320 lines
42 KiB
XML
320 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1129.90535" ID-GBIF-Dataset="8334b5dc-6286-4ccc-b358-f84d2f1c36f6" ID-PMC="PMC9836718" ID-Pensoft-Pub="1313-2970-1129-109" ID-Pensoft-UUID="E06A760431B65AEBAB5B1B370AE54983" ID-PubMed="36761846" ID-ZooBank="3D73FEE944FD4DA98E2BC07536739901" ModsDocID="1313-2970-1129-109" checkinTime="1668554642443" checkinUser="pensoft" docAuthor="Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai & Rujirawan, Attapol" docDate="2022" docId="7E67190E06B0564CB4C8E5A323705A73" docLanguage="en" docName="ZooKeys 1129: 109-162" docOrigin="ZooKeys 1129" docPubDate="2022-11-15" docSource="http://dx.doi.org/10.3897/zookeys.1129.90535" docTitle="Cyrtodactylus kochangensis Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan 2022, sp. nov." docType="treatment" docUuid="96DF655B-BBB7-4C06-9418-744EBCD14703" docUuidSource="ZooBank" docVersion="5" id="E06A760431B65AEBAB5B1B370AE54983" lastPageNumber="109" masterDocId="E06A760431B65AEBAB5B1B370AE54983" masterDocTitle="Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand" masterLastPageNumber="162" masterPageNumber="109" pageNumber="109" updateTime="1732481513498" updateUser="ExternalLinkService">
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<mods:title id="CFD948BFE26FCCE2B6B178D4963B83DC">Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand</mods:title>
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<mods:namePart id="9D384F85E513DD8210B6296CB44698A4">Grismer, L. Lee</mods:namePart>
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<mods:affiliation id="EF9D7E7A4A297242CCA61121D453D2B2">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA & Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA</mods:affiliation>
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<mods:namePart id="DD1F3B9421227D29C0F5DB8BB00024E6">Aowphol, Anchalee</mods:namePart>
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<mods:nameIdentifier id="9B42235D3859787A6109A4E8F389C403" type="ORCID">https://orcid.org/0000-0001-9504-4601</mods:nameIdentifier>
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<mods:affiliation id="5E6E9464C280486459B4FEB0C40099C2">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="6E883CEF5CB8399C7D7632A314A502E7" type="personal">
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<mods:roleTerm id="964CD8D18707F54D8DA2DCBCE23A61DF">Author</mods:roleTerm>
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<mods:namePart id="98E23E60C937F256A0D928ABE735BFF1">Yodthong, Siriporn</mods:namePart>
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<mods:nameIdentifier id="22BAAE2C97BFC9E1970589933BD5B1D9" type="ORCID">https://orcid.org/0000-0001-6577-5358</mods:nameIdentifier>
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<mods:affiliation id="F0BA35F7145F8F6D3BAF3EBFA0A97018">Department of Biology, Faculty of Science, Thaksin University, Pa Phayom, Phattalung 93210, Thailand</mods:affiliation>
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<mods:namePart id="4FADD4FC9786D0FD0F2263506B290CBA">Ampai, Natee</mods:namePart>
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<mods:nameIdentifier id="982CD5E7310BBAC784753DAD9DE8BABA" type="ORCID">https://orcid.org/0000-0001-8562-299X</mods:nameIdentifier>
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<mods:affiliation id="3B3304AFDEEDE4483C828C03E26E1D4D">Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok 10110, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="B60BC002AE0012440FDBFA025D9C51C5" type="personal">
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<mods:roleTerm id="ED366B63AA3F1E29412EF6700B7728CE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9D57E2262977F3FDB57F4061731D3126">Termprayoon, Korkhwan</mods:namePart>
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<mods:affiliation id="3454CD37A249059A00F4E1643CAD5BA0">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="8C4546895214C723F26A4562E2902C6A" type="personal">
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<mods:roleTerm id="8BE26EB11B95BA08227D5BB89D45425C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FA789EFB2CBAE4F0C258D19D0938F74A">Aksornneam, Akrachai</mods:namePart>
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<mods:nameIdentifier id="4C0FE0EB55A15385081A3783CF3A38A5" type="ORCID">https://orcid.org/0000-0003-4780-376X</mods:nameIdentifier>
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<mods:affiliation id="E01BB375987E1BA8D48AB788F443CBBE">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="47495CCC8BCDBFAAEF7377686E97215B" type="personal">
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<mods:namePart id="C938EBD9703E6F673CED38876393234F">Rujirawan, Attapol</mods:namePart>
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<mods:nameIdentifier id="BAEDFE864E9286F7C675CEA2CA91571E" type="ORCID">https://orcid.org/0000-0001-9179-6910</mods:nameIdentifier>
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<mods:affiliation id="185EA2CDA50D973AE90B1ECF63F4A9D7">Animal Systematics and Ecology Speciality Research Unit, Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:nameIdentifier id="F9F07A1F7AFC631299DA7F1B7580751C" type="email">fsciapr@ku.ac.th</mods:nameIdentifier>
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<mods:typeOfResource id="0DFD7B97341E4F9420ECC16EF2BE46FF">text</mods:typeOfResource>
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<mods:title id="7D0B96EEC4C4D81B145CF5945DC0F7FC">ZooKeys</mods:title>
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<mods:date id="8FAF09E38479055CD5E8F5BD39C09F92">2022</mods:date>
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<mods:number id="F260E03C6DD66D60887717EC22337F81">2022-11-15</mods:number>
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<mods:detail id="FD8703F1E890277E4C50BB82BD7EB6AE" type="volume">
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<mods:number id="1CA32231929668A673CCA931C731A5BF">1129</mods:number>
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<mods:start id="C0F0C6ADD16D770A4420BBBB8D07D03C">109</mods:start>
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<mods:url id="287CA5A6BEC4FFF52FB7BEB9907A53D6">http://dx.doi.org/10.3897/zookeys.1129.90535</mods:url>
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<mods:classification id="54ED65C6CC3F9DBAEDBFC7BEFEAA56DF">journal article</mods:classification>
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<mods:identifier id="CF7C6108130F405680E0A5418B3B8803" type="DOI">http://dx.doi.org/10.3897/zookeys.1129.90535</mods:identifier>
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<mods:identifier id="65553F868B962C65778191E690764974" type="Pensoft-Pub">1313-2970-1129-109</mods:identifier>
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<treatment id="7E67190E06B0564CB4C8E5A323705A73" ID-GBIF-Taxon="205021301" LSID="urn:lsid:zoobank.org:act:96DF655B-BBB7-4C06-9418-744EBCD14703" httpUri="http://treatment.plazi.org/id/7E67190E06B0564CB4C8E5A323705A73" lastPageNumber="109" pageId="0" pageNumber="109" scope_class="Squamata" scope_family="Gekkonidae">
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<subSubSection id="505CBA45568948FA763A791DBFAB8EFC" pageId="0" pageNumber="109" type="nomenclature">
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<paragraph id="BCA976E55A0CE05AA65AE267572465B0" pageId="0" pageNumber="109">
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<taxonomicName id="D91F77F9622469923096E8AAB98A641E" ID-CoL="C6X2R" LSID="https://zoobank.org/96DF655B-BBB7-4C06-9418-744EBCD14703" authority="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan, 2022" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis" status="sp. nov.">Cyrtodactylus kochangensis</taxonomicName>
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<taxonomicNameLabel id="5F9B239FB1320E6EDB0FE0D7AD87F633" pageId="0" pageNumber="109">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="865333BAAB96D854A19AD42ABBB8BC2B" pageId="0" pageNumber="109" type="description">
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<paragraph id="0D37497E3C7790D7D2A3AE993B4A11C5" pageId="0" pageNumber="109">
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<figureCitation id="AADBD7D6E333CA080B23ECB16FBDDB0D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Holotype of Cyrtodactylus kochangensis sp. nov. ZMKU R 00945 (field no. AA 04439) from Thailand, Ranong Province, Mueng Ranong District, Ko Phayam Subdistrict, Ko Chang A dorsal view B ventral view C dorsal view of head D ventral view of femoral and precloacal regions E dorsal view of tail F ventral view of tail G holotype in life." figureDoi="10.3897/zookeys.1129.90535.figure10" httpUri="https://binary.pensoft.net/fig/769831" pageId="0" pageNumber="109">Fig. 10 Suggested Common Name: Ko Chang Bent-toed Gecko</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="88AA45BC2A05AA4B5E74DDEF9054AB4F" pageId="0" pageNumber="109" type="holotype">
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<paragraph id="E24009D9AC2631955E7EE9613F7FFC34" pageId="0" pageNumber="109">Holotype.</paragraph>
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<paragraph id="133101E65E1965947D455C7A0C3A70AD" pageId="0" pageNumber="109">
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Adult female ZMKU R 00945 from Ko Chang Island, Ko Phayam Subdistrict, Mueang Ranong District, Ranong Province, Thailand (
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<geoCoordinate id="83224C17AB74BDFDF6C1C0CCC3811F65" degrees="9.82411" direction="north" orientation="latitude" precision="1" value="9.82411">9.82411°N</geoCoordinate>
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,
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<geoCoordinate id="37B47A518A36CA6CD1D48C49A01C7177" degrees="98.43999" direction="east" orientation="longitude" precision="1" value="98.43999">98.43999°E</geoCoordinate>
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, 36 m a.s.l.), collected by Siriporn Yodthong, Natee Ampai, Attapol Rujirawan, and Piyawan Puanprapai on 8 July 2017.
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</paragraph>
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</subSubSection>
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<subSubSection id="CBB05A77B19AC3EED27444462E3E7062" pageId="0" pageNumber="109" type="materials_examined">
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<paragraph id="361980733CD7BDEDB0CA3148B6D64789" pageId="0" pageNumber="109">Additional material examined.</paragraph>
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<paragraph id="71AFA2720079D5F8BA8E75342ABA0A4E" pageId="0" pageNumber="109">
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<materialsCitation id="FA155C200D68F292864A672B45D79B65" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Yodchaiy Chuaynkern" country="Thailand" latitude="9.4596" location="Khlong Naka Wildlife Sanctuary" longLatPrecision="7" longitude="98.5044" municipality="Suk Samran District" specimenCount="1" stateProvince="Ranong Province">
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<taxonomicName id="E61754234580BE4A4D6A6F94A700E2A8" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">Cyrtodactylus cf. kochangensis</taxonomicName>
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sp. nov. adult male THNHM 01667 from
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<location id="D720485A84AD0E819CF7F818FBE4C1AF" LSID="urn:lsid:plazi:treatment:7E67190E06B0564CB4C8E5A323705A73:D720485A84AD0E819CF7F818FBE4C1AF" country="Thailand" latitude="9.4596" longLatPrecision="7" longitude="98.5044" municipality="Suk Samran District" name="Khlong Naka Wildlife Sanctuary" stateProvince="Ranong Province">Khlong Naka Wildlife Sanctuary</location>
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,
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<collectingMunicipality id="18FEB3449F20815FDAFEAA4F5E753B8E">Suk Samran District</collectingMunicipality>
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,
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<collectingRegion id="460320B7C7A940FBEBFC9228292E519C" country="Thailand" name="Ranong">Ranong Province</collectingRegion>
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,
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<collectingCountry id="2C612DEB8A9517B19931AD3479F8FE34" name="Thailand">Thailand</collectingCountry>
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(~
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<geoCoordinate id="5247B95226565A9AD1EC78CA82639B80" degrees="9.4596" direction="north" orientation="latitude" precision="5" value="9.4596">N 9.4596</geoCoordinate>
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,
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<geoCoordinate id="7CC474D79B248A2A751D24144474649D" degrees="98.5044" direction="east" orientation="longitude" precision="5" value="98.5044">E 98.5044</geoCoordinate>
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, elevation unknown), collected by
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<collectorName id="4B76F12CBF928FB011DB1FDCEBD8CD44">Yodchaiy Chuaynkern</collectorName>
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between
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<collectingDate id="FD8D9E03F256DC4E622F8F558B40658F" value="2000-12-28">28 December 2000</collectingDate>
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and
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<collectingDate id="D474D48C473C8913F99270924D833F3A" value="2001-01-02">2 January 2001</collectingDate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="7743EC84E6908FCFCA0A9EAC8272C05C" pageId="0" pageNumber="109" type="diagnosis">
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<paragraph id="BF48A329511E70E8B1314CF1AAC6D873" pageId="0" pageNumber="109">Diagnosis</paragraph>
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<paragraph id="4C127C74616A735D9F5FE6347EFAEC34" pageId="0" pageNumber="109">
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<emphasis id="D76E70E1A2B245300457CD10828924D5" bold="true" pageId="0" pageNumber="109">(based on the holotype).</emphasis>
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<taxonomicName id="750ABA90BC5CBB69FA020F7E9C625686" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">
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<emphasis id="1181DD028226145EC896C22EF8658A5B" italics="true" pageId="0" pageNumber="109">Cyrtodactylus kochangensis</emphasis>
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</taxonomicName>
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sp. nov. can be separated from all other species of the
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<taxonomicName id="0D4FC41BDB23B6E4EBBA13583AD77010" baseAuthorityName="Smith" baseAuthorityYear="1923" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus brevipalmatus" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="brevipalmatus">
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<emphasis id="0BCD2C77B9D6F18D876B2C3B2DB4BD56" italics="true" pageId="0" pageNumber="109">Cyrtodactylus brevipalmatus</emphasis>
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</taxonomicName>
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group by the combination of having 12 or 13 supralabials, nine infralabials, 34 paravertebral tubercles, 14 rows of longitudinally arranged tubercles, 35 transverse rows of ventrals, 172 longitudinal rows of ventrals, 8 or 9 expanded subdigital lamellae on the fourth toe, 11 or 12 unexpanded subdigital lamellae on the fourth toe, 19-21 total subdigital lamellae on the fourth toe; eight expanded subdigital lamellae on the fourth finger, ten unexpanded subdigital lamellae on the fourth finger, 18 total subdigital lamellae on the fourth finger; 12 total enlarged femoral scales; 12 enlarged precloacal scales; enlarged femoral and enlarged precloacal scales not continuous; proximal femoral scales less than one-half the size of the distal femorals; small tubercles on forelimbs and flanks; large dorsolateral caudal tubercles and a wide ventrolateral caudal fringe not composed homogeneous scales; tail square in cross-section; slightly enlarged paired medial subcaudals not posteromedially furrowed; maximum SVL 60.1 mm; five dark transverse body bands (Tables
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<tableCitation id="9B3846FC757F654385DB742926E2E8D5" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary statistics of the normalized morphometric data for the putative species of the Cyrtodactylus brevipalmatus group. SD = + / - standard deviation. Character abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DB30FD29B916930A1B76FCF69ADB7D6D" pageId="0" pageNumber="109" tableUuid="DB30FD29B916930A1B76FCF69ADB7D6D">4</tableCitation>
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,
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<tableCitation id="868CB00C1673C2C91967401FC3DFE9EA" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Sex and raw meristic, categorical, and morphometric data used in the analyses from specimens in the Cyrtodactylus brevipalmatus group. m = male; f = female; R / L = right / left; / = data unavailable." httpUri="http://table.plazi.org/id/188A309A2CA1B482119DD3726E382E32" pageId="0" pageNumber="109" tableUuid="188A309A2CA1B482119DD3726E382E32">5</tableCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="7881F405795268FD0D7705D12D6C8861" pageId="0" pageNumber="109" type="description">
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<paragraph id="0FD4DCF64A98B7A1CE884D7128AB5B50" pageId="0" pageNumber="109">Description of holotype</paragraph>
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<paragraph id="E1CA93902547F93157BF844AEFAEE7C6" pageId="0" pageNumber="109">
|
|
<emphasis id="C476DCD8CBD7FB4F54855849D69F0121" bold="true" pageId="0" pageNumber="109">
|
|
(Fig.
|
|
<figureCitation id="ABF1E9B71871CF668059C332ECD2631A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Holotype of Cyrtodactylus kochangensis sp. nov. ZMKU R 00945 (field no. AA 04439) from Thailand, Ranong Province, Mueng Ranong District, Ko Phayam Subdistrict, Ko Chang A dorsal view B ventral view C dorsal view of head D ventral view of femoral and precloacal regions E dorsal view of tail F ventral view of tail G holotype in life." figureDoi="10.3897/zookeys.1129.90535.figure10" httpUri="https://binary.pensoft.net/fig/769831" pageId="0" pageNumber="109">10</figureCitation>
|
|
).
|
|
</emphasis>
|
|
Adult female SVL 60.1 mm; head moderate in length (HL/SVL 0.29), width (HW/HL 0.67), depth (HD/HL 0.38), distinct from neck, triangular in dorsal profile; lores flat anteriorly, weakly inflated posteriorly; prefrontal region slightly concave; canthus rostralis rounded; snout elongate (ES/HL 0.40), rounded in dorsal profile; eye large (ED/HL 0.24); ear opening subcircular, small; eye to ear distance greater than diameter of eye; rostral rectangular, furrowed dorsally, bordered posteriorly by large left and right supranasals and one slightly smaller azygous internasal, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two slightly smaller postnasals, bordered ventrally by first supralabial; 12R/13L rectangular supralabials tapering smoothly to below eye; 9R/9L infralabials tapering smoothly to below eye; scales of rostrum and lores domed, slightly larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small tubercles; superciliaries subrectangular, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapazoidal postmentals contacting medially for approximately 40% of their length posterior to mental; one row of six (R,L) slightly enlarged sublabials extending posteriorly fifth infralabials, subsequent sublabials much smaller; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales.
|
|
</paragraph>
|
|
<caption id="BA7682B48890D73D05FBB0AB97AFCA0D" doi="10.3897/zookeys.1129.90535.figure10" httpUri="https://binary.pensoft.net/fig/769831" pageId="0" pageNumber="109" start="Figure 10" startId="F10">
|
|
<paragraph id="B8F6677BC1F9DC761FF504A9D4CD0B82" pageId="0" pageNumber="109">
|
|
<emphasis id="2F29980B399DEA2CD4C7661EA98234C5" bold="true" pageId="0" pageNumber="109">Figure 10.</emphasis>
|
|
Holotype of
|
|
<taxonomicName id="8141847226A0FF0950F24E66C9558023" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">
|
|
<emphasis id="C73060F6408852E01624B792AD57AFDF" italics="true" pageId="0" pageNumber="109">Cyrtodactylus kochangensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. ZMKU R 00945 (field no. AA 04439) from Thailand, Ranong Province, Mueng Ranong District, Ko Phayam Subdistrict, Ko Chang
|
|
<emphasis id="AD4D47AF7EDBDF2B67C17EF40EEADA2B" bold="true" pageId="0" pageNumber="109">A</emphasis>
|
|
dorsal view
|
|
<emphasis id="944D3361F2E1A1C8A690B1C2A380E1E5" bold="true" pageId="0" pageNumber="109">B</emphasis>
|
|
ventral view
|
|
<emphasis id="1CD1E127404B9C12828986979586D183" bold="true" pageId="0" pageNumber="109">C</emphasis>
|
|
dorsal view of head
|
|
<emphasis id="0760CF9227CCA05AAC55A01AD3DCA99E" bold="true" pageId="0" pageNumber="109">D</emphasis>
|
|
ventral view of femoral and precloacal regions
|
|
<emphasis id="BAE673474F439BC00561C75D58B038C2" bold="true" pageId="0" pageNumber="109">E</emphasis>
|
|
dorsal view of tail
|
|
<emphasis id="6EE32A775A4962BAFC708993E655EE6E" bold="true" pageId="0" pageNumber="109">F</emphasis>
|
|
ventral view of tail
|
|
<emphasis id="7A1982CAE488BDC6C6BE33BBA52D0379" bold="true" pageId="0" pageNumber="109">G</emphasis>
|
|
holotype in life.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="89A1785FDCBA62C84E375B0AD737B970" pageId="0" pageNumber="109">Body relatively short (AG/SVL 0.48) with well-defined ventrolateral folds; dorsal scales small, granular, interspersed with larger conical, semi-regularly arranged, weakly keeled tubercles; tubercles extend from occipital region onto base of tail and slightly beyond as paravertebral rows; tubercles of nape and occiput smaller than those on body; approximately 14 longitudinal rows of tubercles at midbody; approximately 34 paravertebral tubercles; tubercles on upper flanks smaller than those on dorsum; approximately 35 longitudinal rows of flat, imbricate, ventral scales much larger than dorsal scales; approximately 172 transverse rows of ventral scales; no pore-bearing, precloacal scales; 12 enlarged precloacal scales; no deep precloacal groove or depression; and approximately three rows of post-precloacal scales on midline.</paragraph>
|
|
<paragraph id="5793C6C2E49EE3BC748477110F2152BD" pageId="0" pageNumber="109">Forelimbs moderate in stature, relatively short (ForL/SVL 0.13); granular scales of forearm slightly larger than those on body, interspersed with tubercles; palmar scales rounded, slightly raised; digits well-developed, relatively short, inflected at basal interphalangeal joints; digits narrower distal to inflections; subdigital lamellae wide, transversely expanded proximal to joint inflections, narrower transverse lamellae distal to joint inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; 8R/8L expanded and 10R/10L unexpanded lamellae beneath the fourth finger; hind limbs larger and thicker than forelimbs, moderate in length (TibL/SVL 0.14), covered dorsally by granular scales interspersed with moderately sized, conical tubercles dorsally and posteriorly and anteriorly by flat, slightly larger, subimbricate scales; ventral scales of thigh flat, subimbricate, larger than dorsals; subtibial scales flat, imbricate; no pore-bearing femoral scales; 6R/6L enlarged femoral scales; enlarged femoral scales not contiguous with enlarged precloacal scales, terminating distally at knee; proximal femoral scales smaller than distal femorals, the former forming an abrupt union with much smaller, rounded, ventral scales of posteroventral margin of thigh; plantar scales flat; digits relatively long, well-developed, inflected at basal interphalangeal joints; 9R/8L wide, transversely expanded subdigital lamellae on the fourth toe proximal to joint inflection that extend onto sole, and 12R/11L unexpanded lamellae beneath the fourth toe; and claws well-developed, sheathed by a dorsal and ventral scale at base.</paragraph>
|
|
<paragraph id="73A6DEF1E7523FB3BA0DF0E779FB0DA5" pageId="0" pageNumber="109">Tail original (but in two pieces), long 80.3 mm (TL/SVL 1.34), 4.0 mm in width at base, tapering to a point; nearly square in cross-section; dorsal scales flat, square bearing tubercles forming paravertebral rows and large tubercles forming a dorsolateral longitudinal row; slightly larger, posteriorly directed, semi-spinose tubercles forming wide distinct ventrolateral caudal fringe; scales of ventrolateral fringe generally interspersed at regular intervals with larger spinose scales; medial subcaudal scales paired, slightly enlarged; subcaudals, larger than dorsal caudal scales; base of tail bearing hemipenial swellings; one conical postcloacal tubercle at base of hemipenial swellings; and postcloacal scales flat, imbricate.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D37C97EB0CD0AF506F2670D3D3E1DC7C" pageId="0" pageNumber="109" type="coloration in life">
|
|
<paragraph id="2B7639498E897A2954FC223B912CCD8E" pageId="0" pageNumber="109">Coloration in life</paragraph>
|
|
<paragraph id="844D11E0B6511ED6FB051BDE9F22159E" pageId="0" pageNumber="109">
|
|
<emphasis id="1CBF667D7E22DB863509188CE0167809" bold="true" pageId="0" pageNumber="109">
|
|
(Fig.
|
|
<figureCitation id="6A1D272ADC68B15EBCFC32F4307757DC" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Holotype of Cyrtodactylus kochangensis sp. nov. ZMKU R 00945 (field no. AA 04439) from Thailand, Ranong Province, Mueng Ranong District, Ko Phayam Subdistrict, Ko Chang A dorsal view B ventral view C dorsal view of head D ventral view of femoral and precloacal regions E dorsal view of tail F ventral view of tail G holotype in life." figureDoi="10.3897/zookeys.1129.90535.figure10" httpUri="https://binary.pensoft.net/fig/769831" pageId="0" pageNumber="109">10</figureCitation>
|
|
).
|
|
</emphasis>
|
|
Ground color of the head, body, limbs, and tail pale brown; faint, diffuse mottling on rostrum; lores darkly colored; wide, distinct, pale-colored postorbital stripe; nuchal band faint, bearing two posterior projections; three very faint, wide, irregularly shaped body bands between limb insertions edged in slightly darker brown; band interspaces bearing irregularly shaped, faint, dark-colored markings; dark-colored speckling on limbs and digits; digits bearing pale-colored bands; four wide, faint, dark-colored caudal bands separated by three pale-colored bands on original portion of tail; all caudal bands encircle tail; all ventral surfaces beige, generally immaculate; and iris orangish to coppery in color.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F57B8389C5CA51107342653953511570" pageId="0" pageNumber="109" type="variation">
|
|
<paragraph id="E6DA294352BDFB50B72223559E217C65" pageId="0" pageNumber="109">Variation.</paragraph>
|
|
<paragraph id="4A88B198444444320072858D9D1EEE00" pageId="0" pageNumber="109">
|
|
The additional specimen (THNHM 01667) closely approximates the holotype in overall coloration and pattern except that it is more boldly marked. It has four dark-colored body bands as opposed to three and a complete original tail bearing eight dark-colored and seven pale-colored bands. The pale-colored postorbital stripe is slightly thinner and less distinct. Meristic and morphometric differences are listed in Table
|
|
<tableCitation id="8080857358ADD4853AC31E7E323AA44A" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Sex and raw meristic, categorical, and morphometric data used in the analyses from specimens in the Cyrtodactylus brevipalmatus group. m = male; f = female; R / L = right / left; / = data unavailable." httpUri="http://table.plazi.org/id/188A309A2CA1B482119DD3726E382E32" pageId="0" pageNumber="109" tableUuid="188A309A2CA1B482119DD3726E382E32">5</tableCitation>
|
|
. Given its overall morphological and color pattern similarities and close geographic proximity to the holotype (~ 40 km), we consider this individual as
|
|
<taxonomicName id="D0B049B97E30B8B9619BC670E298E27F" lsidName="C. kochangensis" pageId="0" pageNumber="109" rank="species" species="kochangensis">C. cf. kochangensis</taxonomicName>
|
|
sp. nov. pending genetic data.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B0A2AFB815C8E0BB4156A72FE03528E7" pageId="0" pageNumber="109" type="distribution">
|
|
<paragraph id="CC0647936A19B4D62E593FF89D0B6675" pageId="0" pageNumber="109">Distribution.</paragraph>
|
|
<paragraph id="576D67D1ECF8F8375EACBF21271EE8B5" pageId="0" pageNumber="109">
|
|
<taxonomicName id="1A98538C61340FB82C4BDCABA76FD16A" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">
|
|
<emphasis id="C408354BD81CD58750E07B38BF50A083" italics="true" pageId="0" pageNumber="109">Cyrtodactylus kochangensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is currently known only from the type locality at Ko Chang Island, Ko Phayam Subdistrict, Mueang Ranong District, Ranong Province, Thailand. The additional population of pending species status occurs in the Khlong Naka Wildlife Sanctuary, Suk Samran District Ranong Province.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5C57E00123521FFFC584525F38E468CC" pageId="0" pageNumber="109" type="etymology">
|
|
<paragraph id="89B9AF4364E89D1471A10DFD207F5C2F" pageId="0" pageNumber="109">Etymology.</paragraph>
|
|
<paragraph id="8A05C45BA2B17E2F386685E310AE9235" pageId="0" pageNumber="109">
|
|
The specific epithet
|
|
<emphasis id="69042F6BEA3155F451CEF94522CC8444" italics="true" pageId="0" pageNumber="109">kochangensis</emphasis>
|
|
is in reference to the type locality, Ko Chang, Ranong Province, Thailand
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E9952457184400AD5ACBF75FFC5F2947" pageId="0" pageNumber="109" type="comparisons">
|
|
<paragraph id="303A13D2335EABA0FF9C3B27086A5498" pageId="0" pageNumber="109">Comparisons</paragraph>
|
|
<paragraph id="DD4FA09D3486F2A9D49BCE7C63950634" pageId="0" pageNumber="109">
|
|
<emphasis id="1451130ED398E826078DD7079D548D40" bold="true" pageId="0" pageNumber="109">(based on the holotype)</emphasis>
|
|
.
|
|
<taxonomicName id="9586BCFF705525527A65ED172DD61610" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">
|
|
<emphasis id="70E34F53407BA7480ABCB55DD954AE2A" italics="true" pageId="0" pageNumber="109">Cyrtodactylus kochangensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. forms a clade with the sister species
|
|
<taxonomicName id="91677E3AF90977F67FD58EA422EA61E2" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus rivularis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="rivularis">
|
|
<emphasis id="D54545194E7AE716C0E4104F6068D97F" italics="true" pageId="0" pageNumber="109">Cyrtodactylus rivularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="24328886316E92DB983D6E98E4264606" lsidName="C. rukhadeva" pageId="0" pageNumber="109" rank="species" species="rukhadeva">
|
|
<emphasis id="BC80348A25B2385F63DBD0C530E909AF" italics="true" pageId="0" pageNumber="109">C. rukhadeva</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation id="3111C3EB67805AE16BD718A31B6B311B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum clade credibility BEAST phylogeny of the Cyrtodactylus brevipalmatus group highlighting the new species described herein. Bayesian posterior probabilities (BPP) are listed at the nodes." figureDoi="10.3897/zookeys.1129.90535.figure2" httpUri="https://binary.pensoft.net/fig/769823" pageId="0" pageNumber="109">2</figureCitation>
|
|
) from which it differs by an uncorrected pairwise sequence divergence of 12.00-12.52% and 12.52-13.68%, respectively (Table
|
|
<tableCitation id="ABF2C9567668A58C5433EAA9F43BD4F4" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean (minimum-maximum) percentages of uncorrected pairwise sequence divergence (p - distances) among the putative species of the Cyrtodactylus brevipalmatus group based on 1,386 base pairs of mitochondrial NADH dehydrogenase subunit 2 gene (ND 2) and adjacent tRNAs. Intraspecific p - distance are in bold font. n / a = data not applicable." httpUri="http://table.plazi.org/id/5DF1D514C510FC7E288ECC2377589860" pageId="0" pageNumber="109" tableUuid="5DF1D514C510FC7E288ECC2377589860">2</tableCitation>
|
|
) and it and
|
|
<taxonomicName id="85B4D4C7B93D7C68414236125DB3B744" lsidName="C. kochangensis" pageId="0" pageNumber="109" rank="species" species="kochangensis">C. cf. kochangensis</taxonomicName>
|
|
sp. nov. are separated from
|
|
<taxonomicName id="EDFD0DB05FB7C524C7F77BDA814B6F8B" lsidName="C. rukhadeva" pageId="0" pageNumber="109" rank="species" species="rukhadeva">
|
|
<emphasis id="2B256FB98EC59F8958AC73DE63ACD929" italics="true" pageId="0" pageNumber="109">C. rukhadeva</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="947C8222475E8B58BAA4ED2AF41E8F11" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus rivularis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="rivularis">
|
|
<emphasis id="A82492CDABF6962B2C50C5A0712FE0D9" italics="true" pageId="0" pageNumber="109">Cyrtodactylus rivularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by geographic distance of no less than ~280-470 km (Fig.
|
|
<figureCitation id="125B338B704470D24E28865AE5E97E99" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of nominal species and unnamed populations and specimens of the Cyrtodactylus brevipalmatus group. Stars denote type localities. White circles are literature localities from which specimens were not examined and remain unidentified. Locality data for all material examined is in Table 1." figureDoi="10.3897/zookeys.1129.90535.figure1" httpUri="https://binary.pensoft.net/fig/769822" pageId="0" pageNumber="109">1</figureCitation>
|
|
). The small sample size (
|
|
<emphasis id="E0E63C1792DB13190850C9943590F541" italics="true" pageId="0" pageNumber="109">N</emphasis>
|
|
= 1) precludes it from statistical analyses, however at this point, it differs from
|
|
<taxonomicName id="A9FD33DF73B727C05CAD2DC3538C95A6" lsidName="C. rukhadeva" pageId="0" pageNumber="109" rank="species" species="rukhadeva">
|
|
<emphasis id="8DAF07EBA5A82DBAFC13DD9446E6DCEB" italics="true" pageId="0" pageNumber="109">C. rukhadeva</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="725A89A16A51107C7FCF1901F46CD66F" lsidName="C. rukhadeva" pageId="0" pageNumber="109" rank="species" species="rukhadeva">C. cf. rukhadeva</taxonomicName>
|
|
in having 34 PVT versus 26-30; 14 LRT versus 18-20; 172 VSM versus 152-165; 12 FS versus 13-18; 12 PCS versus 13-17 and 1 PCT versus 2 or 3, collectively. From
|
|
<taxonomicName id="CE75FD1C2C7470BF615AD00400F093EC" lsidName="C. rukhadeva" pageId="0" pageNumber="109" rank="species" species="rukhadeva">
|
|
<emphasis id="D6FAAD5242B1E65F7F82F3DE11A7DE69" italics="true" pageId="0" pageNumber="109">C. rukhadeva</emphasis>
|
|
</taxonomicName>
|
|
by having 12 FS versus 16 or 17; and five BB versus three. From
|
|
<taxonomicName id="83747D141B4F4BBDC261FD888BD54643" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus rivularis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="rivularis">
|
|
<emphasis id="407FF2A810E9FBC529A53AAF74DF9AE1" italics="true" pageId="0" pageNumber="109">Cyrtodactylus rivularis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. it differs in having 14 LRT versus 18-20; 172 VSM versus 160-166; 12 FS versus 14-16; and five BB versus three or four. Discrete differences between
|
|
<taxonomicName id="18B8D32D3D04D0426DCDD1A63A6F6DC3" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">
|
|
<emphasis id="3BD9B9BA90D69C15C69BFD622E6FE660" italics="true" pageId="0" pageNumber="109">Cyrtodactylus kochangensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="24E130B71208DD2EC43C750DA862A08A" lsidName="C. kochangensis" pageId="0" pageNumber="109" rank="species" species="kochangensis">C. cf. kochangensis</taxonomicName>
|
|
sp. nov. and all other species and populations are presented in Tables
|
|
<tableCitation id="AA4BBA80B6BD0BD44E59F2B39B082FAA" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary statistics of the normalized morphometric data for the putative species of the Cyrtodactylus brevipalmatus group. SD = + / - standard deviation. Character abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DB30FD29B916930A1B76FCF69ADB7D6D" pageId="0" pageNumber="109" tableUuid="DB30FD29B916930A1B76FCF69ADB7D6D">4</tableCitation>
|
|
,
|
|
<tableCitation id="1B6AB2D0A5826F58E2D97BCEE5A135A7" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Sex and raw meristic, categorical, and morphometric data used in the analyses from specimens in the Cyrtodactylus brevipalmatus group. m = male; f = female; R / L = right / left; / = data unavailable." httpUri="http://table.plazi.org/id/188A309A2CA1B482119DD3726E382E32" pageId="0" pageNumber="109" tableUuid="188A309A2CA1B482119DD3726E382E32">5</tableCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6CCB435F3CCADF107524ECB8F3AE7A3F" pageId="0" pageNumber="109" type="natural history">
|
|
<paragraph id="76406A037070709A7A2C226282A02A65" pageId="0" pageNumber="109">Natural history.</paragraph>
|
|
<paragraph id="8E864E50B7DF3928F44D570356943343" pageId="0" pageNumber="109">
|
|
The holotype (ZMKU R 00945) was collected at night (2107 h) among branches of a small tree approximately 100 cm above the ground at 36 m elevation with a temperature of 28.6 °C and relative humidity of 83.9%. The surrounding habitat was dry evergreen forest with a rocky stream nearby (Fig.
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<figureCitation id="E78CCF70314AC9C681A6A4F155CE55D5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitat of Cyrtodactylus kochangensis sp. nov. at Thailand, Ranong Province, Mueng Ranong District, Ko Phayam Subdistrict, Ko Chang." figureDoi="10.3897/zookeys.1129.90535.figure11" httpUri="https://binary.pensoft.net/fig/769832" pageId="0" pageNumber="109">11</figureCitation>
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). The new species was found to co-occur with two other species of gekkonid lizards,
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<taxonomicName id="A049D12B9E97B614BD6156C681FD78F1" baseAuthorityName="Theobald" baseAuthorityYear="1876" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus oldhami" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="oldhami">
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<emphasis id="17A99C924CBE012FD3B7EEDDE9A3268C" italics="true" pageId="0" pageNumber="109">Cyrtodactylus oldhami</emphasis>
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</taxonomicName>
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(Theobald, 1876) and
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<taxonomicName id="D6E43EDC6FAC447FAED47F1400811417" baseAuthorityName="Grismer, Wood, Grismer, Quah, Thy, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown" baseAuthorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" lsidName="Gekko tokehos" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="tokehos">
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<emphasis id="B6A77F40B079D6F48C617FBA4043639E" italics="true" pageId="0" pageNumber="109">Gekko tokehos</emphasis>
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</taxonomicName>
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(Grismer, Wood, Grismer, Quah, Thy, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown, 2019).
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</paragraph>
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<caption id="C879CC6632AB27624D334C43E6F4FE39" doi="10.3897/zookeys.1129.90535.figure11" httpUri="https://binary.pensoft.net/fig/769832" pageId="0" pageNumber="109" start="Figure 11" startId="F11">
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<paragraph id="A8E6ACF37ABD4AC4DFD909336A49D9DA" pageId="0" pageNumber="109">
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<emphasis id="B3FBB3A092441C19FBF4ECF69C889084" bold="true" pageId="0" pageNumber="109">Figure 11.</emphasis>
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Habitat of
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<taxonomicName id="BCD88B252243564C142196E1694B15C1" authorityName="Grismer & Aowphol & Yodthong & Ampai & Termprayoon & Aksornneam & Rujirawan" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus kochangensis" order="Squamata" pageId="0" pageNumber="109" phylum="Chordata" rank="species" species="kochangensis">
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<emphasis id="4D784BEB6EA5ACB6179967E603D5F455" italics="true" pageId="0" pageNumber="109">Cyrtodactylus kochangensis</emphasis>
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</taxonomicName>
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sp. nov. at Thailand, Ranong Province, Mueng Ranong District, Ko Phayam Subdistrict, Ko Chang.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |