Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae) Author Zallot, Enrico 365E17AD-6938-4364-A526-F1BDA2663E10 Haagweg 29, 2681 PA Monster, the Netherlands. ezallot@gmail.com Author Mattia, Willy De C74A049D-9D7B-4A1F-878B-5BDFD104219F Central Research Laboratories of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. wdemattia@gmail.com Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 Hungarian Natural History Museum, Baross 13, 1088 Budapest, Hungary. feher.zoltan@nhmus.hu Author Gittenberger, Edmund D786C279-FC92-4D08-AF16-F79A9705E0AE Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, the Netherlands. egittenberger@yahoo.com text European Journal of Taxonomy 2021 2021-08-06 762 1 49 95 http://dx.doi.org/10.5852/ejt.2021.762.1453 journal article 5011 10.5852/ejt.2021.762.1453 b739dd1a-47f7-4e15-a8c6-caf0a89947cb 2118-9773 5176909 D1093CEB-7EBB-44A0-8078-8DC616B857F5 Subgenus Clessiniella Zallot, Groenenberg, De Mattia, Fehér & Gittenberger, 2015 Clessiniella Zallot, Groenenberg, De Mattia, Fehér & Gittenberger, 2015: 80 . Type species Pomatias maculatum var. villae Strobel, 1851 , by original designation. Remarks The classification follows MolluscaBase (2021) except for the subgeneric name. Contrary to ICZN Art. 10.4, Neglecta Wagner, 1897 is used in MolluscaBase as a senior synonym of Clessiniella . Wagner (1897: 6 [570]) introduced Neglecta as “Formenkreis Neglecta ”. It cannot be denied that the German term Formenkreis is “.. a term such as “superspecies””. As a consequence, according to the ICZN ruling, Neglecta “.. is not deemed to be a genus-group name” (ICZN Art. 10.4, see also Zallot et al. 2015 ). Strobel (1851) and De Betta & Martinati (1855) named this entity Pomatias maculatum var. Villae – but referring to Pomatias maculatus Draparnaud, 1805 (= Co. ( Co. ) septemspirale ). Diagnosis of the subgenus SHELL. Ribbed teleoconch with straight ribs, variable in strength and shape. The columellar lobe is curved backwards, covering the umbilicus. The teleoconch spots (if present) form 2 rows on the whorls (often, a 3 rd row of spots is visible on the body whorl): the upper one close to the suture with rectangular spots abapically elongated; the lower one roughly ⅓ up on the whorls ( Fig. 12 ). In some populations the 2 rows of spots merge and there is a continuous vertical darker band on the whorls. The protoconch is dull and robust with fine riblets starting after 1.2 whorls or more from the apex; it is transparent and fragile or the ribbing starts close to the apex in other subgenera. FEMALE GENITALIA. There is an apical connection of the pedunculus to the bursa copulatrix, a long seminal receptacle (normally club-shaped), which runs parallel to the pedunculus. The junction of the uterus gland is situated close to the connection between the pedunculus and the distal oviduct. This Fig. 11. Ribs per mm on the 1 st whorl vs ribs on the 4 th whorls of the species in Clessiniella Zallot et al ., 2015 . Violet dots = Cochlostoma ( Clessiniella ) anomphale Boeckel, 1939 ; red squares = Co. ( Cl. ) stelucarum sp. nov. ; green triangles = Co. ( Cl. ) tergestinum ( Westerlund, 1878 ) ; blue diamonds = Co. ( Cl. ) villae ( Strobel, 1851 ) ; black triangles = Co. ( Cl. ) waldemari (A.J. Wagner, 1897 ) . Analysis performed with the software ‘Past’ (https://palaeo-electronica.org). combination of characters in the female genitalia is most diagnostic to distinguish Clessiniella from the other subgenera. MALE GENITALIA. Apart from its apical part, the penial spermiduct is tortuous. The body spermiduct is protruding (often almost detached from the body) more than in the other subgenera and either straight or more or less tortuous. The posterior side of the penis is wrinkled. Identification key for the species 1. Body whorl inflated shortly before the lip ( Fig. 4A : OB). In the female genitalia, simplified loop path with 1–2 loops ............................................................................ Co. (Cl.) anomphale Boeckel, 1939 – Body whorl not inflated before the lip ( Fig. 4A : FF, AF, II) ............................................................. 2 2. In the female genitalia, the loops of the visceral oviduct are 1–2 ........ Co. (Cl.) stelucarum sp. nov. – In the female genitalia, the loops of the visceral oviduct are more than 2 ....................................... 3 3. In the male genitalia, the body spermiduct is twisted ............... Co. (Cl.) waldemari ( Wagner, 1897 ) – In the male genitalia, the body spermiduct is straight ...................................................................... 4 4. Ribs of the shell widely spaced and barely inclined, protoconch relatively small ( Fig. 13 ). Slim, club-shaped seminal receptacle in the female genitalia ..................... Co. (Cl.) villae ( Strobel, 1851 ) – Ribs less widely spaced and more inclined, relatively larger protoconch. Less slim seminal receptacle .......................................................................... Co. (Cl.) tergestinum ( Westerlund, 1878 )