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<document id="536C765114CB2E0AED3D2FEF6E71650A" ID-CLB-Dataset="5011" ID-DOI="10.5852/ejt.2021.762.1453" ID-GBIF-Dataset="b739dd1a-47f7-4e15-a8c6-caf0a89947cb" ID-ISSN="2118-9773" ID-Zenodo-Dep="5176909" ID-ZooBank="D1093CEB-7EBB-44A0-8078-8DC616B857F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628612905243" checkinUser="felipe" docAuthor="Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán &amp; Gittenberger, Edmund" docDate="2021" docId="038B87913142FFCA7B7E1463F42627B8" docLanguage="en" docName="ejt-762_49-95.pdf" docOrigin="European Journal of Taxonomy 762 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.762.1453" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cochlostoma (Clessiniella) Zallot, Groenenberg, De Mattia, Feher &amp; Gittenberger 2015" docType="treatment" docVersion="6" lastPageNumber="66" masterDocId="FFB2FFE9314DFFDB7A4F1063F1502441" masterDocTitle="Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)" masterLastPageNumber="95" masterPageNumber="49" pageNumber="64" updateTime="1699072062511" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EDB837E060CD1BCEBA92CF0E241D8F3D">Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="A8E1D1DA4952E31BD588AFFAD88F7CC0">Zallot, Enrico</mods:namePart>
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<mods:affiliation id="3DD22BADE163C418402CB0B85F637105">Haagweg 29, 2681 PA Monster, the Netherlands.</mods:affiliation>
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<mods:namePart id="F3A223081981D19B061538F65C8E9277">Mattia, Willy De</mods:namePart>
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<mods:affiliation id="12139709A5D87DDFB015DBB77EE0F322">Central Research Laboratories of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria.</mods:affiliation>
<mods:nameIdentifier id="5794A2D48656B28A7F4064E454CFAC1A" type="email">wdemattia@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5AF014F8C08C50E52A839E4379C5C1B0">Fehér, Zoltán</mods:namePart>
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<mods:affiliation id="275C96A6EF6B60E20A11065799524F38">Hungarian Natural History Museum, Baross 13, 1088 Budapest, Hungary.</mods:affiliation>
<mods:nameIdentifier id="67A8DA0C6ABBF2752AC72F0B9ACAA4AE" type="email">feher.zoltan@nhmus.hu</mods:nameIdentifier>
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<mods:namePart id="7471A139CACB8A50C409A41C48F9A7C0">Gittenberger, Edmund</mods:namePart>
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<mods:affiliation id="64216C5825AE79770307D64086FE0DFF">Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, the Netherlands.</mods:affiliation>
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<mods:title id="63792AEF396F312BE13EDF8683788410">European Journal of Taxonomy</mods:title>
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<mods:date id="5ECB4308CEA70EA96FDD1545FC10FB6A">2021</mods:date>
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<treatment id="038B87913142FFCA7B7E1463F42627B8" ID-DOI="http://doi.org/10.5281/zenodo.5191140" ID-GBIF-Taxon="184096745" ID-Zenodo-Dep="5191140" LSID="urn:lsid:plazi:treatment:038B87913142FFCA7B7E1463F42627B8" httpUri="http://treatment.plazi.org/id/038B87913142FFCA7B7E1463F42627B8" lastPageId="17" lastPageNumber="66" pageId="15" pageNumber="64">
<subSubSection id="C338650C3142FFD47B7E1463F453205A" box="[305,1283,1023,1051]" pageId="15" pageNumber="64" type="nomenclature">
<paragraph id="8B9D36873142FFD47B7E1463F453205A" blockId="15.[305,1283,1023,1051]" box="[305,1283,1023,1051]" pageId="15" pageNumber="64">
<heading id="D0D581EB3142FFD47B7E1463F453205A" box="[305,1283,1023,1051]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="64" reason="2">
Subgenus
<taxonomicName id="4C224D043142FFD47BE5139CF453205A" authority="Zallot, Groenenberg, De Mattia, Feher &amp; Gittenberger, 2015" authorityName="Zallot, Groenenberg, De Mattia, Feher &amp; Gittenberger" authorityYear="2015" box="[426,1283,1023,1051]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="15" pageNumber="64" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA953142FFD47BE5139CF366205B" bold="true" box="[426,566,1023,1050]" italics="true" pageId="15" pageNumber="64">Clessiniella</emphasis>
<bibRefCitation id="EFB34B763142FFD478711463F453205A" author="Zallot E. &amp; Groenenberg D. S. J. &amp; De Mattia W. &amp; Feher Z. &amp; Gittenberger E." box="[574,1283,1024,1051]" pageId="15" pageNumber="64" pagination="63 - 88" refId="ref16559" refString="Zallot E., Groenenberg D. S. J., De Mattia W., Feher Z. &amp; Gittenberger E. 2015. Genera, subgenera and species of the Cochlostomatidae (Gastropoda, Caenogastropoda, Cochlostomatidae). Basteria 78 (4 - 6): 63 - 88." type="journal article" year="2015">Zallot, Groenenberg, De Mattia, Fehér &amp; Gittenberger, 2015</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C338650C3142FFD47AF21422F518201D" box="[189,1096,1089,1116]" pageId="15" pageNumber="64" type="reference_group">
<paragraph id="8B9D36873142FFD47AF21422F518201D" blockId="15.[189,1096,1089,1116]" box="[189,1096,1089,1116]" pageId="15" pageNumber="64">
<treatmentCitationGroup id="AB3211A93142FFD47AF21422F518201D" box="[189,1096,1089,1116]" pageId="15" pageNumber="64">
<treatmentCitation id="0A8310963142FFD47AF21422F514201D" author="Zallot E. &amp; Groenenberg D. S. J. &amp; De Mattia W. &amp; Feher Z. &amp; Gittenberger E." box="[189,1092,1089,1116]" page="80" pageId="15" pageNumber="64" year="2015">
<taxonomicName id="4C224D043142FFD47AF21422F514201D" authority="Zallot, Groenenberg, De Mattia, Feher &amp; Gittenberger, 2015: 80" authorityName="Zallot, Groenenberg, De Mattia, Feher &amp; Gittenberger" authorityYear="2015" box="[189,1092,1089,1116]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="15" pageNumber="64" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA953142FFD47AF21422F018201A" box="[189,328,1089,1115]" italics="true" pageId="15" pageNumber="64">Clessiniella</emphasis>
<bibRefCitation id="EFB34B763142FFD47B001422F514201D" author="Zallot E. &amp; Groenenberg D. S. J. &amp; De Mattia W. &amp; Feher Z. &amp; Gittenberger E." box="[335,1092,1089,1116]" pageId="15" pageNumber="64" pagination="63 - 88" refId="ref16559" refString="Zallot E., Groenenberg D. S. J., De Mattia W., Feher Z. &amp; Gittenberger E. 2015. Genera, subgenera and species of the Cochlostomatidae (Gastropoda, Caenogastropoda, Cochlostomatidae). Basteria 78 (4 - 6): 63 - 88." type="journal article" year="2015">Zallot, Groenenberg, De Mattia, Fehér &amp; Gittenberger, 2015: 80</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C338650C3142FFD47AF214E0F2AB208A" pageId="15" pageNumber="64" type="type_taxon">
<paragraph id="8B9D36873142FFD47AF214E0F00820DC" blockId="15.[189,344,1155,1181]" box="[189,344,1155,1181]" pageId="15" pageNumber="64">
<heading id="D0D581EB3142FFD47AF214E0F00820DC" bold="true" box="[189,344,1155,1181]" fontSize="11" level="3" pageId="15" pageNumber="64" reason="3">
<emphasis id="B956EA953142FFD47AF214E0F00820DC" bold="true" box="[189,344,1155,1181]" pageId="15" pageNumber="64">Type species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873142FFD47AF214D1F2AB208A" blockId="15.[189,1019,1201,1228]" box="[189,1019,1201,1228]" pageId="15" pageNumber="64">
<taxonomicName id="4C224D043142FFD47AF214D1F386208D" authority="Strobel, 1851" authorityName="Strobel" authorityYear="1851" box="[189,726,1201,1228]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="64" phylum="Mollusca" rank="variety" species="maculatum" variety="villae">
<emphasis id="B956EA953142FFD47AF214D1F0E5208A" box="[189,437,1201,1227]" italics="true" pageId="15" pageNumber="64">Pomatias maculatum</emphasis>
var.
<emphasis id="B956EA953142FFD47BA014D2F360208A" box="[495,560,1201,1227]" italics="true" pageId="15" pageNumber="64">villae</emphasis>
<bibRefCitation id="EFB34B763142FFD4787714D2F386208D" author="Strobel P." box="[568,726,1201,1228]" pageId="15" pageNumber="64" refId="ref16289" refString="Strobel P. 1851. Notizie Malacostatiche sul Trentino. Fusi, Pavia. Available from https: // www. biodiversitylibrary. org / page / 46194631 [accessed 12 Jul. 2021]." type="url" year="1851">Strobel, 1851</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C338650C3142FFD47AF21490F535224E" pageId="15" pageNumber="64" type="discussion">
<paragraph id="8B9D36873142FFD47AF21490F07F214C" blockId="15.[189,303,1267,1293]" box="[189,303,1267,1293]" pageId="15" pageNumber="64">
<heading id="D0D581EB3142FFD47AF21490F07F214C" bold="true" box="[189,303,1267,1293]" fontSize="11" level="3" pageId="15" pageNumber="64" reason="3">
