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<document id="A905C9FE93BC2969B788864C6B0AE391" ID-CLB-Dataset="305035" ID-DOI="10.5852/ejt.2024.968.2723" ID-GBIF-Dataset="21b497f6-a2ee-4af8-b2e0-027aadacb4eb" ID-ISSN="2118-9773" ID-Zenodo-Dep="14047314" ID-ZooBank="35322F0D-0A5C-4227-AB4D-D4758ECA6E7A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730916893761" checkinUser="plazi" docAuthor="Poletaev, Vladyslav" docDate="2024" docId="03BA8E19595ED21EFDABC9ECFABDB2EC" docLanguage="en" docName="EJT.2024.968.132-155.pdf" docOrigin="European Journal of Taxonomy 968" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2723/12515" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Johnsoniana oceani" docType="treatment" docVersion="4" lastPageNumber="148" masterDocId="FF83F6615950D20EFF9ACC29FFFCB35A" masterDocTitle="New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia)" masterLastPageNumber="155" masterPageNumber="132" pageNumber="146" updateTime="1731010174111" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="451D92523BFF020CD7D95FC8114E67DE">New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia)</mods:title>
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<mods:affiliation id="16C478AB9E03C626BC9DFDAE32EDBB21">Institute of Geological Sciences of the National Academy of Sciences of Ukraine, 55 - b, Oles Honchar St., Kyiv, 01054, Ukraine.</mods:affiliation>
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<subSubSection id="C3096C84595ED200FDABC9ECFBFEB6BA" box="[561,1026,1477,1504]" pageId="14" pageNumber="146" type="nomenclature">
<paragraph id="8BAC3F0F595ED200FDABC9ECFBFEB6BA" blockId="14.[561,1026,1477,1538]" box="[561,1026,1477,1504]" pageId="14" pageNumber="146">
<heading id="D0E48863595ED200FDABC9ECFBFEB6BA" box="[561,1026,1477,1504]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="146" reason="2">
<taxonomicName id="4C13448C595ED200FDABC9ECFBFEB6BA" authority="(d'Orbigny, 1850)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[561,1026,1477,1504]" class="Rhynchonellata" family="Spiriferidae" genus="Johnsoniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="149" phylum="Brachiopoda" rank="species" species="oceani">
<emphasis id="B967E31D595ED200FDABC9ECFD33B6BA" bold="true" box="[561,719,1477,1504]" italics="true" pageId="14" pageNumber="146">Johnsoniana</emphasis>
<emphasis id="B967E31D595ED200FD4CC9ECFCDAB685" bold="true" box="[726,806,1477,1503]" italics="true" pageId="14" pageNumber="146">oceani</emphasis>
(
<bibRefCitation id="EF8242FE595ED200FCACC9ECFC06B6BA" author="d'Orbigny A." box="[822,1018,1477,1504]" pageId="14" pageNumber="146" pagination="295 - 353" refId="ref11682" refString="d'Orbigny A. 1850. Memoire sur les Brachiopodes. II. Classification des Brachiopodes. Annales des Sciences Naturelles. Zoologie, Serie 3 e 13: 295 - 353. Available from https: // www. biodiversitylibrary. org / page / 13434796 [accessed 7 Oct. 2024]." type="book chapter" year="1850">d’Orbigny, 1850</bibRefCitation>
)
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</heading>
</paragraph>
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<subSubSection id="C3096C84595ED200FD47C9C1FCAAB558" box="[733,854,1512,1538]" pageId="14" pageNumber="146" type="description">
<paragraph id="8BAC3F0F595ED200FD47C9C1FCAAB558" blockId="14.[561,1026,1477,1538]" box="[733,854,1512,1538]" pageId="14" pageNumber="146">
<figureCitation id="1328238A595ED200FD47C9C1FCAAB558" box="[733,854,1512,1538]" captionStart="Fig" captionStartId="16.[189,232,1646,1672]" captionTargetBox="[208,1351,501,1597]" captionTargetId="figure-62@16.[205,1383,488,1606]" captionTargetPageId="16" captionText="Fig. 6. A. Verneuilia cheiropteryx (d’Archiac &amp;de Verneuil, 1842). Lectotype ONCP D568 (Gourvennec 1994 Coll.) from Refrath (Bergisches Land, Germany). Devonian, Frasnian; dorsal, ventral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 1–5). B–E. Johnsoniana oceani (d’Orbigny, 1850). B. Neotype NHM B7923 (Brunton 1984 Coll.) from Visé (Belgium). Visean; ventral, lateral, dorsal, posterior and anterior views (copy from Brunton 1984: text-fig. 160a–e). C. Paratype CNIGR Museum 3085 (Nalivkin 1979 Coll.) from Bashkortostan (Urals, Russia). Upper Tournaisian; ventral, lateral, dorsal and anterior views (copy from Nalivkin 1979: pl. 62 fig. 21). D. Paratype ONCP Museum D306a (Gourvennec 1994 Coll.) from Visé (Belgium). Viséan; ventral, dorsal, lateral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 11–15). E. Paratype ONCP Museum D306e (Gourvennec 1994 Coll.) from same locality; dorsal, ventral, anterior and lateral views (copy from Gourvennec 1994: pl. 1 figs 16–19). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14047333" httpUri="https://zenodo.org/record/14047333/files/figure.png" pageId="14" pageNumber="146">Fig. 6B–E</figureCitation>
</paragraph>
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<subSubSection id="C3096C84595ED200FF27CA03FC29B46C" pageId="14" pageNumber="146" type="reference_group">
<paragraph id="8BAC3F0F595ED200FF27CA03FCE0B51E" blockId="14.[189,796,1578,1605]" box="[189,796,1578,1605]" pageId="14" pageNumber="146">
<treatmentCitationGroup id="AB031821595ED200FF27CA03FCE0B51E" box="[189,796,1578,1605]" pageId="14" pageNumber="146">
<treatmentCitation id="0AB2191E595ED200FF27CA03FD88B51E" author="d'Orbigny A." box="[189,628,1578,1605]" page="149" pageId="14" pageNumber="146" year="1850">
<taxonomicName id="4C13448C595ED200FF27CA03FD88B51E" authority="d'Orbigny, 1850: 149" authorityName="d'Orbigny" authorityPageNumber="149" authorityYear="1850" box="[189,628,1578,1605]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="species" species="oceani">
<emphasis id="B967E31D595ED200FF27CA03FE90B51E" box="[189,364,1578,1604]" italics="true" pageId="14" pageNumber="146">Spirifer oceani</emphasis>
<bibRefCitation id="EF8242FE595ED200FEE9CA03FD88B51E" author="d'Orbigny A." box="[371,628,1578,1605]" pageId="14" pageNumber="146" pagination="295 - 353" refId="ref11682" refString="d'Orbigny A. 1850. Memoire sur les Brachiopodes. II. Classification des Brachiopodes. Annales des Sciences Naturelles. Zoologie, Serie 3 e 13: 295 - 353. Available from https: // www. biodiversitylibrary. org / page / 13434796 [accessed 7 Oct. 2024]." type="book chapter" year="1850">d’Orbigny, 1850: 149</bibRefCitation>
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, pl. 38 fig. 29.
