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<document id="A9F42C2B6C8D81F6C243DE7636A8BA33" ID-DOI="10.5281/zenodo.183541" ID-GBIF-Dataset="7930bb68-74ba-477a-83a8-cde9592287be" ID-ISSN="1175-5326" ID-Zenodo-Dep="183541" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459922778768" checkinUser="plazi" docAuthor="Pilato, Giovanni, Fontoura, Paulo, Lisi, Oscar &amp; Beasley, Clark" docDate="2008" docId="F57F6B0EFF93FFC222E1FCCFFA98612E" docLanguage="en" docName="zt01856p054.pdf" docOrigin="Zootaxa 1856" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Echiniscus scabrospinosus Fontoura 1982" docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="09461376FF92FFCA2276FFF2FF966054" masterDocTitle="New description of Echiniscus scabrospinosus Fontoura, 1982, and description of a new species of Echiniscus (Heterotardigrada) from China" masterLastPageNumber="54" masterPageNumber="41" pageNumber="42" updateTime="1698229302531" updateUser="plazi">
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<mods:title id="09C655AA12CAC1212FAA7A74C7FF8B53">New description of Echiniscus scabrospinosus Fontoura, 1982, and description of a new species of Echiniscus (Heterotardigrada) from China</mods:title>
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<mods:namePart id="9001FC9B8D54CE6542BF887A90CAA22A">Pilato, Giovanni</mods:namePart>
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<mods:namePart id="6C2285F7D1E1663D57BB8021AB877504">Lisi, Oscar</mods:namePart>
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<mods:namePart id="0DF10BCB8EBF234ADE8BD6A526241F30">Beasley, Clark</mods:namePart>
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<treatment id="F57F6B0EFF93FFC222E1FCCFFA98612E" ID-DOI="http://doi.org/10.5281/zenodo.6228933" ID-GBIF-Taxon="119373908" ID-Zenodo-Dep="6228933" LSID="urn:lsid:plazi:treatment:F57F6B0EFF93FFC222E1FCCFFA98612E" httpUri="http://treatment.plazi.org/id/F57F6B0EFF93FFC222E1FCCFFA98612E" lastPageId="8" lastPageNumber="48" pageId="1" pageNumber="42">
<subSubSection id="35CC8993FF93FFCB22E1FCCFFE92632E" pageId="1" pageNumber="42" type="nomenclature">
<paragraph id="7D69DA18FF93FFCB22E1FCCFFD376303" blockId="1.[151,673,829,890]" box="[151,673,829,855]" pageId="1" pageNumber="42">
<heading id="26216D74FF93FFCB22E1FCCFFD376303" bold="true" box="[151,673,829,855]" fontSize="11" level="1" pageId="1" pageNumber="42" reason="1">
<taxonomicName id="BAD6A19BFF93FFCB22E1FCCFFD376303" ID-CoL="38G3C" authority="Fontoura, 1982" authorityName="Fontoura" authorityYear="1982" box="[151,673,829,855]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="42" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF93FFCB22E1FCCFFD376303" bold="true" box="[151,673,829,855]" pageId="1" pageNumber="42">
<emphasis id="4FA2060AFF93FFCB22E1FCCFFE4A6303" bold="true" box="[151,476,829,855]" italics="true" pageId="1" pageNumber="42">Echiniscus scabrospinosus</emphasis>
<bibRefCitation id="1947A7E9FF93FFCB2395FCCFFD376303" author="Fontoura" box="[483,673,829,855]" pageId="1" pageNumber="54" refString="Fontoura, P. (1982) Deux nouvelles especes de Tardigrades muscicoles du Portugal. Publicacoes do Instituto de Zoologia Dr. Augusto Nobre, 165, 4 - 19." type="journal article" year="1982">Fontoura, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="7D69DA18FF93FFCB22E1FC92FE92632E" blockId="1.[151,673,829,890]" box="[151,260,864,890]" pageId="1" pageNumber="42">
<figureCitation id="E5EDC69DFF93FFCB22E1FC92FF71632E" box="[151,231,864,890]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Figs. 1</figureCitation>
, 
<figureCitation id="E5EDC69DFF93FFCB2283FC92FE92632E" box="[245,260,864,890]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="1" pageNumber="42">2</figureCitation>
</paragraph>
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<subSubSection id="35CC8993FF93FFCE22E1FC42FEB5673A" lastPageId="4" lastPageNumber="45" pageId="1" pageNumber="42" type="materials_examined">
<paragraph id="7D69DA18FF93FFCB22E1FC42FD46643E" blockId="1.[151,1437,944,1970]" pageId="1" pageNumber="42">
<emphasis id="4FA2060AFF93FFCB22E1FC42FE18639E" bold="true" box="[151,398,944,970]" pageId="1" pageNumber="42">Material examined:</emphasis>
<collectingCountry id="05C19A88FF93FFCB23E1FC42FE6C639E" box="[407,506,944,970]" name="Portugal" pageId="1" pageNumber="42">Portugal</collectingCountry>
<specimenCount id="6BD01191FF93FFCB2075FC42FD3E639E" box="[515,680,944,970]" pageId="1" pageNumber="42" type="generic">46 specimens</specimenCount>
: one from Serra do Buçaco (
<typeStatus id="A26D64BAFF93FFCB218BFC42FBE6639E" box="[1021,1136,944,970]" pageId="1" pageNumber="42" type="holotype">Holotype</typeStatus>
), and 45 from Amarante; 
<collectingCountry id="05C19A88FF93FFCB22E1FC2AFE9363A6" box="[151,261,984,1010]" name="Tanzania" pageId="1" pageNumber="42">Tanzania</collectingCountry>
: one specimen; Azores (Faial Island: Miradouro da Praia Norte): one specimen. Specimens collected from mosses and lichens. We do not consider here the specimens from 
<collectingCountry id="05C19A88FF93FFCB2197FBF2FBB1644E" box="[993,1063,1024,1050]" name="China" pageId="1" pageNumber="42">China</collectingCountry>
previously attributed to 
<taxonomicName id="BAD6A19BFF93FFCB273CFBF2FEB26415" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="42" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF93FFCB273CFBF2FEB26415" italics="true" pageId="1" pageNumber="42">E. scabrospinosus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="1947A7E9FF93FFCB2324FBDAFD2E6416" author="Beasley" box="[338,696,1064,1090]" pageId="1" pageNumber="54" refString="Beasley, C. W. &amp; Cleveland, A. (1996) Tardigrades of southern Yunnan Province, People's Republic of China. Zoological Journal of the Linnean Society, 116, 239 - 243." type="journal article" year="1996">Beasley and Cleveland (1996)</bibRefCitation>
due to the fact that they do not belong to this species; neither the specimens recently reported by 
<bibRefCitation id="1947A7E9FF93FFCB2040FBA2FD5B643E" author="Yang" box="[566,717,1104,1130]" pageId="1" pageNumber="54" refString="Yang, T. (2008) Tardigrades from some mosses of Wuyi mountain sceneries in east China. Acta Zootaxonomica Sinica. 33, 2, 348 - 353." type="journal article" year="2008">Yang (2008)</bibRefCitation>
.
</paragraph>
<paragraph id="7D69DA18FF93FFCB22B3FB8AFE176566" blockId="1.[151,1437,944,1970]" pageId="1" pageNumber="42">
<emphasis id="4FA2060AFF93FFCB22B3FB8AFE0364C6" bold="true" box="[197,405,1144,1170]" pageId="1" pageNumber="42">
<typeStatus id="A26D64BAFF93FFCB22B3FB8AFE9464C6" box="[197,258,1144,1170]" pageId="1" pageNumber="42">Type</typeStatus>
repository:
</emphasis>
The 
<typeStatus id="A26D64BAFF93FFCB23A3FB8AFDA864C6" box="[469,574,1144,1170]" pageId="1" pageNumber="42" type="holotype">holotype</typeStatus>
, the African specimen, the specimen from the Azores and 3 other specimens from 
<collectingCountry id="05C19A88FF93FFCB236DFB52FEE964EE" box="[283,383,1184,1210]" name="Portugal" pageId="1" pageNumber="42">Portugal</collectingCountry>
are deposited in the collection of Binda and Pilato (Museum of the Department of Animal Biology “Marcello La Greca”, University of Catania); the remaining 
<specimenCount id="6BD01191FF93FFCB21BAFB3AFBFD64B6" box="[972,1131,1224,1250]" pageId="1" pageNumber="42" type="generic">42 specimens</specimenCount>
from 
<collectingCountry id="05C19A88FF93FFCB26C0FB3AFA8F64B6" box="[1206,1305,1224,1250]" name="Portugal" pageId="1" pageNumber="42">Portugal</collectingCountry>
are deposited in the collection of P. Fontoura at the Department of Zoology and Anthropology, Faculty of Sciences, University of Porto.
</paragraph>
<paragraph id="7D69DA18FF93FFCB22B3FACDFE4F65D6" blockId="1.[151,1437,944,1970]" pageId="1" pageNumber="42">
<emphasis id="4FA2060AFF93FFCB22B3FACDFE0F650D" bold="true" box="[197,409,1343,1369]" pageId="1" pageNumber="42">New description:</emphasis>
The new description is based essentially on the 
<typeStatus id="A26D64BAFF93FFCB21BCFAB2FBA5650E" box="[970,1075,1344,1370]" pageId="1" pageNumber="42" type="holotype">holotype</typeStatus>
, but some details are more visible in the other specimens.
</paragraph>
<paragraph id="7D69DA18FF93FFCB22B3FA62FAB76746" blockId="1.[151,1437,944,1970]" pageId="1" pageNumber="42">
Body length 218 µm long. Color reddish; eye spots red (visible in living specimens only). Median plate 3 absent but the corresponding area is sculptured. Cephalic plate with both anterior and posterior margin notched (
<figureCitation id="E5EDC69DFF93FFCB2372FA12FEF665AE" box="[260,352,1504,1530]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Fig. 1C</figureCitation>
); neck plate narrow (
<figureCitation id="E5EDC69DFF93FFCB202CFA12FD2365AE" box="[602,693,1504,1530]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Fig. 1C</figureCitation>
); scapular plate, 42.8 µm long, with a wide central portion and two small lateral portions with different ornamentation (
<figureCitation id="E5EDC69DFF93FFCB2153F9FAFC3A6676" box="[805,940,1544,1570]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Figs. 1A, B</figureCitation>
and 
<figureCitation id="E5EDC69DFF93FFCB2192F9FAFB9F6676" box="[996,1033,1544,1570]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="1" pageNumber="42">2B</figureCitation>
); two small thickened areas (intersegmental lateral plates) are present laterally to the median plate 1, to the median plate 2 and between the paired plate III and the terminal plate (
<figureCitation id="E5EDC69DFF93FFCB201EF9AAFD536626" box="[616,709,1624,1650]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Fig. 1C</figureCitation>
); terminal plate not faceted and with two posterolateral incisions. Median plates 1 and 2 divided into an anterior portion with only the anterior margin sculptured and a sculptured posterior portion (
<figureCitation id="E5EDC69DFF93FFCB239CF95AFDFF6696" box="[490,617,1704,1730]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Fig. 1A, C</figureCitation>
). Paired plates II and III with a wide medial portion and a small lateral part with different ornamentation (
<figureCitation id="E5EDC69DFF93FFCB205DF922FD1966BE" box="[555,655,1744,1770]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Figs. 1B</figureCitation>
and 
<figureCitation id="E5EDC69DFF93FFCB20BEF922FD7B66BE" box="[712,749,1744,1770]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="1" pageNumber="42">2B</figureCitation>
); the medial portion has an anterior part with two alternate sculptured and two smooth transversal bands and a larger sculptured posterior portion (
<figureCitation id="E5EDC69DFF93FFCB26E5F90AFA826746" box="[1171,1300,1784,1810]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Fig. 1A–C</figureCitation>
).
</paragraph>
<paragraph id="7D69DA18FF93FFCB22B3F8D2FB3467E6" blockId="1.[151,1437,944,1970]" pageId="1" pageNumber="42">
Neck plate, intersegmental lateral plates, lateral portions of scapular plate and of paired plates II and III with many very small dots and, at the same optical focus, few small, not well visible, light spots; other plates, or portions of plates, without small dots but with only large circular or elliptical light spots (
<figureCitation id="E5EDC69DFF93FFCB26ADF882FABB67DE" box="[1243,1325,1904,1930]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Figs. 1</figureCitation>
and 
<figureCitation id="E5EDC69DFF93FFCB271EF882FA1B67DE" box="[1384,1421,1904,1930]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="1" pageNumber="42">2B</figureCitation>
); each light spot with a central dark ring from which some dark stripes irradiate (
<figureCitation id="E5EDC69DFF93FFCB2641F86AFB0267E6" box="[1079,1172,1944,1970]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="1" pageNumber="42">Fig. 1D</figureCitation>
).
</paragraph>
<caption id="29A98A90FF90FFC822E1FAEBFE4565EF" httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="2" pageNumber="43" targetBox="[189,1390,201,1279]" targetPageId="2">
<paragraph id="7D69DA18FF90FFC822E1FAEBFE4565EF" blockId="2.[151,1436,1305,1467]" pageId="2" pageNumber="43">
<emphasis id="4FA2060AFF90FFC822E1FAEBFE8C6565" bold="true" box="[151,282,1305,1329]" pageId="2" pageNumber="43">FIGURE 1.</emphasis>
A–D, 
<emphasis id="4FA2060AFF90FFC82312FAE8FD1C6565" box="[356,650,1306,1329]" italics="true" pageId="2" pageNumber="43">
<taxonomicName id="BAD6A19BFF90FFC82312FAE8FD106565" box="[356,646,1306,1329]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="43" phylum="Tardigrada" rank="species" species="scabrospinosus">Echiniscus scabrospinosus</taxonomicName>
.
</emphasis>
A, B, 
<emphasis id="4FA2060AFF90FFC820A4FAE8FCB66565" box="[722,800,1306,1329]" italics="true" pageId="2" pageNumber="43">habitus</emphasis>
(A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm.
</paragraph>
</caption>
<paragraph id="7D69DA18FF90FFC822B3FA1DFC2C6690" blockId="2.[151,1437,1519,2036]" pageId="2" pageNumber="43">
Largest light spots, with a diameter of 2.1 µm, on the scapular, median plates and on the posterior portion of the paired plates II and III. The terminal plate has light spots slightly smaller than those of the scapular plate, of the paired plates and of the median plates 1 and 2 (
<figureCitation id="E5EDC69DFF90FFC82138F9C8FC5B6600" box="[846,973,1594,1620]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="2" pageNumber="43">Fig. 1A, B</figureCitation>
). In some specimens, the terminal plate has three longitudinal and three clearly visible transversal smooth bands appearing as dark stripes (
<figureCitation id="E5EDC69DFF90FFC82740F9ADFA05662D" box="[1334,1427,1631,1657]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="2" pageNumber="43">Fig. 1A</figureCitation>
) but in some specimens these bands, or some of them, are poorly visible (
<figureCitation id="E5EDC69DFF90FFC82189F976FBCD66CA" box="[1023,1115,1668,1694]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="2" pageNumber="43">Fig. 1B</figureCitation>
). Cuticle of legs with very fine dots and without light spots. Ventral surface with very fine dots.
</paragraph>
<paragraph id="7D69DA18FF90FFC822B3F93DFDFF67A0" blockId="2.[151,1437,1519,2036]" pageId="2" pageNumber="43">
External and internal cephalic cirri 14.9 µm and 12.5 µm long respectively; cephalic papilla 6.5 µm long; clava 4.4 µm long (
<figureCitation id="E5EDC69DFF90FFC8230EF906FE40675A" box="[376,470,1780,1806]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="2" pageNumber="43">Fig. 2A</figureCitation>
); in addition to the lateral appendage A, lateral spines C, D and E and dorsal spines D d are present (
<figureCitation id="E5EDC69DFF90FFC8233DF8EFFE3F6763" box="[331,425,1821,1847]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="2" pageNumber="43">Fig. 1A</figureCitation>
). The spines D d are more robust than the others, dagger–shaped with a round tip, and have margins not perfectly smooth. Filament A 33.2 µm long, spine C 11.0 µm, spine D 11.2 µm, spine E 15.6 µm, spine D d 21.8 µm. External and internal claws of the second pair of legs 10.1 µm and 10.9 µm long respectively; on the hind legs they are 12.0 µm and 13.2 µm long respectively. A small spine, 1.8 µm long, is present on the legs I (
<figureCitation id="E5EDC69DFF90FFC823E0F847FD81679B" box="[406,535,1973,1999]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="2" pageNumber="43">Fig. 2A, B</figureCitation>
); leg IV with a dentate fringe with 9 or 10 sharp triangular teeth of different sizes (
<figureCitation id="E5EDC69DFF90FFC82296F828FEAD67A0" box="[224,315,2010,2036]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="2" pageNumber="43">Fig. 1A</figureCitation>
) and a papilla 4 µm long.
</paragraph>
<paragraph id="7D69DA18FF91FFC922B3FF6AFB6A608E" blockId="3.[151,1436,152,467]" pageId="3" pageNumber="44">
External claws smooth; internal claws with a long and thin spur oriented towards the claw base (
<figureCitation id="E5EDC69DFF91FFC92741FF6AFA0560E6" box="[1335,1427,152,178]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="3" pageNumber="44">Fig. 2D</figureCitation>
) (spur 3.3 µm long, inserted at 3.6 µm from the claw base in the largest specimen, 240 µm long).
</paragraph>
<paragraph id="7D69DA18FF91FFC922B3FF1AFB1B6187" blockId="3.[151,1436,152,467]" pageId="3" pageNumber="44">
The other specimens are similar to the 
<typeStatus id="A26D64BAFF91FFC920FFFF1AFD666156" box="[649,752,232,258]" pageId="3" pageNumber="44" type="holotype">holotype</typeStatus>
in both qualitative and metric characters (
<tableCitation id="3054EFA3FF91FFC926ACFF1AFAA46156" box="[1242,1330,232,258]" captionStart="TABLE 1" captionStartId="3.[151,239,520,544]" captionTargetBox="[159,1423,688,1071]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of E. scabrospinosus (% bo = percentage of body length; % sc = percentage of scapular plate length; when appendage asymmetry occurred the two measures are reported; a = absent)." httpUri="http://table.plazi.org/id/29A98A90FF91FFC922E1FDFAFCD26231" pageId="3" pageNumber="44" tableUuid="29A98A90FF91FFC922E1FDFAFCD26231">Table 1</tableCitation>
); but the density of light spots (12–15 light spots / 100 µm 
<quantity id="BA2E77FDFF91FFC920AFFEE2FD6A6178" box="[729,764,272,300]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="44" unit="in" value="2.0">2 in</quantity>
some specimens, up to19–21 (or more, 26 was the maximum observed) light spots / 100 µm 
<quantity id="BA2E77FDFF91FFC92036FEC9FDF56103" box="[576,611,315,343]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="44" unit="in" value="2.0">2 in</quantity>
others) (
<tableCitation id="3054EFA3FF91FFC920B1FECFFCB66103" box="[711,800,317,343]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetBox="[159,1108,208,412]" captionTargetPageId="4" captionText="TABLE 2. Density of light spots in the scapular plate of E. scabrospinosus (estimated in 20 adult specimens)." httpUri="http://table.plazi.org/id/29A98A90FF96FFCE22E1FF65FABD60FB" pageId="3" pageNumber="44" tableUuid="29A98A90FF96FFCE22E1FF65FABD60FB">Table 2</tableCitation>
) and the appendages show a high degree of variability with asymmetries and very different lengths, and some of them (frequently D, very rarely D d) may be absent (
<tableCitation id="3054EFA3FF91FFC922E8FE63FF6E61FF" box="[158,248,401,427]" captionStart="TABLE 3" captionStartId="5.[151,239,151,175]" captionTargetBox="[159,1317,242,567]" captionTargetPageId="5" captionText="TABLE 3. Morphotypes recognizable in E. scabrospinosus as a consequence of the presence or absence of the lateral and dorsal appendages at least in one side of the body (43 adults and 1 larva)." httpUri="http://table.plazi.org/id/29A98A90FF97FFCF22E1FF65FC446085" pageId="3" pageNumber="44" tableUuid="29A98A90FF97FFCF22E1FF65FC446085">Table 3</tableCitation>
). The appendages also show a degree of variability in their shape, in fact some of them may have two or three tips. The number of teeth of the dentate fringe is comprised between 8 and 12.
</paragraph>
<caption id="29A98A90FF91FFC922E1FDFAFCD26231" ID-Table-UUID="29A98A90FF91FFC922E1FDFAFCD26231" httpUri="http://table.plazi.org/id/29A98A90FF91FFC922E1FDFAFCD26231" pageId="3" pageNumber="44" targetBox="[159,1423,688,1071]" targetIsTable="true" targetPageId="3">
<paragraph id="7D69DA18FF91FFC922E1FDFAFCD26231" blockId="3.[151,1436,520,613]" pageId="3" pageNumber="44">
<emphasis id="4FA2060AFF91FFC922E1FDFAFE9E6274" bold="true" box="[151,264,520,544]" pageId="3" pageNumber="44">TABLE 1.</emphasis>
Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of 
<taxonomicName id="BAD6A19BFF91FFC922E1FDD9FECD6216" box="[151,347,555,578]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="44" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF91FFC922E1FDD9FECD6216" box="[151,347,555,578]" italics="true" pageId="3" pageNumber="44">E. scabrospinosus</emphasis>
</taxonomicName>
(% bo = percentage of body length; % sc = percentage of scapular plate length; when appendage asymmetry occurred the two measures are reported; 
<emphasis id="4FA2060AFF91FFC920B3FDBCFD446231" box="[709,722,590,613]" italics="true" pageId="3" pageNumber="44">a</emphasis>
= absent).
</paragraph>
</caption>
<paragraph id="7D69DA18FF91FFCE231BFD74FEB5673A" blockId="3.[365,1267,646,669]" lastBlockId="4.[151,1436,1676,2023]" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
<typeStatus id="A26D64BAFF91FFC9231BFD74FE4662C9" box="[365,464,646,669]" pageId="3" pageNumber="44" type="holotype">Holotype</typeStatus>
smallest specimen largest specimen Papilla on legs IV 4.0 1.8 9.3 2.5 1.5 6.6 4.3 1.8 10.1 
<emphasis id="4FA2060AFF96FFCE22B3F97EFE4A66F2" bold="true" box="[197,476,1676,1702]" pageId="4" pageNumber="45">Differential diagnosis:</emphasis>
Considering the species of 
<taxonomicName id="BAD6A19BFF96FFCE215DF97FFC3C66F2" box="[811,938,1677,1702]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="genus">
<emphasis id="4FA2060AFF96FFCE215DF97FFC3C66F2" box="[811,938,1677,1702]" italics="true" pageId="4" pageNumber="45">Echiniscus</emphasis>
</taxonomicName>
having a similar plate ornamentation and the same lateral and dorsal appendages present in 
<taxonomicName id="BAD6A19BFF96FFCE209FF947FC28669A" box="[745,958,1717,1743]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF96FFCE209FF947FD6D669A" box="[745,763,1717,1742]" italics="true" pageId="4" pageNumber="45">E</emphasis>
. 
<emphasis id="4FA2060AFF96FFCE217CF947FC28669A" box="[778,958,1717,1742]" italics="true" pageId="4" pageNumber="45">scabrospinosus</emphasis>
</taxonomicName>
(also their individual variability considered), we think it opportune to compare to it: 
<taxonomicName id="BAD6A19BFF96FFCE20DCF92FFB1266A3" authority="Murray, 1907" authorityName="Murray" authorityYear="1907" box="[682,1156,1757,1783]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="crassispinosus">
<emphasis id="4FA2060AFF96FFCE20DCF92FFC4D66A2" box="[682,987,1757,1782]" italics="true" pageId="4" pageNumber="45">Echiniscus crassispinosus</emphasis>
<bibRefCitation id="1947A7E9FF96FFCE2194F92FFB1266A3" author="Murray" box="[994,1156,1757,1783]" pageId="4" pageNumber="54" refString="Murray, J. (1907) Some South African Tardigrada. Journal of the Royal Microscopical Society of London, V, 515 - 524." type="book chapter" year="1907">Murray, 1907</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="BAD6A19BFF96FFCE26E6F92FFA0A66A3" authority="Da" authorityName="Da" box="[1168,1436,1757,1783]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="canedoi">
<emphasis id="4FA2060AFF96FFCE26E6F92FFAE566A2" box="[1168,1395,1757,1782]" italics="true" pageId="4" pageNumber="45">Echiniscus canedoi</emphasis>
Da
</taxonomicName>
<collectingCountry id="05C19A88FF96FFCE22E1F8F7FF75674B" box="[151,227,1797,1823]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="45">Cunha</collectingCountry>
&amp; Do Nascimiento 
<bibRefCitation id="1947A7E9FF96FFCE23A3F8F7FDEC674B" author="Ribeiro" box="[469,634,1797,1823]" pageId="4" pageNumber="54" refString="Cunha, A. X. da &amp; Do Nascimento Ribeiro, F. (1962) A fauna de Tardigrados da Ilha da Madeira. Memorias e Etsudos de Museu Zoologico de Universidade de Coimbra, 179, 1 - 24." type="journal article" year="1962">Ribeiro, 1962</bibRefCitation>
; 
<taxonomicName id="BAD6A19BFF96FFCE20FEF8F7FC3A674B" authority="Da" authorityName="Da" box="[648,940,1797,1823]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="manuelae">
<emphasis id="4FA2060AFF96FFCE20FEF8F7FC17674A" box="[648,897,1797,1822]" italics="true" pageId="4" pageNumber="45">Echiniscus manuelae</emphasis>
Da
</taxonomicName>
<collectingCountry id="05C19A88FF96FFCE21C2F8F7FB96674B" box="[948,1024,1797,1823]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="45">Cunha</collectingCountry>
&amp; Do Nascimiento 
<bibRefCitation id="1947A7E9FF96FFCE2684F8F7FA01674B" author="Ribeiro" box="[1266,1431,1797,1823]" pageId="4" pageNumber="54" refString="Cunha, A. X. da &amp; Do Nascimento Ribeiro, F. (1962) A fauna de Tardigrados da Ilha da Madeira. Memorias e Etsudos de Museu Zoologico de Universidade de Coimbra, 179, 1 - 24." type="journal article" year="1962">Ribeiro, 1962</bibRefCitation>
, 
<taxonomicName id="BAD6A19BFF96FFCE22E1F8DFFE526713" authority="Da" authorityName="Da" box="[151,452,1837,1863]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="angolensis">
<emphasis id="4FA2060AFF96FFCE22E1F8DFFE0C6712" box="[151,410,1837,1862]" italics="true" pageId="4" pageNumber="45">Echiniscus angolensis</emphasis>
Da
</taxonomicName>
<collectingCountry id="05C19A88FF96FFCE23BAF8DFFD8E6713" box="[460,536,1837,1863]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="45">Cunha</collectingCountry>
&amp; Do Nascimiento 
<bibRefCitation id="1947A7E9FF96FFCE2173F8DFFC3F6713" author="Ribeiro" box="[773,937,1837,1863]" pageId="4" pageNumber="54" refString="Cunha, A. X. da &amp; Do Nascimento Ribeiro, F. (1964) Tardigrados de Angola. Garcia de Orta (Lisboa). Revista da Junta de Investigacoes do Ultramar, 12, 397 - 406." type="journal article" year="1964">Ribeiro, 1964</bibRefCitation>
; 
<taxonomicName id="BAD6A19BFF96FFCE21C0F8DFFBC46712" box="[950,1106,1837,1862]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF96FFCE21C0F8DFFBC46712" box="[950,1106,1837,1862]" italics="true" pageId="4" pageNumber="45">E. spinulosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="BAD6A19BFF96FFCE2617F8DFFB746712" box="[1121,1250,1837,1862]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF96FFCE2617F8DFFB746712" box="[1121,1250,1837,1862]" italics="true" pageId="4" pageNumber="45">E. spiniger</emphasis>
</taxonomicName>
and, overall, 
<emphasis id="4FA2060AFF96FFCE27F5F8DFFEB5673A" italics="true" pageId="4" pageNumber="45">
<taxonomicName id="BAD6A19BFF96FFCE27F5F8DFFE89673A" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="marginatus">E. marginatus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<paragraph id="7D69DA18FF91FFC922E9FD42FAC5647B" pageId="3" pageNumber="44">
<table id="0FD628B8FF91003522E9FD42FA19647B" box="[159,1423,688,1071]" gridcols="9" gridrows="7" pageId="3" pageNumber="44">
<tr id="C3E6D85AFF91003522E9FD42FA196293" box="[159,1423,688,711]" gridrow="0" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FD42FE0A6293" box="[159,412,688,711]" gridcol="0" gridrow="0" pageId="3" pageNumber="44">Character µm</th>
<th id="8037B126FF9100352396FD42FD876293" box="[480,529,688,711]" gridcol="1" gridrow="0" pageId="3" pageNumber="44">%bo</th>
<th id="8037B126FF9100352028FD42FD1B6293" box="[606,653,688,711]" gridcol="2" gridrow="0" pageId="3" pageNumber="44">%sc</th>
<th id="8037B126FF91003520ABFD42FC8F6293" box="[733,793,688,711]" gridcol="3" gridrow="0" pageId="3" pageNumber="44">µm</th>
<th id="8037B126FF9100352122FD42FC136293" box="[852,901,688,711]" gridcol="4" gridrow="0" pageId="3" pageNumber="44">%bo</th>
<th id="8037B126FF91003521BDFD42FC6C6293" box="[971,1018,688,711]" gridcol="5" gridrow="0" pageId="3" pageNumber="44">%sc</th>
<th id="8037B126FF9100352637FD42FBEB6293" box="[1089,1149,688,711]" gridcol="6" gridrow="0" pageId="3" pageNumber="44">µm</th>
<th id="8037B126FF91003526C5FD42FB726293" box="[1203,1252,688,711]" gridcol="7" gridrow="0" pageId="3" pageNumber="44">%bo</th>
<th id="8037B126FF9100352752FD42FA196293" box="[1316,1423,688,711]" gridcol="8" gridrow="0" pageId="3" pageNumber="44">%sc</th>
</tr>
<tr id="C3E6D85AFF91003522E9FD29FA1962A6" box="[159,1423,731,754]" gridrow="1" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FD29FE0A62A6" box="[159,412,731,754]" gridcol="0" gridrow="1" pageId="3" pageNumber="44">Body length 218</th>
<td id="8037B126FF9100352396FD29FD8762A6" box="[480,529,731,754]" gridcol="1" gridrow="1" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF9100352028FD29FD1B62A6" box="[606,653,731,754]" gridcol="2" gridrow="1" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF91003520ABFD29FC8F62A6" box="[733,793,731,754]" gridcol="3" gridrow="1" pageId="3" pageNumber="44">172.3</td>
<td id="8037B126FF9100352122FD29FC1362A6" box="[852,901,731,754]" gridcol="4" gridrow="1" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF91003521BDFD29FC6C62A6" box="[971,1018,731,754]" gridcol="5" gridrow="1" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF9100352637FD29FBEB62A6" box="[1089,1149,731,754]" gridcol="6" gridrow="1" pageId="3" pageNumber="44">240.2</td>
<td id="8037B126FF91003526C5FD29FB7262A6" box="[1203,1252,731,754]" gridcol="7" gridrow="1" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF9100352752FD29FA1962A6" box="[1316,1423,731,754]" gridcol="8" gridrow="1" pageId="3" pageNumber="44">-</td>
</tr>
<tr id="C3E6D85AFF91003522E9FCF4FA19636B" box="[159,1423,774,831]" gridrow="2" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FCF4FE0A636B" box="[159,412,774,831]" gridcol="0" gridrow="2" pageId="3" pageNumber="44">Scapular plate 42.8 length</th>
<td id="8037B126FF9100352396FCF4FD87636B" box="[480,529,774,831]" gridcol="1" gridrow="2" pageId="3" pageNumber="44">19.6</td>
<td id="8037B126FF9100352028FCF4FD1B636B" box="[606,653,774,831]" gridcol="2" gridrow="2" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF91003520ABFCF4FC8F636B" box="[733,793,774,831]" gridcol="3" gridrow="2" pageId="3" pageNumber="44">37.6</td>
<td id="8037B126FF9100352122FCF4FC13636B" box="[852,901,774,831]" gridcol="4" gridrow="2" pageId="3" pageNumber="44">21.8</td>
<td id="8037B126FF91003521BDFCF4FC6C636B" box="[971,1018,774,831]" gridcol="5" gridrow="2" pageId="3" pageNumber="44">-</td>
<td id="8037B126FF9100352637FCF4FBEB636B" box="[1089,1149,774,831]" gridcol="6" gridrow="2" pageId="3" pageNumber="44">42.6</td>
<td id="8037B126FF91003526C5FCF4FB72636B" box="[1203,1252,774,831]" gridcol="7" gridrow="2" pageId="3" pageNumber="44">17.7</td>
<td id="8037B126FF9100352752FCF4FA19636B" box="[1316,1423,774,831]" gridcol="8" gridrow="2" pageId="3" pageNumber="44">-</td>
</tr>
<tr id="C3E6D85AFF91003522E9FCA1FA1963D9" box="[159,1423,851,909]" gridrow="3" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FCA1FE0A63D9" box="[159,412,851,909]" gridcol="0" gridrow="3" pageId="3" pageNumber="44">Internal cephalic 12.5 cirrus</th>
<td id="8037B126FF9100352396FCA1FD8763D9" box="[480,529,851,909]" gridcol="1" gridrow="3" pageId="3" pageNumber="44">5.7</td>
<td id="8037B126FF9100352028FCA1FD1B63D9" box="[606,653,851,909]" gridcol="2" gridrow="3" pageId="3" pageNumber="44">29.2</td>
<td id="8037B126FF91003520ABFCA1FC8F63D9" box="[733,793,851,909]" gridcol="3" gridrow="3" pageId="3" pageNumber="44">11.7</td>
<td id="8037B126FF9100352122FCA1FC1363D9" box="[852,901,851,909]" gridcol="4" gridrow="3" pageId="3" pageNumber="44">6.8</td>
<td id="8037B126FF91003521BDFCA1FC6C63D9" box="[971,1018,851,909]" gridcol="5" gridrow="3" pageId="3" pageNumber="44">31.1</td>
<td id="8037B126FF9100352637FCA1FBEB63D9" box="[1089,1149,851,909]" gridcol="6" gridrow="3" pageId="3" pageNumber="44">12.9</td>
<td id="8037B126FF91003526C5FCA1FB7263D9" box="[1203,1252,851,909]" gridcol="7" gridrow="3" pageId="3" pageNumber="44">5.4</td>
<td id="8037B126FF9100352752FCA1FA1963D9" box="[1316,1423,851,909]" gridcol="8" gridrow="3" pageId="3" pageNumber="44">30.3</td>
</tr>
<tr id="C3E6D85AFF91003522E9FC52FA19638E" box="[159,1423,928,986]" gridrow="4" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FC52FE0A638E" box="[159,412,928,986]" gridcol="0" gridrow="4" pageId="3" pageNumber="44">External cepha- 14.9 lic cirrus</th>
<td id="8037B126FF9100352396FC52FD87638E" box="[480,529,928,986]" gridcol="1" gridrow="4" pageId="3" pageNumber="44">6.8</td>
<td id="8037B126FF9100352028FC52FD1B638E" box="[606,653,928,986]" gridcol="2" gridrow="4" pageId="3" pageNumber="44">34.8</td>
<td id="8037B126FF91003520ABFC52FC8F638E" box="[733,793,928,986]" gridcol="3" gridrow="4" pageId="3" pageNumber="44">14.3</td>
<td id="8037B126FF9100352122FC52FC13638E" box="[852,901,928,986]" gridcol="4" gridrow="4" pageId="3" pageNumber="44">8.3</td>
<td id="8037B126FF91003521BDFC52FC6C638E" box="[971,1018,928,986]" gridcol="5" gridrow="4" pageId="3" pageNumber="44">38.0</td>
<td id="8037B126FF9100352637FC52FBEB638E" box="[1089,1149,928,986]" gridcol="6" gridrow="4" pageId="3" pageNumber="44">15.9</td>
<td id="8037B126FF91003526C5FC52FB72638E" box="[1203,1252,928,986]" gridcol="7" gridrow="4" pageId="3" pageNumber="44">6.6</td>
<td id="8037B126FF9100352752FC52FA19638E" box="[1316,1423,928,986]" gridcol="8" gridrow="4" pageId="3" pageNumber="44">37.3</td>
</tr>
<tr id="C3E6D85AFF91003522E9FC1CFA196451" box="[159,1423,1006,1029]" gridrow="5" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FC1CFE0A6451" box="[159,412,1006,1029]" gridcol="0" gridrow="5" pageId="3" pageNumber="44">Cephalic papilla 6.5</th>
<td id="8037B126FF9100352396FC1CFD876451" box="[480,529,1006,1029]" gridcol="1" gridrow="5" pageId="3" pageNumber="44">3.0</td>
<td id="8037B126FF9100352028FC1CFD1B6451" box="[606,653,1006,1029]" gridcol="2" gridrow="5" pageId="3" pageNumber="44">15.2</td>
<td id="8037B126FF91003520ABFC1CFC8F6451" box="[733,793,1006,1029]" gridcol="3" gridrow="5" pageId="3" pageNumber="44">6.5</td>
<td id="8037B126FF9100352122FC1CFC136451" box="[852,901,1006,1029]" gridcol="4" gridrow="5" pageId="3" pageNumber="44">3.8</td>
<td id="8037B126FF91003521BDFC1CFC6C6451" box="[971,1018,1006,1029]" gridcol="5" gridrow="5" pageId="3" pageNumber="44">17.3</td>
<td id="8037B126FF9100352637FC1CFBEB6451" box="[1089,1149,1006,1029]" gridcol="6" gridrow="5" pageId="3" pageNumber="44">6.2</td>
<td id="8037B126FF91003526C5FC1CFB726451" box="[1203,1252,1006,1029]" gridcol="7" gridrow="5" pageId="3" pageNumber="44">2.6</td>
<td id="8037B126FF9100352752FC1CFA196451" box="[1316,1423,1006,1029]" gridcol="8" gridrow="5" pageId="3" pageNumber="44">14.6</td>
</tr>
<tr id="C3E6D85AFF91003522E9FBEAFA19647B" box="[159,1423,1048,1071]" gridrow="6" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FBEAFE0A647B" box="[159,412,1048,1071]" gridcol="0" gridrow="6" pageId="3" pageNumber="44">Clava 4.4</th>
<td id="8037B126FF9100352396FBEAFD87647B" box="[480,529,1048,1071]" gridcol="1" gridrow="6" pageId="3" pageNumber="44">2.0</td>
<td id="8037B126FF9100352028FBEAFD1B647B" box="[606,653,1048,1071]" gridcol="2" gridrow="6" pageId="3" pageNumber="44">10.3</td>
<td id="8037B126FF91003520ABFBEAFC8F647B" box="[733,793,1048,1071]" gridcol="3" gridrow="6" pageId="3" pageNumber="44">5.1</td>
<td id="8037B126FF9100352122FBEAFC13647B" box="[852,901,1048,1071]" gridcol="4" gridrow="6" pageId="3" pageNumber="44">3.0</td>
<td id="8037B126FF91003521BDFBEAFC6C647B" box="[971,1018,1048,1071]" gridcol="5" gridrow="6" pageId="3" pageNumber="44">13.6</td>
<td id="8037B126FF9100352637FBEAFBEB647B" box="[1089,1149,1048,1071]" gridcol="6" gridrow="6" pageId="3" pageNumber="44">4.6</td>
<td id="8037B126FF91003526C5FBEAFB72647B" box="[1203,1252,1048,1071]" gridcol="7" gridrow="6" pageId="3" pageNumber="44">1.9</td>
<td id="8037B126FF9100352752FBEAFA19647B" box="[1316,1423,1048,1071]" gridcol="8" gridrow="6" pageId="3" pageNumber="44">10.8</td>
</tr>
</table>
</paragraph>
<paragraph id="7D69DA18FF91FFC922E9FBB1FAC56771" pageId="3" pageNumber="44">
<table id="0FD628B8FF91003522E9FBB1FA196713" box="[159,1423,1091,1863]" gridcols="10" gridrows="12" pageId="3" pageNumber="44">
<tr id="C3E6D85AFF91003522E9FBB1FA196429" box="[159,1423,1091,1149]" gridrow="0" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FBB1FECA6429" box="[159,348,1091,1149]" gridcol="0" gridrow="0" pageId="3" pageNumber="44">Lateral append- age A</th>
<th id="8037B126FF910035231BFBB1FE586429" box="[365,462,1091,1149]" gridcol="1" gridrow="0" pageId="3" pageNumber="44">33.2–37.4</th>
<th id="8037B126FF9100352396FBB1FDD66429" box="[480,576,1091,1149]" gridcol="2" gridrow="0" pageId="3" pageNumber="44">15.2–17.2</th>
<th id="8037B126FF9100352028FBB1FD286429" box="[606,702,1091,1149]" gridcol="3" gridrow="0" pageId="3" pageNumber="44">77.6–87.4</th>
<th id="8037B126FF91003520ABFBB1FCAB6429" box="[733,829,1091,1149]" gridcol="4" gridrow="0" pageId="3" pageNumber="44">35.0</th>
<th id="8037B126FF9100352122FBB1FC316429" box="[852,935,1091,1149]" gridcol="5" gridrow="0" pageId="3" pageNumber="44">20.3</th>
<th id="8037B126FF91003521BCFBB1FBBD6429" box="[970,1067,1091,1149]" gridcol="6" gridrow="0" pageId="3" pageNumber="44">93.1</th>
<th id="8037B126FF9100352637FBB1FB376429" box="[1089,1185,1091,1149]" gridcol="7" gridrow="0" pageId="3" pageNumber="44">43.6</th>
<th id="8037B126FF91003526C5FBB1FA956429" box="[1203,1283,1091,1149]" gridcol="8" gridrow="0" pageId="3" pageNumber="44">18.2</th>
<th id="8037B126FF9100352752FBB1FA196429" box="[1316,1423,1091,1149]" gridcol="9" gridrow="0" pageId="3" pageNumber="44">103.1</th>
</tr>
<tr id="C3E6D85AFF91003522E9FB62FA19649E" box="[159,1423,1168,1226]" gridrow="1" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FB62FECA649E" box="[159,348,1168,1226]" gridcol="0" gridrow="1" pageId="3" pageNumber="44">Lateral append- age C</th>
<td id="8037B126FF910035231BFB62FE58649E" box="[365,462,1168,1226]" gridcol="1" gridrow="1" pageId="3" pageNumber="44">11.0</td>
<td id="8037B126FF9100352396FB62FDD6649E" box="[480,576,1168,1226]" gridcol="2" gridrow="1" pageId="3" pageNumber="44">5.1</td>
<td id="8037B126FF9100352028FB62FD28649E" box="[606,702,1168,1226]" gridcol="3" gridrow="1" pageId="3" pageNumber="44">25.7</td>
<td id="8037B126FF91003520ABFB62FCAB649E" box="[733,829,1168,1226]" gridcol="4" gridrow="1" pageId="3" pageNumber="44">a–5.2</td>
<td id="8037B126FF9100352122FB62FC31649E" box="[852,935,1168,1226]" gridcol="5" gridrow="1" pageId="3" pageNumber="44">a–3.0</td>
<td id="8037B126FF91003521BCFB62FBBD649E" box="[970,1067,1168,1226]" gridcol="6" gridrow="1" pageId="3" pageNumber="44">a–13.8</td>
<td id="8037B126FF9100352637FB62FB37649E" box="[1089,1185,1168,1226]" gridcol="7" gridrow="1" pageId="3" pageNumber="44">6.7–10.0</td>
<td id="8037B126FF91003526C5FB62FA95649E" box="[1203,1283,1168,1226]" gridcol="8" gridrow="1" pageId="3" pageNumber="44">2.8–4.2</td>
<td id="8037B126FF9100352752FB62FA19649E" box="[1316,1423,1168,1226]" gridcol="9" gridrow="1" pageId="3" pageNumber="44">15.7–23.5</td>
</tr>
<tr id="C3E6D85AFF91003522E9FB2CFA196543" box="[159,1423,1246,1303]" gridrow="2" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FB2CFECA6543" box="[159,348,1246,1303]" gridcol="0" gridrow="2" pageId="3" pageNumber="44">Lateral append- age D</th>
<td id="8037B126FF910035231BFB2CFE586543" box="[365,462,1246,1303]" gridcol="1" gridrow="2" pageId="3" pageNumber="44">11.2</td>
<td id="8037B126FF9100352396FB2CFDD66543" box="[480,576,1246,1303]" gridcol="2" gridrow="2" pageId="3" pageNumber="44">5.1</td>
<td id="8037B126FF9100352028FB2CFD286543" box="[606,702,1246,1303]" gridcol="3" gridrow="2" pageId="3" pageNumber="44">26.2</td>
<td id="8037B126FF91003520ABFB2CFCAB6543" box="[733,829,1246,1303]" gridcol="4" gridrow="2" pageId="3" pageNumber="44">a–2.1</td>
<td id="8037B126FF9100352122FB2CFC316543" box="[852,935,1246,1303]" gridcol="5" gridrow="2" pageId="3" pageNumber="44">a–1.2</td>
<td id="8037B126FF91003521BCFB2CFBBD6543" box="[970,1067,1246,1303]" gridcol="6" gridrow="2" pageId="3" pageNumber="44">a–5.6</td>
<td id="8037B126FF9100352637FB2CFB376543" box="[1089,1185,1246,1303]" gridcol="7" gridrow="2" pageId="3" pageNumber="44">5.1–6.8</td>
<td id="8037B126FF91003526C5FB2CFA956543" box="[1203,1283,1246,1303]" gridcol="8" gridrow="2" pageId="3" pageNumber="44">2.1–2.8</td>
<td id="8037B126FF9100352752FB2CFA196543" box="[1316,1423,1246,1303]" gridcol="9" gridrow="2" pageId="3" pageNumber="44">12.0–16.0</td>
</tr>
<tr id="C3E6D85AFF91003522E9FAD9FA196531" box="[159,1423,1323,1381]" gridrow="3" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FAD9FECA6531" box="[159,348,1323,1381]" gridcol="0" gridrow="3" pageId="3" pageNumber="44">Lateral append- age E</th>
<td id="8037B126FF910035231BFAD9FE586531" box="[365,462,1323,1381]" gridcol="1" gridrow="3" pageId="3" pageNumber="44">15.6</td>
<td id="8037B126FF9100352396FAD9FDD66531" box="[480,576,1323,1381]" gridcol="2" gridrow="3" pageId="3" pageNumber="44">7.2</td>
<td id="8037B126FF9100352028FAD9FD286531" box="[606,702,1323,1381]" gridcol="3" gridrow="3" pageId="3" pageNumber="44">36.5</td>
<td id="8037B126FF91003520ABFAD9FCAB6531" box="[733,829,1323,1381]" gridcol="4" gridrow="3" pageId="3" pageNumber="44">?–11.0</td>
<td id="8037B126FF9100352122FAD9FC316531" box="[852,935,1323,1381]" gridcol="5" gridrow="3" pageId="3" pageNumber="44">?–6.4</td>
<td id="8037B126FF91003521BCFAD9FBBD6531" box="[970,1067,1323,1381]" gridcol="6" gridrow="3" pageId="3" pageNumber="44">?–29.3</td>
<td id="8037B126FF9100352637FAD9FB376531" box="[1089,1185,1323,1381]" gridcol="7" gridrow="3" pageId="3" pageNumber="44">9.5–10.9</td>
<td id="8037B126FF91003526C5FAD9FA956531" box="[1203,1283,1323,1381]" gridcol="8" gridrow="3" pageId="3" pageNumber="44">4.0–4.5</td>
<td id="8037B126FF9100352752FAD9FA196531" box="[1316,1423,1323,1381]" gridcol="9" gridrow="3" pageId="3" pageNumber="44">22.3–25.6</td>
</tr>
<tr id="C3E6D85AFF91003522E9FA8AFA1965E1" box="[159,1423,1400,1461]" gridrow="4" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FA8AFECA65E1" box="[159,348,1400,1461]" gridcol="0" gridrow="4" pageId="3" pageNumber="44">Dorsal append- age D d</th>
<td id="8037B126FF910035231BFA8AFE5865E1" box="[365,462,1400,1461]" gridcol="1" gridrow="4" pageId="3" pageNumber="44">21.8</td>
<td id="8037B126FF9100352396FA8AFDD665E1" box="[480,576,1400,1461]" gridcol="2" gridrow="4" pageId="3" pageNumber="44">10.0</td>
<td id="8037B126FF9100352028FA8AFD2865E1" box="[606,702,1400,1461]" gridcol="3" gridrow="4" pageId="3" pageNumber="44">50.9</td>
<td id="8037B126FF91003520ABFA8AFCAB65E1" box="[733,829,1400,1461]" gridcol="4" gridrow="4" pageId="3" pageNumber="44">13.7–19.2</td>
<td id="8037B126FF9100352122FA8AFC3165E1" box="[852,935,1400,1461]" gridcol="5" gridrow="4" pageId="3" pageNumber="44">8.0–11.1</td>
<td id="8037B126FF91003521BCFA8AFBBD65E1" box="[970,1067,1400,1461]" gridcol="6" gridrow="4" pageId="3" pageNumber="44">36.4–51.1</td>
<td id="8037B126FF9100352637FA8AFB3765E1" box="[1089,1185,1400,1461]" gridcol="7" gridrow="4" pageId="3" pageNumber="44">18.3–19.7</td>
<td id="8037B126FF91003526C5FA8AFA9565E1" box="[1203,1283,1400,1461]" gridcol="8" gridrow="4" pageId="3" pageNumber="44">7.6–8.2</td>
<td id="8037B126FF9100352752FA8AFA1965E1" box="[1316,1423,1400,1461]" gridcol="9" gridrow="4" pageId="3" pageNumber="44">43.7–46.2</td>
</tr>
<tr id="C3E6D85AFF91003522E9FA3AFA19658B" box="[159,1423,1480,1503]" gridrow="5" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FA3AFECA658B" box="[159,348,1480,1503]" gridcol="0" gridrow="5" pageId="3" pageNumber="44">Spine leg I</th>
<td id="8037B126FF910035231BFA3AFE58658B" box="[365,462,1480,1503]" gridcol="1" gridrow="5" pageId="3" pageNumber="44">1.8</td>
<td id="8037B126FF9100352396FA3AFDD6658B" box="[480,576,1480,1503]" gridcol="2" gridrow="5" pageId="3" pageNumber="44">0.8</td>
<td id="8037B126FF9100352028FA3AFD28658B" box="[606,702,1480,1503]" gridcol="3" gridrow="5" pageId="3" pageNumber="44">4.2</td>
<td id="8037B126FF91003520ABFA3AFCAB658B" box="[733,829,1480,1503]" gridcol="4" gridrow="5" pageId="3" pageNumber="44">1.6</td>
<td id="8037B126FF9100352122FA3AFC31658B" box="[852,935,1480,1503]" gridcol="5" gridrow="5" pageId="3" pageNumber="44">0.9</td>
<td id="8037B126FF91003521BCFA3AFBBD658B" box="[970,1067,1480,1503]" gridcol="6" gridrow="5" pageId="3" pageNumber="44">4.3</td>
<td id="8037B126FF9100352637FA3AFB37658B" box="[1089,1185,1480,1503]" gridcol="7" gridrow="5" pageId="3" pageNumber="44">2.0</td>
<td id="8037B126FF91003526C5FA3AFA95658B" box="[1203,1283,1480,1503]" gridcol="8" gridrow="5" pageId="3" pageNumber="44">0.8</td>
<td id="8037B126FF9100352752FA3AFA19658B" box="[1316,1423,1480,1503]" gridcol="9" gridrow="5" pageId="3" pageNumber="44">4.7</td>
</tr>
<tr id="C3E6D85AFF91003522E9FA01FA19665E" box="[159,1423,1523,1546]" gridrow="6" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9FA01FECA665E" box="[159,348,1523,1546]" gridcol="0" gridrow="6" pageId="3" pageNumber="44">Internal claws I</th>
<td id="8037B126FF910035231BFA01FE58665E" box="[365,462,1523,1546]" gridcol="1" gridrow="6" pageId="3" pageNumber="44">11.1</td>
<td id="8037B126FF9100352396FA01FDD6665E" box="[480,576,1523,1546]" gridcol="2" gridrow="6" pageId="3" pageNumber="44">5.1</td>
<td id="8037B126FF9100352028FA01FD28665E" box="[606,702,1523,1546]" gridcol="3" gridrow="6" pageId="3" pageNumber="44">25.9</td>
<td id="8037B126FF91003520ABFA01FCAB665E" box="[733,829,1523,1546]" gridcol="4" gridrow="6" pageId="3" pageNumber="44">9.7</td>
<td id="8037B126FF9100352122FA01FC31665E" box="[852,935,1523,1546]" gridcol="5" gridrow="6" pageId="3" pageNumber="44">5.6</td>
<td id="8037B126FF91003521BCFA01FBBD665E" box="[970,1067,1523,1546]" gridcol="6" gridrow="6" pageId="3" pageNumber="44">25.8</td>
<td id="8037B126FF9100352637FA01FB37665E" box="[1089,1185,1523,1546]" gridcol="7" gridrow="6" pageId="3" pageNumber="44">10.7</td>
<td id="8037B126FF91003526C5FA01FA95665E" box="[1203,1283,1523,1546]" gridcol="8" gridrow="6" pageId="3" pageNumber="44">4.5</td>
<td id="8037B126FF9100352752FA01FA19665E" box="[1316,1423,1523,1546]" gridcol="9" gridrow="6" pageId="3" pageNumber="44">25.1</td>
</tr>
<tr id="C3E6D85AFF91003522E9F9ECFA196661" box="[159,1423,1566,1589]" gridrow="7" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9F9ECFECA6661" box="[159,348,1566,1589]" gridcol="0" gridrow="7" pageId="3" pageNumber="44">External claws I</th>
<td id="8037B126FF910035231BF9ECFE586661" box="[365,462,1566,1589]" gridcol="1" gridrow="7" pageId="3" pageNumber="44">10.3</td>
<td id="8037B126FF9100352396F9ECFDD66661" box="[480,576,1566,1589]" gridcol="2" gridrow="7" pageId="3" pageNumber="44">4.7</td>
<td id="8037B126FF9100352028F9ECFD286661" box="[606,702,1566,1589]" gridcol="3" gridrow="7" pageId="3" pageNumber="44">24.1</td>
<td id="8037B126FF91003520ABF9ECFCAB6661" box="[733,829,1566,1589]" gridcol="4" gridrow="7" pageId="3" pageNumber="44">8.7</td>
<td id="8037B126FF9100352122F9ECFC316661" box="[852,935,1566,1589]" gridcol="5" gridrow="7" pageId="3" pageNumber="44">5.0</td>
<td id="8037B126FF91003521BCF9ECFBBD6661" box="[970,1067,1566,1589]" gridcol="6" gridrow="7" pageId="3" pageNumber="44">23.1</td>
<td id="8037B126FF9100352637F9ECFB376661" box="[1089,1185,1566,1589]" gridcol="7" gridrow="7" pageId="3" pageNumber="44">10.1</td>
<td id="8037B126FF91003526C5F9ECFA956661" box="[1203,1283,1566,1589]" gridcol="8" gridrow="7" pageId="3" pageNumber="44">4.2</td>
<td id="8037B126FF9100352752F9ECFA196661" box="[1316,1423,1566,1589]" gridcol="9" gridrow="7" pageId="3" pageNumber="44">23.7</td>
</tr>
<tr id="C3E6D85AFF91003522E9F9BAFA1966D6" box="[159,1423,1608,1666]" gridrow="8" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9F9BAFECA66D6" box="[159,348,1608,1666]" gridcol="0" gridrow="8" pageId="3" pageNumber="44">Internal claws II/ III</th>
<td id="8037B126FF910035231BF9BAFE5866D6" box="[365,462,1608,1666]" gridcol="1" gridrow="8" pageId="3" pageNumber="44">10.9</td>
<td id="8037B126FF9100352396F9BAFDD666D6" box="[480,576,1608,1666]" gridcol="2" gridrow="8" pageId="3" pageNumber="44">5.0</td>
<td id="8037B126FF9100352028F9BAFD2866D6" box="[606,702,1608,1666]" gridcol="3" gridrow="8" pageId="3" pageNumber="44">25.5</td>
<td id="8037B126FF91003520ABF9BAFCAB66D6" box="[733,829,1608,1666]" gridcol="4" gridrow="8" pageId="3" pageNumber="44">9.4</td>
<td id="8037B126FF9100352122F9BAFC3166D6" box="[852,935,1608,1666]" gridcol="5" gridrow="8" pageId="3" pageNumber="44">5.5</td>
<td id="8037B126FF91003521BCF9BAFBBD66D6" box="[970,1067,1608,1666]" gridcol="6" gridrow="8" pageId="3" pageNumber="44">25.0</td>
<td id="8037B126FF9100352637F9BAFB3766D6" box="[1089,1185,1608,1666]" gridcol="7" gridrow="8" pageId="3" pageNumber="44">11.5</td>
<td id="8037B126FF91003526C5F9BAFA9566D6" box="[1203,1283,1608,1666]" gridcol="8" gridrow="8" pageId="3" pageNumber="44">4.8</td>
<td id="8037B126FF9100352752F9BAFA1966D6" box="[1316,1423,1608,1666]" gridcol="9" gridrow="8" pageId="3" pageNumber="44">27.0</td>
</tr>
<tr id="C3E6D85AFF91003522E9F964FA19669B" box="[159,1423,1686,1743]" gridrow="9" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9F964FECA669B" box="[159,348,1686,1743]" gridcol="0" gridrow="9" pageId="3" pageNumber="44">External claws II/ III</th>
<td id="8037B126FF910035231BF964FE58669B" box="[365,462,1686,1743]" gridcol="1" gridrow="9" pageId="3" pageNumber="44">10.1</td>
<td id="8037B126FF9100352396F964FDD6669B" box="[480,576,1686,1743]" gridcol="2" gridrow="9" pageId="3" pageNumber="44">4.6</td>
<td id="8037B126FF9100352028F964FD28669B" box="[606,702,1686,1743]" gridcol="3" gridrow="9" pageId="3" pageNumber="44">23.6</td>
<td id="8037B126FF91003520ABF964FCAB669B" box="[733,829,1686,1743]" gridcol="4" gridrow="9" pageId="3" pageNumber="44">8.5</td>
<td id="8037B126FF9100352122F964FC31669B" box="[852,935,1686,1743]" gridcol="5" gridrow="9" pageId="3" pageNumber="44">4.9</td>
<td id="8037B126FF91003521BCF964FBBD669B" box="[970,1067,1686,1743]" gridcol="6" gridrow="9" pageId="3" pageNumber="44">22.6</td>
<td id="8037B126FF9100352637F964FB37669B" box="[1089,1185,1686,1743]" gridcol="7" gridrow="9" pageId="3" pageNumber="44">10.2</td>
<td id="8037B126FF91003526C5F964FA95669B" box="[1203,1283,1686,1743]" gridcol="8" gridrow="9" pageId="3" pageNumber="44">4.2</td>
<td id="8037B126FF9100352752F964FA19669B" box="[1316,1423,1686,1743]" gridcol="9" gridrow="9" pageId="3" pageNumber="44">23.9</td>
</tr>
<tr id="C3E6D85AFF91003522E9F911FA1966AE" box="[159,1423,1763,1786]" gridrow="10" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9F911FECA66AE" box="[159,348,1763,1786]" gridcol="0" gridrow="10" pageId="3" pageNumber="44">Internal claw IV</th>
<td id="8037B126FF910035231BF911FE5866AE" box="[365,462,1763,1786]" gridcol="1" gridrow="10" pageId="3" pageNumber="44">13.2</td>
<td id="8037B126FF9100352396F911FDD666AE" box="[480,576,1763,1786]" gridcol="2" gridrow="10" pageId="3" pageNumber="44">5.6</td>
<td id="8037B126FF9100352028F911FD2866AE" box="[606,702,1763,1786]" gridcol="3" gridrow="10" pageId="3" pageNumber="44">30.8</td>
<td id="8037B126FF91003520ABF911FCAB66AE" box="[733,829,1763,1786]" gridcol="4" gridrow="10" pageId="3" pageNumber="44">11.2</td>
<td id="8037B126FF9100352122F911FC3166AE" box="[852,935,1763,1786]" gridcol="5" gridrow="10" pageId="3" pageNumber="44">6.5</td>
<td id="8037B126FF91003521BCF911FBBD66AE" box="[970,1067,1763,1786]" gridcol="6" gridrow="10" pageId="3" pageNumber="44">29.8</td>
<td id="8037B126FF9100352637F911FB3766AE" box="[1089,1185,1763,1786]" gridcol="7" gridrow="10" pageId="3" pageNumber="44">13.3</td>
<td id="8037B126FF91003526C5F911FA9566AE" box="[1203,1283,1763,1786]" gridcol="8" gridrow="10" pageId="3" pageNumber="44">5.5</td>
<td id="8037B126FF9100352752F911FA1966AE" box="[1316,1423,1763,1786]" gridcol="9" gridrow="10" pageId="3" pageNumber="44">31.2</td>
</tr>
<tr id="C3E6D85AFF91003522E9F8FCFA196713" box="[159,1423,1806,1863]" gridrow="11" pageId="3" pageNumber="44">
<th id="8037B126FF91003522E9F8FCFECA6713" box="[159,348,1806,1863]" gridcol="0" gridrow="11" pageId="3" pageNumber="44">External claws IV</th>
<td id="8037B126FF910035231BF8FCFE586713" box="[365,462,1806,1863]" gridcol="1" gridrow="11" pageId="3" pageNumber="44">12.0</td>
<td id="8037B126FF9100352396F8FCFDD66713" box="[480,576,1806,1863]" gridcol="2" gridrow="11" pageId="3" pageNumber="44">5.5</td>
<td id="8037B126FF9100352028F8FCFD286713" box="[606,702,1806,1863]" gridcol="3" gridrow="11" pageId="3" pageNumber="44">28.0</td>
<td id="8037B126FF91003520ABF8FCFCAB6713" box="[733,829,1806,1863]" gridcol="4" gridrow="11" pageId="3" pageNumber="44">10.0</td>
<td id="8037B126FF9100352122F8FCFC316713" box="[852,935,1806,1863]" gridcol="5" gridrow="11" pageId="3" pageNumber="44">5.8</td>
<td id="8037B126FF91003521BCF8FCFBBD6713" box="[970,1067,1806,1863]" gridcol="6" gridrow="11" pageId="3" pageNumber="44">26.6</td>
<td id="8037B126FF9100352637F8FCFB376713" box="[1089,1185,1806,1863]" gridcol="7" gridrow="11" pageId="3" pageNumber="44">11.7</td>
<td id="8037B126FF91003526C5F8FCFA956713" box="[1203,1283,1806,1863]" gridcol="8" gridrow="11" pageId="3" pageNumber="44">4.9</td>
<td id="8037B126FF9100352752F8FCFA196713" box="[1316,1423,1806,1863]" gridcol="9" gridrow="11" pageId="3" pageNumber="44">27.5</td>
</tr>
</table>
</paragraph>
<caption id="29A98A90FF96FFCE22E1FF65FABD60FB" ID-Table-UUID="29A98A90FF96FFCE22E1FF65FABD60FB" box="[151,1323,151,175]" httpUri="http://table.plazi.org/id/29A98A90FF96FFCE22E1FF65FABD60FB" pageId="4" pageNumber="45" targetBox="[159,1108,208,412]" targetIsTable="true" targetPageId="4">
<paragraph id="7D69DA18FF96FFCE22E1FF65FABD60FB" blockId="4.[151,1323,151,175]" box="[151,1323,151,175]" pageId="4" pageNumber="45">
<emphasis id="4FA2060AFF96FFCE22E1FF65FE9460FB" bold="true" box="[151,258,151,175]" pageId="4" pageNumber="45">TABLE 2</emphasis>
. Density of light spots in the scapular plate of 
<taxonomicName id="BAD6A19BFF96FFCE2083FF6AFC2060FB" box="[757,950,152,175]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF96FFCE2083FF6AFC2060FB" box="[757,950,152,175]" italics="true" pageId="4" pageNumber="45">E. scabrospinosus</emphasis>
</taxonomicName>
(estimated in 20 adult specimens).
</paragraph>
</caption>
<paragraph id="7D69DA18FF96FFCE22E9FF22FCFC61CD" pageId="4" pageNumber="45">
<table id="0FD628B8FF96003522E9FF22FBC261C8" box="[159,1108,208,412]" gridcols="5" gridrows="5" pageId="4" pageNumber="45">
<tr id="C3E6D85AFF96003522E9FF22FBC260B3" box="[159,1108,208,231]" gridrow="0" pageId="4" pageNumber="45">
<th id="8037B126FF96003522E9FF22FE5360B3" box="[159,453,208,231]" gridcol="0" gridrow="0" pageId="4" pageNumber="45">Density on scapular plate</th>
<th id="8037B126FF9600352157FF22FCEE60B3" box="[801,888,208,231]" gridcol="1" gridrow="0" pageId="4" pageNumber="45">Number</th>
<th id="8037B126FF9600352109FF22FB9C60B3" box="[895,1034,208,231]" gridcol="2" gridrow="0" pageId="4" pageNumber="45">of specimens</th>
<th id="8037B126FF9600352667FF22FBA160B3" box="[1041,1079,208,231]" gridcol="3" gridrow="0" pageId="4" pageNumber="45">and</th>
<th id="8037B126FF9600352648FF22FBC260B3" box="[1086,1108,208,231]" gridcol="4" gridrow="0" pageId="4" pageNumber="45">%</th>
</tr>
<tr id="C3E6D85AFF96003522E9FF08FBC26140" box="[159,1108,250,276]" gridrow="1" pageId="4" pageNumber="45" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="8037B126FF96003522E9FF08FE536140" box="[159,453,250,276]" gridcol="0" gridrow="1" pageId="4" pageNumber="45">12–15 light spots / 100 µm2</th>
<td id="8037B126FF9600352157FF08FCEE6140" box="[801,888,250,276]" gridcol="1" gridrow="1" pageId="4" pageNumber="45">8 (40%)</td>
</tr>
<tr id="C3E6D85AFF96003522E9FEDAFBC26115" box="[159,1108,296,321]" gridrow="2" pageId="4" pageNumber="45" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="8037B126FF96003522E9FEDAFE536115" box="[159,453,296,321]" gridcol="0" gridrow="2" pageId="4" pageNumber="45">16–18 light spots / 100 µm2</th>
<td id="8037B126FF9600352157FEDAFCEE6115" box="[801,888,296,321]" gridcol="1" gridrow="2" pageId="4" pageNumber="45">7 (35%)</td>
</tr>
<tr id="C3E6D85AFF96003522E9FEA7FBC2613B" box="[159,1108,341,367]" gridrow="3" pageId="4" pageNumber="45" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="8037B126FF96003522E9FEA7FE53613B" box="[159,453,341,367]" gridcol="0" gridrow="3" pageId="4" pageNumber="45">19–21 light spots / 100 µm2</th>
<td id="8037B126FF9600352157FEA7FCEE613B" box="[801,888,341,367]" gridcol="1" gridrow="3" pageId="4" pageNumber="45">4 (20%)</td>
</tr>
<tr id="C3E6D85AFF96003522E9FE70FBC261C8" box="[159,1108,386,412]" gridrow="4" pageId="4" pageNumber="45" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="8037B126FF96003522E9FE70FE5361C8" box="[159,453,386,412]" gridcol="0" gridrow="4" pageId="4" pageNumber="45">&gt; 21 light spots / 100 µm2</th>
<td id="8037B126FF9600352157FE70FCEE61C8" box="[801,888,386,412]" gridcol="1" gridrow="4" pageId="4" pageNumber="45">1 (5%)</td>
</tr>
</table>
</paragraph>
<caption id="29A98A90FF96FFCE22E1FA28FCE2660D" httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="4" pageNumber="45" targetBox="[205,1386,498,1472]" targetPageId="4">
<paragraph id="7D69DA18FF96FFCE22E1FA28FCE2660D" blockId="4.[151,1436,1498,1625]" pageId="4" pageNumber="45">
<emphasis id="4FA2060AFF96FFCE22E1FA28FE8465A6" bold="true" box="[151,274,1498,1522]" pageId="4" pageNumber="45">FIGURE 2</emphasis>
. A–D, 
<emphasis id="4FA2060AFF96FFCE2317FA28FD1065A5" box="[353,646,1498,1521]" italics="true" pageId="4" pageNumber="45">
<taxonomicName id="BAD6A19BFF96FFCE2317FA28FD1465A5" box="[353,642,1498,1521]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="scabrospinosus">Echiniscus scabrospinosus</taxonomicName>
.
</emphasis>
A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm.
</paragraph>
</caption>
<subSubSection id="35CC8993FF96FFCF22B3F88FFEF46729" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45" type="reference_group">
<paragraph id="7D69DA18FF96FFCE22B3F88FFEAA67B3" blockId="4.[151,1436,1676,2023]" pageId="4" pageNumber="45">
<taxonomicName id="BAD6A19BFF96FFCE22B3F88FFE6967C2" ID-CoL="38G3C" box="[197,511,1917,1942]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF96FFCE22B3F88FFE6967C2" box="[197,511,1917,1942]" italics="true" pageId="4" pageNumber="45">Echiniscus scabrospinosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="BAD6A19BFF96FFCE20ECF88FFCF367C2" ID-CoL="38FXX" box="[666,869,1917,1943]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="45" phylum="Tardigrada" rank="species" species="crassispinosus">
<emphasis id="4FA2060AFF96FFCE20ECF88FFD3A67C2" box="[666,684,1917,1942]" italics="true" pageId="4" pageNumber="45">E</emphasis>
. 
<emphasis id="4FA2060AFF96FFCE20CCF88FFCF367C2" box="[698,869,1917,1942]" italics="true" pageId="4" pageNumber="45">crassispinosus</emphasis>
</taxonomicName>
in details of the plate ornamentation and in having, according to the literature (
<bibRefCitation id="1947A7E9FF96FFCE207BF857FD2267EB" author="Marcus" box="[525,692,1957,1983]" pageId="4" pageNumber="54" refString="Marcus, E. (1936) Tardigrada. In: De Gruyter W. (Ed.), Das Tierreichs, 66,1 - 340." type="book" year="1936">Marcus, 1936</bibRefCitation>
; 
<bibRefCitation id="1947A7E9FF96FFCE20B7F857FBB367EB" author="Ramazzotti" box="[705,1061,1957,1983]" pageId="4" pageNumber="54" refString="Ramazzotti, G. &amp; Maucci, W. (1983) Il phylum Tardigrada. Memorie dell'Istituto Italiano di Idrobiologia Marco De Marchi, Pallanza, 41, 1 - 1012." type="journal article" year="1983">Ramazzotti and Maucci, 1983</bibRefCitation>
) internal claws with longer and straight spurs.
</paragraph>
<caption id="29A98A90FF97FFCF22E1FF65FC446085" ID-Table-UUID="29A98A90FF97FFCF22E1FF65FC446085" httpUri="http://table.plazi.org/id/29A98A90FF97FFCF22E1FF65FC446085" pageId="5" pageNumber="46" targetBox="[159,1317,242,567]" targetIsTable="true" targetPageId="5">
<paragraph id="7D69DA18FF97FFCF22E1FF65FC446085" blockId="5.[151,1436,151,209]" pageId="5" pageNumber="46">
<emphasis id="4FA2060AFF97FFCF22E1FF65FE9C60FB" bold="true" box="[151,266,151,175]" pageId="5" pageNumber="46">TABLE 3.</emphasis>
Morphotypes recognizable in 
<taxonomicName id="BAD6A19BFF97FFCF202FFF6AFC8D60FB" ID-CoL="38G3C" box="[601,795,152,175]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="46" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF97FFCF202FFF6AFC8D60FB" box="[601,795,152,175]" italics="true" pageId="5" pageNumber="46">E. scabrospinosus</emphasis>
</taxonomicName>
as a consequence of the presence or absence of the lateral and dorsal appendages at least in one side of the body (43 adults and 1 larva).
</paragraph>
</caption>
<paragraph id="7D69DA18FF97FFCF22E9FF00FA826263" pageId="5" pageNumber="46">
<table id="0FD628B8FF97003522E9FF00FAB36263" box="[159,1317,242,567]" gridcols="6" gridrows="8" pageId="5" pageNumber="46">
<tr id="C3E6D85AFF97003522E9FF00FAB36158" box="[159,1317,242,268]" gridrow="0" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FF00FDA76158" box="[159,561,242,268]" gridcol="0" gridrow="0" pageId="5" pageNumber="46">Morphotype Specimens n (%)</th>
<th id="8037B126FF97003520D3FF00FD2E6158" box="[677,696,242,268]" gridcol="1" gridrow="0" pageId="5" pageNumber="46">A</th>
<th id="8037B126FF9700352149FF00FCC76158" box="[831,849,242,268]" gridcol="2" gridrow="0" pageId="5" pageNumber="46">C</th>
<th id="8037B126FF97003521AEFF00FC7D6158" box="[984,1003,242,268]" gridcol="3" gridrow="0" pageId="5" pageNumber="46">D</th>
<th id="8037B126FF9700352607FF00FB176158" box="[1137,1153,242,268]" gridcol="4" gridrow="0" pageId="5" pageNumber="46">E</th>
<th id="8037B126FF970035277DFF00FAB36158" box="[1291,1317,242,268]" gridcol="5" gridrow="0" pageId="5" pageNumber="46">D d</th>
</tr>
<tr id="C3E6D85AFF97003522E9FED2FAB36163" box="[159,1317,288,311]" gridrow="1" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FED2FDA76163" box="[159,561,288,311]" gridcol="0" gridrow="1" pageId="5" pageNumber="46">1 27 (62.8%)</th>
<td id="8037B126FF97003520D3FED2FD2E6163" box="[677,696,288,311]" gridcol="1" gridrow="1" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FED2FCC76163" box="[831,849,288,311]" gridcol="2" gridrow="1" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF97003521AEFED2FC7D6163" box="[984,1003,288,311]" gridcol="3" gridrow="1" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352607FED2FB176163" box="[1137,1153,288,311]" gridcol="4" gridrow="1" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF970035277DFED2FAB36163" box="[1291,1317,288,311]" gridcol="5" gridrow="1" pageId="5" pageNumber="46">+</td>
</tr>
<tr id="C3E6D85AFF97003522E9FEB8FAB36135" box="[159,1317,330,353]" gridrow="2" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FEB8FDA76135" box="[159,561,330,353]" gridcol="0" gridrow="2" pageId="5" pageNumber="46">2 8 (18.6%)</th>
<td id="8037B126FF97003520D3FEB8FD2E6135" box="[677,696,330,353]" gridcol="1" gridrow="2" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FEB8FCC76135" box="[831,849,330,353]" gridcol="2" gridrow="2" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF97003521AEFEB8FC7D6135" box="[984,1003,330,353]" gridcol="3" gridrow="2" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF9700352607FEB8FB176135" box="[1137,1153,330,353]" gridcol="4" gridrow="2" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF970035277DFEB8FAB36135" box="[1291,1317,330,353]" gridcol="5" gridrow="2" pageId="5" pageNumber="46">+</td>
</tr>
<tr id="C3E6D85AFF97003522E9FE87FAB361D8" box="[159,1317,373,396]" gridrow="3" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FE87FDA761D8" box="[159,561,373,396]" gridcol="0" gridrow="3" pageId="5" pageNumber="46">3 3 (7.0%)</th>
<td id="8037B126FF97003520D3FE87FD2E61D8" box="[677,696,373,396]" gridcol="1" gridrow="3" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FE87FCC761D8" box="[831,849,373,396]" gridcol="2" gridrow="3" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF97003521AEFE87FC7D61D8" box="[984,1003,373,396]" gridcol="3" gridrow="3" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF9700352607FE87FB1761D8" box="[1137,1153,373,396]" gridcol="4" gridrow="3" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF970035277DFE87FAB361D8" box="[1291,1317,373,396]" gridcol="5" gridrow="3" pageId="5" pageNumber="46">+</td>
</tr>
<tr id="C3E6D85AFF97003522E9FE52FAB361E3" box="[159,1317,416,439]" gridrow="4" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FE52FDA761E3" box="[159,561,416,439]" gridcol="0" gridrow="4" pageId="5" pageNumber="46">4 3 (7.0%)</th>
<td id="8037B126FF97003520D3FE52FD2E61E3" box="[677,696,416,439]" gridcol="1" gridrow="4" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FE52FCC761E3" box="[831,849,416,439]" gridcol="2" gridrow="4" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF97003521AEFE52FC7D61E3" box="[984,1003,416,439]" gridcol="3" gridrow="4" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF9700352607FE52FB1761E3" box="[1137,1153,416,439]" gridcol="4" gridrow="4" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF970035277DFE52FAB361E3" box="[1291,1317,416,439]" gridcol="5" gridrow="4" pageId="5" pageNumber="46">+</td>
</tr>
<tr id="C3E6D85AFF97003522E9FE38FAB361B5" box="[159,1317,458,481]" gridrow="5" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FE38FDA761B5" box="[159,561,458,481]" gridcol="0" gridrow="5" pageId="5" pageNumber="46">5 1 (2.3%)</th>
<td id="8037B126FF97003520D3FE38FD2E61B5" box="[677,696,458,481]" gridcol="1" gridrow="5" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FE38FCC761B5" box="[831,849,458,481]" gridcol="2" gridrow="5" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF97003521AEFE38FC7D61B5" box="[984,1003,458,481]" gridcol="3" gridrow="5" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF9700352607FE38FB1761B5" box="[1137,1153,458,481]" gridcol="4" gridrow="5" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF970035277DFE38FAB361B5" box="[1291,1317,458,481]" gridcol="5" gridrow="5" pageId="5" pageNumber="46">+</td>
</tr>
<tr id="C3E6D85AFF97003522E9FE07FAB36258" box="[159,1317,501,524]" gridrow="6" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FE07FDA76258" box="[159,561,501,524]" gridcol="0" gridrow="6" pageId="5" pageNumber="46">6 1 (2.3%)</th>
<td id="8037B126FF97003520D3FE07FD2E6258" box="[677,696,501,524]" gridcol="1" gridrow="6" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FE07FCC76258" box="[831,849,501,524]" gridcol="2" gridrow="6" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF97003521AEFE07FC7D6258" box="[984,1003,501,524]" gridcol="3" gridrow="6" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF9700352607FE07FB176258" box="[1137,1153,501,524]" gridcol="4" gridrow="6" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF970035277DFE07FAB36258" box="[1291,1317,501,524]" gridcol="5" gridrow="6" pageId="5" pageNumber="46">-</td>
</tr>
<tr id="C3E6D85AFF97003522E9FDD2FAB36263" box="[159,1317,544,567]" gridrow="7" pageId="5" pageNumber="46">
<th id="8037B126FF97003522E9FDD2FDA76263" box="[159,561,544,567]" gridcol="0" gridrow="7" pageId="5" pageNumber="46">larva 1</th>
<td id="8037B126FF97003520D3FDD2FD2E6263" box="[677,696,544,567]" gridcol="1" gridrow="7" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF9700352149FDD2FCC76263" box="[831,849,544,567]" gridcol="2" gridrow="7" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF97003521AEFDD2FC7D6263" box="[984,1003,544,567]" gridcol="3" gridrow="7" pageId="5" pageNumber="46">-</td>
<td id="8037B126FF9700352607FDD2FB176263" box="[1137,1153,544,567]" gridcol="4" gridrow="7" pageId="5" pageNumber="46">+</td>
<td id="8037B126FF970035277DFDD2FAB36263" box="[1291,1317,544,567]" gridcol="5" gridrow="7" pageId="5" pageNumber="46">-</td>
</tr>
</table>
</paragraph>
<paragraph id="7D69DA18FF97FFCF22ECF909FEF46729" blockId="5.[151,1436,1787,1917]" pageId="5" pageNumber="46">
<emphasis id="4FA2060AFF97FFCF22ECF909FE8E6747" bold="true" box="[154,280,1787,1811]" pageId="5" pageNumber="46">FIGURE 3</emphasis>
. 
<taxonomicName id="BAD6A19BFF97FFCF235EF909FD836746" ID-CoL="38G3L" box="[296,533,1787,1810]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="46" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF97FFCF235EF909FD836746" box="[296,533,1787,1810]" italics="true" pageId="5" pageNumber="46">Echiniscus spinulosus</emphasis>
</taxonomicName>
. A, anterior portion of the body; the narrow neck plate and the scapular plate are shown; the arrow indicates the lateral portion of scapular plate with light spots; B, the dorsal appendages C d and D d are shown; arrow indicates light spots present on the legs; C, claws of the first pair of legs; D, claws of the fourth pair of legs. Scale bar = 10 µm.
</paragraph>
</subSubSection>
<subSubSection id="35CC8993FF97FFCD22B3F859FB9B617E" lastPageId="7" lastPageNumber="48" pageId="5" pageNumber="46" type="description">
<paragraph id="7D69DA18FF97FFCC22B3F859FB1D60E6" blockId="5.[151,1436,1963,2029]" lastBlockId="6.[151,1436,152,300]" lastPageId="6" lastPageNumber="47" pageId="5" pageNumber="46">
It is also similar to 
<taxonomicName id="BAD6A19BFF97FFCF23D2F85EFDB76791" box="[420,545,1964,1989]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="46" phylum="Tardigrada" rank="species" species="canedoi">
<emphasis id="4FA2060AFF97FFCF23D2F85EFDB76791" box="[420,545,1964,1989]" italics="true" pageId="5" pageNumber="46">E. canedoi</emphasis>
</taxonomicName>
but it differs from this species in lacking spine E. Unfortunately the description of 
<taxonomicName id="BAD6A19BFF97FFCF229AF826FEFC67B9" box="[236,362,2004,2029]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="46" phylum="Tardigrada" rank="species" species="canedoi">
<emphasis id="4FA2060AFF97FFCF229AF826FEFC67B9" box="[236,362,2004,2029]" italics="true" pageId="5" pageNumber="46">E. canedoi</emphasis>
</taxonomicName>
is not detailed as to the plates’ ornamentation and it is not specified whether a spine on the first pair of legs, a papilla on the hind leg and spurs on the claws are present or absent.
</paragraph>
<paragraph id="7D69DA18FF94FFCC22B3FF31FCDB6178" blockId="6.[151,1436,152,300]" pageId="6" pageNumber="47">
<taxonomicName id="BAD6A19BFF94FFCC22B3FF31FE0C6088" box="[197,410,195,220]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF94FFCC22B3FF31FE0C6088" box="[197,410,195,220]" italics="true" pageId="6" pageNumber="47">E. scabrospinosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="BAD6A19BFF94FFCC204EFF31FD5D6088" box="[568,715,195,220]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="manuelae">
<emphasis id="4FA2060AFF94FFCC204EFF31FD5D6088" box="[568,715,195,220]" italics="true" pageId="6" pageNumber="47">E. manuelae</emphasis>
</taxonomicName>
in lacking C d, in having filament A slightly longer and spine E smooth; unfortunately the description of 
<taxonomicName id="BAD6A19BFF94FFCC20F8FF19FCB76150" box="[654,801,235,260]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="manuelae">
<emphasis id="4FA2060AFF94FFCC20F8FF19FCB76150" box="[654,801,235,260]" italics="true" pageId="6" pageNumber="47">E. manuelae</emphasis>
</taxonomicName>
is not detailed as to the plates ornamentation, and it is not specified whether spurs are present or not on the claws.
</paragraph>
<paragraph id="7D69DA18FF94FFCC22E1FAFDFBA265C5" blockId="6.[151,1436,1295,1425]" pageId="6" pageNumber="47">
<emphasis id="4FA2060AFF94FFCC22E1FAFDFE846573" bold="true" box="[151,274,1295,1319]" pageId="6" pageNumber="47">FIGURE 4</emphasis>
. 
<taxonomicName id="BAD6A19BFF94FFCC2369FAE2FE796573" box="[287,495,1296,1319]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC2369FAE2FE796573" box="[287,495,1296,1319]" italics="true" pageId="6" pageNumber="47">Echiniscus spiniger</emphasis>
</taxonomicName>
. A, anterior portion of the body; the narrow neck plate and the scapular plate are shown; the arrow indicates the lateral portion of scapular plate with light spots; B, the dorsal appendages C d and D d are shown; C; terminal plate; its light spots are not smaller than those of the other plates (photos A, B and C from the same specimen); D, claws of the first pair of legs; E, claws of the fourth pair of legs. Scale bar = 10 µm.
</paragraph>
<paragraph id="7D69DA18FF94FFCC22B3FA32FB156651" blockId="6.[151,1437,1472,2027]" pageId="6" pageNumber="47">
<taxonomicName id="BAD6A19BFF94FFCC22B3FA32FE0A658D" box="[197,412,1472,1497]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF94FFCC22B3FA32FE0A658D" box="[197,412,1472,1497]" italics="true" pageId="6" pageNumber="47">E. scabrospinosus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="BAD6A19BFF94FFCC204BFA32FD4B658D" box="[573,733,1472,1498]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="angolensis">
<emphasis id="4FA2060AFF94FFCC204BFA32FDD9658D" box="[573,591,1472,1497]" italics="true" pageId="6" pageNumber="47">E</emphasis>
. 
<emphasis id="4FA2060AFF94FFCC2016FA32FD4B658D" box="[608,733,1472,1497]" italics="true" pageId="6" pageNumber="47">angolensis</emphasis>
</taxonomicName>
in lacking the median plate 3, in details of the plate ornamentation and, according to the description but not to the figure, in having a spine C d.
</paragraph>
<paragraph id="7D69DA18FF94FFCC22B3F9E1FCED6601" blockId="6.[151,1437,1472,2027]" pageId="6" pageNumber="47">
We noted the presence of a dark ring, from which some dark stripes irradiate, in the center of the light spots also in 
<taxonomicName id="BAD6A19BFF94FFCC2359F9C9FE296600" box="[303,447,1595,1621]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosu">
<emphasis id="4FA2060AFF94FFCC2359F9C9FED76600" box="[303,321,1595,1620]" italics="true" pageId="6" pageNumber="47">E</emphasis>
. 
<emphasis id="4FA2060AFF94FFCC2339F9C9FE296600" box="[335,447,1595,1620]" italics="true" pageId="6" pageNumber="47">spinulosu</emphasis>
</taxonomicName>
s, in 
<taxonomicName id="BAD6A19BFF94FFCC238EF9C9FDEF6600" box="[504,633,1595,1621]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC238EF9C9FD9C6600" box="[504,522,1595,1620]" italics="true" pageId="6" pageNumber="47">E</emphasis>
. 
<emphasis id="4FA2060AFF94FFCC206FF9C9FDEF6600" box="[537,633,1595,1620]" italics="true" pageId="6" pageNumber="47">spiniger</emphasis>
</taxonomicName>
and in 
<taxonomicName id="BAD6A19BFF94FFCC20A6F9C9FCE26600" box="[720,884,1595,1620]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="marginatus">
<emphasis id="4FA2060AFF94FFCC20A6F9C9FCE26600" box="[720,884,1595,1620]" italics="true" pageId="6" pageNumber="47">E. marginatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7D69DA18FF94FFCD22B3F994FB9B617E" blockId="6.[151,1437,1472,2027]" lastBlockId="7.[151,1436,152,298]" lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47">
<taxonomicName id="BAD6A19BFF94FFCC22B3F994FEF4662B" box="[197,354,1638,1663]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC22B3F994FEF4662B" box="[197,354,1638,1663]" italics="true" pageId="6" pageNumber="47">E. spinulosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="BAD6A19BFF94FFCC23EDF994FD8D662B" box="[411,539,1638,1663]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC23EDF994FD8D662B" box="[411,539,1638,1663]" italics="true" pageId="6" pageNumber="47">E. spiniger</emphasis>
</taxonomicName>
, when the lateral appendage B and the dorsal appendage C d are absent, have the same appendages as 
<taxonomicName id="BAD6A19BFF94FFCC23C2F97CFD1E66F3" box="[436,648,1678,1703]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF94FFCC23C2F97CFD1E66F3" box="[436,648,1678,1703]" italics="true" pageId="6" pageNumber="47">E. scabrospinosus</emphasis>
</taxonomicName>
. But this species differs from 
<taxonomicName id="BAD6A19BFF94FFCC219FF97CFB1366F3" box="[1001,1157,1678,1704]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC219FF97CFC6D66F3" box="[1001,1019,1678,1703]" italics="true" pageId="6" pageNumber="47">E</emphasis>
. 
<emphasis id="4FA2060AFF94FFCC267FF97CFB1366F3" box="[1033,1157,1678,1703]" italics="true" pageId="6" pageNumber="47">spinulosus</emphasis>
</taxonomicName>
and from 
<taxonomicName id="BAD6A19BFF94FFCC268BF97CFAE866F3" box="[1277,1406,1678,1703]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC268BF97CFAE866F3" box="[1277,1406,1678,1703]" italics="true" pageId="6" pageNumber="47">E. spiniger</emphasis>
</taxonomicName>
in having small dots only in the neck plate, in the lateral portion of the scapular plate and of paired plates, and on the intersegmental lateral plates (in 
<taxonomicName id="BAD6A19BFF94FFCC2049F92CFD4A66A3" box="[575,732,1758,1784]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC2049F92CFDC766A3" box="[575,593,1758,1783]" italics="true" pageId="6" pageNumber="47">E</emphasis>
. 
<emphasis id="4FA2060AFF94FFCC2016F92CFD4A66A3" box="[608,732,1758,1783]" italics="true" pageId="6" pageNumber="47">spinulosus</emphasis>
</taxonomicName>
and in 
<taxonomicName id="BAD6A19BFF94FFCC2140F92CFC2166A3" box="[822,951,1758,1784]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC2140F92CFCDE66A3" box="[822,840,1758,1783]" italics="true" pageId="6" pageNumber="47">E</emphasis>
. 
<emphasis id="4FA2060AFF94FFCC2121F92CFC2166A3" box="[855,951,1758,1783]" italics="true" pageId="6" pageNumber="47">spiniger</emphasis>
</taxonomicName>
they are present in all plates); in lacking light spots on the neck plate (
<figureCitation id="E5EDC69DFF94FFCC2383F8F4FDD86774" box="[501,590,1798,1824]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="6" pageNumber="47">Fig. 1C</figureCitation>
) and on lateral portions of the scapular plate and of the paired plates II and III (
<figureCitation id="E5EDC69DFF94FFCC2283F8DCFEC6671C" box="[245,336,1838,1864]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="6" pageNumber="47">Fig. 1B</figureCitation>
; 2B) while in 
<taxonomicName id="BAD6A19BFF94FFCC2380F8DCFD046713" box="[502,658,1838,1863]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC2380F8DCFD046713" box="[502,658,1838,1863]" italics="true" pageId="6" pageNumber="47">E. spinulosus</emphasis>
</taxonomicName>
(Fig. 3A) and in 
<taxonomicName id="BAD6A19BFF94FFCC212AF8DCFC4A6713" box="[860,988,1838,1863]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC212AF8DCFC4A6713" box="[860,988,1838,1863]" italics="true" pageId="6" pageNumber="47">E. spiniger</emphasis>
</taxonomicName>
(Fig. 4A, C) they are present and well visible; in having light spots of the terminal plate smaller than those of other plates; in lacking light spots on the cuticle of the legs (present and well visible in 
<taxonomicName id="BAD6A19BFF94FFCC2099F88CFC1A67C3" box="[751,908,1918,1943]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC2099F88CFC1A67C3" box="[751,908,1918,1943]" italics="true" pageId="6" pageNumber="47">E. spinulosus</emphasis>
</taxonomicName>
and in 
<taxonomicName id="BAD6A19BFF94FFCC219FF88CFBFD67C3" box="[1001,1131,1918,1943]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC219FF88CFBFD67C3" box="[1001,1131,1918,1943]" italics="true" pageId="6" pageNumber="47">E. spiniger</emphasis>
</taxonomicName>
); in having longer lateral spines than 
<taxonomicName id="BAD6A19BFF94FFCC2355F85BFE566796" box="[291,448,1961,1986]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC2355F85BFE566796" box="[291,448,1961,1986]" italics="true" pageId="6" pageNumber="47">E. spinulosus</emphasis>
</taxonomicName>
; dorsal appendages D d (only exceptionally absent) wider than the other appendages (
<figureCitation id="E5EDC69DFF94FFCC22E9F823FE8B67BF" box="[159,285,2001,2027]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="6" pageNumber="47">Fig. 1A, B</figureCitation>
) while in 
<taxonomicName id="BAD6A19BFF94FFCC23E2F823FDA667BE" box="[404,560,2001,2026]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF94FFCC23E2F823FDA667BE" box="[404,560,2001,2026]" italics="true" pageId="6" pageNumber="47">E. spinulosus</emphasis>
</taxonomicName>
(Fig. 3B) and in 
<taxonomicName id="BAD6A19BFF94FFCC208FF823FCEC67BE" box="[761,890,2001,2026]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="47" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF94FFCC208FF823FCEC67BE" box="[761,890,2001,2026]" italics="true" pageId="6" pageNumber="47">E. spiniger</emphasis>
</taxonomicName>
(Fig. 4B) they are similar to the other appendages; internal claws with a thin spur (those of the legs I–III are slightly smaller than those of the hind legs, 
<figureCitation id="E5EDC69DFF95FFCD271BFF6AFF77608E" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="7" pageNumber="48">Fig. 2C, D</figureCitation>
) while in 
<taxonomicName id="BAD6A19BFF95FFCD2315FF32FD92608D" box="[355,516,192,217]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF95FFCD2315FF32FD92608D" box="[355,516,192,217]" italics="true" pageId="7" pageNumber="48">E. spinulosus</emphasis>
</taxonomicName>
(Fig. 3C, D) and in 
<taxonomicName id="BAD6A19BFF95FFCD2171FF32FC1A608D" box="[775,908,192,217]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF95FFCD2171FF32FC1A608D" box="[775,908,192,217]" italics="true" pageId="7" pageNumber="48">E. spiniger</emphasis>
</taxonomicName>
(Fig. 4D, E) the difference in size is more marked; in addition in 
<taxonomicName id="BAD6A19BFF95FFCD23D8FF1AFD106155" box="[430,646,232,257]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF95FFCD23D8FF1AFD106155" box="[430,646,232,257]" italics="true" pageId="7" pageNumber="48">E. scabrospinosus</emphasis>
</taxonomicName>
the spurs are clearly oriented towards the claw base, while in 
<taxonomicName id="BAD6A19BFF95FFCD27F5FF1AFE85617D" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="spinulosus">
<emphasis id="4FA2060AFF95FFCD27F5FF1AFE85617D" italics="true" pageId="7" pageNumber="48">E. spinulosus</emphasis>
</taxonomicName>
and in 
<taxonomicName id="BAD6A19BFF95FFCD231CFEE2FE7C617D" box="[362,490,272,297]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="4FA2060AFF95FFCD231CFEE2FE7C617D" box="[362,490,272,297]" italics="true" pageId="7" pageNumber="48">E. spiniger</emphasis>
</taxonomicName>
they form an almost right angle with the claw.
</paragraph>
</subSubSection>
<caption id="29A98A90FF95FFCD22E1F991FE0B66B7" httpUri="https://zenodo.org/record/183544/files/figure.png" pageId="7" pageNumber="48" targetBox="[194,1391,342,1619]" targetPageId="7">
<paragraph id="7D69DA18FF95FFCD22E1F991FE0B66B7" blockId="7.[151,1436,1635,1763]" pageId="7" pageNumber="48">
<emphasis id="4FA2060AFF95FFCD22E1F991FE85662F" bold="true" box="[151,275,1635,1659]" pageId="7" pageNumber="48">FIGURE 5</emphasis>
. 
<taxonomicName id="BAD6A19BFF95FFCD2357F996FD83662F" box="[289,533,1636,1659]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="marginatus">
<emphasis id="4FA2060AFF95FFCD2357F996FD83662F" box="[289,533,1636,1659]" italics="true" pageId="7" pageNumber="48">Echiniscus marginatus</emphasis>
</taxonomicName>
. A, anterior portion of the body of the holotype; B, posterior portion of the body of the holotype; C, detail of the cuticular ornamentation (paratype); arrow indicates light spots present also in the lateral portions of the paired plates; D, claws of the first pair of legs of a paratype; E, claws of the fourth pair of legs of the holotype. Scale bar = 10 µm.
</paragraph>
</caption>
<subSubSection id="35CC8993FF95FFC222B3F8E0FA98612E" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48" type="reference_group">
<paragraph id="7D69DA18FF95FFC222B3F8E0FA98612E" blockId="7.[151,1437,1810,2036]" lastBlockId="8.[151,1437,152,378]" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48">
<taxonomicName id="BAD6A19BFF95FFCD22B3F8E0FE69677F" ID-CoL="38G3C" box="[197,511,1810,1835]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF95FFCD22B3F8E0FE69677F" box="[197,511,1810,1835]" italics="true" pageId="7" pageNumber="48">Echiniscus scabrospinosus</emphasis>
</taxonomicName>
is extremely similar to 
<taxonomicName id="BAD6A19BFF95FFCD216CF8E0FB526778" ID-CoL="38FZH" authority="Binda &amp; Pilato, 1994" authorityName="Binda &amp; Pilato" authorityYear="1994" box="[794,1220,1810,1836]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="marginatus">
<emphasis id="4FA2060AFF95FFCD216CF8E0FC29677F" box="[794,959,1810,1835]" italics="true" pageId="7" pageNumber="48">E. marginatus</emphasis>
<bibRefCitation id="1947A7E9FF95FFCD21BEF8E0FB526778" author="Binda" box="[968,1220,1810,1836]" pageId="7" pageNumber="54" refString="Binda, M. G. &amp; Pilato, G. (1994) Notizie sui tardigradi delle isole Hawaii con descrizione di due specie nuove. Animalia, 21, 57 - 62." type="journal article" year="1994">Binda &amp; Pilato, 1994</bibRefCitation>
</taxonomicName>
that generally has the same appendages as 
<taxonomicName id="BAD6A19BFF95FFCD23C0F8C8FD1D6707" ID-CoL="38G3C" box="[438,651,1850,1875]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF95FFCD23C0F8C8FD1D6707" box="[438,651,1850,1875]" italics="true" pageId="7" pageNumber="48">E. scabrospinosus</emphasis>
</taxonomicName>
, and only a careful comparison confirmed that this species is not a synonym of 
<taxonomicName id="BAD6A19BFF95FFCD235AF890FD97672F" ID-CoL="38G3C" box="[300,513,1890,1915]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF95FFCD235AF890FD97672F" box="[300,513,1890,1915]" italics="true" pageId="7" pageNumber="48">E. scabrospinosus</emphasis>
</taxonomicName>
. We noted that the description of 
<taxonomicName id="BAD6A19BFF95FFCD21E3F890FBAD672F" ID-CoL="38FZH" box="[917,1083,1890,1915]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="48" phylum="Tardigrada" rank="species" species="marginatus">
<emphasis id="4FA2060AFF95FFCD21E3F890FBAD672F" box="[917,1083,1890,1915]" italics="true" pageId="7" pageNumber="48">E. marginatus</emphasis>
</taxonomicName>
includes a misunderstanding: the anterior margin of the paired plates II and III, of the median plate 2 and the area corresponding to the absent median plate 3 are strongly sculptured, but due to the partial overlapping of the plates, they had been erroneously considered as a marginal crenate stripe of the median plates 1 and 2 and of the paired plates II and III respectively (
<figureCitation id="E5EDC69DFF9AFFC2232DFF6AFE3760E6" box="[347,417,152,178]" captionStart="FIGURE 5" captionStartId="7.[151,255,1635,1659]" captionTargetBox="[194,1391,342,1619]" captionTargetId="figure@7.[189,1397,338,1626]" captionTargetPageId="7" captionText="FIGURE 5. Echiniscus marginatus. A, anterior portion of the body of the holotype; B, posterior portion of the body of the holotype; C, detail of the cuticular ornamentation (paratype); arrow indicates light spots present also in the lateral portions of the paired plates; D, claws of the first pair of legs of a paratype; E, claws of the fourth pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183544/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
A, B). We also noted that in 
<taxonomicName id="BAD6A19BFF9AFFC22081FF6AFC0B60E5" ID-CoL="38FZH" box="[759,925,152,177]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="49" phylum="Tardigrada" rank="species" species="marginatus">
<emphasis id="4FA2060AFF9AFFC22081FF6AFC0B60E5" box="[759,925,152,177]" italics="true" pageId="8" pageNumber="49">E. marginatus</emphasis>
</taxonomicName>
the light spots have a central dark ring not recorded in the description of the species. Notwithstanding these resemblances, we confirm that 
<taxonomicName id="BAD6A19BFF9AFFC2268EFF32FA0D608D" ID-CoL="38FZH" box="[1272,1435,192,217]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="49" phylum="Tardigrada" rank="species" species="marginatus">
<emphasis id="4FA2060AFF9AFFC2268EFF32FA0D608D" box="[1272,1435,192,217]" italics="true" pageId="8" pageNumber="49">E. marginatus</emphasis>
</taxonomicName>
is a 
<emphasis id="4FA2060AFF9AFFC222B1FF1AFEC86155" box="[199,350,232,257]" italics="true" pageId="8" pageNumber="49">bona species</emphasis>
; it differs from 
<taxonomicName id="BAD6A19BFF9AFFC2206AFF1AFD676155" ID-CoL="38G3C" box="[540,753,232,257]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="49" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF9AFFC2206AFF1AFD676155" box="[540,753,232,257]" italics="true" pageId="8" pageNumber="49">E. scabrospinosus</emphasis>
</taxonomicName>
in lacking a neck plate (
<figureCitation id="E5EDC69DFF9AFFC22665FF1AFBF46156" box="[1043,1122,232,258]" captionStart="FIGURE 5" captionStartId="7.[151,255,1635,1659]" captionTargetBox="[194,1391,342,1619]" captionTargetId="figure@7.[189,1397,338,1626]" captionTargetPageId="7" captionText="FIGURE 5. Echiniscus marginatus. A, anterior portion of the body of the holotype; B, posterior portion of the body of the holotype; C, detail of the cuticular ornamentation (paratype); arrow indicates light spots present also in the lateral portions of the paired plates; D, claws of the first pair of legs of a paratype; E, claws of the fourth pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183544/files/figure.png" pageId="8" pageNumber="49">Figs. 5</figureCitation>
A and 1C); in having light spots more variable in size (
<figureCitation id="E5EDC69DFF9AFFC2239BFEE2FDA9617E" box="[493,575,272,298]" captionStart="FIGURE 5" captionStartId="7.[151,255,1635,1659]" captionTargetBox="[194,1391,342,1619]" captionTargetId="figure@7.[189,1397,338,1626]" captionTargetPageId="7" captionText="FIGURE 5. Echiniscus marginatus. A, anterior portion of the body of the holotype; B, posterior portion of the body of the holotype; C, detail of the cuticular ornamentation (paratype); arrow indicates light spots present also in the lateral portions of the paired plates; D, claws of the first pair of legs of a paratype; E, claws of the fourth pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183544/files/figure.png" pageId="8" pageNumber="49">Figs. 5</figureCitation>
A–C and 1A–C); light spots of terminal plate not smaller than those of other plates (
<figureCitation id="E5EDC69DFF9AFFC22344FECAFEEF6106" box="[306,377,312,338]" captionStart="FIGURE 5" captionStartId="7.[151,255,1635,1659]" captionTargetBox="[194,1391,342,1619]" captionTargetId="figure@7.[189,1397,338,1626]" captionTargetPageId="7" captionText="FIGURE 5. Echiniscus marginatus. A, anterior portion of the body of the holotype; B, posterior portion of the body of the holotype; C, detail of the cuticular ornamentation (paratype); arrow indicates light spots present also in the lateral portions of the paired plates; D, claws of the first pair of legs of a paratype; E, claws of the fourth pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183544/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
A, B and 
<figureCitation id="E5EDC69DFF9AFFC22399FECAFDE76106" box="[495,625,312,338]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="8" pageNumber="49">Fig. 1A, B</figureCitation>
); less evident (probably absent) intersegmental lateral plates; lateral portions of paired plates with light spots (
<figureCitation id="E5EDC69DFF9AFFC22009FE92FD52612E" box="[639,708,352,378]" captionStart="FIGURE 5" captionStartId="7.[151,255,1635,1659]" captionTargetBox="[194,1391,342,1619]" captionTargetId="figure@7.[189,1397,338,1626]" captionTargetPageId="7" captionText="FIGURE 5. Echiniscus marginatus. A, anterior portion of the body of the holotype; B, posterior portion of the body of the holotype; C, detail of the cuticular ornamentation (paratype); arrow indicates light spots present also in the lateral portions of the paired plates; D, claws of the first pair of legs of a paratype; E, claws of the fourth pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183544/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
C) absent in 
<taxonomicName id="BAD6A19BFF9AFFC22116FE92FBA3612D" ID-CoL="38G3C" box="[864,1077,352,377]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="49" phylum="Tardigrada" rank="species" species="scabrospinosus">
<emphasis id="4FA2060AFF9AFFC22116FE92FBA3612D" box="[864,1077,352,377]" italics="true" pageId="8" pageNumber="49">E. scabrospinosus</emphasis>
</taxonomicName>
(
<figureCitation id="E5EDC69DFF9AFFC22632FE92FB59612E" box="[1092,1231,352,378]" captionStart="FIGURE 1. A – D" captionStartId="2.[151,255,1305,1329]" captionTargetBox="[189,1390,201,1279]" captionTargetId="figure@2.[187,1398,197,1279]" captionTargetPageId="2" captionText="FIGURE 1. A – D, Echiniscus scabrospinosus. A, B, habitus (A, specimen from Amarante, Portugal; B, specimen from Faial Island, Azores); arrows indicate the lateral portions of scapular plate and paired plates lacking light spots; C, anterior portion of the body of a specimen from Amarante (Portugal); the arrow indicates a pair of intersegmental lateral plates; D, detail of the plate ornamentation of the holotype; arrows indicate the dark ring present in the centre of each light spot. Scale bar = 10 µm." httpUri="https://zenodo.org/record/183542/files/figure.png" pageId="8" pageNumber="49">Figs. 1A, B</figureCitation>
, 
<figureCitation id="E5EDC69DFF9AFFC226ADFE92FA97612E" box="[1243,1281,352,378]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1498,1522]" captionTargetBox="[205,1386,498,1472]" captionTargetId="figure@4.[200,1394,493,1472]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Echiniscus scabrospinosus. A, B, anterior portion of the body of two specimens from Amarante (Portugal); arrows indicate the small spine on the first pair of legs and the lateral portion, lacking light spots, of scapular plate and paired plates; C, claws of the third pair of legs of a specimen from Amarante (Portugal); D, claws of the fourth pair of legs of a specimen from Faial Island (Azores). Scale bar = 10 µm." httpUri="https://zenodo.org/record/183543/files/figure.png" pageId="8" pageNumber="49">2B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>