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<mods:title id="BAAC7559EC6E69C9620AF0B06AE72D31">Redescription of Arcotheres pernicola (Bürger, 1895) (Crustacea: Decapoda Brachyura: Pinnotheridae) from the oyster Magallana gryphoides (Schlotheim 1820) in India</mods:title>
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<mods:namePart id="26CA4105DC97AAC4F88D52B9F584CA0C">Trivedi, Jigneshkumar N.</mods:namePart>
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<mods:namePart id="1194DC45947AC1F103807812ED0D8841">Vachhrajani, Kauresh D.</mods:namePart>
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<mods:namePart id="0B852D277429FCDA6D0AEEF7BEA7D135">Ng, Peter K. L.</mods:namePart>
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<mods:date id="414A886913D3FA5D6208AABE80EDB8CE">2019</mods:date>
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<paragraph id="F829A779FF83FFD0FF3B6EF6FDD8B580" blockId="1.[151,591,909,971]" box="[151,591,909,936]" pageId="1" pageNumber="588">
<heading id="A3611015FF83FFD0FF3B6EF6FDD8B580" bold="true" box="[151,591,909,936]" fontSize="11" level="1" pageId="1" pageNumber="588" reason="1">
<taxonomicName id="3F96DCFAFF83FFD0FF3B6EF6FDD8B580" ID-CoL="5W29W" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[151,591,909,936]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF83FFD0FF3B6EF6FDD8B580" bold="true" box="[151,591,909,936]" pageId="1" pageNumber="588">
<emphasis id="CAE27B6BFF83FFD0FF3B6EF6FE02B58F" bold="true" box="[151,405,909,936]" italics="true" pageId="1" pageNumber="588">Arcotheres pernicola</emphasis>
(
<bibRefCitation id="9C07DA88FF83FFD0FE086EF6FDD0B580" author="Burger, O." box="[420,583,909,935]" pageId="1" pageNumber="588" pagination="361 - 390" refId="ref3337" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="B08CF4F2FF83FFD0FF3B6ECAFE85B5EC" box="[151,274,945,971]" pageId="1" pageNumber="588" type="description">
<paragraph id="F829A779FF83FFD0FF3B6ECAFE85B5EC" blockId="1.[151,591,909,971]" box="[151,274,945,971]" pageId="1" pageNumber="588">
(
<figureCitation id="60ADBBFCFF83FFD0FF336ECAFE9EB5EC" box="[159,265,945,971]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1181,1206]" captionStartId-1="3.[151,250,1963,1988]" captionStartId-2="4.[151,250,1945,1970]" captionTargetBox-0="[340,1247,594,1152]" captionTargetBox-1="[161,1424,604,1927]" captionTargetBox-2="[193,1386,192,1904]" captionTargetId-0="figure@2.[340,1247,594,1152]" captionTargetId-1="figure@3.[151,1436,597,1938]" captionTargetId-2="figure@4.[186,1402,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128). Colour in life, dorsal view." captionText-1="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." captionText-2="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi-0="http://doi.org/10.5281/zenodo.3617045" figureDoi-1="http://doi.org/10.5281/zenodo.3617049" figureDoi-2="http://doi.org/10.5281/zenodo.3617051" httpUri-0="https://zenodo.org/record/3617045/files/figure.png" httpUri-1="https://zenodo.org/record/3617049/files/figure.png" httpUri-2="https://zenodo.org/record/3617051/files/figure.png" pageId="1" pageNumber="588">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B08CF4F2FF83FFD0FF3B6E8EFA0BB280" pageId="1" pageNumber="588" type="reference_group">
<paragraph id="F829A779FF83FFD0FF3B6E8EFA0BB22A" blockId="1.[151,1436,1012,1221]" box="[151,1436,1012,1038]" pageId="1" pageNumber="588">
<treatmentCitationGroup id="D8868057FF83FFD0FF3B6E8EFA0BB22A" box="[151,1436,1012,1038]" pageId="1" pageNumber="588">
<taxonomicName id="3F96DCFAFF83FFD0FF3B6E8EFDA7B22A" ID-CoL="4HZ8F" authority="Burger 1895: 375" authorityName="Burger" authorityPageNumber="375" authorityYear="1895" box="[151,560,1012,1038]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF83FFD0FF3B6E8EFEE0B22A" box="[151,375,1013,1037]" italics="true" pageId="1" pageNumber="588">Pinnotheres pernicola</emphasis>
<treatmentCitation id="79378168FF83FFD0FED16E8DFDA7B22A" author="Burger, O." box="[381,560,1012,1038]" page="375" pageId="1" pageNumber="588" year="1895">
<bibRefCitation id="9C07DA88FF83FFD0FED16E8DFDA7B22A" author="Burger, O." box="[381,560,1012,1038]" pageId="1" pageNumber="588" pagination="361 - 390" refId="ref3337" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger 1895: 375</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, 376, pl. 9 fig. 17, pl. 10 fig. 16;
<treatmentCitation id="79378168FF83FFD0FCD56E8FFBB2B22A" author="Nobili, G." box="[889,1061,1012,1037]" page="264" pageId="1" pageNumber="588" year="1899">
<bibRefCitation id="9C07DA88FF83FFD0FCD56E8FFBB2B22A" author="Nobili, G." box="[889,1061,1012,1037]" pageId="1" pageNumber="588" pagination="230 - 282" refId="ref3941" refString="Nobili, G. (1899) Contribuzioni alla conoscienza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali dei Museo Civico di Storia Naturale di Genova, Series 2, 20, 230 - 282." type="journal article" year="1899">Nobili 1899: 264</bibRefCitation>
</treatmentCitation>
(discussion);
<treatmentCitation id="79378168FF83FFD0FB1E6E8FFAF7B22A" author="Laurie, R. D." box="[1202,1376,1012,1037]" page="415" pageId="1" pageNumber="588" year="1915">
<bibRefCitation id="9C07DA88FF83FFD0FB1E6E8FFAF7B22A" author="Laurie, R. D." box="[1202,1376,1012,1037]" pageId="1" pageNumber="588" pagination="407 - 475" refId="ref3578" refString="Laurie, R. D. (1915) Reports on the marine biology of the Sudanese Red Sea, from collections made by Cyril Crossland, M. A., B. Sc., F. Z. S. XXI. On the Brachyura. Journal of the Linnaean Society of London, Zoology, 31 (209), 407 - 475, pls. 42 - 45. https: // doi. org / 10.1111 / j. 1096 - 3642.1915. tb 00463. x" type="journal article" year="1915">Laurie 1915: 415</bibRefCitation>
</treatmentCitation>
(list);
</treatmentCitationGroup>
</paragraph>
<paragraph id="F829A779FF83FFD0FF6B6968FA0BB280" blockId="1.[151,1436,1012,1221]" pageId="1" pageNumber="588">
<bibRefCitation id="9C07DA88FF83FFD0FF6B6968FE39B20B" author="Schmitt, W. L. &amp; McCain, J. C. &amp; Davidson, E. S." box="[199,430,1043,1068]" pageId="1" pageNumber="588" pagination="1 - 160" refId="ref4030" refString="Schmitt, W. L., McCain, J. C. &amp; Davidson, E. S. (1973) Decapoda. I. Brachyura. I. Fam. Pinnotheridae. In: Gruner, H. E. &amp; Holthuis, L. B. (Eds.), Crustaceorum Catalogus, 3, pp. 1 - 160." type="journal article" year="1973">
Schmitt
<emphasis id="CAE27B6BFF83FFD0FEB0696EFEC7B20B" box="[284,336,1044,1068]" italics="true" pageId="1" pageNumber="588">et al.</emphasis>
1973: 66
</bibRefCitation>
(discussion).
