Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae) Author Zallot, Enrico 365E17AD-6938-4364-A526-F1BDA2663E10 Haagweg 29, 2681 PA Monster, the Netherlands. ezallot@gmail.com Author Mattia, Willy De C74A049D-9D7B-4A1F-878B-5BDFD104219F Central Research Laboratories of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. wdemattia@gmail.com Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 Hungarian Natural History Museum, Baross 13, 1088 Budapest, Hungary. feher.zoltan@nhmus.hu Author Gittenberger, Edmund D786C279-FC92-4D08-AF16-F79A9705E0AE Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, the Netherlands. egittenberger@yahoo.com text European Journal of Taxonomy 2021 2021-08-06 762 1 49 95 http://dx.doi.org/10.5852/ejt.2021.762.1453 journal article 5011 10.5852/ejt.2021.762.1453 b739dd1a-47f7-4e15-a8c6-caf0a89947cb 2118-9773 5176909 D1093CEB-7EBB-44A0-8078-8DC616B857F5 Cochlostoma (Clessiniella) stelucarum sp. nov. urn:lsid:zoobank.org:act: 4609B41C-3611-4F8A-9048-5F97EA0446E6 Figs 1 , 3E , 14B , 15B , 16B, 15–27 Diagnosis Slender shell with 8½–10½ rather widely ribbed whorls. Spots always visible. Poorly developed lip with FF or AF external lobe. Visceral oviduct simplified and seminal receptacle sometimes oval. Etymology The epithet ‘ steluca ’ is an acronym noun formed from the initial syllables of the names of the wife and two daughters (Stefania, Ludovica and Camilla Zallot) of the senior author. Type material ( Fig. 27 ) Holotype ITALY • ♀ (in ethanol); Friuli Venezia Giulia , Udine , Cornino ; 46.2297° N , 13.0250° E ; 180 m a.s.l. ; 5 Jun. 2002 ; Zallot leg.; rocks surrounding the lake ; RMNH.MOL.347617. Paratypes ITALY • 1 ♂ (in ethanol); same collection data as for holotype; RMNH.MOL.347618 • 1 ♀ (empty shell); same collection data as for holotype; GenBank voucher H3: KX120830 ; RMNH.MOL.347619 . Other material ITALY • Anduins-cliffs ; 46.23755° N , 12.97106° E ; 1997; Zallot leg.; EZ0102 • Campone ; 46.26194° N , 12.83333° E ; 2010; Zallot leg.; EZ0098 • Castelmonte ; 46.094° N , 13.52095° E ; Oct. 2010 ; Margelli leg.; EZ1020 • Cjanet-Peonis ; 46.25917° N , 13.04111° E ; 2002; Zallot leg.; EZ0105 • Cornino ; 46.22972° N , 13.025° E ; 2002; Zallot leg.; EZ0104 • F. Piccola Mt Valinins ; 46.245° N , 12.80111° E ; 2000; Zallot leg.; EZ0095 • Mt Bernadia ; 46.23361° N , 13.25556° E ; 2005; Zallot leg.; EZ0107 • Mt Ciaurlec ; 46.21226° N , 12.88018° E ; 2005; Zallot leg.; EZ0099 • Mt Prat ; 46.24577° N , 12.9907° E ; 2002; Zallot leg.; EZ0103 • Mt Valinis ; 46.23417° N , 12.8075° E ; 2002; Zallot leg.; EZ0096 • San Vito D’asio ; 46.2337° N , 12.94531° E ; 1997; Zallot leg.; EZ0101 • Tramonti-Campone ; 46.27833° N , 12.81917° E ; 2000; Zallot leg.; EZ0097 • Val di Torre-all’imbocco ; 46.23861° N , 13.23111° E ; 1998; Zallot leg.; EZ0106 • Valle D’Arzino ; 46.30278° N , 12.92972° E ; 2000; Zallot leg.; EZ0100 . Fig 25. Shell variability in Cochlostoma ( Clessiniella ) stelucarum sp. nov. A . Cornino, Italy (EZ0104). B . Mt Valinis, Italy (EZ0096). C . Anduins, Italy (EZ0102). D . Mt Prat, Italy (EZ0103). E . Val di Torre, Italy (EZ0106). Description SHELL. The shell has prominent spots and strong and rounded ribs, which are regularly and widely spaced and coloured like the background. The ribs become less prominent on the body whorl. The body whorl slightly and gradually enlarges while approaching the aperture ( Fig. 4A : FF, AF). In comparison with the other species of the subgenus, the whorls increase more slowly in size so that, given the same shell height, there is a higher number of whorls ( Fig. 9 ). FEMALE GENITALIA. The visceral oviduct has only 1–2 superficial loops, what makes it one of the simplest in the genus Cochlostoma . The seminal receptacle is thin and may be oval, thus not always club-shaped as in the other species. MALE GENITALIA. The penis is longer than the body and inflated, although less so than in the other species of Clessiniella . The penial spermiduct is gently twisted. The body spermiduct is straight. There is a long groove, ending in a well-formed sperm pocket, delimited at the frontal side by an ascending sperm funnel. Distribution The species inhabits a restricted area in the eastern Southern Pre-Alps. Our sampling localities are located between Val Tramontina to the west and Val di Torre to the east ( Friuli region , Italy ). Habitat preference Like in most species of Cochlostoma , these are obligate rock-dwelling snails but, rather than on the surface of limestone cliffs, their main habitat is on smaller surfaces of stones and boulders which are normally found in large numbers at the foot of the cliffs. Fig 26. Variation of the female genitalia in Cochlostoma ( Clessiniella ) stelucarum sp. nov. A . Mt Valinis, Italy (EZ0096). B . Campone, Italy (EZ0098). C . Anduins, Italy (EZ0102). D1–D3 . Cornino, Italy (EZ0104). E . Cjanet-Peonis, Italy (EZ0105). F . Mt Bernadia, Italy (EZ0107). Remarks Cochlostoma ( Clessiniella ) stelucarum sp. nov. was introduced as Cochlostoma sp. by Zallot (2002: 98) . At most of the localities, it lives syntopically with either Co. ( Eupomatias ) philippianum (Gredler, 1853) or Co. ( Eupomatias ) henricae ( Strobel, 1851 ) . Near the village of Anduins it has been found a few centimeters apart from both Co. ( Eu. ) henricae and Co. ( Cochlostoma ) septemspirale on stones and boulders. This is one of the rare cases where 3 species of Cochlostoma , belonging to different subgenera, live syntopically. The range of the species is situated in between the range of Co. ( Cl. ) villae in the west and that of the other species of Clessiniella in the east.