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<mods:title id="9D285E5F3BAA6A9C2FBA89169FA330C9">Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="2C79D07A7FDFD342F40BE1A2A71A2EDC">Zallot, Enrico</mods:namePart>
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<mods:affiliation id="4A4B80B7A2AC2714C6D249C584B95E69">Haagweg 29, 2681 PA Monster, the Netherlands.</mods:affiliation>
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<mods:namePart id="D22F2CD01869127F5C64028F7F836A30">Mattia, Willy De</mods:namePart>
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<mods:affiliation id="06C754E665672CFD9999651D23DDD464">Central Research Laboratories of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria.</mods:affiliation>
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<mods:namePart id="1FCC16ECBFFC66CC03416935E846C845">Fehér, Zoltán</mods:namePart>
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<mods:affiliation id="2EB010AD07EBF6F585B4D2989641162D">Hungarian Natural History Museum, Baross 13, 1088 Budapest, Hungary.</mods:affiliation>
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<mods:namePart id="D4B70D4D480746F19C5D51A147324963">Gittenberger, Edmund</mods:namePart>
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<mods:affiliation id="D0BEE839C9DBF32C73B33B4601D76ABF">Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, the Netherlands.</mods:affiliation>
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<treatment id="038B87913150FFFB7BB014C8F26D2664" ID-DOI="http://doi.org/10.5281/zenodo.5188571" ID-GBIF-Taxon="184096744" ID-Zenodo-Dep="5188571" LSID="urn:lsid:zoobank.org:act:4609B41C-3611-4F8A-9048-5F97EA0446E6" httpUri="http://treatment.plazi.org/id/038B87913150FFFB7BB014C8F26D2664" lastPageId="32" lastPageNumber="81" pageId="29" pageNumber="78">
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<heading id="D0D581EB3150FFC67BB014C8F5652087" box="[511,1077,1195,1222]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="78" reason="2">
<taxonomicName id="4C224D043150FFC67BB014C8F2832087" ID-CoL="8VQSP" authority="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger, 2021" authorityName="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger" authorityYear="2021" box="[511,979,1195,1222]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="29" pageNumber="78" phylum="Mollusca" rank="species" species="stelucarum" status="sp. nov." subGenus="Clessiniella">
<emphasis id="B956EA953150FFC67BB014C8F2832087" bold="true" box="[511,979,1195,1222]" italics="true" pageId="29" pageNumber="78">Cochlostoma (Clessiniella) stelucarum</emphasis>
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<paragraph id="8B9D36873150FFC6781D1491F2B1214D" blockId="29.[377,1211,1195,1292]" box="[594,993,1265,1292]" pageId="29" pageNumber="78">
<figureCitation id="13192A023150FFC6781D1491F3CB214D" box="[594,667,1266,1292]" captionStart="Fig" captionStartId="1.[189,232,1913,1939]" captionTargetBox="[189,1398,1172,1873]" captionTargetId="figure-331@1.[189,1398,1172,1873]" captionTargetPageId="1" captionText="Fig. 1. Map of the sampling localities. Blue dots = Cochlostoma (Clessiniella) villae (Strobel, 1851); red dots = Co. (Cl.) stelucarum sp. nov.; green dots = Co. (Cl.) tergestinum (Westerlund, 1878); yellow dots = Co. (Cl.) waldemari (A.J. Wagner, 1897); orange dots = Co. (Cl.) anomphale Boeckel, 1939." figureDoi="http://doi.org/10.5281/zenodo.5176913" httpUri="https://zenodo.org/record/5176913/files/figure.png" pageId="29" pageNumber="78">
Figs
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<figureCitation id="13192A023150FFC678E51491F39D214D" box="[682,717,1266,1292]" captionStart="Fig" captionStartId="4.[189,232,1854,1880]" captionTargetBox="[199,1389,1004,1815]" captionTargetId="figure-265@4.[199,1389,1004,1815]" captionTargetPageId="4" captionText="Fig. 3. Examples of protoconchs: the red dots mark the end of the protoconchs. A. Cochlostoma (Clessiniella) anomphale Boeckel, 1939 (EZ0034). A1. Co. (Cl.) anomphale (EZ0034): 1 = end of the first smooth part of the protoconch; 2 = end of the protoconch. B. Co. (Cl.) tergestinum (Westerlund, 1878) (EZ0799). C. Co. (Cl.) villae (Strobel, 1851) (EZ0885). D. Co. (Cl.) waldemari (A.J. Wagner, 1897) (WdM6697). E. Co. (Cl.) stelucarum sp. nov. (EZ0104)." figureDoi="http://doi.org/10.5281/zenodo.5176917" httpUri="https://zenodo.org/record/5176917/files/figure.png" pageId="29" pageNumber="78">3E</figureCitation>
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<figureCitation id="13192A023150FFC678961491F25D214D" box="[729,781,1266,1292]" captionStart="Fig" captionStartId="19.[189,232,1814,1840]" captionTargetBox="[189,1398,743,1775]" captionTargetId="figure-148@19.[189,1399,743,1775]" captionTargetPageId="19" captionText="Fig. 14. The five species of Clessiniella Zallot et al., 2015. A–D. Picture of a female (top) and male (bottom) specimen from the type localities. A. Cochlostoma (Clessiniella) anomphale Boeckel, 1939. Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Specimens from a locality close to the type one: Co. (Cl.) waldemari (A.J.Wagner, 1897), Mt Klek, Croatia (EZ0905)." figureDoi="http://doi.org/10.5281/zenodo.5176939" httpUri="https://zenodo.org/record/5176939/files/figure.png" pageId="29" pageNumber="78">14B</figureCitation>
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<figureCitation id="13192A023150FFC679571492F21C214D" box="[792,844,1265,1292]" captionStart="Fig" captionStartId="20.[189,231,960,986]" captionTargetBox="[406,1228,270,878]" captionTargetId="figure-14@20.[406,1232,265,882]" captionTargetPageId="20" captionText="Fig. 15.Comparison of femalegenitalia.A. Cochlostoma (Clessiniella) anomphaleBoeckel, 1939.Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Co. (Cl.) waldemari (A.J.Wagner, 1897), Mt Klek, Croatia (EZ0905)." figureDoi="http://doi.org/10.5281/zenodo.5176941" httpUri="https://zenodo.org/record/5176941/files/figure.png" pageId="29" pageNumber="78">15B</figureCitation>
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<figureCitation id="13192A023150FFC679171491F2B1214D" box="[856,993,1265,1292]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig 25" captionStart-11="Fig 26" captionStart-12="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="20.[189,232,1879,1905]" captionStartId-1="20.[189,231,960,986]" captionStartId-10="30.[189,230,1922,1948]" captionStartId-11="31.[189,230,1925,1951]" captionStartId-12="32.[189,232,1854,1880]" captionStartId-2="21.[189,232,1911,1937]" captionStartId-3="22.[190,233,1911,1937]" captionStartId-4="23.[189,232,1953,1979]" captionStartId-5="24.[189,232,1953,1979]" captionStartId-6="25.[189,232,1958,1984]" captionStartId-7="26.[189,232,1920,1946]" captionStartId-8="27.[189,232,1920,1946]" captionStartId-9="28.[189,232,1958,1984]" captionTargetBox-0="[401,1174,1156,1820]" captionTargetBox-1="[406,1228,270,878]" captionTargetBox-10="[193,1395,794,1884]" captionTargetBox-11="[189,1370,1213,1868]" captionTargetBox-12="[189,1398,656,1802]" captionTargetBox-2="[189,1398,980,1873]" captionTargetBox-3="[196,1362,1131,1762]" captionTargetBox-4="[275,1312,265,1914]" captionTargetBox-5="[189,1398,694,1914]" captionTargetBox-6="[293,1295,265,1920]" captionTargetBox-7="[189,1398,1002,1882]" captionTargetBox-8="[209,1378,1136,1869]" captionTargetBox-9="[376,1212,265,1920]" captionTargetId-0="figure-84@20.[401,1187,1137,1828]" captionTargetId-1="figure-14@20.[406,1232,265,882]" captionTargetId-10="figure-171@30.[193,1395,794,1884]" captionTargetId-11="figure-274@31.[189,1398,1185,1888]" captionTargetId-12="figure-136@32.[189,1398,656,1802]" captionTargetId-2="figure-168@21.[189,1398,980,1873]" captionTargetId-3="figure-221@22.[189,1398,1092,1762]" captionTargetId-4="figure-11@23.[275,1312,265,1914]" captionTargetId-5="figure-152@24.[189,1398,694,1914]" captionTargetId-6="figure-11@25.[293,1295,265,1920]" captionTargetId-7="figure-134@26.[189,1398,1002,1882]" captionTargetId-8="figure-245@27.[189,1398,1129,1883]" captionTargetId-9="figure-15@28.[376,1212,265,1920]" captionTargetPageId-0="20" captionTargetPageId-1="20" captionTargetPageId-10="30" captionTargetPageId-11="31" captionTargetPageId-12="32" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="23" captionTargetPageId-5="24" captionTargetPageId-6="25" captionTargetPageId-7="26" captionTargetPageId-8="27" captionTargetPageId-9="28" captionText-0="Fig. 16.Comparisonof male genitalia.A. Cochlostoma (Clessiniella)anomphaleBoeckel, 1939.Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Co. (Cl.) waldemari (A.J.Wagner, 1897), Tisovec, Croatia (WdM6750)." captionText-1="Fig. 15.Comparison of femalegenitalia.A. Cochlostoma (Clessiniella) anomphaleBoeckel, 1939.Vellach, Austria (EZ0991). B. Co. (Cl.) stelucarum sp. nov., Cornino, Italy (EZ0104). C. Co. (Cl.) tergestinum (Westerlund, 1878), Val Rosandra, Italy (EZ0840). D. Co. (Cl.) villae (Strobel, 1851), Mizzole, Italy (EZ0878). E. Co. (Cl.) waldemari (A.J.Wagner, 1897), Mt Klek, Croatia (EZ0905)." captionText-10="Fig 25. Shell variability in Cochlostoma (Clessiniella) stelucarum sp. nov. A. Cornino, Italy (EZ0104). B. Mt Valinis, Italy (EZ0096). C. Anduins, Italy (EZ0102). D. Mt Prat, Italy (EZ0103). E. Val di Torre, Italy (EZ0106)." captionText-11="Fig 26.Variation of the female genitalia in Cochlostoma (Clessiniella) stelucarum sp. nov.A. Mt Valinis, Italy (EZ0096). B. Campone, Italy (EZ0098). C. Anduins, Italy (EZ0102). D1–D3. Cornino, Italy (EZ0104). E. Cjanet-Peonis, Italy (EZ0105). F. Mt Bernadia, Italy (EZ0107)." captionText-12="Fig. 27. Cochlostoma (Clessiniella) stelucarum sp. nov. Top left: holotype, ♀ (RMNH.MOL.347617), H = 8 mm. Top right: paratype, ♂ (RMNH.MOL.347618), H = 7 mm. Bottom: paratype, ♀ (RMNH. MOL.347619), empty shell, H = 8.7 mm." captionText-2="Fig. 17. Shell variability in Cochlostoma (Clessiniella) villae (Strobel, 1851). A. Mizzole, Italy (EZ0878). B. Covoli di Velo, Italy (EZ0879). C. Ferrazza, Italy (EZ0881). D. Mt delle Piume, Italy (EZ0889). E. Perarolo, Italy (EZ0891). F. Lumignano, Italy (EZ0893). G. Ca’ Gottolo, Italy (EZ1015)." captionText-3="Fig. 18. Variation of the female genitalia in Cochlostoma (Clessiniella) villae (Strobel, 1851). A. Mizzole, Italy (EZ0878). B. Ferrazza, Italy (EZ0881). C. Mt delle Piume, Italy (EZ0889). D1– D3. Perarolo, Italy (EZ0891). E. Mt Rusta, Italy (EZ0894). F. Ca’ Gottolo, Italy (EZ1015)." captionText-4="Fig. 19. Cochlostoma (Clessiniella) villae (Strobel, 1851), syntype, ♀, at the Museo di Storia Naturale of Parma University, Italy (PU1610-C1)." captionText-5="Fig. 20. Cochlostoma (Clessiniella) villae (Strobel, 1851), syntypes, ♂♂, at the Museo di Storia Naturale of Parma University, Italy (1610-C2, left; 1610-C3, right)." captionText-6="Fig. 21. Shell of Cochlostoma (Clessiniella) villae (Strobel, 1851). Collezione De Betta, Museo di Storia Naturale di Verona (photo Folco Giusti)." captionText-7="Fig. 22.Shell variability in Cochlostoma (Clessiniella) anomphale Boeckel, 1939.A. Savinjska Dolina, Slovenia (WdM1579). B. Eisenkappel, Austria (EZ0034).C. Stahovika, Slovenia (EZ0035).D. Vellach, Austria (EZ0991). E. Solčava, Slovenia (WdM7010). F. Logarska Dolina, Slovenia (EZ0994)." captionText-8="Fig. 23. Variation of the female genitalia in Cochlostoma (Clesssiniella) anomphale Boeckel, 1939. A. Eisenkappel, Austria (EZ0034). B. Savinjska Dolina, Slovenia (WdM1579). C. Vellach, Austria (EZ0991). D. Logarska Dolina, Slovenia (EZ0994). E1–E2. Solčava, Slovenia (WdM7010)." captionText-9="Fig. 24. Cochlostoma (Clessiniella) anomphale Boeckel, 1939. Paratypes at the Senckenberg Museum Frankfurt (SMF42967/2)." figureDoi-0="http://doi.org/10.5281/zenodo.5176943" figureDoi-1="http://doi.org/10.5281/zenodo.5176941" figureDoi-10="http://doi.org/10.5281/zenodo.5176963" figureDoi-11="http://doi.org/10.5281/zenodo.5176965" figureDoi-12="http://doi.org/10.5281/zenodo.5176967" figureDoi-2="http://doi.org/10.5281/zenodo.5176945" figureDoi-3="http://doi.org/10.5281/zenodo.5176947" figureDoi-4="http://doi.org/10.5281/zenodo.5176949" figureDoi-5="http://doi.org/10.5281/zenodo.5176951" figureDoi-6="http://doi.org/10.5281/zenodo.5176953" figureDoi-7="http://doi.org/10.5281/zenodo.5176955" figureDoi-8="http://doi.org/10.5281/zenodo.5176957" figureDoi-9="http://doi.org/10.5281/zenodo.5176961" httpUri-0="https://zenodo.org/record/5176943/files/figure.png" httpUri-1="https://zenodo.org/record/5176941/files/figure.png" httpUri-10="https://zenodo.org/record/5176963/files/figure.png" httpUri-11="https://zenodo.org/record/5176965/files/figure.png" httpUri-12="https://zenodo.org/record/5176967/files/figure.png" httpUri-2="https://zenodo.org/record/5176945/files/figure.png" httpUri-3="https://zenodo.org/record/5176947/files/figure.png" httpUri-4="https://zenodo.org/record/5176949/files/figure.png" httpUri-5="https://zenodo.org/record/5176951/files/figure.png" httpUri-6="https://zenodo.org/record/5176953/files/figure.png" httpUri-7="https://zenodo.org/record/5176955/files/figure.png" httpUri-8="https://zenodo.org/record/5176957/files/figure.png" httpUri-9="https://zenodo.org/record/5176961/files/figure.png" pageId="29" pageNumber="78">16B, 15–27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C338650C3150FFC67AF21550F5BA21DE" pageId="29" pageNumber="78" type="diagnosis">
<paragraph id="8B9D36873150FFC67AF21550F066210C" blockId="29.[189,310,1331,1357]" box="[189,310,1331,1357]" pageId="29" pageNumber="78">
<heading id="D0D581EB3150FFC67AF21550F066210C" bold="true" box="[189,310,1331,1357]" fontSize="11" level="3" pageId="29" pageNumber="78" reason="3">
<emphasis id="B956EA953150FFC67AF21550F066210C" bold="true" box="[189,310,1331,1357]" pageId="29" pageNumber="78">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873150FFC67AF21501F5BA21DE" blockId="29.[188,1398,1376,1439]" pageId="29" pageNumber="78">Slender shell with 8½–10½ rather widely ribbed whorls. Spots always visible. Poorly developed lip with FF or AF external lobe. Visceral oviduct simplified and seminal receptacle sometimes oval.</paragraph>
</subSubSection>
<subSubSection id="C338650C3150FFC67AF215A5F5612273" pageId="29" pageNumber="78" type="etymology">
<paragraph id="8B9D36873150FFC67AF215A5F01421A1" blockId="29.[189,324,1478,1504]" box="[189,324,1478,1504]" pageId="29" pageNumber="78">
<heading id="D0D581EB3150FFC67AF215A5F01421A1" bold="true" box="[189,324,1478,1504]" fontSize="11" level="3" pageId="29" pageNumber="78" reason="3">
<emphasis id="B956EA953150FFC67AF215A5F01421A1" bold="true" box="[189,324,1478,1504]" pageId="29" pageNumber="78">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873150FFC67AF21596F5612273" blockId="29.[189,1398,1525,1587]" pageId="29" pageNumber="78">
The epithet ‘
<emphasis id="B956EA953150FFC67B1C1596F0F6224E" box="[339,422,1525,1551]" italics="true" pageId="29" pageNumber="78">steluca</emphasis>
’ is an acronym noun formed from the initial syllables of the names of the wife and two daughters (Stefania, Ludovica and Camilla Zallot) of the senior author.
</paragraph>
</subSubSection>
<subSubSection id="C338650C3150FFC57AF21639F52726CF" lastPageId="30" lastPageNumber="79" pageId="29" pageNumber="78" type="materials_examined">
<paragraph id="8B9D36873150FFC67AF21639F08A2235" blockId="29.[189,474,1625,1652]" box="[189,474,1625,1652]" pageId="29" pageNumber="78">
<emphasis id="B956EA953150FFC67AF21639F03C2232" bold="true" box="[189,364,1625,1652]" pageId="29" pageNumber="78">Type material</emphasis>
(
<figureCitation id="13192A023150FFC67B33163AF0812235" box="[380,465,1625,1652]" captionStart="Fig" captionStartId="32.[189,232,1854,1880]" captionTargetBox="[189,1398,656,1802]" captionTargetId="figure-136@32.[189,1398,656,1802]" captionTargetPageId="32" captionText="Fig. 27. Cochlostoma (Clessiniella) stelucarum sp. nov. Top left: holotype, ♀ (RMNH.MOL.347617), H = 8 mm. Top right: paratype, ♂ (RMNH.MOL.347618), H = 7 mm. Bottom: paratype, ♀ (RMNH. MOL.347619), empty shell, H = 8.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5176967" httpUri="https://zenodo.org/record/5176967/files/figure.png" pageId="29" pageNumber="78">Fig. 27</figureCitation>
)
</paragraph>
<paragraph id="8B9D36873150FFC67AA516EBF57C22A9" blockId="29.[189,1398,1672,1769]" pageId="29" pageNumber="78">
<materialsCitation id="3B4A3CDA3150FFC67AA516EBF57C22A9" ID-GBIF-Occurrence="3333861325" collectedFrom="rocks surrounding the lake" collectingDate="2002-06-05" collectionCode="RMNH" collectorName="Zallot" country="Italy" county="Venezia Giulia" elevation="180" latitude="46.2297" location="Friuli Venezia Giulia" longLatPrecision="6" longitude="13.025" municipality="Udine" pageId="29" pageNumber="78" specimenCode="RMNH.MOL.347617." specimenCount="1" specimenCount-female="1" stateProvince="Friuli-Venezia Giulia" typeStatus="holotype">
<emphasis id="B956EA953150FFC67AA516EBF00C22E3" bold="true" box="[234,348,1672,1698]" pageId="29" pageNumber="78">
<typeStatus id="549988253150FFC67AA516EBF00C22E3" box="[234,348,1672,1698]" pageId="29" pageNumber="78" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F33576173150FFC67AF216C8F05F2284" box="[189,271,1707,1733]" name="Italy" pageId="29" pageNumber="78">ITALY</collectingCountry>
•
<specimenCount id="9D24FD0E3150FFC67B6516CFF0102287" box="[298,320,1708,1734]" pageId="29" pageNumber="78" type="female">♀</specimenCount>
(in ethanol);
<location id="8EFD605C3150FFC67BAC16C8F38E2284" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3150FFC67BAC16C8F38E2284" box="[483,734,1707,1733]" country="Italy" county="Venezia Giulia" latitude="46.2297" longLatPrecision="6" longitude="13.025" municipality="Udine" name="Friuli Venezia Giulia" pageId="29" pageNumber="78" stateProvince="Friuli-Venezia Giulia">Friuli Venezia Giulia</location>
,
<collectingMunicipality id="6BF9ACFD3150FFC678A416C8F2662284" box="[747,822,1707,1733]" pageId="29" pageNumber="78">Udine</collectingMunicipality>
,
<location id="8EFD605C3150FFC6790C16C8F2F72284" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3150FFC6790C16C8F2F72284" box="[835,935,1707,1733]" country="Italy" county="Venezia Giulia" latitude="46.2297" longLatPrecision="6" longitude="13.025" municipality="Udine" name="Cornino" pageId="29" pageNumber="78" stateProvince="Friuli-Venezia Giulia">Cornino</location>
;
<geoCoordinate id="EE1650403150FFC679FB16C8F5102287" box="[948,1088,1707,1734]" degrees="46.2297" direction="north" orientation="latitude" pageId="29" pageNumber="78" precision="5" value="46.2297">46.2297° N</geoCoordinate>
,
<geoCoordinate id="EE1650403150FFC67E0216C8F5862287" box="[1101,1238,1707,1734]" degrees="13.0250" direction="east" orientation="longitude" pageId="29" pageNumber="78" precision="5" value="13.025">13.0250° E</geoCoordinate>
;
<elevation id="000FD1B43150FFC67EAB16C8F4202284" box="[1252,1392,1707,1734]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="78" unit="m" value="180.0">
<quantity id="4CDA9B623150FFC67EAB16C8F47F2287" box="[1252,1327,1707,1734]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="78" unit="m" value="180.0">180 m</quantity>
a.s.l.
</elevation>
;
<date id="FF9C10473150FFC67AF216ADF01822A9" box="[189,328,1742,1769]" pageId="29" pageNumber="78" value="2002-06-05">
<collectingDate id="EFD8E9AF3150FFC67AF216ADF01822A9" box="[189,328,1742,1769]" pageId="29" pageNumber="78" value="2002-06-05">5 Jun. 2002</collectingDate>
</date>
;
<collectorName id="26D753513150FFC67B1A16ADF0CB22A9" box="[341,411,1742,1768]" pageId="29" pageNumber="78">Zallot</collectorName>
leg.;
<collectedFrom id="383E757A3150FFC67B9216ADF24922A9" box="[477,793,1742,1768]" pageId="29" pageNumber="78">rocks surrounding the lake</collectedFrom>
;
<specimenCode id="DB849EFC3150FFC6796B16ADF57C22A9" box="[804,1068,1742,1768]" pageId="29" pageNumber="78">RMNH.MOL.347617.</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="8B9D36873150FFC67AA51773F43D2331" blockId="29.[189,1398,1808,1905]" pageId="29" pageNumber="78">
<materialsCitation id="3B4A3CDA3150FFC67AA51773F59C230C" ID-GBIF-Occurrence="3333861437" collectedFrom="rocks surrounding the lake" collectingDate="2002-06-05" collectionCode="RMNH" collectorName="Zallot" country="Italy" county="Venezia Giulia" elevation="180" latitude="46.2297" location="ITALY" longLatPrecision="6" longitude="13.025" municipality="Udine" pageId="29" pageNumber="78" specimenCode="RMNH.MOL.347618" specimenCount="1" specimenCount-male="1" stateProvince="Friuli-Venezia Giulia" typeStatus="paratype">
<emphasis id="B956EA953150FFC67AA51773F037236B" bold="true" box="[234,359,1808,1834]" pageId="29" pageNumber="78">
<typeStatus id="549988253150FFC67AA51773F037236B" box="[234,359,1808,1834]" pageId="29" pageNumber="78" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F33576173150FFC67AF21750F05F230C" box="[189,271,1843,1869]" name="Italy" pageId="29" pageNumber="78">ITALY</collectingCountry>
•
<specimenCount id="9D24FD0E3150FFC67B621750F00C230F" box="[301,348,1843,1870]" pageId="29" pageNumber="78" type="male">1 ♂</specimenCount>
(in ethanol); same collection data as for holotype;
<specimenCode id="DB849EFC3150FFC679831750F59C230C" box="[972,1228,1843,1869]" pageId="29" pageNumber="78">RMNH.MOL.347618</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3150FFC67EA51750F4382331" ID-GBIF-Occurrence="3333861315" accessionNumber="KX120830" collectedFrom="rocks surrounding the lake" collectingDate="2002-06-05" collectionCode="RMNH" collectorName="Zallot" country="Italy" county="Venezia Giulia" elevation="180" latitude="46.2297" location="Cornino" longLatPrecision="6" longitude="13.025" municipality="Udine" pageId="29" pageNumber="78" specimenCode="RMNH.MOL.347619" specimenCount="1" specimenCount-female="1" stateProvince="Friuli-Venezia Giulia" typeStatus="paratype">
<specimenCount id="9D24FD0E3150FFC67EA51750F449230F" box="[1258,1305,1843,1870]" pageId="29" pageNumber="78" type="female">1 ♀</specimenCount>
(empty shell); same collection data as for holotype; GenBank voucher H3:
<accessionNumber id="9471AB643150FFC6799C1735F5092331" box="[979,1113,1878,1904]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX120830" pageId="29" pageNumber="78">KX120830</accessionNumber>
;
<specimenCode id="DB849EFC3150FFC67E2A1735F4382331" box="[1125,1384,1878,1904]" pageId="29" pageNumber="78">RMNH.MOL.347619</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9D36873153FFC57AF2116BF0292563" blockId="30.[189,377,264,290]" box="[189,377,264,290]" pageId="30" pageNumber="79">
<heading id="D0D581EB3153FFC57AF2116BF0292563" bold="true" box="[189,377,264,290]" fontSize="11" level="3" pageId="30" pageNumber="79" reason="3">
<emphasis id="B956EA953153FFC57AF2116BF0292563" bold="true" box="[189,377,264,290]" pageId="30" pageNumber="79">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873153FFC57AF21154F52726CF" blockId="30.[189,1399,311,654]" pageId="30" pageNumber="79">
<materialsCitation id="3B4A3CDA3153FFC57AF21154F5172510" ID-GBIF-Occurrence="3333861422" box="[189,1095,311,338]" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.23755" location="Anduins-cliffs" longLatPrecision="1" longitude="12.97106" pageId="30" pageNumber="79" specimenCode="EZ0102" specimenCount="1">
<collectingCountry id="F33576173153FFC57AF21154F05F2510" box="[189,271,311,337]" name="Italy" pageId="30" pageNumber="79">ITALY</collectingCountry>
•
<location id="8EFD605C3153FFC57B6C1154F0802510" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57B6C1154F0802510" box="[291,464,311,337]" country="Italy" latitude="46.23755" longLatPrecision="1" longitude="12.97106" name="Anduins-cliffs" pageId="30" pageNumber="79">Anduins-cliffs</location>
;
<geoCoordinate id="EE1650403153FFC57B951154F3212513" box="[474,625,311,338]" degrees="46.23755" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.23755">46.23755° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC578341154F2402513" box="[635,784,311,338]" degrees="12.97106" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="12.97106">12.97106° E</geoCoordinate>
; 1997;
<collectorName id="26D753513153FFC5792C1154F2F92510" box="[867,937,311,337]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC579A71154F5172510" box="[1000,1095,311,337]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0102</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57E131154F3D52535" ID-GBIF-Occurrence="3333861372" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.26194" location="Campone" longLatPrecision="1" longitude="12.83333" pageId="30" pageNumber="79" specimenCode="EZ0098" specimenCount="1">
<location id="8EFD605C3153FFC57E131154F5802510" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57E131154F5802510" box="[1116,1232,311,337]" country="Italy" latitude="46.26194" longLatPrecision="1" longitude="12.83333" name="Campone" pageId="30" pageNumber="79">Campone</location>
;
<geoCoordinate id="EE1650403153FFC57E951154F4222513" box="[1242,1394,311,338]" degrees="46.26194" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.26194">46.26194° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC57AF21139F0012534" box="[189,337,346,373]" degrees="12.83333" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="12.83333">12.83333° E</geoCoordinate>
; 2010;
<collectorName id="26D753513153FFC57BEC1139F0B92535" box="[419,489,346,372]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC578681139F3D52535" box="[551,645,346,372]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0098</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC578D51139F04C25D9" ID-GBIF-Occurrence="3333861385" collectingDate="2010-10" collectionCode="EZ" collectorName="Margelli" country="Italy" latitude="46.094" location="Castelmonte" longLatPrecision="55" longitude="13.52095" pageId="30" pageNumber="79" specimenCode="EZ1020" specimenCount="1">
<location id="8EFD605C3153FFC578D51139F2612535" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC578D51139F2612535" box="[666,817,346,372]" country="Italy" latitude="46.094" longLatPrecision="55" longitude="13.52095" name="Castelmonte" pageId="30" pageNumber="79">Castelmonte</location>
;
<geoCoordinate id="EE1650403153FFC579751139F2E32534" box="[826,947,346,373]" degrees="46.094" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="55" value="46.094">46.094° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC579F21139F5012534" box="[957,1105,346,373]" degrees="13.52095" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="13.52095">13.52095° E</geoCoordinate>
;
<date id="FF9C10473153FFC57E141139F5802535" box="[1115,1232,346,372]" pageId="30" pageNumber="79" value="2010-10">
<collectingDate id="EFD8E9AF3153FFC57E141139F5802535" box="[1115,1232,346,372]" pageId="30" pageNumber="79" value="2010-10">Oct. 2010</collectingDate>
</date>
;
<collectorName id="26D753513153FFC57E951139F46E2535" box="[1242,1342,346,372]" pageId="30" pageNumber="79">Margelli</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC57AF2111DF04C25D9" box="[189,284,382,408]" collectionCode="EZ" pageId="30" pageNumber="79">EZ1020</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57B7D111EF50425D9" ID-GBIF-Occurrence="3333861330" box="[306,1108,381,408]" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.25917" location="Cjanet-Peonis" longLatPrecision="1" longitude="13.04111" pageId="30" pageNumber="79" specimenCode="EZ0105" specimenCount="1">
<location id="8EFD605C3153FFC57B7D111EF08B25D6" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57B7D111EF08B25D6" box="[306,475,381,407]" country="Italy" latitude="46.25917" longLatPrecision="1" longitude="13.04111" name="Cjanet-Peonis" pageId="30" pageNumber="79">Cjanet-Peonis</location>
;
<geoCoordinate id="EE1650403153FFC57BAA111EF32E25D9" box="[485,638,381,408]" degrees="46.25917" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.25917">46.25917° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC578C7111DF24B25D9" box="[648,795,382,408]" degrees="13.04111" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="13.04111">13.04111° E</geoCoordinate>
; 2002;
<collectorName id="26D753513153FFC5793F111EF2E625D6" box="[880,950,381,407]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC579B9111EF50425D9" box="[1014,1108,381,408]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0105</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57E24111EF32D25FA" ID-GBIF-Occurrence="3333861363" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.22972" location="Cornino" longLatPrecision="38" longitude="13.025" pageId="30" pageNumber="79" specimenCode="EZ0104" specimenCount="1">
<location id="8EFD605C3153FFC57E24111EF59F25D6" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57E24111EF59F25D6" box="[1131,1231,381,407]" country="Italy" latitude="46.22972" longLatPrecision="38" longitude="13.025" name="Cornino" pageId="30" pageNumber="79">Cornino</location>
;
<geoCoordinate id="EE1650403153FFC57E95111EF42225D9" box="[1242,1394,381,408]" degrees="46.22972" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.22972">46.22972° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC57AF211C3F06825FA" box="[189,312,416,443]" degrees="13.025" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="55" value="13.025">13.025° E</geoCoordinate>
; 2002;
<collectorName id="26D753513153FFC57BDD11C2F08825FA" box="[402,472,417,443]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC5785111C2F32D25FA" box="[542,637,417,443]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0104</specimenCode>
</materialsCitation>
•
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F. Piccola
<location id="8EFD605C3153FFC5795711C2F2F725FA" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC5795711C2F2F725FA" box="[792,935,417,443]" country="Italy" latitude="46.245" longLatPrecision="55" longitude="12.80111" name="Mt Valinins" pageId="30" pageNumber="79">Mt Valinins</location>
;
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,
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; 2000;
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leg.;
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•
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<location id="8EFD605C3153FFC57B3311A7F347259F" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57B3311A7F347259F" box="[380,535,452,478]" country="Italy" latitude="46.23361" longLatPrecision="1" longitude="13.25556" name="Mt Bernadia" pageId="30" pageNumber="79">Mt Bernadia</location>
;
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,
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; 2005;
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leg.;
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</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57E9211A7F2612640" ID-GBIF-Occurrence="3333861347" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.21226" location="Mt Ciaurlec" longLatPrecision="1" longitude="12.88018" pageId="30" pageNumber="79" specimenCode="EZ0099" specimenCount="1">
<location id="8EFD605C3153FFC57E9211A7F422259F" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57E9211A7F422259F" box="[1245,1394,452,478]" country="Italy" latitude="46.21226" longLatPrecision="1" longitude="12.88018" name="Mt Ciaurlec" pageId="30" pageNumber="79">Mt Ciaurlec</location>
;
<geoCoordinate id="EE1650403153FFC57AF21184F0052643" box="[189,341,487,514]" degrees="46.21226" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.21226">46.21226° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC57B2E1184F0A62643" box="[353,502,487,514]" degrees="12.88018" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="12.88018">12.88018° E</geoCoordinate>
; 2005;
<collectorName id="26D753513153FFC578031184F3C22640" box="[588,658,487,513]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
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</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57906118BF0092665" ID-GBIF-Occurrence="3333861399" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.24577" location="Mt Prat" longLatPrecision="3" longitude="12.9907" pageId="30" pageNumber="79" specimenCode="EZ0103" specimenCount="1">
<location id="8EFD605C3153FFC57906118BF2F52643" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57906118BF2F52643" box="[841,933,488,514]" country="Italy" latitude="46.24577" longLatPrecision="3" longitude="12.9907" name="Mt Prat" pageId="30" pageNumber="79">Mt Prat</location>
;
<geoCoordinate id="EE1650403153FFC579FF1184F5192643" box="[944,1097,487,514]" degrees="46.24577" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.24577">46.24577° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC57E1B1184F58B2643" box="[1108,1243,487,514]" degrees="12.9907" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="5" value="12.9907">12.9907° E</geoCoordinate>
; 2002;
<collectorName id="26D753513153FFC57F7F1184F4262640" box="[1328,1398,487,513]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
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</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57B351268F5DD2665" ID-GBIF-Occurrence="3333861308" box="[378,1165,522,549]" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.23417" location="Mt Valinis" longLatPrecision="3" longitude="12.8075" pageId="30" pageNumber="79" specimenCode="EZ0096" specimenCount="1">
<location id="8EFD605C3153FFC57B351268F0AB2665" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57B351268F0AB2665" box="[378,507,522,549]" country="Italy" latitude="46.23417" longLatPrecision="3" longitude="12.8075" name="Mt Valinis" pageId="30" pageNumber="79">Mt Valinis</location>
;
<geoCoordinate id="EE1650403153FFC578441269F3F72664" box="[523,679,522,549]" degrees="46.23417" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.23417">46.23417° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC578F81269F2112664" box="[695,833,522,549]" degrees="12.8075" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="5" value="12.8075">12.8075° E</geoCoordinate>
; 2002;
<collectorName id="26D753513153FFC579D01269F2B52665" box="[927,997,522,548]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
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</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57EE21269F2782609" ID-GBIF-Occurrence="3333861445" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.2337" location="San Vito D'asio" longLatPrecision="5" longitude="12.94531" pageId="30" pageNumber="79" specimenCode="EZ0101" specimenCount="1">
<location id="8EFD605C3153FFC57EE21269F4222665" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57EE21269F4222665" box="[1197,1394,522,548]" country="Italy" latitude="46.2337" longLatPrecision="5" longitude="12.94531" name="San Vito D'asio" pageId="30" pageNumber="79">San Vito D’asio</location>
;
<geoCoordinate id="EE1650403153FFC57AF2124EF0182609" box="[189,328,557,584]" degrees="46.2337" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="5" value="46.2337">46.2337° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC57B1B124EF0BA2609" box="[340,490,557,584]" degrees="12.94531" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="12.94531">12.94531° E</geoCoordinate>
; 1997;
<collectorName id="26D753513153FFC5780E124EF3D72606" box="[577,647,557,583]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
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</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC5790F124EF0A8262A" ID-GBIF-Occurrence="3333861357" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.27833" location="Tramonti-Campone" longLatPrecision="1" longitude="12.81917" pageId="30" pageNumber="79" specimenCode="EZ0097" specimenCount="1">
<location id="8EFD605C3153FFC5790F124EF57A2606" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC5790F124EF57A2606" box="[832,1066,557,583]" country="Italy" latitude="46.27833" longLatPrecision="1" longitude="12.81917" name="Tramonti-Campone" pageId="30" pageNumber="79">Tramonti-Campone</location>
;
<geoCoordinate id="EE1650403153FFC57E79124EF59F2609" box="[1078,1231,557,584]" degrees="46.27833" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.27833">46.27833° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC57E94124DF4212609" box="[1243,1393,558,584]" degrees="12.81917" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="12.81917">12.81917° E</geoCoordinate>
; 2000;
<collectorName id="26D753513153FFC57B441232F001262A" box="[267,337,593,619]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC57BD61232F0A8262A" box="[409,504,593,619]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0097</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC578581232F04C26CF" ID-GBIF-Occurrence="3333861433" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.23861" location="Val di Torre-all'imbocco" longLatPrecision="1" longitude="13.23111" pageId="30" pageNumber="79" specimenCode="EZ0106" specimenCount="1">
<location id="8EFD605C3153FFC578581232F213262A" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC578581232F213262A" box="[535,835,593,619]" country="Italy" latitude="46.23861" longLatPrecision="1" longitude="13.23111" name="Val di Torre-all'imbocco" pageId="30" pageNumber="79">Val di Torre-all’imbocco</location>
;
<geoCoordinate id="EE1650403153FFC5791D1233F2BE262A" box="[850,1006,592,619]" degrees="46.23861" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.23861">46.23861° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC579B11232F5C5262A" box="[1022,1173,593,619]" degrees="13.23111" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="13.23111">13.23111° E</geoCoordinate>
; 1998;
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leg.;
<specimenCode id="DB849EFC3153FFC57AF21217F04C26CF" box="[189,284,628,654]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0106</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B4A3CDA3153FFC57B7B1217F52326CF" ID-GBIF-Occurrence="3333861381" box="[308,1139,628,654]" collectionCode="EZ" collectorName="Zallot" country="Italy" latitude="46.30278" location="Valle D'Arzino" longLatPrecision="1" longitude="12.92972" pageId="30" pageNumber="79" specimenCode="EZ0100" specimenCount="1">
<location id="8EFD605C3153FFC57B7B1217F0BC26CF" LSID="urn:lsid:plazi:treatment:038B87913150FFFB7BB014C8F26D2664:8EFD605C3153FFC57B7B1217F0BC26CF" box="[308,492,628,654]" country="Italy" latitude="46.30278" longLatPrecision="1" longitude="12.92972" name="Valle D'Arzino" pageId="30" pageNumber="79">Valle D’Arzino</location>
;
<geoCoordinate id="EE1650403153FFC57BB81217F3C026CF" box="[503,656,628,654]" degrees="46.30278" direction="north" orientation="latitude" pageId="30" pageNumber="79" precision="1" value="46.30278">46.30278° N</geoCoordinate>
,
<geoCoordinate id="EE1650403153FFC578D31217F26226CF" box="[668,818,628,654]" degrees="12.92972" direction="east" orientation="longitude" pageId="30" pageNumber="79" precision="1" value="12.92972">12.92972° E</geoCoordinate>
; 2000;
<collectorName id="26D753513153FFC579C61217F29F26CF" box="[905,975,628,654]" pageId="30" pageNumber="79">Zallot</collectorName>
leg.;
<specimenCode id="DB849EFC3153FFC57E5E1217F52326CF" box="[1041,1139,628,654]" collectionCode="EZ" pageId="30" pageNumber="79">EZ0100</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5D660F3153FFC57AF217E1F02523A2" ID-DOI="http://doi.org/10.5281/zenodo.5176963" ID-Zenodo-Dep="5176963" httpUri="https://zenodo.org/record/5176963/files/figure.png" pageId="30" pageNumber="79" startId="30.[189,230,1922,1948]" targetBox="[193,1395,794,1884]" targetPageId="30">
<paragraph id="8B9D36873153FFC57AF217E1F02523A2" blockId="30.[188,1399,1922,2019]" pageId="30" pageNumber="79">
<emphasis id="B956EA953153FFC57AF217E1F04223DD" bold="true" box="[189,274,1922,1948]" pageId="30" pageNumber="79">Fig 25.</emphasis>
Shell variability in
<taxonomicName id="4C224D043153FFC57BB717E1F29623DD" authority="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger, 2021" authorityName="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger" authorityYear="2021" box="[504,966,1922,1948]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="30" pageNumber="79" phylum="Mollusca" rank="species" species="stelucarum" status="sp. nov." subGenus="Clessiniella">
<emphasis id="B956EA953153FFC57BB717E1F3C423DD" box="[504,660,1922,1948]" italics="true" pageId="30" pageNumber="79">Cochlostoma</emphasis>
(
<emphasis id="B956EA953153FFC578E917E1F26223DD" box="[678,818,1922,1948]" italics="true" pageId="30" pageNumber="79">Clessiniella</emphasis>
)
<emphasis id="B956EA953153FFC5790D17E1F29623DD" box="[834,966,1922,1948]" italics="true" pageId="30" pageNumber="79">stelucarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557EE3153FFC5798317E1F57723DD" box="[972,1063,1922,1948]" pageId="30" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B956EA953153FFC57E6317E1F51123DD" bold="true" box="[1068,1089,1922,1948]" pageId="30" pageNumber="79">A</emphasis>
. Cornino, Italy (EZ0104).
<emphasis id="B956EA953153FFC57AF317C6F18023FE" bold="true" box="[188,208,1957,1983]" pageId="30" pageNumber="79">B</emphasis>
. Mt Valinis, Italy (EZ0096).
<emphasis id="B956EA953153FFC5786917C6F36B23FE" bold="true" box="[550,571,1957,1983]" pageId="30" pageNumber="79">C</emphasis>
. Anduins, Italy (EZ0102).
<emphasis id="B956EA953153FFC5793617C6F2DE23FE" bold="true" box="[889,910,1957,1983]" pageId="30" pageNumber="79">D</emphasis>
. Mt Prat, Italy (EZ0103).
<emphasis id="B956EA953153FFC57E8D17C6F58623FE" bold="true" box="[1218,1238,1957,1983]" pageId="30" pageNumber="79">E</emphasis>
. Val di Torre, Italy (EZ0106).
</paragraph>
</caption>
<subSubSection id="C338650C3152FFC47AF2116BF05F2750" pageId="31" pageNumber="80" type="description">
<paragraph id="8B9D36873152FFC47AF2116BF01E2563" blockId="31.[189,334,264,290]" box="[189,334,264,290]" pageId="31" pageNumber="80">
<heading id="D0D581EB3152FFC47AF2116BF01E2563" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="31" pageNumber="80" reason="3">
<emphasis id="B956EA953152FFC47AF2116BF01E2563" bold="true" box="[189,334,264,290]" pageId="31" pageNumber="80">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873152FFC47AF21154F21F259F" blockId="31.[189,1399,311,478]" pageId="31" pageNumber="80">
SHELL. The shell has prominent spots and strong and rounded ribs, which are regularly and widely spaced and coloured like the background. The ribs become less prominent on the body whorl. The body whorl slightly and gradually enlarges while approaching the aperture (
<figureCitation id="13192A023152FFC479B7111EF50725D9" box="[1016,1111,381,408]" captionStart="Fig" captionStartId="5.[189,232,1854,1880]" captionTargetBox="[200,1387,265,1815]" captionTargetId="figure-11@5.[200,1387,265,1815]" captionTargetPageId="5" captionText="Fig. 4. Features of the shell of Clessiniella Zallot et al., 2015. A. Shape of the body whorl approaching the aperture. B. Strength of the spots on the shell (0 = spotless shell to 2 = really evident spots). C. Strength of the ribs (1 = weak ribs to 3 = really prominent ribs). D. Strength of the ribs on the external side of the body whorl approaching the aperture (W= smooth shell; I = intermediate; S = prominent ribs)." figureDoi="http://doi.org/10.5281/zenodo.5176919" httpUri="https://zenodo.org/record/5176919/files/figure.png" pageId="31" pageNumber="80">Fig. 4A</figureCitation>
: FF, AF). In comparison with the other species of the subgenus, the whorls increase more slowly in size so that, given the same shell height, there is a higher number of whorls (
<figureCitation id="13192A023152FFC478B711A7F210259F" box="[760,832,452,478]" captionStart="Fig" captionStartId="14.[188,231,925,951]" captionTargetBox="[254,1297,271,867]" captionTargetId="figure-15@14.[242,1347,262,889]" captionTargetPageId="14" captionText="Fig. 9. Number of whorls vs height of the species in Clessiniella Zallot et al., 2015. Violet dots = Cochlostoma (Clessiniella) anomphale Boeckel, 1939; red squares = Co. (Cl.) stelucarum sp. nov.; green triangles = Co. (Cl.) tergestinum (Westerlund, 1878); blue diamonds = Co. (Cl.) villae (Strobel, 1851); black triangles =Co. (Cl.) waldemari (A.J. Wagner, 1897). Analysis performed with the software ‘Past’ (https://palaeo-electronica.org)." figureDoi="http://doi.org/10.5281/zenodo.5176929" httpUri="https://zenodo.org/record/5176929/files/figure.png" pageId="31" pageNumber="80">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B9D36873152FFC47AF21265F0962627" blockId="31.[189,1399,517,614]" pageId="31" pageNumber="80">
FEMALE GENITALIA. The visceral oviduct has only 1–2 superficial loops, what makes it one of the simplest in the genus
<taxonomicName id="4C224D043152FFC47B1E124BF0BC2603" authorityName="Jan" authorityYear="1830" box="[337,492,552,578]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="31" pageNumber="80" phylum="Mollusca" rank="genus">
<emphasis id="B956EA953152FFC47B1E124BF0BC2603" box="[337,492,552,578]" italics="true" pageId="31" pageNumber="80">Cochlostoma</emphasis>
</taxonomicName>
. The seminal receptacle is thin and may be oval, thus not always club-shaped as in the other species.
</paragraph>
<paragraph id="8B9D36873152FFC47AF212EDF05F2750" blockId="31.[189,1399,653,785]" pageId="31" pageNumber="80">
MALE GENITALIA. The penis is longer than the body and inflated, although less so than in the other species of
<taxonomicName id="4C224D043152FFC47AAF12D3F03B268B" box="[224,363,688,714]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="31" pageNumber="64" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA953152FFC47AAF12D3F03B268B" box="[224,363,688,714]" italics="true" pageId="31" pageNumber="80">Clessiniella</emphasis>
</taxonomicName>
. The penial spermiduct is gently twisted. The body spermiduct is straight. There is a long groove, ending in a well-formed sperm pocket, delimited at the frontal side by an ascending sperm funnel.
</paragraph>
</subSubSection>
<subSubSection id="C338650C3152FFC47AF2135BF5AA27E4" pageId="31" pageNumber="80" type="distribution">
<paragraph id="8B9D36873152FFC47AF2135BF0062713" blockId="31.[189,342,824,850]" box="[189,342,824,850]" pageId="31" pageNumber="80">
<heading id="D0D581EB3152FFC47AF2135BF0062713" bold="true" box="[189,342,824,850]" fontSize="11" level="3" pageId="31" pageNumber="80" reason="3">
<emphasis id="B956EA953152FFC47AF2135BF0062713" bold="true" box="[189,342,824,850]" pageId="31" pageNumber="80">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873152FFC47AF21304F5AA27E4" blockId="31.[189,1399,871,933]" pageId="31" pageNumber="80">
The species inhabits a restricted area in the eastern Southern Pre-Alps. Our sampling localities are located between Val Tramontina to the west and Val di Torre to the east (
<collectingRegion id="49E6F8653152FFC47E4013E9F5F727E5" box="[1039,1191,906,932]" country="Italy" name="Friuli-Venezia Giulia" pageId="31" pageNumber="80">Friuli region</collectingRegion>
,
<collectingCountry id="F33576173152FFC47EFC13E9F5BE27E4" box="[1203,1262,906,933]" name="Italy" pageId="31" pageNumber="80">Italy</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C338650C3152FFC47AF213A8F205201B" pageId="31" pageNumber="80" type="description">
<paragraph id="8B9D36873152FFC47AF213A8F0F827A4" blockId="31.[189,424,971,997]" box="[189,424,971,997]" pageId="31" pageNumber="80">
<heading id="D0D581EB3152FFC47AF213A8F0F827A4" bold="true" box="[189,424,971,997]" fontSize="11" level="3" pageId="31" pageNumber="80" reason="3">
<emphasis id="B956EA953152FFC47AF213A8F0F827A4" bold="true" box="[189,424,971,997]" pageId="31" pageNumber="80">Habitat preference</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873152FFC47AF21399F205201B" blockId="31.[189,1399,1018,1115]" pageId="31" pageNumber="80">
Like in most species of
<taxonomicName id="4C224D043152FFC47BAC1399F32E2055" authorityName="Jan" authorityYear="1830" box="[483,638,1018,1044]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="31" pageNumber="80" phylum="Mollusca" rank="genus">
<emphasis id="B956EA953152FFC47BAC1399F32E2055" box="[483,638,1018,1044]" italics="true" pageId="31" pageNumber="80">Cochlostoma</emphasis>
</taxonomicName>
, these are obligate rock-dwelling snails but, rather than on the surface of limestone cliffs, their main habitat is on smaller surfaces of stones and boulders which are normally found in large numbers at the foot of the cliffs.
</paragraph>
</subSubSection>
<caption id="DF5D660F3152FFC47AF217E6F50523A7" ID-DOI="http://doi.org/10.5281/zenodo.5176965" ID-Zenodo-Dep="5176965" httpUri="https://zenodo.org/record/5176965/files/figure.png" pageId="31" pageNumber="80" startId="31.[189,230,1925,1951]" targetBox="[189,1370,1213,1868]" targetPageId="31">
<paragraph id="8B9D36873152FFC47AF217E6F50523A7" blockId="31.[186,1398,1925,2023]" pageId="31" pageNumber="80">
<emphasis id="B956EA953152FFC47AF217E6F04023DE" bold="true" box="[189,272,1925,1951]" pageId="31" pageNumber="80">Fig 26.</emphasis>
Variation of the female genitalia in
<taxonomicName id="4C224D043152FFC478E917E5F52323E1" authority="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger, 2021" authorityName="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger" authorityYear="2021" box="[678,1139,1926,1952]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="31" pageNumber="80" phylum="Mollusca" rank="species" species="stelucarum" status="sp. nov." subGenus="Clessiniella">
<emphasis id="B956EA953152FFC478E917E5F21223E1" box="[678,834,1926,1952]" italics="true" pageId="31" pageNumber="80">Cochlostoma</emphasis>
(
<emphasis id="B956EA953152FFC4791B17E5F2B023E1" box="[852,992,1926,1952]" italics="true" pageId="31" pageNumber="80">Clessiniella</emphasis>
)
<emphasis id="B956EA953152FFC479A017E5F52323E1" box="[1007,1139,1926,1952]" italics="true" pageId="31" pageNumber="80">stelucarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557EE3152FFC47E3817E5F58023E1" box="[1143,1232,1926,1952]" pageId="31" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B956EA953152FFC47E9D17E6F5B723DE" bold="true" box="[1234,1255,1925,1951]" pageId="31" pageNumber="80">A</emphasis>
. Mt Valinis, Italy (EZ0096).
<emphasis id="B956EA953152FFC47BC917CAF0CA2382" bold="true" box="[390,410,1961,1987]" pageId="31" pageNumber="80">B</emphasis>
. Campone, Italy (EZ0098).
<emphasis id="B956EA953152FFC478B517CAF25F2382" bold="true" box="[762,783,1961,1987]" pageId="31" pageNumber="80">C</emphasis>
. Anduins, Italy (EZ0102).
<emphasis id="B956EA953152FFC47E2E17CAF5E92382" bold="true" box="[1121,1209,1961,1987]" pageId="31" pageNumber="80">D1–D3</emphasis>
. Cornino, Italy (EZ0104).
<emphasis id="B956EA953152FFC47B7017AFF00323A7" bold="true" box="[319,339,1996,2022]" pageId="31" pageNumber="80">E</emphasis>
. Cjanet-Peonis, Italy (EZ0105).
<emphasis id="B956EA953152FFC4789717AFF3BA23A7" bold="true" box="[728,746,1996,2022]" pageId="31" pageNumber="80">F</emphasis>
. Mt Bernadia, Italy (EZ0107).
</paragraph>
</caption>
<subSubSection id="C338650C316DFFFB7AF2116AF26D2664" pageId="32" pageNumber="81" type="discussion">
<paragraph id="8B9D3687316DFFFB7AF2116AF07F2562" blockId="32.[189,303,265,291]" box="[189,303,265,291]" pageId="32" pageNumber="81">
<heading id="D0D581EB316DFFFB7AF2116AF07F2562" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="32" pageNumber="81" reason="3">
<emphasis id="B956EA95316DFFFB7AF2116AF07F2562" bold="true" box="[189,303,265,291]" pageId="32" pageNumber="81">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D3687316DFFFB7AF21154F26D2664" blockId="32.[188,1399,311,549]" pageId="32" pageNumber="81">
<taxonomicName id="4C224D04316DFFFB7AF21154F3DC2510" authority="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger, 2021" authorityName="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger" authorityYear="2021" box="[189,652,311,338]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="stelucarum" status="sp. nov." subGenus="Clessiniella">
<emphasis id="B956EA95316DFFFB7AF21154F0082510" box="[189,344,311,337]" italics="true" pageId="32" pageNumber="81">Cochlostoma</emphasis>
(
<emphasis id="B956EA95316DFFFB7B231154F0A72510" box="[364,503,311,337]" italics="true" pageId="32" pageNumber="81">Clessiniella</emphasis>
)
<emphasis id="B956EA95316DFFFB78451154F3DC2510" box="[522,652,311,337]" italics="true" pageId="32" pageNumber="81">stelucarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557EE316DFFFB78D8115BF3A52513" box="[663,757,312,338]" pageId="32" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
was introduced as
<taxonomicName id="4C224D04316DFFFB79AE1154F5F72513" box="[993,1191,311,338]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B956EA95316DFFFB79AE1154F52C2510" box="[993,1148,311,337]" italics="true" pageId="32" pageNumber="81">Cochlostoma</emphasis>
sp.
</taxonomicName>
by
<bibRefCitation id="EFB34B76316DFFFB7E961154F1B72535" author="Zallot E." pageId="32" pageNumber="81" pagination="93 - 113" refId="ref16527" refString="Zallot E. 2002. Alcune note sul genere Cochlostoma Jan, 1830 (Gastropoda, Prosobranchia) in Friuli (Italia nord-orientale). Gortania 24: 93 - 113." type="journal article" year="2002">Zallot (2002: 98)</bibRefCitation>
. At most of the localities, it lives syntopically with either
<taxonomicName id="4C224D04316DFFFB79DE1138F05125D9" authority="(Gredler, 1853)" baseAuthorityName="Gredler" baseAuthorityYear="1853" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="philippianum" subGenus="Eupomatias">
<emphasis id="B956EA95316DFFFB79DE1138F2EA2534" box="[913,954,347,373]" italics="true" pageId="32" pageNumber="81">Co.</emphasis>
(
<emphasis id="B956EA95316DFFFB79841138F5072535" box="[971,1111,347,372]" italics="true" pageId="32" pageNumber="81">Eupomatias</emphasis>
)
<emphasis id="B956EA95316DFFFB7E261139F4552535" box="[1129,1285,346,372]" italics="true" pageId="32" pageNumber="81">philippianum</emphasis>
(Gredler, 1853)
</taxonomicName>
or
<taxonomicName id="4C224D04316DFFFB7B61111DF27F25D9" authority="(Strobel, 1851)" baseAuthorityName="Strobel" baseAuthorityYear="1851" box="[302,815,381,408]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="henricae" subGenus="Eupomatias">
<emphasis id="B956EA95316DFFFB7B61111DF00825D9" box="[302,344,382,408]" italics="true" pageId="32" pageNumber="81">Co.</emphasis>
(
<emphasis id="B956EA95316DFFFB7B26111DF0A525D6" box="[361,501,382,407]" italics="true" pageId="32" pageNumber="81">Eupomatias</emphasis>
)
<emphasis id="B956EA95316DFFFB7847111EF33F25D6" box="[520,623,381,407]" italics="true" pageId="32" pageNumber="81">henricae</emphasis>
(
<bibRefCitation id="EFB34B76316DFFFB78CF111EF27625D9" author="Strobel P." box="[640,806,381,408]" pageId="32" pageNumber="81" refId="ref16289" refString="Strobel P. 1851. Notizie Malacostatiche sul Trentino. Fusi, Pavia. Available from https: // www. biodiversitylibrary. org / page / 46194631 [accessed 12 Jul. 2021]." type="url" year="1851">Strobel, 1851</bibRefCitation>
)
</taxonomicName>
. Near the village of Anduins it has been found a few centimeters apart from both
<taxonomicName id="4C224D04316DFFFB787711C2F24225FA" baseAuthorityName="Strobel" baseAuthorityYear="1851" box="[568,786,417,443]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="henricae" subGenus="Eupomatias">
<emphasis id="B956EA95316DFFFB787711C2F33225FA" box="[568,610,417,443]" italics="true" pageId="32" pageNumber="81">Co.</emphasis>
(
<emphasis id="B956EA95316DFFFB783C11C2F3CB25FB" box="[627,667,417,442]" italics="true" pageId="32" pageNumber="81">Eu.</emphasis>
)
<emphasis id="B956EA95316DFFFB78E411C2F24225FA" box="[683,786,417,443]" italics="true" pageId="32" pageNumber="81">henricae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C224D04316DFFFB790611C2F58225FA" baseAuthorityName="Razoumowsky" baseAuthorityYear="1789" box="[841,1234,417,443]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="septemspirale" subGenus="Cochlostoma">
<emphasis id="B956EA95316DFFFB790611C2F22225FA" box="[841,882,417,443]" italics="true" pageId="32" pageNumber="81">Co.</emphasis>
(
<emphasis id="B956EA95316DFFFB79CC11C2F54E25FA" box="[899,1054,417,443]" italics="true" pageId="32" pageNumber="81">Cochlostoma</emphasis>
)
<emphasis id="B956EA95316DFFFB7E6011C2F58225FA" box="[1071,1234,417,443]" italics="true" pageId="32" pageNumber="81">septemspirale</emphasis>
</taxonomicName>
on stones and boulders. This is one of the rare cases where 3 species of
<taxonomicName id="4C224D04316DFFFB790311A7F2B7259F" authorityName="Jan" authorityYear="1830" box="[844,999,452,478]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="genus">
<emphasis id="B956EA95316DFFFB790311A7F2B7259F" box="[844,999,452,478]" italics="true" pageId="32" pageNumber="81">Cochlostoma</emphasis>
</taxonomicName>
, belonging to different subgenera, live syntopically. The range of the species is situated in between the range of
<taxonomicName id="4C224D04316DFFFB7E0D1184F5A12640" baseAuthorityName="Strobel" baseAuthorityYear="1851" box="[1090,1265,487,514]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="81" phylum="Mollusca" rank="species" species="villae" subGenus="Clessiniella">
<emphasis id="B956EA95316DFFFB7E0D1184F53B2640" box="[1090,1131,487,513]" italics="true" pageId="32" pageNumber="81">Co.</emphasis>
(
<emphasis id="B956EA95316DFFFB7E331184F5CF2640" box="[1148,1183,487,513]" italics="true" pageId="32" pageNumber="81">Cl.</emphasis>
)
<emphasis id="B956EA95316DFFFB7EFF1184F5A12640" box="[1200,1265,487,513]" italics="true" pageId="32" pageNumber="81">villae</emphasis>
</taxonomicName>
in the west and that of the other species of
<taxonomicName id="4C224D04316DFFFB78631269F3E72665" box="[556,695,522,548]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="32" pageNumber="64" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA95316DFFFB78631269F3E72665" box="[556,695,522,548]" italics="true" pageId="32" pageNumber="81">Clessiniella</emphasis>
</taxonomicName>
in the east.
</paragraph>
</subSubSection>
</treatment>
</document>
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