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</paragraph>
<paragraph id="8BAC3F0F595ED200FF27CA45FA99B5DC" blockId="14.[189,1399,1643,1846]" box="[189,1381,1643,1670]" pageId="14" pageNumber="146">
<treatmentCitationGroup id="AB031821595ED200FF27CA45FA99B5DC" box="[189,1381,1643,1670]" pageId="14" pageNumber="146">
<taxonomicName id="4C13448C595ED200FF27CA45FE4DB5DF" authorityName="d'Archiac &amp; de Verneuil" authorityYear="1842" box="[189,433,1643,1670]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="species" species="cheiropteryx">
<emphasis id="B967E31D595ED200FF27CA45FE4DB5DF" box="[189,433,1643,1670]" italics="true" pageId="14" pageNumber="146">Spirifer cheiropteryx</emphasis>
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– de Koninck 1843: 245, pl. 15 fig. 9. —
<treatmentCitation id="0AB2191E595ED200FC06CA42FB68B5DC" author="" box="[924,1172,1643,1670]" page="510" pageId="14" pageNumber="146">
<bibRefCitation id="EF8242FE595ED200FC06CA42FB68B5DC" box="[924,1172,1643,1670]" firstAuthor="Quenstedt" pageId="14" pageNumber="146" pagination="465 - 748" refId="ref12259" refString="465 - 748 (1870); 25 pl. (1871)." type="book" year="1870">Quenstedt 1871: 510</bibRefCitation>
</treatmentCitation>
, pl 53 figs 71–73.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAC3F0F595ED200FF27CAA6FC95B5F2" blockId="14.[189,1399,1643,1846]" box="[189,873,1678,1705]" pageId="14" pageNumber="146">
<treatmentCitationGroup id="AB031821595ED200FF27CAA6FC95B5F2" box="[189,873,1678,1705]" pageId="14" pageNumber="146">
<taxonomicName id="4C13448C595ED200FF27CAA6FE90B5F2" authorityName="d'Orbigny" authorityYear="1850" box="[189,364,1679,1705]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="species" species="oceani">
<emphasis id="B967E31D595ED200FF27CAA6FE90B5F2" box="[189,364,1679,1705]" italics="true" pageId="14" pageNumber="146">Spirifer oceani</emphasis>
</taxonomicName>
– de
<treatmentCitation id="0AB2191E595ED200FE36CAA6FD6EB5F3" author="di Koninck L. G." box="[428,658,1679,1705]" page="132" pageId="14" pageNumber="146" year="1887">
<bibRefCitation id="EF8242FE595ED200FE36CAA6FD6EB5F3" author="di Koninck L. G." box="[428,658,1679,1705]" pageId="14" pageNumber="146" pagination="1 - 159" refId="ref11114" refString="di Koninck L. G. 1887. Faune du Calcaire Carbonifere de la Belgique. Brachiopodes. Annales du Musee royal d ' histoire naturelle de Belgique 14: 1 - 159. https: // doi. org / 10.5962 / bhl. title. 149304" type="journal article" year="1887">Koninck 1887: 132</bibRefCitation>
</treatmentCitation>
, pl. 28 figs 11–16.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAC3F0F595ED200FF27CA9BFD39B448" blockId="14.[189,1399,1643,1846]" pageId="14" pageNumber="146">
<treatmentCitationGroup id="AB031821595ED200FF27CA9BFD39B448" pageId="14" pageNumber="146">
<taxonomicName id="4C13448C595ED200FF27CA9BFE77B596" authority="(d'Orbigny, 1850)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[189,395,1714,1740]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="species" species="oceani">
<emphasis id="B967E31D595ED200FF27CA9BFE77B596" box="[189,395,1714,1740]" italics="true" pageId="14" pageNumber="146">Verneuilia oceani</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AB2191E595ED200FE33CA9BFD11B596" author="Hall J. &amp; Clarke J. M." box="[425,749,1714,1740]" page="1" pageId="14" pageNumber="146" year="1893">
<bibRefCitation id="EF8242FE595ED200FE33CA9BFD11B596" author="Hall J. &amp; Clarke J. M." box="[425,749,1714,1740]" pageId="14" pageNumber="146" refId="ref10739" refString="Hall J. &amp; Clarke J. M. 1893. An Introduction to the Study of the Genera of Paleozoic Brachiopoda: intended as a Handbook for the Use of Students. Palaeontology of New York 8 Pt. 2. James B. Lyon, Albany. https: // doi. org / 10.5962 / bhl. title. 15111" type="book" year="1893">Hall &amp; Clarke 1893: 1–317</bibRefCitation>
</treatmentCitation>
. —
<bibRefCitation id="EF8242FE595ED200FC87CA9BFC3CB596" author="Ivanova E. A." box="[797,960,1714,1740]" pageId="14" pageNumber="146" pagination="264 - 280" refId="ref10886" refString="Ivanova E. A. 1960. Otriad Spiriferida. In: Orlov Y. A. (ed.) Osnovy Paleontologii. Mshanki, Brachiopody. Vol. 7: 264 - 280. Izdatel'stvo Akademii Nauk SSSR, Moscow." type="book chapter" year="1960">Ivanova 1960</bibRefCitation>
, pl. 58 figs 14–15. —
<bibRefCitation id="EF8242FE595ED200FB5CCA9BFA8BB596" author="Nalivkin D. V." box="[1222,1399,1714,1740]" pageId="14" pageNumber="146" refId="ref11601" refString="Nalivkin D. V. 1979. Brakhiopody turneyskogo yarusa Urala. Nauka, Leningrad." type="book" year="1979">Nalivkin 1979:</bibRefCitation>
143, pl. 62 figs 19–24. —
<treatmentCitation id="0AB2191E595ED200FD89CAFFFD13B5B5" author="Brunton C. H. C." box="[531,751,1749,1776]" page="101" pageId="14" pageNumber="146" year="1984">
<bibRefCitation id="EF8242FE595ED200FD89CAFFFD13B5B5" author="Brunton C. H. C." box="[531,751,1749,1776]" pageId="14" pageNumber="146" pagination="27 - 127" refId="ref9844" refString="Brunton C. H. C. 1984. Silicified brachiopods from the Visean of County Fermanagh, Ireland. (III). Rhynchonellids, spiriferids and terebratulids. Bulletin of the British Museum (Natural History) Geology 38 (2): 27 - 127. Available from https: // www. biodiversitylibrary. org / page / 2273038 [accessed 7 Oct. 2024]." type="journal article" year="1984">Brunton 1984: 101</bibRefCitation>
</treatmentCitation>
, text-fig. 160. —
<treatmentCitation id="0AB2191E595ED200FC2DCAFCFB3FB5AA" author="Gourvennec R." box="[951,1219,1749,1776]" page="578" pageId="14" pageNumber="146" year="1994">
<bibRefCitation id="EF8242FE595ED200FC2DCAFCFB3FB5AA" author="Gourvennec R." box="[951,1219,1749,1776]" pageId="14" pageNumber="146" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec 1994: 578</bibRefCitation>
</treatmentCitation>
, pl. 1 figs 7–31. —
<bibRefCitation id="EF8242FE595ED200FE95CAD1FDD7B448" author="Mottequin B. &amp; Poty E." box="[271,555,1784,1811]" pageId="14" pageNumber="146" pagination="351 - 371" refId="ref11541" refString="Mottequin B. &amp; Poty E. 2022. Brachiopods from the historical type area of the Visean Stage (Carboniferous, Mississippian; Belgium) and the Vise fauna: preliminary remarks. Palaeobiodiversity and Palaeoenvironments 102: 351 - 371. https: // doi. org / 10.1007 / s 12549 - 021 - 00498 - 9" type="journal article" year="2022">Mottequin &amp; Poty 2022</bibRefCitation>
: text-fig. 11j.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAC3F0F595ED200FF27CB35FC29B46C" blockId="14.[189,1399,1643,1846]" box="[189,981,1819,1846]" pageId="14" pageNumber="146">
<treatmentCitationGroup id="AB031821595ED200FF27CB35FC29B46C" box="[189,981,1819,1846]" pageId="14" pageNumber="146">
<taxonomicName id="4C13448C595ED200FF27CB35FE8AB46F" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[189,374,1820,1846]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="14" pageNumber="146" phylum="Cnidaria" rank="species" species="oceani">
<emphasis id="B967E31D595ED200FF27CB35FE8AB46F" box="[189,374,1820,1846]" italics="true" pageId="14" pageNumber="146">Oceania oceani</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AB2191E595ED200FE09CB32FD6AB46C" author="Poletaev V. I." box="[403,662,1819,1846]" page="17" pageId="14" pageNumber="146" year="2015">
<bibRefCitation id="EF8242FE595ED200FE09CB32FD6AB46C" author="Poletaev V. I." box="[403,662,1819,1846]" pageId="14" pageNumber="146" pagination="17 - 18" refId="ref12092" refString="Poletaev V. I. 2015. Novyy rod i semeystvo kamennougol'nykh spiriferid Yevropy. In: Gozhyk P. F. (ed.) Stratotypovi ta oporni rozrizy fanerozois'kykh vidkladiv Ukrainy: suchasnyi stan ta perspektyvy podal'shykh doslidzhen'. Materialy 36 sesii Paleontologichnogo tovarustva Ukrainy: 17 - 18. Kyiv." type="book chapter" year="2015">Poletaev 2015: 17–18</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB2191E595ED200FD39CB35FCF1B46C" author="Poletaev V. I." box="[675,781,1819,1846]" page="95" pageId="14" pageNumber="146" year="2018">
<bibRefCitation id="EF8242FE595ED200FD39CB35FCF1B46C" author="Poletaev V. I." box="[675,781,1819,1846]" pageId="14" pageNumber="146" refId="ref12191" refString="Poletaev V. I. 2018. Atlas-opredelitel' kamennougol'nykh spiriferid Vostochnoy Yevropy. LAT &amp; K, Kyiv." type="book" year="2018">2018: 95</bibRefCitation>
</treatmentCitation>
, pl. 117 figs 1–7.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3096C84595ED200FF27CB74FEA7B4FC" box="[189,347,1885,1958]" pageId="14" pageNumber="146" type="diagnosis">
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<emphasis id="B967E31D595ED200FF27CB74FECAB42D" bold="true" box="[189,310,1885,1911]" pageId="14" pageNumber="146">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3F0F595ED200FF27CBA5FEA7B4FC" blockId="14.[189,347,1931,1958]" box="[189,347,1931,1958]" pageId="14" pageNumber="146">As for genus.</paragraph>
</subSubSection>
<subSubSection id="C3096C84595FD201FF27CD21FC54B1DE" pageId="15" pageNumber="147" type="materials_examined">
<paragraph id="8BAC3F0F595FD201FF27CD21FE56B278" blockId="15.[189,426,264,290]" box="[189,426,264,290]" pageId="15" pageNumber="147">
<heading id="D0E48863595FD201FF27CD21FE56B278" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="15" pageNumber="147" reason="3">
<emphasis id="B967E31D595FD201FF27CD21FE56B278" bold="true" box="[189,426,264,290]" pageId="15" pageNumber="147">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3F0F595FD201FF70CD1EFD34B2C2" blockId="15.[189,1398,311,408]" pageId="15" pageNumber="147">
<materialsCitation id="3B7B3552595FD201FF70CD1EFD34B2C2" collectionCode="NHM" collectorName="Davidson" country="United Kingdom" location="Visean" pageId="15" pageNumber="147" specimenCode="NHM B7923" specimenCount="1" stateProvince="Visean" typeStatus="neotype">
<emphasis id="B967E31D595FD201FF70CD1EFEADB20B" bold="true" box="[234,337,311,337]" pageId="15" pageNumber="147">
<typeStatus id="54A881AD595FD201FF70CD1EFEADB20B" box="[234,337,311,337]" pageId="15" pageNumber="147" type="neotype">Neotype</typeStatus>
</emphasis>
(
<figureCitation id="1328238A595FD201FEFACD1EFE48B20B" box="[352,436,311,337]" captionStart="Fig" captionStartId="16.[189,232,1646,1672]" captionTargetBox="[208,1351,501,1597]" captionTargetId="figure-62@16.[205,1383,488,1606]" captionTargetPageId="16" captionText="Fig. 6. A. Verneuilia cheiropteryx (d’Archiac &amp;de Verneuil, 1842). Lectotype ONCP D568 (Gourvennec 1994 Coll.) from Refrath (Bergisches Land, Germany). Devonian, Frasnian; dorsal, ventral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 1–5). B–E. Johnsoniana oceani (d’Orbigny, 1850). B. Neotype NHM B7923 (Brunton 1984 Coll.) from Visé (Belgium). Visean; ventral, lateral, dorsal, posterior and anterior views (copy from Brunton 1984: text-fig. 160a–e). C. Paratype CNIGR Museum 3085 (Nalivkin 1979 Coll.) from Bashkortostan (Urals, Russia). Upper Tournaisian; ventral, lateral, dorsal and anterior views (copy from Nalivkin 1979: pl. 62 fig. 21). D. Paratype ONCP Museum D306a (Gourvennec 1994 Coll.) from Visé (Belgium). Viséan; ventral, dorsal, lateral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 11–15). E. Paratype ONCP Museum D306e (Gourvennec 1994 Coll.) from same locality; dorsal, ventral, anterior and lateral views (copy from Gourvennec 1994: pl. 1 figs 16–19). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14047333" httpUri="https://zenodo.org/record/14047333/files/figure.png" pageId="15" pageNumber="147">Fig 6B</figureCitation>
)
<collectingCountry id="F3047F9F595FD201FF27CD73FE6AB22E" box="[189,406,346,372]" name="United Kingdom" pageId="15" pageNumber="147">GREAT BRITAIN</collectingCountry>
•
<collectingRegion id="49D7F1ED595FD201FE36CD73FE03B22E" box="[428,511,346,372]" pageId="15" pageNumber="147">Viséan</collectingRegion>
,
<geologicalTimeScale id="F933BFA4595FD201FD93CD72FD24B22E" box="[521,728,346,373]" pageId="15" pageNumber="147">upper Warnantian</geologicalTimeScale>
(late Asbian);
<specimenCount id="9D15F486595FD201FC18CD73FC0DB22E" box="[898,1009,346,372]" pageId="15" pageNumber="147" type="generic">specimen</specimenCount>
from
<collectorName id="26E65AD9595FD201FBACCD73FB5AB22E" box="[1078,1190,346,372]" pageId="15" pageNumber="147">Davidson</collectorName>
Coll., selected by
<bibRefCitation id="EF8242FE595FD201FF27CD57FE5AB2C2" author="Brunton C. H. C." box="[189,422,382,408]" pageId="15" pageNumber="147" pagination="27 - 127" refId="ref9844" refString="Brunton C. H. C. 1984. Silicified brachiopods from the Visean of County Fermanagh, Ireland. (III). Rhynchonellids, spiriferids and terebratulids. Bulletin of the British Museum (Natural History) Geology 38 (2): 27 - 127. Available from https: // www. biodiversitylibrary. org / page / 2273038 [accessed 7 Oct. 2024]." type="journal article" year="1984">Brunton (1984: 101</bibRefCitation>
, fig. 160);
<specimenCode id="DBB59774595FD201FDBCCD57FD38B2C2" box="[550,708,382,408]" pageId="15" pageNumber="147">NHM B7923</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAC3F0F595FD201FF70CD96FC81B145" blockId="15.[189,1399,447,544]" pageId="15" pageNumber="147">
<materialsCitation id="3B7B3552595FD201FF70CD96FC84B145" collectionCode="ONCP" country="France" location="Visean" pageId="15" pageNumber="147" specimenCode="ONCP D306a, ONCP D306e" specimenCount="2" stateProvince="Visean" typeStatus="paratype">
<emphasis id="B967E31D595FD201FF70CD96FE9BB283" bold="true" box="[234,359,447,473]" pageId="15" pageNumber="147">
<typeStatus id="54A881AD595FD201FF70CD96FE9BB283" box="[234,359,447,473]" pageId="15" pageNumber="147" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3047F9F595FD201FF27CDCBFECDB2A6" box="[189,305,482,508]" name="France" pageId="15" pageNumber="147">FRANCE</collectingCountry>
•
<specimenCount id="9D15F486595FD201FECCCDCBFE48B2A7" box="[342,436,482,509]" pageId="15" pageNumber="147">2 specs</specimenCount>
;
<collectingRegion id="49D7F1ED595FD201FE5FCDCBFDEBB2A6" box="[453,535,482,508]" pageId="15" pageNumber="147">Viséan</collectingRegion>
; Office National des Collections Paléontologiques (
<collectionCode id="ED02A7CA595FD201FB0ACDCBFB1FB2A6" box="[1168,1251,482,508]" pageId="15" pageNumber="147">ONCP</collectionCode>
), University C. Bernard, Lyon;
<specimenCode id="DBB59774595FD201FE0ECE2CFDCBB145" box="[404,567,517,543]" pageId="15" pageNumber="147">ONCP D306a</specimenCode>
,
<specimenCode id="DBB59774595FD201FDD8CE2CFD1FB145" box="[578,739,517,543]" pageId="15" pageNumber="147">ONCP D306e</specimenCode>
(
<figureCitation id="1328238A595FD201FD68CE2CFC8CB145" box="[754,880,517,543]" captionStart="Fig" captionStartId="16.[189,232,1646,1672]" captionTargetBox="[208,1351,501,1597]" captionTargetId="figure-62@16.[205,1383,488,1606]" captionTargetPageId="16" captionText="Fig. 6. A. Verneuilia cheiropteryx (d’Archiac &amp;de Verneuil, 1842). Lectotype ONCP D568 (Gourvennec 1994 Coll.) from Refrath (Bergisches Land, Germany). Devonian, Frasnian; dorsal, ventral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 1–5). B–E. Johnsoniana oceani (d’Orbigny, 1850). B. Neotype NHM B7923 (Brunton 1984 Coll.) from Visé (Belgium). Visean; ventral, lateral, dorsal, posterior and anterior views (copy from Brunton 1984: text-fig. 160a–e). C. Paratype CNIGR Museum 3085 (Nalivkin 1979 Coll.) from Bashkortostan (Urals, Russia). Upper Tournaisian; ventral, lateral, dorsal and anterior views (copy from Nalivkin 1979: pl. 62 fig. 21). D. Paratype ONCP Museum D306a (Gourvennec 1994 Coll.) from Visé (Belgium). Viséan; ventral, dorsal, lateral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 11–15). E. Paratype ONCP Museum D306e (Gourvennec 1994 Coll.) from same locality; dorsal, ventral, anterior and lateral views (copy from Gourvennec 1994: pl. 1 figs 16–19). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14047333" httpUri="https://zenodo.org/record/14047333/files/figure.png" pageId="15" pageNumber="147">Fig. 6D–E</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC3F0F595FD201FF27CE6EFC54B1DE" blockId="15.[189,1399,583,645]" pageId="15" pageNumber="147">
<materialsCitation id="3B7B3552595FD201FF27CE6EFC58B1DE" collectingDate="1979" collectionCode="CNIGR" collectorName="Nalivkin" country="Russia" county="Southern Urals" location="Ussuily River" pageId="15" pageNumber="147" specimenCode="3085" specimenCount="17" stateProvince="Bashkortostan">
<collectingCountry id="F3047F9F595FD201FF27CE6EFED9B13B" box="[189,293,583,609]" name="Russia" pageId="15" pageNumber="147">RUSSIA</collectingCountry>
•
<specimenCount id="9D15F486595FD201FEDACE6EFE56B13B" box="[320,426,583,609]" pageId="15" pageNumber="147">17 specs</specimenCount>
;
<collectingCounty id="62CD4783595FD201FE22CE6EFD97B13B" box="[440,619,583,609]" pageId="15" pageNumber="147">Southern Urals</collectingCounty>
(
<collectingRegion id="49D7F1ED595FD201FDE6CE6EFCD6B13B" box="[636,810,583,609]" country="Russia" name="Bashkortostan" pageId="15" pageNumber="147">Bashkortostan</collectingRegion>
),
<location id="8ECC69D4595FD201FCDACE6EFC14B13B" LSID="urn:lsid:plazi:treatment:03BA8E19595ED21EFDABC9ECFABDB2EC:8ECC69D4595FD201FCDACE6EFC14B13B" box="[832,1000,583,609]" country="Russia" county="Southern Urals" name="Ussuily River" pageId="15" pageNumber="147" stateProvince="Bashkortostan">Ussuily River</location>
;
<geologicalTimeScale id="F933BFA4595FD201FC6FCE6EFB32B13B" box="[1013,1230,583,609]" pageId="15" pageNumber="147">upper Tournaisian</geologicalTimeScale>
, Kizelovskian Horizon;
<bibRefCitation id="EF8242FE595FD201FEB7CE43FE2AB1DE" author="Nalivkin D. V." box="[301,470,618,644]" pageId="15" pageNumber="147" refId="ref11601" refString="Nalivkin D. V. 1979. Brakhiopody turneyskogo yarusa Urala. Nauka, Leningrad." type="book" year="1979">
<collectorName id="26E65AD9595FD201FEB7CE43FE68B1DE" box="[301,404,618,644]" pageId="15" pageNumber="147">Nalivkin</collectorName>
<collectingDate id="EFE9E027595FD201FE01CE43FE2AB1DE" box="[411,470,618,644]" pageId="15" pageNumber="147">1979</collectingDate>
</bibRefCitation>
Coll.;
<collectionCode id="ED02A7CA595FD201FDBCCE43FD7DB1DE" box="[550,641,618,644]" collectionName="CNIGR" pageId="15" pageNumber="147">CNIGR</collectionCode>
Museum
<specimenCode id="DBB59774595FD201FD6DCE43FCCDB1DF" box="[759,817,618,645]" pageId="15" pageNumber="147">3085</specimenCode>
(
<figureCitation id="1328238A595FD201FCDACE43FC67B1DE" box="[832,923,618,644]" captionStart="Fig" captionStartId="16.[189,232,1646,1672]" captionTargetBox="[208,1351,501,1597]" captionTargetId="figure-62@16.[205,1383,488,1606]" captionTargetPageId="16" captionText="Fig. 6. A. Verneuilia cheiropteryx (d’Archiac &amp;de Verneuil, 1842). Lectotype ONCP D568 (Gourvennec 1994 Coll.) from Refrath (Bergisches Land, Germany). Devonian, Frasnian; dorsal, ventral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 1–5). B–E. Johnsoniana oceani (d’Orbigny, 1850). B. Neotype NHM B7923 (Brunton 1984 Coll.) from Visé (Belgium). Visean; ventral, lateral, dorsal, posterior and anterior views (copy from Brunton 1984: text-fig. 160a–e). C. Paratype CNIGR Museum 3085 (Nalivkin 1979 Coll.) from Bashkortostan (Urals, Russia). Upper Tournaisian; ventral, lateral, dorsal and anterior views (copy from Nalivkin 1979: pl. 62 fig. 21). D. Paratype ONCP Museum D306a (Gourvennec 1994 Coll.) from Visé (Belgium). Viséan; ventral, dorsal, lateral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 11–15). E. Paratype ONCP Museum D306e (Gourvennec 1994 Coll.) from same locality; dorsal, ventral, anterior and lateral views (copy from Gourvennec 1994: pl. 1 figs 16–19). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14047333" httpUri="https://zenodo.org/record/14047333/files/figure.png" pageId="15" pageNumber="147">Fig. 6C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3096C84595FD201FF27CE82FBA1B53A" pageId="15" pageNumber="147" type="description">
<paragraph id="8BAC3F0F595FD201FF27CE82FEB2B19F" blockId="15.[189,334,683,709]" box="[189,334,683,709]" pageId="15" pageNumber="147">
<heading id="D0E48863595FD201FF27CE82FEB2B19F" bold="true" box="[189,334,683,709]" fontSize="11" level="3" pageId="15" pageNumber="147" reason="3">
<emphasis id="B967E31D595FD201FF27CE82FEB2B19F" bold="true" box="[189,334,683,709]" pageId="15" pageNumber="147">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3F0F595FD201FF27CEF3FC32B09D" blockId="15.[188,1400,730,968]" pageId="15" pageNumber="147">The shell is small with trapezoidal outline; the cardinal extremities are angular but not mucronate. The maximum width coincides with the hinge line. The convexity of the shell is regular but the ventral valve is slightly more convex than the dorsal. The ventral interarea is high, triangular, regularly curved, usually apsacline to catacline; beak is small and curved. The delthyrium is narrow and much higher than the width, the upper half is covered by a convex pseudodeltidium. The surface of the interarea bears strong vertical crenulations that are never branching. These crenulations produce a fine denticulation on the hinge, corresponding to equivalent cupules on the dorsal valve.</paragraph>
<paragraph id="8BAC3F0F595FD201FF27CFC7FBA1B53A" blockId="15.[188,1401,1006,1632]" pageId="15" pageNumber="147">
The dorsal interarea is very narrow, anacline. The angle between ventral and dorsal interareas is 90°– 120°. The ventral sulcus is shallow, widely rounded, tends to become deeper and wider towards the anterior margin. The dorsal sulcus corresponds to the ventral one. The sulci are bordered by low angular plications, that never become carina. The sulci widen regularly during the early stages of growth but bounding plications tend to become parallel or almost convergent. The anterior edge of the plications does not significantly protrude beyond the general anterior outline. The commissure is usually bent towards the ventral valve before abruptly turning towards the dorsal one near the sulci. In this case a more or less developed tongue is present. The flanks are smooth. The microornament consists of faint, regularly spaced growth lines coarser than in
<taxonomicName id="4C13448C595FD201FD4BC921FC18B678" baseAuthorityName="d'Archiac &amp; de Verneuil" baseAuthorityYear="1842" box="[721,996,1288,1314]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="15" pageNumber="147" phylum="Brachiopoda" rank="species" species="cheiropteryx">
<emphasis id="B967E31D595FD201FD4BC921FC18B678" box="[721,996,1288,1314]" italics="true" pageId="15" pageNumber="147">Verneuilia cheiropteryx</emphasis>
</taxonomicName>
. In the ventral valve are very long divergent dental flanges, bordering the inner side of the delthyrium (
<bibRefCitation id="EF8242FE595FD201FC43C902FB16B61C" author="Gourvennec R." box="[985,1258,1323,1350]" pageId="15" pageNumber="147" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec 1994: 580</bibRefCitation>
, fig. 2). The teeth are small. In the apical part of the ventral valve sometimes is a short, low, flat median ridge. In the dorsal valve, the very faint striated ctenophoridium is located on an elevated cardinal platform anteriorly and extended by a narrower median shelf at about ¼ of the length of the shell. The ctenophoridium is bounded by weakly developed very short and narrow sockets. Under the sockets are located the narrow, slightly oblique crural bases, supported by small “columnal” thickenings of the shell united the cardinal platform with crural bases and joining them at the floor of the valve. These very poorly developed thickenings do not properly constitute the crural plates or only rudimentary ones. The direction of the spires is approximately parallel to the plications (after
<bibRefCitation id="EF8242FE595FD201FCA5CA6CFBADB53A" author="Gourvennec R." box="[831,1105,1605,1632]" pageId="15" pageNumber="147" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec 1994: 578</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3096C84595FD21EFF27CAAFFAABB21D" lastPageId="16" lastPageNumber="148" pageId="15" pageNumber="147" type="discussion">
<paragraph id="8BAC3F0F595FD201FF27CAAFFED3B5FA" blockId="15.[189,303,1670,1696]" box="[189,303,1670,1696]" pageId="15" pageNumber="147">
<heading id="D0E48863595FD201FF27CAAFFED3B5FA" bold="true" box="[189,303,1670,1696]" fontSize="11" level="3" pageId="15" pageNumber="147" reason="3">
<emphasis id="B967E31D595FD201FF27CAAFFED3B5FA" bold="true" box="[189,303,1670,1696]" pageId="15" pageNumber="147">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3F0F595FD21EFF27CA9CFAABB21D" blockId="15.[188,1400,1717,1990]" lastBlockId="16.[189,1399,265,327]" lastPageId="16" lastPageNumber="148" pageId="15" pageNumber="147">
The external appearance of
<emphasis id="B967E31D595FD201FD9DCA9CFD08B595" box="[519,756,1717,1743]" italics="true" pageId="15" pageNumber="147">
<taxonomicName id="4C13448C595FD201FD9DCA9CFD61B595" authorityName="Poletaev" authorityYear="2024" box="[519,669,1717,1743]" genus="Johnsoniana" pageId="15" pageNumber="149" rank="genus">Johnsoniana</taxonomicName>
<taxonomicName id="4C13448C595FD201FD3CCA9FFD08B595" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[678,756,1718,1743]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="147" phylum="Cnidaria" rank="species" species="oceani">oceani</taxonomicName>
</emphasis>
is similar to that of
<taxonomicName id="4C13448C595FD201FC7CCA9CFB05B595" baseAuthorityName="d'Archiac &amp; de Verneuil" baseAuthorityYear="1842" box="[998,1273,1717,1743]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="15" pageNumber="147" phylum="Brachiopoda" rank="species" species="cheiropteryx">
<emphasis id="B967E31D595FD201FC7CCA9CFB05B595" box="[998,1273,1717,1743]" italics="true" pageId="15" pageNumber="147">Verneuilia cheiropteryx</emphasis>
</taxonomicName>
(
<figureCitation id="1328238A595FD201FA90CA9CFA95B595" box="[1290,1385,1717,1743]" captionStart="Fig" captionStartId="16.[189,232,1646,1672]" captionTargetBox="[208,1351,501,1597]" captionTargetId="figure-62@16.[205,1383,488,1606]" captionTargetPageId="16" captionText="Fig. 6. A. Verneuilia cheiropteryx (d’Archiac &amp;de Verneuil, 1842). Lectotype ONCP D568 (Gourvennec 1994 Coll.) from Refrath (Bergisches Land, Germany). Devonian, Frasnian; dorsal, ventral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 1–5). B–E. Johnsoniana oceani (d’Orbigny, 1850). B. Neotype NHM B7923 (Brunton 1984 Coll.) from Visé (Belgium). Visean; ventral, lateral, dorsal, posterior and anterior views (copy from Brunton 1984: text-fig. 160a–e). C. Paratype CNIGR Museum 3085 (Nalivkin 1979 Coll.) from Bashkortostan (Urals, Russia). Upper Tournaisian; ventral, lateral, dorsal and anterior views (copy from Nalivkin 1979: pl. 62 fig. 21). D. Paratype ONCP Museum D306a (Gourvennec 1994 Coll.) from Visé (Belgium). Viséan; ventral, dorsal, lateral, anterior and posterior views (copy from Gourvennec 1994: pl. 1 figs 11–15). E. Paratype ONCP Museum D306e (Gourvennec 1994 Coll.) from same locality; dorsal, ventral, anterior and lateral views (copy from Gourvennec 1994: pl. 1 figs 16–19). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14047333" httpUri="https://zenodo.org/record/14047333/files/figure.png" pageId="15" pageNumber="147">Fig. 6A</figureCitation>
), but the former can be distinguished by its smaller, less transverse shell outline, lack of mucronation of the cardinal extremities, denticulate hinge line, much narrower sulcus bordered by low angled but never carinate plications. The plications bordering the sulci of
<taxonomicName id="4C13448C595FD201FCC3CB36FC3FB462" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[857,963,1823,1848]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="147" phylum="Cnidaria" rank="species" species="oceani">
<emphasis id="B967E31D595FD201FCC3CB36FC3FB462" box="[857,963,1823,1848]" italics="true" pageId="15" pageNumber="147">J. oceani</emphasis>
</taxonomicName>
tend to become parallel during adult growth, whereas they are regularly divergent in
<taxonomicName id="4C13448C595FD201FC9BCB6BFC48B406" baseAuthorityName="d'Archiac &amp; de Verneuil" baseAuthorityYear="1842" box="[769,948,1858,1884]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="15" pageNumber="147" phylum="Brachiopoda" rank="species" species="cheiropteryx">
<emphasis id="B967E31D595FD201FC9BCB6BFC48B406" box="[769,948,1858,1884]" italics="true" pageId="15" pageNumber="147">V. cheiropteryx</emphasis>
</taxonomicName>
.
<emphasis id="B967E31D595FD201FC5FCB6BFB4FB401" box="[965,1203,1858,1884]" italics="true" pageId="15" pageNumber="147">
<taxonomicName id="4C13448C595FD201FC5FCB6BFBA7B406" authorityName="Poletaev" authorityYear="2024" box="[965,1115,1858,1884]" genus="Johnsoniana" pageId="15" pageNumber="149" rank="genus">Johnsoniana</taxonomicName>
<taxonomicName id="4C13448C595FD201FBFFCB6BFB4FB401" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[1125,1203,1858,1883]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="147" phylum="Cnidaria" rank="species" species="oceani">oceani</taxonomicName>
</emphasis>
is distinguished from
<taxonomicName id="4C13448C595FD201FF65CB4FFE4CB425" baseAuthorityName="d'Archiac &amp; de Verneuil" baseAuthorityYear="1842" box="[255,432,1893,1919]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="15" pageNumber="147" phylum="Brachiopoda" rank="species" species="cheiropteryx">
<emphasis id="B967E31D595FD201FF65CB4FFE4CB425" box="[255,432,1893,1919]" italics="true" pageId="15" pageNumber="147">V. cheiropteryx</emphasis>
</taxonomicName>
by the absence of short dental plates in the ventral valve.
<emphasis id="B967E31D595FD201FBF4CB4CFAA7B425" box="[1134,1371,1893,1919]" italics="true" pageId="15" pageNumber="147">
<taxonomicName id="4C13448C595FD201FBF4CB4CFAF8B425" authorityName="Poletaev" authorityYear="2024" box="[1134,1284,1893,1919]" genus="Johnsoniana" pageId="15" pageNumber="149" rank="genus">Johnsoniana</taxonomicName>
<taxonomicName id="4C13448C595FD201FA97CB4FFAA7B425" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[1293,1371,1894,1919]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="15" pageNumber="147" phylum="Cnidaria" rank="species" species="oceani">oceani</taxonomicName>
</emphasis>
is distinguished from
<taxonomicName id="4C13448C595FD201FE3ECBA1FD63B4F8" box="[420,671,1928,1954]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="147" phylum="Brachiopoda" rank="species" species="yangi">
<emphasis id="B967E31D595FD201FE3ECBA1FD63B4F8" box="[420,671,1928,1954]" italics="true" pageId="15" pageNumber="147">Changshunella yangi</emphasis>
</taxonomicName>
by the much larger size, lack of prominent median plications in sulcus and opposite rib on dorsal valve, lack of sticks on anterior ends of ribs, bordered sulci. The
<typeStatus id="54A881AD5940D21EFF27CD23FF0CB27E" box="[189,240,266,292]" pageId="16" pageNumber="148">type</typeStatus>
species of
<taxonomicName id="4C13448C5940D21EFEEFCD20FE12B279" authorityName="Hall &amp; Clarke" authorityYear="1893" box="[373,494,265,291]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="16" pageNumber="148" phylum="Brachiopoda" rank="genus">
<emphasis id="B967E31D5940D21EFEEFCD20FE12B279" box="[373,494,265,291]" italics="true" pageId="16" pageNumber="148">Verneuilia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C13448C5940D21EFDB2CD20FD42B279" authorityName="Poletaev" authorityYear="2024" box="[552,702,265,291]" genus="Johnsoniana" pageId="16" pageNumber="149" rank="genus">
<emphasis id="B967E31D5940D21EFDB2CD20FD42B279" box="[552,702,265,291]" italics="true" pageId="16" pageNumber="148">Johnsoniana</emphasis>
</taxonomicName>
also differ in the micro-ornamentation of the shell surface: the presence of fine randomly arranged pits on
<taxonomicName id="4C13448C5940D21EFD7FCD04FC69B21C" baseAuthorityName="d'Archiac &amp; de Verneuil" baseAuthorityYear="1842" box="[741,917,300,326]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="16" pageNumber="148" phylum="Brachiopoda" rank="species" species="cheiropteryx">
<emphasis id="B967E31D5940D21EFD7FCD04FC69B21C" box="[741,917,300,326]" italics="true" pageId="16" pageNumber="148">V. cheiropteryx</emphasis>
</taxonomicName>
and obscure radial striae on
<taxonomicName id="4C13448C5940D21EFB7CCD04FAACB21C" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[1254,1360,301,326]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="16" pageNumber="148" phylum="Cnidaria" rank="species" species="oceani">
<emphasis id="B967E31D5940D21EFB7CCD04FAACB21C" box="[1254,1360,301,326]" italics="true" pageId="16" pageNumber="148">J. oceani</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3096C845940D21EFF27CD47FABDB2EC" pageId="16" pageNumber="148" type="distribution">
<paragraph id="8BAC3F0F5940D21EFF27CD47FEAAB2DD" blockId="16.[189,342,365,392]" box="[189,342,365,392]" pageId="16" pageNumber="148">
<heading id="D0E488635940D21EFF27CD47FEAAB2DD" bold="true" box="[189,342,365,392]" fontSize="11" level="3" pageId="16" pageNumber="148" reason="3">
<emphasis id="B967E31D5940D21EFF27CD47FEAAB2DD" bold="true" box="[189,342,365,392]" pageId="16" pageNumber="148">Dіstribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3F0F5940D21EFF27CDB5FABDB2EC" blockId="16.[189,1345,412,438]" box="[189,1345,412,438]" pageId="16" pageNumber="148">
Calcaire de Visé (Visean) in
<collectingCountry id="F3047F9F5940D21EFD97CDB5FD8AB2EC" box="[525,630,412,438]" name="Belgium" pageId="16" pageNumber="148">Belgium</collectingCountry>
; late Tournaisian of the South Urals (
<collectingRegion id="49D7F1ED5940D21EFBB3CDB5FB2BB2EC" box="[1065,1239,412,438]" country="Russia" name="Bashkortostan" pageId="16" pageNumber="148">Bashkortostan</collectingRegion>
),
<collectingCountry id="F3047F9F5940D21EFB71CDB5FAC1B2EC" box="[1259,1341,412,438]" name="Russia" pageId="16" pageNumber="148">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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