<emphasis id="B956EA953142FFD47AF21490F07F214C" bold="true" box="[189,303,1267,1293]" pageId="15" pageNumber="64">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873142FFD47AF21542F535224E" blockId="15.[188,1400,1313,1551]" pageId="15" pageNumber="64">
The classification follows MolluscaBase (2021) except for the subgeneric name. Contrary to ICZN Art. 10.4,
<emphasis id="B956EA953142FFD47B7B1527F0CC211F" box="[308,412,1348,1374]" italics="true" pageId="15" pageNumber="64">Neglecta</emphasis>
<bibRefCitation id="EFB34B763142FFD47BEC1526F316211E" author="Wagner A. J." box="[419,582,1349,1375]" pageId="15" pageNumber="64" pagination="565 - 632" refId="ref16375" refString="Wagner A. J. 1897. Monographie der Gattung Pomatias Studer. Denkschriften der kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 64: 565 - 632. https: // doi. org / 10.5962 / bhl. title. 10792" type="journal article" year="1897">Wagner, 1897</bibRefCitation>
is used in MolluscaBase as a senior synonym of
<taxonomicName id="4C224D043142FFD47ECC1527F45E211F" authorityName="Zallot, Groenenberg, De Mattia, Feher &amp; Gittenberger" authorityYear="2015" box="[1155,1294,1348,1374]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="15" pageNumber="64" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA953142FFD47ECC1527F45E211F" box="[1155,1294,1348,1374]" italics="true" pageId="15" pageNumber="64">Clessiniella</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB34B763142FFD47F531526F07121C3" author="Wagner A. J." pageId="15" pageNumber="64" pagination="565 - 632" refId="ref16375" refString="Wagner A. J. 1897. Monographie der Gattung Pomatias Studer. Denkschriften der kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 64: 565 - 632. https: // doi. org / 10.5962 / bhl. title. 10792" type="journal article" year="1897">Wagner (1897: 6</bibRefCitation>
[570]) introduced
<emphasis id="B956EA953142FFD47849150BF33E21C3" box="[518,622,1384,1410]" italics="true" pageId="15" pageNumber="64">Neglecta</emphasis>
as “Formenkreis
<emphasis id="B956EA953142FFD4790C150BF2F921C3" box="[835,937,1384,1410]" italics="true" pageId="15" pageNumber="64">Neglecta</emphasis>
”. It cannot be denied that the German term Formenkreis is “.. a term such as “superspecies””. As a consequence, according to the ICZN ruling,
<emphasis id="B956EA953142FFD47AF315CDF0742189" box="[188,292,1454,1480]" italics="true" pageId="15" pageNumber="64">Neglecta</emphasis>
“.. is not deemed to be a genus-group name” (ICZN Art. 10.4, see also
<bibRefCitation id="EFB34B763142FFD47EDC15CDF43B2188" author="Zallot E. &amp; Groenenberg D. S. J. &amp; De Mattia W. &amp; Feher Z. &amp; Gittenberger E." box="[1171,1387,1454,1481]" pageId="15" pageNumber="64" pagination="63 - 88" refId="ref16559" refString="Zallot E., Groenenberg D. S. J., De Mattia W., Feher Z. &amp; Gittenberger E. 2015. Genera, subgenera and species of the Cochlostomatidae (Gastropoda, Caenogastropoda, Cochlostomatidae). Basteria 78 (4 - 6): 63 - 88." type="journal article" year="2015">
Zallot
<emphasis id="B956EA953142FFD47EAC15CCF4712189" box="[1251,1313,1454,1480]" italics="true" pageId="15" pageNumber="64">et al.</emphasis>
2015
</bibRefCitation>
).
<bibRefCitation id="EFB34B763142FFD47AF215B2F03721AD" author="Strobel P." box="[189,359,1489,1516]" pageId="15" pageNumber="64" refId="ref16289" refString="Strobel P. 1851. Notizie Malacostatiche sul Trentino. Fusi, Pavia. Available from https: // www. biodiversitylibrary. org / page / 46194631 [accessed 12 Jul. 2021]." type="url" year="1851">Strobel (1851)</bibRefCitation>
and
<bibRefCitation id="EFB34B763142FFD47BD115B1F3BB21AD" author="De Betta E. &amp; Martinati P." box="[414,747,1489,1516]" pageId="15" pageNumber="64" refId="ref15858" refString="De Betta E. &amp; Martinati P. 1855. Catalogo dei Molluschi Terrestri e Fluviatili Viventi nelle Provincie Venete di Edoardo de Betta e Pietropaolo Martinati. G. Antonelli, Verona. https: // doi. org / 10.5962 / bhl. title. 13113" type="book" year="1855">De Betta &amp; Martinati (1855)</bibRefCitation>
named this entity
<taxonomicName id="4C224D043142FFD4798F15B1F46521AA" authorityName="Strobel" authorityYear="1851" box="[960,1333,1489,1516]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="71" phylum="Mollusca" rank="variety" species="maculatum" variety="villae">
<emphasis id="B956EA953142FFD4798F15B1F5E821AA" box="[960,1208,1489,1515]" italics="true" pageId="15" pageNumber="64">Pomatias maculatum</emphasis>
var.
<emphasis id="B956EA953142FFD47EBE15B2F46521AA" box="[1265,1333,1489,1515]" italics="true" pageId="15" pageNumber="64">Villae</emphasis>
</taxonomicName>
<emphasis id="B956EA953142FFD47F7315B1F41B21AA" box="[1340,1355,1490,1515]" italics="true" pageId="15" pageNumber="64">–</emphasis>
but referring to
<taxonomicName id="4C224D043142FFD47B061596F243224E" authority="Draparnaud, 1805" authorityName="Draparnaud" authorityYear="1805" box="[329,787,1524,1551]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="64" phylum="Mollusca" rank="species" species="maculatus">
<emphasis id="B956EA953142FFD47B061596F368224F" box="[329,568,1524,1550]" italics="true" pageId="15" pageNumber="64">Pomatias maculatus</emphasis>
Draparnaud, 1805
</taxonomicName>
(=
<taxonomicName id="4C224D043142FFD479731596F504224F" baseAuthorityName="Razoumowsky" baseAuthorityYear="1789" box="[828,1108,1524,1551]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="15" pageNumber="64" phylum="Mollusca" rank="species" species="septemspirale" subGenus="Cochlostoma">
<emphasis id="B956EA953142FFD479731596F235224E" box="[828,869,1525,1551]" italics="true" pageId="15" pageNumber="64">Co.</emphasis>
(
<emphasis id="B956EA953142FFD479381596F2F0224E" box="[887,928,1525,1551]" italics="true" pageId="15" pageNumber="64">Co.</emphasis>
)
<emphasis id="B956EA953142FFD479FE1597F504224F" box="[945,1108,1524,1550]" italics="true" pageId="15" pageNumber="64">septemspirale</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C338650C3142FFD47AF21655F2F92313" pageId="15" pageNumber="64" type="diagnosis">
<paragraph id="8B9D36873142FFD47AF21655F0AF2211" blockId="15.[189,511,1590,1616]" box="[189,511,1590,1616]" pageId="15" pageNumber="64">
<heading id="D0D581EB3142FFD47AF21655F0AF2211" bold="true" box="[189,511,1590,1616]" fontSize="11" level="3" pageId="15" pageNumber="64" reason="3">
<emphasis id="B956EA953142FFD47AF21655F0AF2211" bold="true" box="[189,511,1590,1616]" pageId="15" pageNumber="64">Diagnosis of the subgenus</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36873142FFD47AF21606F2F92313" blockId="15.[188,1399,1636,1874]" pageId="15" pageNumber="64">
SHELL. Ribbed teleoconch with straight ribs, variable in strength and shape. The columellar lobe is curved backwards, covering the umbilicus. The teleoconch spots (if present) form 2 rows on the whorls (often, a 3
<superScript id="7C579BCF3142FFD47B7C16C9F01122F9" attach="left" box="[307,321,1706,1720]" fontSize="6" pageId="15" pageNumber="64">rd</superScript>
row of spots is visible on the body whorl): the upper one close to the suture with rectangular spots abapically elongated; the lower one roughly ⅓ up on the whorls (
<figureCitation id="13192A023142FFD47E5B16ADF53D22A9" box="[1044,1133,1742,1768]" captionStart="Fig" captionStartId="17.[189,232,1990,2016]" captionTargetBox="[215,1371,1099,1923]" captionTargetId="figure-238@17.[189,1399,1082,1953]" captionTargetPageId="17" captionText="Fig. 12. Most common spot patterns on the shells of the species of Clessiniella Zallot et al., 2015." figureDoi="http://doi.org/10.5281/zenodo.5176935" httpUri="https://zenodo.org/record/5176935/files/figure.png" pageId="15" pageNumber="64">Fig. 12</figureCitation>
). In some populations the 2 rows of spots merge and there is a continuous vertical darker band on the whorls. The protoconch is dull and robust with fine riblets starting after 1.2 whorls or more from the apex; it is transparent and fragile or the ribbing starts close to the apex in other subgenera.
</paragraph>
</subSubSection>
<subSubSection id="C338650C3142FFCA7AF21719F2E7258F" lastPageId="17" lastPageNumber="66" pageId="15" pageNumber="64" type="description">
<paragraph id="8B9D36873142FFD47AF21719F4282398" blockId="15.[189,1400,1913,2010]" pageId="15" pageNumber="64">FEMALE GENITALIA. There is an apical connection of the pedunculus to the bursa copulatrix, a long seminal receptacle (normally club-shaped), which runs parallel to the pedunculus. The junction of the uterus gland is situated close to the connection between the pedunculus and the distal oviduct. This</paragraph>
<caption id="DF5D660F315DFFCB7AF2175BF28123A1" ID-DOI="http://doi.org/10.5281/zenodo.5176933" ID-Zenodo-Dep="5176933" httpUri="https://zenodo.org/record/5176933/files/figure.png" pageId="16" pageNumber="65" startId="16.[189,232,1848,1874]" targetBox="[291,1294,297,1793]" targetPageId="16">
<paragraph id="8B9D3687315DFFCB7AF2175BF28123A1" blockId="16.[187,1399,1848,2016]" pageId="16" pageNumber="65">
<emphasis id="B956EA95315DFFCB7AF2175BF04B2313" bold="true" box="[189,283,1848,1874]" pageId="16" pageNumber="65">Fig. 11.</emphasis>
Ribs per mm on the 1
<superScript id="7C579BCF315DFFCB787D175BF36D2307" attach="left" box="[562,573,1848,1862]" fontSize="6" pageId="16" pageNumber="65">st</superScript>
whorl vs ribs on the 4
<superScript id="7C579BCF315DFFCB7919175BF2332307" attach="left" box="[854,867,1848,1862]" fontSize="6" pageId="16" pageNumber="65">th</superScript>
whorls of the species in
<taxonomicName id="4C224D04315DFFCB7ED5175AF0012336" authority="Zallot et al., 2015" authorityName="Zallot" authorityYear="2015" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="61" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA95315DFFCB7ED5175AF4762312" box="[1178,1318,1849,1875]" italics="true" pageId="16" pageNumber="65">Clessiniella</emphasis>
<bibRefCitation id="EFB34B76315DFFCB7F7F175AF0012336" author="Zallot E. &amp; Groenenberg D. S. J. &amp; De Mattia W. &amp; Feher Z. &amp; Gittenberger E." pageId="16" pageNumber="65" pagination="63 - 88" refId="ref16559" refString="Zallot E., Groenenberg D. S. J., De Mattia W., Feher Z. &amp; Gittenberger E. 2015. Genera, subgenera and species of the Cochlostomatidae (Gastropoda, Caenogastropoda, Cochlostomatidae). Basteria 78 (4 - 6): 63 - 88." type="journal article" year="2015">
Zallot
<emphasis id="B956EA95315DFFCB7AF2173EF1A72337" box="[189,247,1884,1910]" italics="true" pageId="16" pageNumber="65">et al</emphasis>
., 2015
</bibRefCitation>
</taxonomicName>
. Violet dots =
<taxonomicName id="4C224D04315DFFCB7857173FF5E82337" authority="Boeckel, 1939" authorityName="Boeckel" authorityYear="1939" box="[536,1208,1884,1910]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="65" phylum="Mollusca" rank="species" species="anomphale" subGenus="Clessiniella">
<emphasis id="B956EA95315DFFCB7857173FF3E42337" box="[536,692,1884,1910]" italics="true" pageId="16" pageNumber="65">Cochlostoma</emphasis>
(
<emphasis id="B956EA95315DFFCB7883173FF2082337" box="[716,856,1884,1910]" italics="true" pageId="16" pageNumber="65">Clessiniella</emphasis>
)
<emphasis id="B956EA95315DFFCB793E173FF2A52337" box="[881,1013,1884,1910]" italics="true" pageId="16" pageNumber="65">anomphale</emphasis>
<bibRefCitation id="EFB34B76315DFFCB7E4C173FF5E82337" author="Boeckel W." box="[1027,1208,1884,1910]" pageId="16" pageNumber="65" pagination="41 - 50" refId="ref15670" refString="Boeckel W. 1939. Cochlostoma - Arten aus den Karawanken. Archiv fur Molluskenkunde 71: 41 - 50." type="journal article" year="1939">Boeckel, 1939</bibRefCitation>
</taxonomicName>
; red squares =
<taxonomicName id="4C224D04315DFFCB7AF2171CF0FD23D8" authority="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger, 2021" authorityName="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger" authorityYear="2021" box="[189,429,1919,1946]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="65" phylum="Mollusca" rank="species" species="stelucarum" status="sp. nov." subGenus="Clessiniella">
<emphasis id="B956EA95315DFFCB7AF2171CF1B723D8" box="[189,231,1919,1945]" italics="true" pageId="16" pageNumber="65">Co.</emphasis>
(
<emphasis id="B956EA95315DFFCB7AB9171CF04923D8" box="[246,281,1919,1945]" italics="true" pageId="16" pageNumber="65">Cl.</emphasis>
)
<emphasis id="B956EA95315DFFCB7B66171CF0FD23D8" box="[297,429,1919,1945]" italics="true" pageId="16" pageNumber="65">stelucarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557EE315DFFCB7BE017E3F35C23DB" box="[431,524,1920,1946]" pageId="16" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
; green triangles =
<taxonomicName id="4C224D04315DFFCB7891171CF59123D8" authority="(Westerlund, 1878)" baseAuthorityName="Westerlund" baseAuthorityYear="1878" box="[734,1217,1919,1946]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="65" phylum="Mollusca" rank="species" species="tergestinum" subGenus="Clessiniella">
<emphasis id="B956EA95315DFFCB7891171CF25823D8" box="[734,776,1919,1945]" italics="true" pageId="16" pageNumber="65">Co.</emphasis>
(
<emphasis id="B956EA95315DFFCB7956171CF26C23D8" box="[793,828,1919,1945]" italics="true" pageId="16" pageNumber="65">Cl.</emphasis>
)
<emphasis id="B956EA95315DFFCB790317E3F28723D8" box="[844,983,1920,1945]" italics="true" pageId="16" pageNumber="65">tergestinum</emphasis>
(
<bibRefCitation id="EFB34B76315DFFCB79AB171CF5E923D8" author="Westerlund C. A." box="[996,1209,1919,1945]" pageId="16" pageNumber="65" pagination="108 - 109" refId="ref16440" refString="Westerlund C. A. 1878. Diagnosen neuer Mollusken. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 10: 108 - 109." type="journal article" year="1878">Westerlund, 1878</bibRefCitation>
)
</taxonomicName>
; blue diamonds =
<taxonomicName id="4C224D04315DFFCB7A9A17C0F31423FC" authority="(Strobel, 1851)" baseAuthorityName="Strobel" baseAuthorityYear="1851" box="[213,580,1954,1981]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="65" phylum="Mollusca" rank="species" species="villae" subGenus="Clessiniella">
<emphasis id="B956EA95315DFFCB7A9A17C0F1AF23FC" box="[213,255,1955,1981]" italics="true" pageId="16" pageNumber="65">Co.</emphasis>
(
<emphasis id="B956EA95315DFFCB7B5F17C1F06323FD" box="[272,307,1954,1980]" italics="true" pageId="16" pageNumber="65">Cl.</emphasis>
)
<emphasis id="B956EA95315DFFCB7B0A17C1F0D723FD" box="[325,391,1954,1980]" italics="true" pageId="16" pageNumber="65">villae</emphasis>
(
<bibRefCitation id="EFB34B76315DFFCB7BD917C1F36B23FC" author="Strobel P." box="[406,571,1954,1981]" pageId="16" pageNumber="65" refId="ref16289" refString="Strobel P. 1851. Notizie Malacostatiche sul Trentino. Fusi, Pavia. Available from https: // www. biodiversitylibrary. org / page / 46194631 [accessed 12 Jul. 2021]." type="url" year="1851">Strobel, 1851</bibRefCitation>
)
</taxonomicName>
; black triangles =
<taxonomicName id="4C224D04315DFFCB795217C0F45523FD" authority="(A. J. Wagner, 1897)" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1897" box="[797,1285,1954,1981]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="65" phylum="Mollusca" rank="species" species="waldemari" subGenus="Clessiniella">
<emphasis id="B956EA95315DFFCB795217C0F21723FC" box="[797,839,1955,1981]" italics="true" pageId="16" pageNumber="65">Co.</emphasis>
(
<emphasis id="B956EA95315DFFCB791617C1F22C23FD" box="[857,892,1954,1980]" italics="true" pageId="16" pageNumber="65">Cl.</emphasis>
)
<emphasis id="B956EA95315DFFCB79C217C1F55C23FD" box="[909,1036,1954,1980]" italics="true" pageId="16" pageNumber="65">waldemari</emphasis>
(A.J.
<bibRefCitation id="EFB34B76315DFFCB7E1B17C0F5AD23FD" author="Wagner A. J." box="[1108,1277,1954,1981]" pageId="16" pageNumber="65" pagination="565 - 632" refId="ref16375" refString="Wagner A. J. 1897. Monographie der Gattung Pomatias Studer. Denkschriften der kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 64: 565 - 632. https: // doi. org / 10.5962 / bhl. title. 10792" type="journal article" year="1897">Wagner, 1897</bibRefCitation>
)
</taxonomicName>
. Analysis performed with the software ‘Past’ (https://palaeo-electronica.org).
</paragraph>
</caption>
<paragraph id="8B9D3687315CFFCA7AF2116AF02F2507" blockId="17.[189,1399,265,326]" pageId="17" pageNumber="66">
combination of characters in the female genitalia is most diagnostic to distinguish
<taxonomicName id="4C224D04315CFFCA7ECE116AF45C2562" box="[1153,1292,265,291]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="64" phylum="Mollusca" rank="subGenus" subGenus="Clessiniella">
<emphasis id="B956EA95315CFFCA7ECE116AF45C2562" box="[1153,1292,265,291]" italics="true" pageId="17" pageNumber="66">Clessiniella</emphasis>
</taxonomicName>
from the other subgenera.
</paragraph>
<paragraph id="8B9D3687315CFFCA7AF2110DF2E7258F" blockId="17.[188,1400,366,463]" pageId="17" pageNumber="66">MALE GENITALIA. Apart from its apical part, the penial spermiduct is tortuous. The body spermiduct is protruding (often almost detached from the body) more than in the other subgenera and either straight or more or less tortuous. The posterior side of the penis is wrinkled.</paragraph>
</subSubSection>
<subSubSection id="C338650C315CFFCA7AF21196F300264E" box="[189,592,501,528]" pageId="17" pageNumber="66" type="discussion">
<paragraph id="8B9D3687315CFFCA7AF21196F300264E" blockId="17.[189,592,501,528]" box="[189,592,501,528]" pageId="17" pageNumber="66">
<heading id="D0D581EB315CFFCA7AF21196F300264E" bold="true" box="[189,592,501,528]" fontSize="11" level="3" pageId="17" pageNumber="66" reason="3">
<emphasis id="B956EA95315CFFCA7AF21196F300264E" bold="true" box="[189,592,501,528]" pageId="17" pageNumber="66">Identification key for the species</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338650C315CFFCA7AF21247F42627B8" pageId="17" pageNumber="66" type="key">
<key id="FFB361B6315CFFCA7AF21247F42627B8" pageId="17" pageNumber="66">
<keyStep id="30D62E22315CFFCA7AF21247F42726C4" pageId="17" pageNumber="66">
<paragraph id="8B9D3687315CFFCA7AF21247F4272620" blockId="17.[189,1399,548,1017]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7AF21247F4272620" pageId="17" pageNumber="66">
1. Body whorl inflated shortly before the lip (
<figureCitation id="13192A02315CFFCA78971247F265267F" box="[728,821,548,574]" captionStart="Fig" captionStartId="5.[189,232,1854,1880]" captionTargetBox="[200,1387,265,1815]" captionTargetId="figure-11@5.[200,1387,265,1815]" captionTargetPageId="5" captionText="Fig. 4. Features of the shell of Clessiniella Zallot et al., 2015. A. Shape of the body whorl approaching the aperture. B. Strength of the spots on the shell (0 = spotless shell to 2 = really evident spots). C. Strength of the ribs (1 = weak ribs to 3 = really prominent ribs). D. Strength of the ribs on the external side of the body whorl approaching the aperture (W= smooth shell; I = intermediate; S = prominent ribs)." figureDoi="http://doi.org/10.5281/zenodo.5176919" httpUri="https://zenodo.org/record/5176919/files/figure.png" pageId="17" pageNumber="66">Fig. 4A</figureCitation>
: OB). In the female genitalia, simplified loop path with 1–2 loops ............................................................................
<taxonomicName id="4C224D04315CFFCA799F1224F4272620" ID-CoL="WNMX" authority="Boeckel, 1939" authorityName="Boeckel" authorityYear="1939" box="[976,1399,583,610]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="66" phylum="Mollusca" rank="species" species="anomphale" subGenus="Clessiniella">
<emphasis id="B956EA95315CFFCA799F1224F5952623" bold="true" box="[976,1221,583,610]" italics="true" pageId="17" pageNumber="66">Co. (Cl.) anomphale</emphasis>
<bibRefCitation id="EFB34B76315CFFCA7E821224F4272620" author="Boeckel W." box="[1229,1399,583,609]" pageId="17" pageNumber="66" pagination="41 - 50" refId="ref15670" refString="Boeckel W. 1939. Cochlostoma - Arten aus den Karawanken. Archiv fur Molluskenkunde 71: 41 - 50." type="journal article" year="1939">Boeckel, 1939</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9D3687315CFFCA7A8E1208F42726C4" blockId="17.[189,1399,548,1017]" box="[193,1399,618,645]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7A8E1208F42726C4" box="[193,1399,618,645]" pageId="17" pageNumber="66">
– Body whorl not inflated before the lip (
<figureCitation id="13192A02315CFFCA78F91209F24426C4" box="[694,788,618,645]" captionStart="Fig" captionStartId="5.[189,232,1854,1880]" captionTargetBox="[200,1387,265,1815]" captionTargetId="figure-11@5.[200,1387,265,1815]" captionTargetPageId="5" captionText="Fig. 4. Features of the shell of Clessiniella Zallot et al., 2015. A. Shape of the body whorl approaching the aperture. B. Strength of the spots on the shell (0 = spotless shell to 2 = really evident spots). C. Strength of the ribs (1 = weak ribs to 3 = really prominent ribs). D. Strength of the ribs on the external side of the body whorl approaching the aperture (W= smooth shell; I = intermediate; S = prominent ribs)." figureDoi="http://doi.org/10.5281/zenodo.5176919" httpUri="https://zenodo.org/record/5176919/files/figure.png" pageId="17" pageNumber="66">Fig. 4A</figureCitation>
: FF, AF, II) ............................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E22315CFFCA7AF212CFF42726A8" pageId="17" pageNumber="66">
<paragraph id="8B9D3687315CFFCA7AF212CFF4272687" blockId="17.[189,1399,548,1017]" box="[189,1399,683,710]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7AF212CFF4272687" box="[189,1399,683,710]" pageId="17" pageNumber="66">
2. In the female genitalia, the loops of the visceral oviduct are 1–2 ........
<taxonomicName id="4C224D04315CFFCA7E5012CFF4452687" ID-CoL="8VQSP" authority="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger, 2021" authorityName="Zallot &amp; Mattia &amp; Fehér &amp; Gittenberger" authorityYear="2021" box="[1055,1301,683,710]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="66" phylum="Mollusca" rank="species" species="stelucarum" status="sp. nov." subGenus="Clessiniella">
<emphasis id="B956EA95315CFFCA7E5012CFF4452687" bold="true" box="[1055,1301,683,710]" italics="true" pageId="17" pageNumber="66">Co. (Cl.) stelucarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26557EE315CFFCA7F5312CFF4272687" box="[1308,1399,684,710]" pageId="17" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B9D3687315CFFCA7A8E12B3F42726A8" blockId="17.[189,1399,548,1017]" box="[193,1399,719,746]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7A8E12B3F42726A8" box="[193,1399,719,746]" pageId="17" pageNumber="66">– In the female genitalia, the loops of the visceral oviduct are more than 2 ....................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E22315CFFCA7AF21372F427270F" pageId="17" pageNumber="66">
<paragraph id="8B9D3687315CFFCA7AF21372F426276A" blockId="17.[189,1399,548,1017]" box="[189,1398,784,811]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7AF21372F426276A" box="[189,1398,784,811]" pageId="17" pageNumber="66">
3. In the male genitalia, the body spermiduct is twisted ...............
<taxonomicName id="4C224D04315CFFCA79841373F426276A" ID-CoL="WNPL" authority="(Wagner, 1897)" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1897" box="[971,1398,784,811]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="66" phylum="Mollusca" rank="species" species="waldemari" subGenus="Clessiniella">
<emphasis id="B956EA95315CFFCA79841373F5E9276A" bold="true" box="[971,1209,784,811]" italics="true" pageId="17" pageNumber="66">Co. (Cl.) waldemari</emphasis>
(
<bibRefCitation id="EFB34B76315CFFCA7E871372F43E276A" author="Wagner A. J." box="[1224,1390,785,811]" pageId="17" pageNumber="66" pagination="565 - 632" refId="ref16375" refString="Wagner A. J. 1897. Monographie der Gattung Pomatias Studer. Denkschriften der kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 64: 565 - 632. https: // doi. org / 10.5962 / bhl. title. 10792" type="journal article" year="1897">Wagner, 1897</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9D3687315CFFCA7A8E1357F427270F" blockId="17.[189,1399,548,1017]" box="[193,1399,820,846]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7A8E1357F427270F" box="[193,1399,820,846]" pageId="17" pageNumber="66">– In the male genitalia, the body spermiduct is straight ...................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D62E22315CFFCA7AF21316F42627B8" pageId="17" pageNumber="66">
<paragraph id="8B9D3687315CFFCA7AF21316F42727F2" blockId="17.[189,1399,548,1017]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7AF21316F42727F2" pageId="17" pageNumber="66">
4. Ribs of the shell widely spaced and barely inclined, protoconch relatively small (
<figureCitation id="13192A02315CFFCA7E8A1316F44F27CE" box="[1221,1311,885,911]" captionStart="Fig" captionStartId="18.[189,232,888,914]" captionTargetBox="[419,1117,288,814]" captionTargetId="figure-255@18.[404,1184,262,852]" captionTargetPageId="18" captionText="Fig. 13. Shell characters (ribs inclination and relative size of the protoconch) which could be helpful to distinguish Cochlostoma (Clessiniella) villae (Strobel, 1851)(blue diamonds) fromCo. (Cl.) tergestinum (Westerlund, 1878) (green triangles)." figureDoi="http://doi.org/10.5281/zenodo.5176937" httpUri="https://zenodo.org/record/5176937/files/figure.png" pageId="17" pageNumber="66">Fig. 13</figureCitation>
). Slim, club-shaped seminal receptacle in the female genitalia .....................
<taxonomicName id="4C224D04315CFFCA7E4113FBF42727F2" ID-CoL="WNPK" authority="(Strobel, 1851)" baseAuthorityName="Strobel" baseAuthorityYear="1851" box="[1038,1399,920,947]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="66" phylum="Mollusca" rank="species" species="villae" subGenus="Clessiniella">
<emphasis id="B956EA95315CFFCA7E4113FBF5ED27F2" bold="true" box="[1038,1213,920,947]" italics="true" pageId="17" pageNumber="66">Co. (Cl.) villae</emphasis>
(
<bibRefCitation id="EFB34B76315CFFCA7E8213FBF43E27F2" author="Strobel P." box="[1229,1390,920,947]" pageId="17" pageNumber="66" refId="ref16289" refString="Strobel P. 1851. Notizie Malacostatiche sul Trentino. Fusi, Pavia. Available from https: // www. biodiversitylibrary. org / page / 46194631 [accessed 12 Jul. 2021]." type="url" year="1851">Strobel, 1851</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9D3687315CFFCA7A8E13DFF42627B8" blockId="17.[189,1399,548,1017]" pageId="17" pageNumber="66">
<keyLead id="30D395B2315CFFCA7A8E13DFF42627B8" pageId="17" pageNumber="66">
– Ribs less widely spaced and more inclined, relatively larger protoconch. Less slim seminal receptacle ..........................................................................
<taxonomicName id="4C224D04315CFFCA79C013BCF42627B8" ID-CoL="WNPG" authority="(Westerlund, 1878)" baseAuthorityName="Westerlund" baseAuthorityYear="1878" box="[911,1398,990,1017]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="66" phylum="Mollusca" rank="species" species="tergestinum" subGenus="Clessiniella">
<emphasis id="B956EA95315CFFCA79C013BCF5DD27B8" bold="true" box="[911,1165,990,1017]" italics="true" pageId="17" pageNumber="66">Co. (Cl.) tergestinum</emphasis>
(
<bibRefCitation id="EFB34B76315CFFCA7ED313BCF43E27B8" author="Westerlund C. A." box="[1180,1390,991,1017]" pageId="17" pageNumber="66" pagination="108 - 109" refId="ref16440" refString="Westerlund C. A. 1878. Diagnosen neuer Mollusken. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 10: 108 - 109." type="journal article" year="1878">Westerlund, 1878</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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