<taxonomicName id="3F96DCFAFF83FFD0FF3B6948FDA3B26C" authority="- Nobili 1906: 303" authorityName="Nobili" authorityPageNumber="303" authorityYear="1906" box="[151,564,1074,1099]" class="Malacostraca" family="Pinnotheridae" genus="Pinnoteres" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF83FFD0FF3B6948FEFCB26D" box="[151,363,1074,1098]" italics="true" pageId="1" pageNumber="588">Pinnoteres pernicola</emphasis>
—
<bibRefCitation id="9C07DA88FF83FFD0FE296949FDA3B26C" author="Nobili, G." box="[389,564,1074,1099]" pageId="1" pageNumber="588" pagination="393 - 411" refId="ref3981" refString="Nobili, G. (1906) Diagnoses preliminaires de 34 especes et varietes nouvelles, et de 2 genres nouveaux de Decapodes de la Mer Rouge. Bulletin du Museum national d'Histoire naturelle, Paris, 11 (6), 393 - 411. [1906]" type="journal article" year="1906">Nobili 1906: 303</bibRefCitation>
</taxonomicName>
, 304.
<taxonomicName id="3F96DCFAFF83FFD0FF3B692AFDD3B24E" ID-CoL="7Y7YF" authority="- Tesch 1918: 250" authorityName="Tesch" authorityPageNumber="250" authorityYear="1918" box="[151,580,1104,1129]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pernicolus">
<emphasis id="CAE27B6BFF83FFD0FF3B692AFE15B24E" box="[151,386,1105,1129]" italics="true" pageId="1" pageNumber="588">Pinnotheres pernicolus</emphasis>
—
<bibRefCitation id="9C07DA88FF83FFD0FE37692BFDD3B24E" author="Tesch, J. J." box="[411,580,1104,1129]" pageId="1" pageNumber="588" pagination="149 - 295" refId="ref4187" refString="Tesch, J. J. (1918) Goneplacidae and Pinnotheridae. The Decapoda Brachyura of the Siboga-Expedition. II. Siboga Expeditie Monografie, 39 c (84), 149 - 295, pls. 7 - 18. https: // doi. org / 10.5962 / bhl. title. 10267" type="journal article" year="1918">Tesch 1918: 250</bibRefCitation>
</taxonomicName>
, 254, 287 (list, key);
<bibRefCitation id="9C07DA88FF83FFD0FCB1692BFBF3B24E" author="Silas, E. G. &amp; Algarswami, K." box="[797,1124,1104,1129]" pageId="1" pageNumber="588" pagination="1161 - 1227" refId="ref4096" refString="Silas, E. G. &amp; Algarswami, K. (1967) On an instance of parasitisation by the pea crab (Pinnotheres sp.) on the backwater clam Meretrix casta (Chemnitz) from India with a review of the work on the systematics, ecology, biology and ethology of the pea-crabs of the genus Pinnotheres Latreille. In: Proceeding of Symposium in Crustacea. Pt. III. Marine Biological Association of India, Cochin, Kerala, pp. 1161 - 1227." type="book chapter" year="1967">Silas &amp; Algarswami 1967: 1206</bibRefCitation>
, 1216, 1222 (list)
<taxonomicName id="3F96DCFAFF83FFD0FF3B690AFDD6B2AF" authority="- Guinot 1967: 279" authorityName="Guinot" authorityPageNumber="279" authorityYear="1967" box="[151,577,1135,1160]" class="Malacostraca" family="Pinnotheridae" genus="Pinnoteres" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pernicolus">
<emphasis id="CAE27B6BFF83FFD0FF3B690AFEE2B2AF" box="[151,373,1136,1160]" italics="true" pageId="1" pageNumber="588">Pinnoteres pernicolus</emphasis>
—
<bibRefCitation id="9C07DA88FF83FFD0FE236914FDD6B2AF" author="Guinot, D." box="[399,577,1135,1160]" pageId="1" pageNumber="588" pagination="235 - 352" refId="ref3516" refString="Guinot, D. (1967) La faune carcinologique (Crustacea Brachyura) de l'Ocean Indien occidental et de la mer Rouge. Catalogue, remarques biogeographiques et bibliographie. In: Reunion de Specialistes C. S. A. sur les Crustaces, Zanzibar 1964. Memoires de l'Institute fundamental d'Afrique Noire, 77, 235 - 352." type="journal article" year="1967">Guinot 1967: 279</bibRefCitation>
</taxonomicName>
(list)
<taxonomicName id="3F96DCFAFF83FFD0FF3B69F5FD06B280" ID-CoL="5W29W" authority="- Ahyong &amp; Ng 2007: 202" authorityName="Ahyong &amp; Ng" authorityPageNumber="202" authorityYear="2007" box="[151,657,1166,1191]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF83FFD0FF3B69F5FEF9B281" box="[151,366,1166,1190]" italics="true" pageId="1" pageNumber="588">Arcotheres pernicola</emphasis>
—
<bibRefCitation id="9C07DA88FF83FFD0FE2469F5FD06B280" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[392,657,1166,1191]" pageId="1" pageNumber="588" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007: 202</bibRefCitation>
</taxonomicName>
, 203, fig. 10;
<bibRefCitation id="9C07DA88FF83FFD0FC8469F4FC60B280" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[808,1015,1166,1191]" pageId="1" pageNumber="588" pagination="1 - 286" refId="ref3887" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CAE27B6BFF83FFD0FCFC69F4FC11B281" box="[848,902,1166,1190]" italics="true" pageId="1" pageNumber="588">et al.</emphasis>
2008: 248
</bibRefCitation>
(list);
<bibRefCitation id="9C07DA88FF83FFD0FBEC69F4FA85B280" author="Ng, P. K. L. &amp; Clark, P. F. &amp; Mitra, S. &amp; Kumar, A. B." box="[1088,1298,1166,1191]" pageId="1" pageNumber="588" pagination="1079 - 1097" refId="ref3796" refString="Ng, P. K. L., Clark, P. F., Mitra, S. &amp; Kumar, A. B. (2017) Arcotheres borradailei (Nobili, 1906) and Pinnotheres ridgewayi Southwell, 1911: a reassessment of characters and generic assignment of species to Arcotheres Manning, 1993 (Decapoda, Brachyura, Pinnotheridae). Crustaceana, 90 (7 - 10), 1079 - 1097. https: // doi. org / 10.1163 / 15685403 - 00003613" type="journal article" year="2017">
Ng
<emphasis id="CAE27B6BFF83FFD0FBCB69F4FB0AB281" box="[1127,1181,1166,1190]" italics="true" pageId="1" pageNumber="588">et al.</emphasis>
2017: 1094
</bibRefCitation>
(discussion);
</paragraph>
</subSubSection>
<subSubSection id="B08CF4F2FF83FFD0FF6B69D7FDDDB2E2" box="[199,586,1196,1221]" pageId="1" pageNumber="588" type="description">
<paragraph id="F829A779FF83FFD0FF6B69D7FDDDB2E2" blockId="1.[151,1436,1012,1221]" box="[199,586,1196,1221]" pageId="1" pageNumber="588">
<bibRefCitation id="9C07DA88FF83FFD0FF6B69D7FE56B2E2" author="Trivedi, J. N. &amp; Campos, E. &amp; Vachhrajani, K. D." box="[199,449,1196,1221]" pageId="1" pageNumber="588" pagination="195 - 200" refId="ref4243" refString="Trivedi, J. N., Campos, E. &amp; Vachhrajani, K. D. (2018 a) A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa, 4433 (1), 195 - 200. https: // doi. org / 10.11646 / zootaxa. 4433.1.13" type="journal article" year="2018">
Trivedi
<emphasis id="CAE27B6BFF83FFD0FEBB69D5FEDCB2E2" box="[279,331,1197,1221]" italics="true" pageId="1" pageNumber="588">et al.</emphasis>
2018a: 198
</bibRefCitation>
(discussion).
