="[948,1032,2002,2027]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,264,1907,1929]" captionTargetBox="[151,1436,303,1885]" captionTargetId="figure@13.[151,1436,303,1885]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Calapnita lehi sp. nov., SEM micrographs (ZFMK, Ar 15965 – 66). 26 – 27. Male cheliceral apophyses. 28. Right procursus, prolateral view. 29. Tip of bulbal appendix. 30. Comb-hair on male tarsus 4. 31. Male ALS. 32. Epigynum, ventral view. Scale lines: 10 µm (26, 27, 30, 31), 50 µm (28, 29, 32)." httpUri="https://zenodo.org/record/273092/files/figure.png" pageId="12" pageNumber="13">Fig. 32</figureCitation>
|
||
); internal genitalia as in
|
||
<figureCitation id="1324C2FB853E073CFAE1F8572E19F848" box="[1308,1390,2002,2027]" captionStart="FIGURES 20 – 25" captionStartId="12.[151,264,1735,1757]" captionTargetBox="[151,1431,193,1708]" captionTargetId="figure@12.[151,1436,193,1714]" captionTargetPageId="12" captionText="FIGURES 20 – 25. Calapnita lehi sp. nov. (ZFMK, Ar 15965 – 66). 20 – 21. Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘ knee’). 22. Procursus, dorsal view. 23. Male chelicerae, frontal view. 24 – 25. Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23 – 25), 0.5 mm (20 – 22)." httpUri="https://zenodo.org/record/273091/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
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.
|
||
</paragraph>
|
||
<paragraph id="8BA0DE7E853F073DFF3AFF12289AFF5B" blockId="13.[151,1436,151,248]" pageId="13" pageNumber="14">
|
||
<emphasis id="B96B026C853F073DFF3AFF122AF7FF13" bold="true" box="[199,384,151,176]" pageId="13" pageNumber="14">Natural history</emphasis>
|
||
. At Lambir, most specimens were collected from palm leaves; at Gunung Mulu, they were found on large leaves of various monocot and dicot species. Two egg-sacs contained 8 and 16 eggs respectively.
|
||
<emphasis id="B96B026C853F073DFF3AFF5A2A2EFF5B" bold="true" box="[199,345,223,248]" pageId="13" pageNumber="14">Distribution</emphasis>
|
||
. Known from two localities in eastern Sarawak (
|
||
<figureCitation id="1324C2FB853F073DFC83FF5A2897FF5B" box="[894,992,223,248]" captionStart="FIGURE 281" captionStartId="54.[151,250,1037,1059]" captionTargetBox="[150,1436,267,1016]" captionTargetId="figure@54.[150,1436,267,1016]" captionTargetPageId="54" captionText="FIGURE 281. Known distribution of the Calapnita phyllicola group, part 1 (cf. Fig. 282). The western Java record is based on a single female of uncertain identity. For details on the undescribed species in RMNH see C. phasmoides." httpUri="https://zenodo.org/record/273116/files/figure.png" pageId="13" pageNumber="14">Fig. 281</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |