Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae) Author Bernhard A. Huber text Zootaxa 2017 4219 1 1 63 journal article 37319 10.5281/zenodo.273086 0d2332a5-1968-49d7-a570-f31d386eccb3 1175-5326 273086 0FA0F51A-3868-4F13-A93D-E34CA5A689F8 Calapnita lehi sp. nov. Figs 3–6 , 20–32 Diagnosis. Males are easily distinguished from congeners by modified male palpal femur (distinctive ventral process; Fig. 21 ), by shape of appendix (bifid, i.e. with long subdistal branch; Figs 20 , 29 ) and by shape of procursus tip (with seemingly hinged prolatero-ventral process; Figs 20–22 ); females are difficult to distinguish externally from other representatives of the phyllicola group; internally they can be distinguished by almost round pore plates and absence of membranous ‘sac’ ( Fig. 25 ). Etymology. The species is named for Dr. Charles Leh Moi Ung from Sarawak Museum Kuching, in recognition of his help before and during the Borneo expedition 2014. Material examined. Holotype . MALAYSIA-BORNEO: ♂ , ZFMK ( Ar 15964), Sarawak , Lambir Hills National Park (4.198– 4.207°N , 114.034– 114.045°E ), 60–150 m a.s.l. , underside of leaves, 22.vii.2014 ( B.A. Huber , S.B. Huber ). Other material . MALAYSIA-BORNEO: 4♂ 9♀ , ZFMK (Ar 15965–66), and 1♂ 1♀ , SMK , same data as holotype ; 2♀ 1 juv. in absolute ethanol, ZFMK ( Bor 199), same data . 8♂ 5♀ 1 juv. , ZFMK ( Ar 15967–68), Sarawak , Gunung Mulu National Park , forest near Lagang Cave ( 4.051°N , 114.822°E ), 60 m a.s.l. , undersides of leaves, 24.vii.2014 ( B.A. Huber , S.B. Huber ) ; 1♂ 1♀ 7 juvs in absolute ethanol, ZFMK ( Bor 236), same data . Description. Male ( holotype ) MEASUREMENTS. Total body length 5.0, carapace width 0.9. Leg 1: 41.7 (9.6 + 0.4 + 9.8 + 19.9 + 2.0), tibia 2: 6.9, tibia 3: 3.9, tibia 4: 5.9; tibia 1 L/d: 122. Distance PME-PME 240 µm, diameter PME 120 µm, distance PME- ALE ~30 µm; no trace of AME. COLOR. Entire animal mostly whitish to pale ochre-yellow, patellae and tibia-metatarsus joints slightly darker brown, abdomen monochromous whitish. BODY. Habitus as in Fig. 3 ; ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum as wide as long (0.56), unmodified. CHELICERAE. As in Fig. 23 , with pair of simple scaly apophyses near lamellae ( Figs 26, 27 ) and pair of distinct but whitish proximal humps; without modified hairs; without stridulatory ridges. PALPS. As in Figs 20–21 ; coxa unmodified; trochanter with simple retrolatero-ventral apophysis; femur with distinctive ventral process proximally; tarsus with short conical whitish process, procursus with small transparent rounded process from ‘knee’, with apparently hinged distal element; bulb with long weakly sclerotized embolus, large appendix with distinctive bifid tip ( Fig. 29 ), small membranous basal process. LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments not visible in dissecting microscope. Male (variation). Tibia 1 in 12 other males: 8.4–10.0 (mean 9.2). Males from Gunung Mulu tend to be smaller than males from Lambir Hills (tibia 1 lengths: 8.4–9.2 vs. 9.4–10.0). FIGURES 20–25 . Calapnita lehi sp. nov. (ZFMK, Ar 15965–66). 20–21 . Left male palp, prolateral and retrolateral views (asterisk: membranous bulbal process; arrow points at short membranous process from ‘knee’). 22 . Procursus, dorsal view. 23 . Male chelicerae, frontal view. 24–25 . Cleared female genitalia, ventral and dorsal views. a, appendix; b, genital bulb; e, embolus; f, femur; p, procursus; tr, trochanter. Scale lines: 0.3 mm (23–25), 0.5 mm (20–22). Female . In general similar to male; eye triads slightly closer together (distance PME-PME 200 µm). Tibia 1 in 14 females : 7.0–8.0 (mean 7.5). Females from Gunung Mulu tend to be smaller than females from Lambir Hills (tibia 1 lengths: 7.0–7.4 vs. 7.3–8.0). Epigynum weakly sclerotized, internal anterior arch visible through cuticle ( Fig. 24 ); with simple short posterior ‘knob’ on extensible wide scape ( Fig. 32 ); internal genitalia as in Fig. 25 . Natural history . At Lambir, most specimens were collected from palm leaves; at Gunung Mulu, they were found on large leaves of various monocot and dicot species. Two egg-sacs contained 8 and 16 eggs respectively. Distribution . Known from two localities in eastern Sarawak ( Fig. 281 ).