</paragraph>
</subSubSection>
<subSubSection id="B08CF4F2FF83FFD0FF3B6988FD30B35D" pageId="1" pageNumber="588" type="materials_examined">
<paragraph id="F829A779FF83FFD0FF3B6988FD30B35D" blockId="1.[151,1437,1267,2014]" pageId="1" pageNumber="588">
<emphasis id="CAE27B6BFF83FFD0FF3B6988FE17B329" bold="true" box="[151,384,1267,1294]" pageId="1" pageNumber="588">Material examined.</emphasis>
<specimenCount id="EE906CF0FF83FFD0FE2A698FFE66B32A" box="[390,497,1267,1294]" pageId="1" pageNumber="588" type="female">8 females</specimenCount>
(CW
<quantity id="3F6E0A9CFF83FFD0FD98698FFD1CB329" box="[564,651,1268,1294]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="588" unit="mm" value="4.1">4.1 mm</quantity>
, CL
<quantity id="3F6E0A9CFF83FFD0FD13698FFC82B329" box="[703,789,1268,1294]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="588" unit="mm" value="3.1">3.1 mm</quantity>
to CW
<quantity id="3F6E0A9CFF83FFD0FCC6698FFC56B329" box="[874,961,1268,1294]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="588" unit="mm" value="7.3">7.3 mm</quantity>
, CL
<quantity id="3F6E0A9CFF83FFD0FC596988FBDCB329" box="[1013,1099,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="588" unit="mm" value="5.9">5.9 mm</quantity>
) (ZL-AR-CR-128), in
<taxonomicName id="3F96DCFAFF83FFD0FAE6698FFD22B315" authority="(Schlotheim, 1820)" baseAuthorityName="Schlotheim" baseAuthorityYear="1820" class="Bivalvia" family="Ostreidae" genus="Magallana" higherTaxonomySource="CoL" kingdom="Plantae" order="Ostreida" pageId="1" pageNumber="588" phylum="Mollusca" rank="species" species="gryphoides">
<emphasis id="CAE27B6BFF83FFD0FAE6698FFEDBB315" italics="true" pageId="1" pageNumber="588">Magallana gryphoides</emphasis>
(Schlotheim, 1820) (Ostreidae)
</taxonomicName>
, from Alibaugh,
<collectingRegion id="3A52699BFF83FFD0FCDF686CFB95B316" box="[883,1026,1303,1329]" country="India" name="Maharashtra" pageId="1" pageNumber="588">Maharashtra</collectingRegion>
,
<collectingCountry id="8081E7E9FF83FFD0FBA2686CFBDDB316" box="[1038,1098,1303,1329]" name="India" pageId="1" pageNumber="588">India</collectingCountry>
,
<geoCoordinate id="9DA2C1BEFF83FFD0FBFA686CFB64B315" box="[1110,1267,1303,1330]" degrees="18" direction="north" minutes="38" orientation="latitude" pageId="1" pageNumber="588" precision="1" seconds="11.5" value="18.636528">18°38’11.5”N</geoCoordinate>
<geoCoordinate id="9DA2C1BEFF83FFD0FB56686CFA0FB315" box="[1274,1432,1303,1330]" degrees="72" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="588" precision="1" seconds="52.2" value="72.8645">72°51’52.2”E</geoCoordinate>
, coll. S. Gosavi,
<date id="8C2881B9FF83FFD0FEE06847FD8BB372" box="[332,540,1339,1366]" pageId="1" pageNumber="588" value="2016-11-08">8 November 2016</date>
. 1 damaged female (ca. CW
<quantity id="3F6E0A9CFF83FFD0FCCE6847FC50B371" box="[866,967,1340,1366]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="588" unit="mm" value="11.7">11.7 mm</quantity>
, CL
<quantity id="3F6E0A9CFF83FFD0FC526840FBC1B371" box="[1022,1110,1339,1366]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="588" unit="mm" value="8.5">8.5 mm</quantity>
) (NHM 1955.6.22.55), coast of
<collectingCountry id="8081E7E9FF83FFD0FF1A681BFE95B35D" box="[182,258,1376,1402]" name="Kenya" pageId="1" pageNumber="588">Kenya</collectingCountry>
, east Africa, coll. S. H. Copley, 1955.
</paragraph>
</subSubSection>
<subSubSection id="B08CF4F2FF83FFD3FF6B68F8FC3AB361" lastPageId="2" lastPageNumber="589" pageId="1" pageNumber="588" type="description">
<paragraph id="F829A779FF83FFD0FF6B68F8FC4FB10D" blockId="1.[151,1437,1267,2014]" pageId="1" pageNumber="588">
<emphasis id="CAE27B6BFF83FFD0FF6B68F8FECEB3B9" bold="true" box="[199,345,1411,1438]" pageId="1" pageNumber="588">Description.</emphasis>
Female: carapace subelliptical (
<figureCitation id="60ADBBFCFF83FFD0FD1268F8FC9DB3B9" box="[702,778,1411,1438]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1206]" captionTargetBox="[340,1247,594,1152]" captionTargetId="figure@2.[340,1247,594,1152]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128). Colour in life, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3617045" httpUri="https://zenodo.org/record/3617045/files/figure.png" pageId="1" pageNumber="588">Figs. 1</figureCitation>
,
<figureCitation id="60ADBBFCFF83FFD0FCBB68FFFCAAB3B9" box="[791,829,1412,1438]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">2A</figureCitation>
), wider than long, regions poorly defined, antero-lateral margin bluntly rounded, front curved, posterior margin concave (
<figureCitation id="60ADBBFCFF83FFD0FC2468DCFC42B3E5" box="[904,981,1447,1474]" captionStart="FIGURE 1" captionStartId="2.[151,250,1181,1206]" captionTargetBox="[340,1247,594,1152]" captionTargetId="figure@2.[340,1247,594,1152]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128). Colour in life, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3617045" httpUri="https://zenodo.org/record/3617045/files/figure.png" pageId="1" pageNumber="588">Figs. 1</figureCitation>
,
<figureCitation id="60ADBBFCFF83FFD0FC4F68D3FB9EB3E5" box="[995,1033,1448,1474]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">2A</figureCitation>
). Eyes small, pigmented, not visible in dorsal view; orbits ovate. Antennules folded obliquely, antennular fossa slightly larger than ocular cavities. Antennae with 6 articles; basal article with large elliptical tubercle, fused with epistome, distal segment setose, scarcely overreaching orbito-antennular fossae (
<figureCitation id="60ADBBFCFF83FFD0FDF96B68FD27B009" box="[597,688,1555,1582]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">Fig. 2B</figureCitation>
). margin of first sternite of thoracic sternum emarginated. MXP3 (
<figureCitation id="60ADBBFCFF83FFD0FF0C6B4CFF64B075" box="[160,243,1591,1618]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">Fig. 2F</figureCitation>
) obliquely placed in buccal cavity, outer surface with scattered setae on mesial margin, inner margin with long scattered setae; propodus about 1.7 times as long as wide, stout, longer than carpus, anterior margin rounded with long setae; digitiform dactylus distally setose, inserted medially in notch on ventral margin of propodus, distally not overreaching tip of propodus but overreaching anteromesial angle of merus; ischiomerus completely fused, 1.7 times as long as wide, lateral margin convex with short scattered setae, mesial margin angular at the widest point forming antero mesial angle, inner margin almost straight with long slender setae. Exopod longer than half length of ischiomerus, flagellum with 2 articles, distally with fine setae (
<figureCitation id="60ADBBFCFF83FFD0FCDA6A74FC5CB10D" box="[886,971,1807,1834]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="F829A779FF83FFD0FF6B6A4FFBA9B1F9" blockId="1.[151,1437,1267,2014]" pageId="1" pageNumber="588">
P1 (cheliped) stout, merus slightly longer than carpus; chela (
<figureCitation id="60ADBBFCFF83FFD0FCC16A48FC75B169" box="[877,994,1843,1870]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">Fig. 2D, E</figureCitation>
) slightly globose, ventral margin gently convex, outer margin slightly curved, palm about 1.8 times as long as high; dactylus and pollex shorter than dorsal margin of palm, longer than greatest width of palm, palm relatively stout; dactylus cutting edge with one large tooth proximally, ending with sharp claw. Pollex inner surface with patch of long setae, outer surface smooth, cutting edge with 3 large proximal teeth separated by notch with denticulate margin (
<figureCitation id="60ADBBFCFF83FFD0FC136AB8FBA6B1F9" box="[959,1073,1987,2014]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="1" pageNumber="588">Fig. 2D,E</figureCitation>
).
</paragraph>
<paragraph id="F829A779FF80FFD3FF3B6DE3FE08B418" blockId="2.[151,1437,152,575]" pageId="2" pageNumber="589">
P2–P5 subequal (
<figureCitation id="60ADBBFCFF80FFD3FEF66DE2FE0CB694" box="[346,411,153,179]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="2" pageNumber="589">Fig. 3</figureCitation>
A–H), those on right side longer than left side. P2, P3 (
<figureCitation id="60ADBBFCFF80FFD3FBAB6DE2FBD0B694" box="[1031,1095,153,179]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="2" pageNumber="589">Fig. 3</figureCitation>
A–D) similar in shape; merus, carpus, propodus glabrous, propodus longer than carpus, slightly curved on both margins; dactylus conical, shorter than carpus and propodus, ventral margin with short fine setae, scattered setae on outer margin. P4 (
<figureCitation id="60ADBBFCFF80FFD3FAB16D9AFA03B6DC" box="[1309,1428,225,251]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="2" pageNumber="589">Fig. 3E, F</figureCitation>
) markedly asymmetrical, distinctly longer than P2, P3, P5; articles (except dactylus) similar in shape to those of P2, P3. Longer P4 (
<figureCitation id="60ADBBFCFF80FFD3FEE96C52FE0EB764" box="[325,409,297,323]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="2" pageNumber="589">Fig. 3E</figureCitation>
) more slender than P2, P3, carpus outer margin slightly curved; dactylus sickle-like, slightly shorter than propodus, ending in small curved tip, longer than dactyli of P2, P3, P5, ventral margin of P4 dactylus with long fine setae, few short setae on outer margin. P5 (
<figureCitation id="60ADBBFCFF80FFD3FCBF6C0AFC1DB7AC" box="[787,906,369,395]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="2" pageNumber="589">Fig. 3G, H</figureCitation>
) more slender than P2–P4; articles (except dactylus) similar in shape to those of P2, P3; dactylus longer, sickle-like, much longer than those of P2, P3 but shorter than that of P4; ventral margin of P5 dactylus with long fine setae, 2 rows of scattered setae present on outer margin, short setae present on dorsal side in distal half, dorsal and ventral margins of tip with short setae; distal part with 2 rows of short, 6–9 just visible very short spinules (
<figureCitation id="60ADBBFCFF80FFD3FD616F7AFC8CB43C" box="[717,795,513,539]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="2" pageNumber="589">
Fig.
<date id="8C2881B9FF80FFD3FCAD6F7AFC8CB43C" box="[769,795,513,539]" pageId="2" pageNumber="589">3I</date>
</figureCitation>
). Relative length of ambulatory legs, P4&gt;P3&gt;P5&gt;P2, and dactyli, P4&gt;P5&gt;P3&gt;P2.
</paragraph>
<caption id="ACE9F7F1FF80FFD3FF3B69E6FF59B2FD" ID-DOI="http://doi.org/10.5281/zenodo.3617045" ID-Zenodo-Dep="3617045" httpUri="https://zenodo.org/record/3617045/files/figure.png" pageId="2" pageNumber="589" startId="2.[151,250,1181,1206]" targetBox="[340,1247,594,1152]" targetPageId="2">
<paragraph id="F829A779FF80FFD3FF3B69E6FF59B2FD" blockId="2.[151,1436,1181,1242]" pageId="2" pageNumber="589">
<emphasis id="CAE27B6BFF80FFD3FF3B69E6FE83B291" bold="true" box="[151,276,1181,1206]" pageId="2" pageNumber="589">FIGURE 1.</emphasis>
<taxonomicName id="3F96DCFAFF80FFD3FEB069E6FD05B291" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[284,658,1181,1206]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF80FFD3FEB069E6FE66B292" box="[284,497,1181,1205]" italics="true" pageId="2" pageNumber="589">Arcotheres pernicola</emphasis>
(
<bibRefCitation id="9C07DA88FF80FFD3FDAC69E6FD1DB291" author="Burger, O." box="[512,650,1181,1206]" pageId="2" pageNumber="589" pagination="361 - 390" refId="ref3337" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128). Colour in life, dorsal view.
</paragraph>
</caption>
<paragraph id="F829A779FF80FFD3FF6B6873FC3AB361" blockId="2.[151,1437,1287,2034]" pageId="2" pageNumber="589">
Pleon wide (
<figureCitation id="60ADBBFCFF80FFD3FEF86873FE3AB305" box="[340,429,1288,1314]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="2" pageNumber="589">Fig. 2C</figureCitation>
), with 6 free somites and telson, covering bases of ambulatory legs; somites increasing in length distally, widening from somites 1–5, decreasing towards telson.
</paragraph>
</subSubSection>
<subSubSection id="B08CF4F2FF80FFD3FF6B682BFCDFB089" pageId="2" pageNumber="589" type="biology_ecology">
<paragraph id="F829A779FF80FFD3FF6B682BFCDFB089" blockId="2.[151,1437,1287,2034]" pageId="2" pageNumber="589">
<emphasis id="CAE27B6BFF80FFD3FF6B682BFE91B34D" bold="true" box="[199,262,1360,1386]" pageId="2" pageNumber="589">Host.</emphasis>
<bibRefCitation id="9C07DA88FF80FFD3FEBD682BFE67B34D" author="Burger, O." box="[273,496,1359,1386]" pageId="2" pageNumber="589" pagination="361 - 390" refId="ref3337" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger (1895: 376)</bibRefCitation>
recorded the species were from “
<taxonomicName id="3F96DCFAFF80FFD3FCD1682BFC56B34E" authorityName="Bruguiere" authorityYear="1789" box="[893,961,1360,1385]" class="Bivalvia" family="Mytilidae" genus="Perna" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FCD1682BFC56B34E" box="[893,961,1360,1385]" italics="true" pageId="2" pageNumber="589">Perna</emphasis>
</taxonomicName>
”.
<bibRefCitation id="9C07DA88FF80FFD3FC4C682BFB7DB34D" author="Schmitt, W. L. &amp; McCain, J. C. &amp; Davidson, E. S." box="[992,1258,1360,1386]" pageId="2" pageNumber="589" pagination="1 - 160" refId="ref4030" refString="Schmitt, W. L., McCain, J. C. &amp; Davidson, E. S. (1973) Decapoda. I. Brachyura. I. Fam. Pinnotheridae. In: Gruner, H. E. &amp; Holthuis, L. B. (Eds.), Crustaceorum Catalogus, 3, pp. 1 - 160." type="journal article" year="1973">Schmitt et al. (1973: 9</bibRefCitation>
, 66) noted that Bürger’s “
<taxonomicName id="3F96DCFAFF80FFD3FEA2680FFEC5B3AA" authorityName="Bruguiere" authorityYear="1789" box="[270,338,1396,1421]" class="Bivalvia" family="Mytilidae" genus="Perna" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FEA2680FFEC5B3AA" box="[270,338,1396,1421]" italics="true" pageId="2" pageNumber="589">Perna</emphasis>
</taxonomicName>
” is not the mytilid (Mytilida:
<taxonomicName id="3F96DCFAFF80FFD3FD1E680FFCB6B3A8" authorityName="Rafinesque" authorityYear="1815" box="[690,801,1396,1423]" class="Bivalvia" family="Mytilidae" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="family">Mytilidae</taxonomicName>
) but was actually
<taxonomicName id="3F96DCFAFF80FFD3FC59680FFAA7B3AA" authority="Lightfoot, 1786" authorityName="Lightfoot" authorityYear="1786" box="[1013,1328,1395,1422]" class="Bivalvia" family="Pteriidae" genus="Isognomon" kingdom="Animalia" order="Pterioida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FC59680FFBE6B3AA" box="[1013,1137,1396,1421]" italics="true" pageId="2" pageNumber="589">Isognomon</emphasis>
Lightfoot, 1786
</taxonomicName>
, a pteriid oyster (Ostreida:
<taxonomicName id="3F96DCFAFF80FFD3FEF568E3FE56B394" box="[345,449,1432,1459]" class="Bivalvia" family="Pteriidae" kingdom="Animalia" order="Pterioida" pageId="2" pageNumber="589" phylum="Mollusca" rank="family">Pteriidae</taxonomicName>
), instead. This was probably because at that time, there was confusion in the genus name (Bürger himself recorded other host mytilids as
<taxonomicName id="3F96DCFAFF80FFD3FD6D68C7FC82B3F1" authorityName="Linnaeus" authorityYear="1758" box="[705,789,1468,1494]" class="Bivalvia" family="Mytilidae" genus="Mytilus" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FD6D68C7FC82B3F1" box="[705,789,1468,1494]" italics="true" pageId="2" pageNumber="589">Mytilus</emphasis>
</taxonomicName>
); with Bürger likely to be referring to
<taxonomicName id="3F96DCFAFF80FFD3FB7B68C7FF45B3DD" authority="Bruguiere, 1789" authorityName="Bruguiere" authorityYear="1789" class="Bivalvia" family="Mytilidae" genus="Perna" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FB7B68C7FA8CB3F2" box="[1239,1307,1468,1493]" italics="true" pageId="2" pageNumber="589">Perna</emphasis>
Bruguière, 1789
</taxonomicName>
, which is now regarded as a junior synonym of
<taxonomicName id="3F96DCFAFF80FFD3FD5C689BFCFBB3DE" authorityName="Lightfoot" authorityYear="1786" box="[752,876,1504,1529]" class="Bivalvia" family="Pteriidae" genus="Isognomon" kingdom="Animalia" order="Pterioida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FD5C689BFCFBB3DE" box="[752,876,1504,1529]" italics="true" pageId="2" pageNumber="589">Isognomon</emphasis>
</taxonomicName>
. True
<taxonomicName id="3F96DCFAFF80FFD3FC1A689BFB56B3DD" authority="Philipsson, 1788" authorityName="Philipsson" authorityYear="1788" box="[950,1217,1504,1530]" class="Bivalvia" family="Mytilidae" genus="Perna" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FC1A689BFC6DB3DE" box="[950,1018,1504,1529]" italics="true" pageId="2" pageNumber="589">Perna</emphasis>
Philipsson, 1788
</taxonomicName>
, is a mytilid (Mytilida) while the modern understanding of
<taxonomicName id="3F96DCFAFF80FFD3FDF16B7FFCF0B039" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[605,871,1539,1566]" class="Bivalvia" family="Mytilidae" genus="Mytilus" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="CAE27B6BFF80FFD3FDF16B7FFD26B039" box="[605,689,1540,1566]" italics="true" pageId="2" pageNumber="589">Mytilus</emphasis>
Linnaeus, 1758
</taxonomicName>
, places it in the Ostreida (see
<bibRefCitation id="9C07DA88FF80FFD3FB1E6B7FFA06B039" author="Bouchet, P. &amp; Rocroi, J. - P. &amp; Bieler, R. &amp; Carter, J. G. &amp; Coan, E. V." box="[1202,1425,1540,1566]" pageId="2" pageNumber="589" pagination="1 - 184" refId="ref3271" refString="Bouchet, P., Rocroi, J. - P., Bieler, R., Carter, J. G. &amp; Coan, E. V. (2010) Nomenclator of bivalve families with a classification of bivalve families. Malacologia, 52 (2), 1 - 184. https: // doi. org / 10.4002 / 040.052.0201" type="journal article" year="2010">
Bouchet
<emphasis id="CAE27B6BFF80FFD3FABA6B7FFAD9B039" box="[1302,1358,1540,1566]" italics="true" pageId="2" pageNumber="589">et al.</emphasis>
2010
</bibRefCitation>
). As such, the species name of the pinnotherid, “
<taxonomicName id="3F96DCFAFF80FFD3FD0E6B53FC9AB065" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[674,781,1576,1602]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF80FFD3FD0E6B53FC9AB065" box="[674,781,1576,1602]" italics="true" pageId="2" pageNumber="589">pernicola</emphasis>
</taxonomicName>
” is rather unfortunate as it suggests it lives in
<taxonomicName id="3F96DCFAFF80FFD3FABC6B53FAC3B066" authority="Trivedi &amp; Gosavi &amp; Vachhrajani &amp; Ng, 2019" authorityName="Trivedi &amp; Gosavi &amp; Vachhrajani &amp; Ng" authorityYear="2019" box="[1296,1364,1576,1601]" class="Bivalvia" family="Mytilidae" genus="Perna" kingdom="Animalia" order="Mytiloida" pageId="2" pageNumber="589" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="CAE27B6BFF80FFD3FABC6B53FAC3B066" box="[1296,1364,1576,1601]" italics="true" pageId="2" pageNumber="589">Perna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1D1C610FF80FFD3FAF76B53FA0FB065" box="[1371,1432,1576,1602]" pageId="2" pageNumber="589" sensu="stricto">s. str.</taxonomicNameLabel>
, when it does not.
<bibRefCitation id="9C07DA88FF80FFD3FEF06B37FDBAB041" author="Nobili, G." box="[348,557,1611,1638]" pageId="2" pageNumber="589" pagination="230 - 282" refId="ref3941" refString="Nobili, G. (1899) Contribuzioni alla conoscienza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali dei Museo Civico di Storia Naturale di Genova, Series 2, 20, 230 - 282." type="journal article" year="1899">Nobili (1899: 264)</bibRefCitation>
recorded the species from Katau in
<collectingCountry id="8081E7E9FF80FFD3FC126B37FB01B041" box="[958,1174,1612,1638]" name="Papua New Guinea" pageId="2" pageNumber="589">Papua New Guinea</collectingCountry>
but did not indicate the host.
<bibRefCitation id="9C07DA88FF80FFD3FF796B0BFE4FB0AD" author="Nobili, G." box="[213,472,1647,1674]" pageId="2" pageNumber="589" pagination="393 - 411" refId="ref3981" refString="Nobili, G. (1906) Diagnoses preliminaires de 34 especes et varietes nouvelles, et de 2 genres nouveaux de Decapodes de la Mer Rouge. Bulletin du Museum national d'Histoire naturelle, Paris, 11 (6), 393 - 411. [1906]" type="journal article" year="1906">Nobili (1906:303–304)</bibRefCitation>
later recorded this species from oysters from
<collectingCountry id="8081E7E9FF80FFD3FC726B0BFBA8B0AD" box="[990,1087,1648,1674]" name="Djibouti" pageId="2" pageNumber="589">Djibouti</collectingCountry>
. The present record from
<collectingCountry id="8081E7E9FF80FFD3FACF6B0BFA0AB0AD" box="[1379,1437,1648,1674]" name="India" pageId="2" pageNumber="589">India</collectingCountry>
is from the oyster
<taxonomicName id="3F96DCFAFF80FFD3FECF6BEFFCD3B089" authority="(Schlotheim, 1820)" baseAuthorityName="Schlotheim" baseAuthorityYear="1820" box="[355,836,1684,1710]" class="Bivalvia" family="Ostreidae" genus="Magallana" higherTaxonomySource="CoL" kingdom="Plantae" order="Ostreida" pageId="2" pageNumber="589" phylum="Mollusca" rank="species" species="gryphoides">
<emphasis id="CAE27B6BFF80FFD3FECF6BEFFDF6B089" box="[355,609,1684,1710]" italics="true" pageId="2" pageNumber="589">Magallana gryphoides</emphasis>
(Schlotheim, 1820)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B08CF4F2FF80FFD2FF6B6BC3FB5CB418" lastPageId="3" lastPageNumber="590" pageId="2" pageNumber="589" type="discussion">
<paragraph id="F829A779FF80FFD2FF6B6BC3FDA6B788" blockId="2.[151,1437,1287,2034]" lastBlockId="3.[151,1437,152,575]" lastPageId="3" lastPageNumber="590" pageId="2" pageNumber="589">
<emphasis id="CAE27B6BFF80FFD3FF6B6BC3FEACB0F5" bold="true" box="[199,315,1720,1746]" pageId="2" pageNumber="589">Remarks.</emphasis>
<bibRefCitation id="9C07DA88FF80FFD3FEEE6BC3FD8CB0F5" author="Burger, O." box="[322,539,1719,1746]" pageId="2" pageNumber="589" pagination="361 - 390" refId="ref3337" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger (1895: 376)</bibRefCitation>
described
<taxonomicName id="3F96DCFAFF80FFD3FD386BC3FCE8B0F5" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[660,895,1720,1746]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF80FFD3FD386BC3FCE8B0F5" box="[660,895,1720,1746]" italics="true" pageId="2" pageNumber="589">Arcotheres pernicola</emphasis>
</taxonomicName>
from two ovigerous females measuring CW
<quantity id="3F6E0A9CFF80FFD3FAD56BCCFF52B0D1" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="589" unit="mm" value="7.5">7.5 mm</quantity>
, CL
<quantity id="3F6E0A9CFF80FFD3FF506BA0FEC4B0D1" box="[252,339,1755,1782]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="589" unit="mm" value="5.5">5.5 mm</quantity>
and CW
<quantity id="3F6E0A9CFF80FFD3FE126BA0FDB2B0D1" box="[446,549,1755,1782]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" pageId="2" pageNumber="589" unit="mm" value="8.75">8.75 mm</quantity>
, CL
<quantity id="3F6E0A9CFF80FFD3FDF06BA0FD22B0D1" box="[604,693,1755,1782]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="589" unit="mm" value="6.5">6.5 mm</quantity>
.
<bibRefCitation id="9C07DA88FF80FFD3FD136BA7FC7AB0D1" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[703,1005,1756,1782]" pageId="2" pageNumber="589" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng (2007: 202)</bibRefCitation>
found only one extant specimen measuring CW 6.0 mm, CL 5.0 mm and designated it as the
<typeStatus id="272D19DBFF80FFD3FCA16A7BFCE3B13D" box="[781,884,1792,1818]" pageId="2" pageNumber="589" type="lectotype">lectotype</typeStatus>
of the species. The discrepancy in measurements with the smaller specimen listed by Bürger and the
<typeStatus id="272D19DBFF80FFD3FD796A5FFCABB119" box="[725,828,1828,1854]" pageId="2" pageNumber="589" type="lectotype">lectotype</typeStatus>
can easily be explained by the soft and poor condition of the specimen. The present specimens differ from the redescription of
<taxonomicName id="3F96DCFAFF80FFD3FC1B6A33FBD6B145" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[951,1089,1864,1890]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF80FFD3FC1B6A33FBD6B145" box="[951,1089,1864,1890]" italics="true" pageId="2" pageNumber="589">A. pernicola</emphasis>
</taxonomicName>
given by
<bibRefCitation id="9C07DA88FF80FFD3FB006A33FA0BB145" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[1196,1436,1864,1890]" pageId="2" pageNumber="589" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng (2007)</bibRefCitation>
in several aspects: the frontal margin is distinctly convex (
<figureCitation id="60ADBBFCFF80FFD3FC846A17FC16B1A1" box="[808,897,1900,1926]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="2" pageNumber="589">Fig. 2A</figureCitation>
) (
<emphasis id="CAE27B6BFF80FFD3FC356A17FC76B1A2" box="[921,993,1900,1925]" italics="true" pageId="2" pageNumber="589">versus</emphasis>
curved gently; cf.
<bibRefCitation id="9C07DA88FF80FFD3FB186A17FA0FB1A1" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[1204,1432,1900,1926]" pageId="2" pageNumber="589" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10F); the MXP3 propodus not tapering distally and dactylus does not reach the propodal apex (
<figureCitation id="60ADBBFCFF80FFD3FB456AEBFAAAB18D" box="[1257,1341,1936,1962]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="2" pageNumber="589">Fig. 2F</figureCitation>
) (
<emphasis id="CAE27B6BFF80FFD3FAF96AEBFA0AB18E" box="[1365,1437,1936,1961]" italics="true" pageId="2" pageNumber="589">versus</emphasis>
propodus tapering distally and dactylus reaching propodal apex; cf.
<bibRefCitation id="9C07DA88FF80FFD3FC0E6ACFFB1CB1E9" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[930,1163,1972,1998]" pageId="2" pageNumber="589" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10G); the cheliped dactylus is slightly longer than half the length of the palm (
<figureCitation id="60ADBBFCFF80FFD3FC8F6AA3FC00B1D5" box="[803,919,2008,2034]" captionStart="FIGURE 2" captionStartId="3.[151,250,1963,1988]" captionTargetBox="[161,1424,604,1927]" captionTargetId="figure@3.[151,1436,597,1938]" captionTargetPageId="3" captionText="FIGURE 2 Arcotheres pernicola (Bürger, 1895), female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128).A, carapace; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." figureDoi="http://doi.org/10.5281/zenodo.3617049" httpUri="https://zenodo.org/record/3617049/files/figure.png" pageId="2" pageNumber="589">Fig. 2D, E</figureCitation>
) (
<emphasis id="CAE27B6BFF80FFD3FC036AA3FC60B1D6" box="[943,1015,2008,2033]" italics="true" pageId="2" pageNumber="589">versus</emphasis>
dactylus is half the length of the palm cf.
<bibRefCitation id="9C07DA88FF81FFD2FF156DE2FE0CB694" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[185,411,153,179]" pageId="3" pageNumber="590" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10A) and the longer P4 dactylus is 1.13 times as long as that on the longer P5 (
<figureCitation id="60ADBBFCFF81FFD2FAEE6DE2FF3BB6F0" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[193,1386,192,1904]" captionTargetId="figure@4.[186,1402,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres pernicola (Bürger, 1895), A–H, female (CW 6.2 mm, CL 4.8 mm) (ZL-AR-CR-128); I, female (CW 7.3 mm, CL 5.5 mm) (ZL-AR-CR-128).A, C, E, G, right P2–P5 dorsal view; B, D, F, H, left P2–P5 dorsal view; I, right P5 dactylus tip (drawn at an angle, setae not drawn)." figureDoi="http://doi.org/10.5281/zenodo.3617051" httpUri="https://zenodo.org/record/3617051/files/figure.png" pageId="3" pageNumber="590">Fig. 3E, G</figureCitation>
) (
<emphasis id="CAE27B6BFF81FFD2FF696DC5FE9AB6F0" box="[197,269,190,215]" italics="true" pageId="3" pageNumber="590">versus</emphasis>
dactylus of the longer P4 dactylus about twice as long as dactylus of longer P5; cf.
<bibRefCitation id="9C07DA88FF81FFD2FB196DC6FA0FB6F0" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[1205,1432,189,215]" pageId="3" pageNumber="590" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10E, F).The African specimen (NHM 955.6.22.55), although larger than the present series from
<collectingCountry id="8081E7E9FF81FFD2FB5D6D9AFABAB6DC" box="[1265,1325,225,251]" name="India" pageId="3" pageNumber="590">India</collectingCountry>
, agrees in all the key characters. It is hard to ascertain the significance of these differences because the
<typeStatus id="272D19DBFF81FFD2FB076C7EFA85B738" box="[1195,1298,261,287]" pageId="3" pageNumber="590" type="lectotype">lectotype</typeStatus>
female is in poor condition, and what is observed may merely be an artifact of preservation. As such, we prefer to treat the Indian Ocean material as
<taxonomicName id="3F96DCFAFF81FFD2FECA6C36FE67B740" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[358,496,333,359]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF81FFD2FECA6C36FE67B740" box="[358,496,333,359]" italics="true" pageId="3" pageNumber="590">A. pernicola</emphasis>
</taxonomicName>
for the time being. When fresh specimens of
<taxonomicName id="3F96DCFAFF81FFD2FC416C36FBE0B740" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1005,1143,333,359]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF81FFD2FC416C36FBE0B740" box="[1005,1143,333,359]" italics="true" pageId="3" pageNumber="590">A. pernicola</emphasis>
</taxonomicName>
become available from or near the type locality in the
<collectingCountry id="8081E7E9FF81FFD2FE606C0AFDDAB7AC" box="[460,589,369,395]" name="Philippines" pageId="3" pageNumber="590">Philippines</collectingCountry>
, the material from the Indian and Pacific Oceans must be compared again to determine if they are all conspecific.
</paragraph>
<paragraph id="F829A779FF81FFD2FF6B6CC2FB5CB418" blockId="3.[151,1437,152,575]" pageId="3" pageNumber="590">
Following
<bibRefCitation id="9C07DA88FF81FFD2FEED6CC2FDA3B7F4" author="Trivedi, J. N. &amp; Campos, E. &amp; Vachhrajani, K. D." box="[321,564,441,467]" pageId="3" pageNumber="590" pagination="195 - 200" refId="ref4243" refString="Trivedi, J. N., Campos, E. &amp; Vachhrajani, K. D. (2018 a) A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa, 4433 (1), 195 - 200. https: // doi. org / 10.11646 / zootaxa. 4433.1.13" type="journal article" year="2018">
Trivedi
<emphasis id="CAE27B6BFF81FFD2FE366CC1FE44B7F4" box="[410,467,441,467]" italics="true" pageId="3" pageNumber="590">et al.</emphasis>
(2018a)
</bibRefCitation>
,
<taxonomicName id="3F96DCFAFF81FFD2FDEC6CC2FCEDB7F4" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[576,890,440,467]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="CAE27B6BFF81FFD2FDEC6CC2FD5CB7F4" box="[576,715,441,467]" italics="true" pageId="3" pageNumber="590">A. pernicola</emphasis>
(
<bibRefCitation id="9C07DA88FF81FFD2FD776CC2FCE5B7F4" author="Burger, O." box="[731,882,440,467]" pageId="3" pageNumber="590" pagination="361 - 390" refId="ref3337" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
, belongs to a small subgroup of
<taxonomicName id="3F96DCFAFF81FFD2FB466CC2FAF4B7F4" authorityName="Manning" authorityYear="1993" box="[1258,1379,441,467]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="CAE27B6BFF81FFD2FB466CC2FAF4B7F4" box="[1258,1379,441,467]" italics="true" pageId="3" pageNumber="590">Arcotheres</emphasis>
</taxonomicName>
having the longer P4 dactylus longer than that of the P5. There are three other species in this subgroup:
<taxonomicName id="3F96DCFAFF81FFD2FB5F6CA6FEDEB43B" authority="(Gordon, 1936)" baseAuthorityName="Gordon" baseAuthorityYear="1936" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="CAE27B6BFF81FFD2FB5F6CA6FA0BB7D0" box="[1267,1436,477,503]" italics="true" pageId="3" pageNumber="590">A. winckworthi</emphasis>
(
<bibRefCitation id="9C07DA88FF81FFD2FF336F7AFED6B43B" author="Gordon, I." box="[159,321,513,540]" pageId="3" pageNumber="590" pagination="163 - 180" refId="ref3455" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article" year="1936">Gordon, 1936</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="3F96DCFAFF81FFD2FEFA6F7AFD00B43C" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[342,663,513,539]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="CAE27B6BFF81FFD2FEFA6F7AFE34B43C" box="[342,419,513,539]" italics="true" pageId="3" pageNumber="590">A. rayi</emphasis>
<bibRefCitation id="9C07DA88FF81FFD2FE066F7AFD00B43C" author="Ahyong, S. T. &amp; Ng, P. K. L." box="[426,663,513,539]" pageId="3" pageNumber="590" pagination="191 - 226" refId="ref3214" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng, 2007</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="3F96DCFAFF81FFD2FD7F6F7AFB70B43C" authority="Trivedi, Campos &amp; Vachhrajani, 2018" authorityName="Trivedi, Campos &amp; Vachhrajani" authorityYear="2018" box="[723,1255,513,539]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="shahi">
<emphasis id="CAE27B6BFF81FFD2FD7F6F7AFCA6B43C" box="[723,817,513,539]" italics="true" pageId="3" pageNumber="590">A. shahi</emphasis>
<bibRefCitation id="9C07DA88FF81FFD2FC946F7AFB70B43C" author="Trivedi, J. N. &amp; Trivedi, D. J. &amp; Vachhrajani, K. D. &amp; Ng, P. K. L." box="[824,1255,513,539]" pageId="3" pageNumber="590" pagination="1 - 83" refId="ref4386" refString="Trivedi, J. N., Trivedi, D. J., Vachhrajani, K. D. &amp; Ng, P. K. L. (2018 c) An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502 (1), 1 - 83. https: // doi. org / 10.11646 / zootaxa. 4502.1.1" type="journal article" year="2018">Trivedi, Campos &amp; Vachhrajani, 2018</bibRefCitation>
</taxonomicName>
(Campos 2001;
<bibRefCitation id="9C07DA88FF81FFD2FF3B6F5EFEEFB418" author="Trivedi, J. N. &amp; Campos, E. &amp; Vachhrajani, K. D." box="[151,376,549,575]" pageId="3" pageNumber="590" pagination="195 - 200" refId="ref4243" refString="Trivedi, J. N., Campos, E. &amp; Vachhrajani, K. D. (2018 a) A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa, 4433 (1), 195 - 200. https: // doi. org / 10.11646 / zootaxa. 4433.1.13" type="journal article" year="2018">
Trivedi
<emphasis id="CAE27B6BFF81FFD2FF5C6F5DFEBFB418" box="[240,296,549,575]" italics="true" pageId="3" pageNumber="590">et al.</emphasis>
2018a
</bibRefCitation>
). The morphological differences between these species are given in
<tableCitation id="B51492C2FF81FFD2FBDD6F5EFB52B418" box="[1137,1221,549,575]" captionStart="TABLE 1" captionStartId="5.[151,244,152,179]" captionTargetBox="[163,1415,230,940]" captionText="TABLE 1. Morphological comparsion between female specimens of A. pernicola (Bürger, 1895), A. winckworthi (Gordon, 1936), A. rayi Ahyong &amp; Ng, 2007, and A. shahi Trivedi, Campos &amp; Vachhrajani, 2018." httpUri="http://table.plazi.org/id/ACE9F7F1FF87FFD4FF3B6DE3FB4AB6F0" pageId="3" pageNumber="590" tableUuid="ACE9F7F1FF87FFD4FF3B6DE3FB4AB6F0">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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