366 lines
54 KiB
XML
366 lines
54 KiB
XML
<document ID-CLB-Dataset="122795" ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-GBIF-Dataset="5f938945-980c-433c-942a-f05d39529a25" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo" docDate="2022" docId="F527066760BD5643AE59B52817A99A28" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Metasarcus Roewer 1913" docType="treatment" docVersion="5" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1732651490011" updateUser="ExternalLinkService">
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<mods:titleInfo id="6BB9F83230C58083D0682774DE9246EC">
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<mods:title id="9D2D03247E28EC5E43A14EF21236FE0C">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:name id="7B0A1BAC24DCBA73168E4519CD83B5E9" type="personal">
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<mods:roleTerm id="1694374B73DFE241FF270C9D569E9842">Author</mods:roleTerm>
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<mods:namePart id="3393AE86970939B64F95E6144B81C886">Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation id="361DB94458FE6EA70FEC7FC346A15853">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier id="61AD776A85373DF45BADCF1A612AFD36" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FBD3101357AD325A6A2FE8BB5C9D3A58">Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier id="922BFE4415FB475FA60AEF1C92694FCD" type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation id="F9667BEDFADF10809192BC20B5D27B5A">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="FBB0C38AC6EC534976E44B88FC882BBB">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="60E36D09A5A013CB236570AF8A3FB393">
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<mods:date id="03637A7D3BD5D0C304AA49E60D7026C2">2022</mods:date>
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<mods:number id="E84D7B35F02A00FB8410675F03047C6C">2022-08-16</mods:number>
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<mods:number id="64020BB8E743FDCB91AE12052C775A3B">80</mods:number>
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<mods:classification id="ADE12C0905B7649CE2FD6F3EF25F8CAA">journal article</mods:classification>
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<treatment id="F527066760BD5643AE59B52817A99A28" ID-GBIF-Taxon="199468660" LSID="urn:lsid:plazi:treatment:F527066760BD5643AE59B52817A99A28" httpUri="http://treatment.plazi.org/id/F527066760BD5643AE59B52817A99A28" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
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<subSubSection id="D09C3B6F5A199AB15CC07BED93B79338" pageId="0" pageNumber="309" type="nomenclature">
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<paragraph id="EC10FEEC3E8B7D61383FAC307442A37D" pageId="0" pageNumber="309">
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3.39.
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<taxonomicName id="9FA7E57BA55B7F805E220A32ADF6FF41" ID-CoL="7PNZ9" LSID="F5270667-60BD-5643-AE59-B52817A99A28" authority="Roewer, 1913" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Metasarcus Roewer, 1913</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="F2D105DEB5B1209EA355E02F414BEB37" pageId="0" pageNumber="309" type="description">
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<paragraph id="88B5170B5DFE2FCE4A00DFAA31A3711A" pageId="0" pageNumber="309">
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<figureCitation id="650F10EA349583D4D4AE643FC84D1591" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5</figureCitation>
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<figureCitation id="2B8C74E0C4A7C777490B4E03DEDFF2E3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">, 10</figureCitation>
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<figureCitation id="81876388F0A3BC3DB3B353DBB1E48B69" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">, 14C-H</figureCitation>
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<figureCitation id="83F91377FB0636F6725FF1668ECFA996" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. bergmani sp. nov.; D - F M. clavifemur (Roewer, 1929); G - I M. fellinii sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309">, 19</figureCitation>
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<figureCitation id="D32EC71A4C13E6F7E2E7D0C42E258A10" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. trispinosus sp. nov.; D - F M. vacafloresae sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure20" httpUri="https://binary.pensoft.net/fig/729894" pageId="0" pageNumber="309">, 20</figureCitation>
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<figureCitation id="2BB0560923E4B59D0312F213E6D96E13" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">, 25</figureCitation>
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<figureCitation id="E37EED7F39EE0CEEFB0D8106607FEE5B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Live specimens of Metasarcus. A M. limachii sp. nov., male; B female; C Metasarcus trispinosus sp. nov., male; D female; E M. vacafloresae sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309">, 26</figureCitation>
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<figureCitation id="398BC506F6A445DC083194D451B9B33B" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">, 31</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4EF554A31F2268B27B2BAAEF774D847E" pageId="0" pageNumber="309" type="reference_group">
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<paragraph id="7281321C1E055C9921A8892C37BF7A39" pageId="0" pageNumber="309">
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<taxonomicName id="203A1B2C8FE99D56810B585D566F75AD" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Metasarcus</taxonomicName>
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Roewer, 1913: 304 (desc);
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<bibRefCitation id="798EA6F575A69EACE14E240B3D7F995A" author="Roewer, CF" journalOrPublisher="Gustav Fischer, Jena, 1" pageId="0" pageNumber="309" refId="B98" refString="Roewer, CF, 1923. Die Weberknechte der Erde. Systematische Bear-beitung der bisher bekannten Opiliones. Gustav Fischer, Jena, 1" title="Die Weberknechte der Erde. Systematische Bear-beitung der bisher bekannten Opiliones." year="1923">Roewer 1923</bibRefCitation>
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: 508 (key), 517 (rdesc);
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<bibRefCitation id="BD00506A376143EE8DC363D6732B3DE4" author="Mello-Leitao, CFd" journalOrPublisher="Revista do Museu Paulista" pageId="0" pageNumber="309" pagination="327 - 383" refId="B72" refString="Mello-Leitao, CFd, 1926. Notas sobre Opiliones Laniatores sul-americanos. Revista do Museu Paulista 14: 327 - 383" title="Notas sobre Opiliones Laniatores sul-americanos." volume="14" year="1926">
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<normalizedToken id="74EF52962BBAEEE66CBF4F3378B50DF8" originalValue="Mello-leitão">Mello-leitao</normalizedToken>
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1926
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</bibRefCitation>
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: 34 (key); 1932: 408 (rdesc); 1935: 108 (cit.);
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<bibRefCitation id="C9A01649A2326BF2AC1339E5BECEDB1F" author="Soares, BAM" journalOrPublisher="Arquivos de Zoologia do Estado de Sao Paulo" pageId="0" pageNumber="309" pagination="149 - 240" refId="B115" refString="Soares, BAM, Soares, HEM, 1949. Monografia dos generos de opilioes neotropicos II. Arquivos de Zoologia do Estado de Sao Paulo 7 (2): 149 - 240" title="Monografia dos generos de opilioes neotropicos II." volume="7" year="1949">Soares and Soares 1949</bibRefCitation>
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: 234 (rdesc);
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<bibRefCitation id="D8364191C06C87FCF2000CF3F908E1E2" DOI="https://doi.org/10.1080/03946975.1994.10539264" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" pagination="343 - 353" refId="B50" refString="Kury, AB, 1994. Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores). Tropical Zoology 7 (2): 343 - 353, DOI: https://doi.org/10.1080/03946975.1994.10539264" title="Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores)." url="https://doi.org/10.1080/03946975.1994.10539264" volume="7" year="1994">Kury 1994</bibRefCitation>
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: 349 (syst);
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<bibRefCitation id="08A4B4DD15A3FDD56B3353619C082647" DOI="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" author="Kury, AB" journalOrPublisher="Zoo-logical Journal of Linnean Society" pageId="0" pageNumber="309" pagination="143 - 162" refId="B58" refString="Kury, AB, Maury, EA, 1998. A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae). Zoo-logical Journal of Linnean Society 123: 143 - 162, DOI: https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" title="A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae)." url="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" volume="123" year="1998">Kury and Maury 1998</bibRefCitation>
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: 145 (cit);
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<bibRefCitation id="1B5E78EA78FFBDC62D81082B098619AF" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 145 (cat): Kury & Villarreal, 2015: 5, 14 (syst).
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<emphasis id="BAC8ED977B7E08DEA713B8D85D8F87FA" bold="true" pageId="0" pageNumber="309">Type-species</emphasis>
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:
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<taxonomicName id="01A8AF6E5DC2B2D4AB17EB94227D376D" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">Metasarcus bolivianus</taxonomicName>
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Roewer, 1913 (by monotypy).
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</paragraph>
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<paragraph id="E10F90E90145AD2B624106DA1FC3808F" pageId="0" pageNumber="309">
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<taxonomicName id="3FE94EAA28948FBD2C1D0E325D8BE3D6" authorityName="Roewer" authorityYear="1931" class="Arachnida" family="Metasarcidae" genus="Metarascus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metarascus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Metarascus</taxonomicName>
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[lapsus calami]:
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<bibRefCitation id="588C53875832DEC79BF5ACC9B5D526EE" author="Roewer, CF" journalOrPublisher="Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen" pageId="0" pageNumber="309" pagination="101 - 164" refId="B101" refString="Roewer, CF, 1931. Weitere Weberknechte V. (5. Ergaenzung der Weber-knechte der Erde, 1923). Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 28 (2-3): 101 - 164" title="Weitere Weberknechte V. (5. Ergaenzung der Weber-knechte der Erde, 1923)." volume="28" year="1931">Roewer 1931</bibRefCitation>
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: 107 (key).
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</paragraph>
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<paragraph id="9A373E3D9068AB4B78D6E02C98E538BD" pageId="0" pageNumber="309">
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<taxonomicName id="886FFBC2926EEFAC1C34A36B3094428E" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chaconatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chaconatus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Chaconatus</taxonomicName>
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Roewer, 1929: 275 (key), 276 (desc);
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<bibRefCitation id="E75DE037901BBAC228EE60A9B8EC1837" author="Mello-Leitao, CFd" journalOrPublisher="Revista do Museu Paulista" pageId="0" pageNumber="309" pagination="1 - 505" refId="B73" refString="Mello-Leitao, CFd, 1932. Opilioes do Brasil. Revista do Museu Paulista 17 (2): 1 - 505" title="Opilioes do Brasil." volume="17" year="1932">
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<normalizedToken id="E538408B359282654CFA530CBDC9139E" originalValue="Mello-leitão">Mello-leitao</normalizedToken>
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1932
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</bibRefCitation>
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: 108 (rdesc);
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<bibRefCitation id="277EDFC736DEB86E7BC178DCED26D16D" author="Juberthie, C" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="309" refId="B43" refString="Juberthie, C, 1970. IX Opilions des Galapagos: Galanomma microphtalma gen. nov.sp. nov. (Gonyleptidae). In. Resultats scientifiques de la mission zoologique belge aux iles Galapagos et en Ecuador (Leleup, 1964-5), Royal Museum for Central Africa - Tervuren 2: 137-153." title="IX Opilions des Galapagos: Galanomma microphtalma gen. nov. sp. nov. (Gonyleptidae). In. Resultats scientifiques de la mission zoologique belge aux iles Galapagos et en Ecuador (Leleup, 1964 - 5), Royal Museum for Central Africa - Tervuren 2: 137 - 153." year="1970">Juberthie 1970</bibRefCitation>
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: 142 (cit);
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<bibRefCitation id="B738939C8FE44997ADAF6C1DA145C2DE" author="Soares, HEM" journalOrPublisher="Revista Brasileira de Entomologia" pageId="0" pageNumber="309" pagination="1 - 14" refId="B116" refString="Soares, HEM, Soares, BAM, Jim, RLS, 1992. Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae). Revista Brasileira de Entomologia 36 (1): 1 - 14" title="Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae)." volume="36" year="1992">Soares, Soares and Jim 1992</bibRefCitation>
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: 5 (cat);
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<bibRefCitation id="AEA6737E188D5D53CECF6566F2A2E034" DOI="https://doi.org/10.1080/03946975.1994.10539264" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" pagination="343 - 353" refId="B50" refString="Kury, AB, 1994. Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores). Tropical Zoology 7 (2): 343 - 353, DOI: https://doi.org/10.1080/03946975.1994.10539264" title="Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores)." url="https://doi.org/10.1080/03946975.1994.10539264" volume="7" year="1994">Kury 1994</bibRefCitation>
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: 349 (syst);
|
|
<bibRefCitation id="82BFE8E2CEC65A2C5602A844E44B18CF" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
|
|
: 145 (cat).
|
|
<emphasis id="B5BCDB0A0A5D8EE52168EAF7A8B53F19" bold="true" pageId="0" pageNumber="309">Type-species</emphasis>
|
|
:
|
|
<taxonomicName id="C02BD2D65A7F87144B0BFDDCEB279CA6" class="Arachnida" family="Metasarcidae" genus="Chaconatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chaconatus armatipalpus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="armatipalpus">Chaconatus armatipalpus</taxonomicName>
|
|
Roewer, 1929 (by monotypy). Synonymy established by
|
|
<bibRefCitation id="7A77D0E51C066985DA5EA5BC040AD94D" DOI="https://doi.org/10.1080/03946975.1994.10539264" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" pagination="343 - 353" refId="B50" refString="Kury, AB, 1994. Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores). Tropical Zoology 7 (2): 343 - 353, DOI: https://doi.org/10.1080/03946975.1994.10539264" title="Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores)." url="https://doi.org/10.1080/03946975.1994.10539264" volume="7" year="1994">Kury 1994</bibRefCitation>
|
|
: 349.
|
|
</paragraph>
|
|
<paragraph id="956E46156FA0C9FE48EA266243260E20" pageId="0" pageNumber="309">
|
|
<taxonomicName id="1053F83E31B40C0BDDA146969D95D21D" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Chacoikeontus</taxonomicName>
|
|
Roewer, 1929: 275 (key), 278 (desc);
|
|
<bibRefCitation id="6600CEA4890BD8074B215DD1152ED328" author="Mello-Leitao, CFd" journalOrPublisher="Revista do Museu Paulista" pageId="0" pageNumber="309" pagination="1 - 505" refId="B73" refString="Mello-Leitao, CFd, 1932. Opilioes do Brasil. Revista do Museu Paulista 17 (2): 1 - 505" title="Opilioes do Brasil." volume="17" year="1932">
|
|
<normalizedToken id="B5B4EBB49B1A23B066F5B03CD850EC52" originalValue="Mello-leitão">Mello-leitao</normalizedToken>
|
|
1932
|
|
</bibRefCitation>
|
|
: 104 (key), 108 (rdesc);
|
|
<bibRefCitation id="42BECB5D4C70AB161D0FF0012819A2A1" author="Juberthie, C" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="309" refId="B43" refString="Juberthie, C, 1970. IX Opilions des Galapagos: Galanomma microphtalma gen. nov.sp. nov. (Gonyleptidae). In. Resultats scientifiques de la mission zoologique belge aux iles Galapagos et en Ecuador (Leleup, 1964-5), Royal Museum for Central Africa - Tervuren 2: 137-153." title="IX Opilions des Galapagos: Galanomma microphtalma gen. nov. sp. nov. (Gonyleptidae). In. Resultats scientifiques de la mission zoologique belge aux iles Galapagos et en Ecuador (Leleup, 1964 - 5), Royal Museum for Central Africa - Tervuren 2: 137 - 153." year="1970">Juberthie 1970</bibRefCitation>
|
|
: 142 (cit);
|
|
<bibRefCitation id="5E8040EB203242C5AB34D70BFA4F41BF" author="Soares, HEM" journalOrPublisher="Revista Brasileira de Entomologia" pageId="0" pageNumber="309" pagination="1 - 14" refId="B116" refString="Soares, HEM, Soares, BAM, Jim, RLS, 1992. Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae). Revista Brasileira de Entomologia 36 (1): 1 - 14" title="Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae)." volume="36" year="1992">Soares, Soares and Jim 1992</bibRefCitation>
|
|
: 3 (key), 5 (rdesc, cat);
|
|
<bibRefCitation id="FBD7CABE5C77CEF6186E2F3F82F9CE67" DOI="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" author="Kury, AB" journalOrPublisher="Zoo-logical Journal of Linnean Society" pageId="0" pageNumber="309" pagination="143 - 162" refId="B58" refString="Kury, AB, Maury, EA, 1998. A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae). Zoo-logical Journal of Linnean Society 123: 143 - 162, DOI: https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" title="A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae)." url="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" volume="123" year="1998">Kury and Maury 1998</bibRefCitation>
|
|
: 145 (syst);
|
|
<bibRefCitation id="EA8F701A285C6C6B2ACEFE6C6E081CDD" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
|
|
: 144 (cat);
|
|
<bibRefCitation id="BDCF246066C671A19A52C715C050025F" DOI="https://doi.org/10.1111/j.1096-3642.2008.00484.x" author="Yamaguti, HY" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="309" pagination="319 - 362" refId="B127" refString="Yamaguti, HY, Pinto-da-Rocha, R, 2009. Taxonomic review of Bourguyiinae, cladistic analysis, and a new hypothesis of biogeographic relantioships of the Brazilian Atlantic Rainforest (Arachnida: Opiliones, Gonyleptidae). Zoological Journal of the Linnean Society 156: 319 - 362, DOI: https://doi.org/10.1111/j.1096-3642.2008.00484.x" title="Taxonomic review of Bourguyiinae, cladistic analysis, and a new hypothesis of biogeographic relantioships of the Brazilian Atlantic Rainforest (Arachnida: Opiliones, Gonyleptidae)." url="https://doi.org/10.1111/j.1096-3642.2008.00484.x" volume="156" year="2009">Yamaguti and Pinto-da-Rocha 2009</bibRefCitation>
|
|
: 321 (syst); Kury & Villarreal, 2015: 23, 26 (cit).
|
|
<emphasis id="32FBAAB11ACDCC405AF6054182EA4AED" bold="true" pageId="0" pageNumber="309">Type-species</emphasis>
|
|
:
|
|
<taxonomicName id="43EB6C7890EB05DF2C6B4B902BAB3D41" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">Chacoikeontus clavifemur</taxonomicName>
|
|
Roewer, 1929 (by monotypy). Synonymy established by
|
|
<bibRefCitation id="C51349350321E8B1ED8567C0315270FD" DOI="https://doi.org/10.1016/j.jcz.2018.12.005" author="Townsend, VR" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="309" pagination="101 - 109" refId="B120" refString="Townsend, VR, Perez-Gonzalez, A, Proud, DN, 2019. Putative mating plugs of harvestmen (Opiliones, Laniatores). Zoologischer Anzeiger 278: 101 - 109, DOI: https://doi.org/10.1016/j.jcz.2018.12.005" title="Putative mating plugs of harvestmen (Opiliones, Laniatores)." url="https://doi.org/10.1016/j.jcz.2018.12.005" volume="278" year="2019">Townsend et al. 2019</bibRefCitation>
|
|
: 102
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="92A7EDC50510AB5F225DEF5382311BE2" pageId="0" pageNumber="309" type="diagnosis">
|
|
<paragraph id="4EB23958574F598D6BE665375A813A1F" pageId="0" pageNumber="309">Diagnosis.</paragraph>
|
|
<paragraph id="5ADAAFA9874E51A837F400EE7551F23C" pageId="0" pageNumber="309">
|
|
<taxonomicName id="70F794409CC60385F412C362ED107DAB" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="A39D5A285D8E296664D273034BE73B83" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
|
|
</taxonomicName>
|
|
can be diagnosed from other
|
|
<taxonomicName id="3325EA9B6AD9F50A314D45A123E72EEC" authorityName="Kury" authorityYear="1994" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="family">Metasarcidae</taxonomicName>
|
|
genera with long legs (except
|
|
<taxonomicName id="17598AE84D64120AB5E5840457DBC8AB" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="5459E731542D847FA9BCB18E200658DF" italics="true" pageId="0" pageNumber="309">Ayacucho</emphasis>
|
|
</taxonomicName>
|
|
) by the combination of the following features: gamma-type; gamma-P-type or kappa-type DSS; ocularium unarmed; area III with two spines. Half of
|
|
<taxonomicName id="0DAFBE603B141EBD9C6ACE942415EEE8" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="0F9BFEDD981288B3566D48A6D127206A" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
|
|
</taxonomicName>
|
|
species present a spine, long projection or trifid projection in free tergite III, present in no other
|
|
<taxonomicName id="C9B8FC7A2F0BDFD2DD10E3E4F59050CC" authorityName="Kury" authorityYear="1994" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="family">Metasarcidae</taxonomicName>
|
|
species; coxa III with two apophyses; coxa IV reaching posterior margin of dorsal scutum or surpassing it; femur IV smooth or slightly granulate; penis stylus and VP thin thickness, with less than 10 MS C
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="AD2121E36D603E107D83250F2C8D7D9C" pageId="0" pageNumber="309" type="redescription">
|
|
<paragraph id="61D057ABBD725BF541AE6298CCC779F1" pageId="0" pageNumber="309">Redescription.</paragraph>
|
|
<paragraph id="2484BDCC5AD78B636B684BA65DC7AE34" pageId="0" pageNumber="309">
|
|
Gamma-type DSS (
|
|
<taxonomicName id="5AF27AF6887703CEA341E169B915B979" class="Arachnida" kingdom="Animalia" lsidName="M. bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">
|
|
<emphasis id="A7F67C32B218555F1F75CC5E63DCB20F" italics="true" pageId="0" pageNumber="309">M. bolivianus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="A55D95C91DA939F41A275ECC84B990EB" class="Arachnida" kingdom="Animalia" lsidName="M. fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
|
|
<emphasis id="9CB274B17F4313B321E90F79C053F6FD" italics="true" pageId="0" pageNumber="309">M. fellinii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="C492A2F0523F6CE6DB4DB80E5710D508" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
; Fig.
|
|
<figureCitation id="C36CF452A19B7F47FB839518C1EDB88B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6E</figureCitation>
|
|
), gamma-P-type DSS (
|
|
<taxonomicName id="21B6170563A63DF6526AF8E45CBF5F3B" class="Arachnida" kingdom="Animalia" lsidName="M. beni" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="beni">
|
|
<emphasis id="0BD7B10587B67960958A786F9660132D" italics="true" pageId="0" pageNumber="309">M. beni</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="84F502CD3DF39E9790DF98F288F10D17" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
,
|
|
<taxonomicName id="C7737389934CCB43A84F5F83C8E5CA50" class="Arachnida" kingdom="Animalia" lsidName="M. bergmani" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bergmani">
|
|
<emphasis id="625418E8AD276B7ED64A069C8537C5F2" italics="true" pageId="0" pageNumber="309">M. bergmani</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="51B6649B7C2CD58421596EC3082EEF10" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
,
|
|
<taxonomicName id="65F507C0F60B30907ED20B7873AD38BD" class="Arachnida" kingdom="Animalia" lsidName="M. limachii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="limachii">
|
|
<emphasis id="3CC20371F593DD6C54B2ACC89B757633" italics="true" pageId="0" pageNumber="309">M. limachii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="4001A057F9687736BF0616B6ABD39F88" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
and
|
|
<taxonomicName id="0006B2E15F4FFEB0B0BB9A3CAB0F84D8" class="Arachnida" kingdom="Animalia" lsidName="M. trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
|
|
<emphasis id="E518C70C998C06A0F10D522CD18B4FD5" italics="true" pageId="0" pageNumber="309">M. trispinosus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="511B7340AE7A1C20BC76E5B6DFF9FDE6" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
; Fig.
|
|
<figureCitation id="BE5AA32E558D266D80F7A91D69870D46" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6A, B, G, H</figureCitation>
|
|
) or kappa-type DSS (
|
|
<taxonomicName id="4E404FDEA75CAE81C977D8100FE9DF37" class="Arachnida" kingdom="Animalia" lsidName="M. clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
|
|
<emphasis id="C7C8276AC0DEFDEBD4704C27BB452B48" italics="true" pageId="0" pageNumber="309">M. clavifemur</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="FAAF28B078529D9216E548A6323772BD" class="Arachnida" kingdom="Animalia" lsidName="M. kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
|
|
<emphasis id="BB91A1B74EFD409C5EE9A64F2EF52017" italics="true" pageId="0" pageNumber="309">M. kurosawai</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B488C8A19357EDAB1632E886ADE937A0" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
and
|
|
<taxonomicName id="F73DDA4CA0A920925E33D01F1F3C9D53" class="Arachnida" kingdom="Animalia" lsidName="M. vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
|
|
<emphasis id="461C1F08D3D752277A5A2AA1AC0F810D" italics="true" pageId="0" pageNumber="309">M. vacafloresae</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="1868F3E2B399982594D957253566FA14" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
; Fig.
|
|
<figureCitation id="77E476FE68ED613E4431D623E42744A2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6D, F, I</figureCitation>
|
|
). Ocularium low, medially depressed. Ocularium unarmed, small tuberculate or smooth (Fig.
|
|
<figureCitation id="B14A7AC4B62D579713C8A1B11D844199" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6</figureCitation>
|
|
). Areas of dorsal scutum moderately or densely tuberculate (
|
|
<taxonomicName id="929C63697EAB92AE645B4868A7431A1C" class="Arachnida" kingdom="Animalia" lsidName="M. fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
|
|
<emphasis id="B3F055C50053EE40128618E23E8945E8" italics="true" pageId="0" pageNumber="309">M. fellinii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="A2E383289604F36243663EFD269D87B2" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
). Area I undivided (Fig.
|
|
<figureCitation id="1F7C9287DCB2610184038E66978E3AC9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6G-I</figureCitation>
|
|
) or divided in two halves (Fig.
|
|
<figureCitation id="4F09BFCCA98983602094375BF0899CEF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6A-F</figureCitation>
|
|
). Area III armed with two high spines in most species (e.g. Fig.
|
|
<figureCitation id="BBE9B1124D5E1172CD9224EF0069BCE8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6D</figureCitation>
|
|
), with two tubercles (
|
|
<taxonomicName id="D92BAD7B660CDC5E115E8491A1DA4D20" class="Arachnida" kingdom="Animalia" lsidName="M. trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
|
|
<emphasis id="9BC0A723F8E2245FB3B20EF87CDD00D1" italics="true" pageId="0" pageNumber="309">M. trispinosus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="ED69D3AA9F54DFCD8EE414E57ADFF2BA" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
; Fig.
|
|
<figureCitation id="C3BE3767B0FCEEE7B698C884DD2A7C83" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6H</figureCitation>
|
|
) or unarmed (
|
|
<taxonomicName id="0625ED09EEF57D590D78C70F505A268C" class="Arachnida" kingdom="Animalia" lsidName="M. vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
|
|
<emphasis id="C5B3D4203CEA86A4E7C041BDE4A52005" italics="true" pageId="0" pageNumber="309">M. vacafloresae</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="AD12C8A155D13A643A19A4EA8B3771D1" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
; Fig.
|
|
<figureCitation id="0E2AF4B2756CEBF01F6A16D75EF79913" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6I</figureCitation>
|
|
). Posterior margin unarmed. Coxa IV reaching area III-IV or posterior margin. Coxa IV unarmed or with retrolateral armature (
|
|
<taxonomicName id="00420B90B4180EBAF0E10A22A9E9F4EC" class="Arachnida" kingdom="Animalia" lsidName="M. bergmani" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bergmani">
|
|
<emphasis id="6778FA264178C03A2EC1217C90AF3433" italics="true" pageId="0" pageNumber="309">M. bergmani</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="AD7BE0EF739BAF2F572FE542FE72D967" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
,
|
|
<taxonomicName id="85FE98760EF1AA94B5275C3E95C62B9D" lsidName="A. limachii" pageId="0" pageNumber="309" rank="species" species="limachii">
|
|
<emphasis id="A18C88AB1AB8B0A6FB7C48126CBCB1AD" italics="true" pageId="0" pageNumber="309">A. limachii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="5A9F509C452AEFC2208088B3E9F2D7F6" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
; Fig.
|
|
<figureCitation id="ED9DAE704208D5CEF2ACCA6EEFE5E060" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6B, G</figureCitation>
|
|
). Femur IV much longer than dorsal scutum length. (Figs
|
|
<figureCitation id="D25EC6FA143A521CDC7F5B58D1EDF601" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10</figureCitation>
|
|
,
|
|
<figureCitation id="668891DC624AE52C054F55A641D2B74F" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">25</figureCitation>
|
|
,
|
|
<figureCitation id="4F5A75893B9C0CD971A70BDCAFA6DF2B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Live specimens of Metasarcus. A M. limachii sp. nov., male; B female; C Metasarcus trispinosus sp. nov., male; D female; E M. vacafloresae sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309">26</figureCitation>
|
|
) Less than 10 MS C. Penial stylus thin. Penial ventral plate thin (Figs
|
|
<figureCitation id="52BB1796F2177C842654658F9681B86B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14C-H</figureCitation>
|
|
,
|
|
<figureCitation id="2905520E9D18E0AB0B17CEB8F86910AA" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. bergmani sp. nov.; D - F M. clavifemur (Roewer, 1929); G - I M. fellinii sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309">19</figureCitation>
|
|
,
|
|
<figureCitation id="3EAEDFE9A341CB2B9BC31AC057A77BCD" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. trispinosus sp. nov.; D - F M. vacafloresae sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure20" httpUri="https://binary.pensoft.net/fig/729894" pageId="0" pageNumber="309">20</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="662CC2EAF41CEFE4D3A9F0B53B24A81B" pageId="0" pageNumber="309" type="distribution">
|
|
<paragraph id="6BA99D7F0DB86B9B84A4D269B281F8D1" pageId="0" pageNumber="309">Distribution.</paragraph>
|
|
<paragraph id="555F3B7485AA69397835945B5E039B85" pageId="0" pageNumber="309">
|
|
(Fig.
|
|
<figureCitation id="DCBE4C8190181BCC4E6628A835A3D2D0" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">31</figureCitation>
|
|
) BOLIVIA. Beni, Cochabamba, La Paz and Tarija.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="55AFE419EAC0ADEC0B73F36EA9B23F05" pageId="0" pageNumber="309" type="remarks">
|
|
<paragraph id="CA90A9F4A522796AAA6F7B60BEFC83F9" pageId="0" pageNumber="309">Remarks.</paragraph>
|
|
<paragraph id="8CC4C73D7A2C945BD8672D67775AC91A" pageId="0" pageNumber="309">
|
|
<bibRefCitation id="DAF18461D32E82FA30AF169A2F0B3755" DOI="https://doi.org/10.1016/j.jcz.2018.12.005" author="Townsend, VR" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="309" pagination="101 - 109" refId="B120" refString="Townsend, VR, Perez-Gonzalez, A, Proud, DN, 2019. Putative mating plugs of harvestmen (Opiliones, Laniatores). Zoologischer Anzeiger 278: 101 - 109, DOI: https://doi.org/10.1016/j.jcz.2018.12.005" title="Putative mating plugs of harvestmen (Opiliones, Laniatores)." url="https://doi.org/10.1016/j.jcz.2018.12.005" volume="278" year="2019">Townsend et al. (2019</bibRefCitation>
|
|
: pp. 102-103, 105-106), in a non-taxonomic or systematic study, cited
|
|
<taxonomicName id="B94FBEBB9A92DED91C527365964A62E6" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
|
|
<emphasis id="87413BE316D7A7EDF066EBF19657C698" italics="true" pageId="0" pageNumber="309">Chacoikeontus clavifemur</emphasis>
|
|
</taxonomicName>
|
|
as
|
|
<taxonomicName id="0A4D3E72CFB7C9860B28B9E7CC091772" baseAuthorityName="Roewer" baseAuthorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
|
|
<emphasis id="4E9EED1EBD0AC2B4438C8D91A4A90BF6" italics="true" pageId="0" pageNumber="309">Metasarcus clavifemur</emphasis>
|
|
</taxonomicName>
|
|
(Roewer, 1929). The authors did not provide justification for or specify it was a new combination or nomenclatural act, did not mention the genus
|
|
<taxonomicName id="5E5C0657501BA6E4DD8B5FCAD42B3797" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="F67994389E8B37C7ABD6E7BF389643B3" italics="true" pageId="0" pageNumber="309">Chacoikeontus</emphasis>
|
|
</taxonomicName>
|
|
(and its consequent status as a junior subjective synonym) and did not discuss the genus
|
|
<taxonomicName id="221831C705407F45575527B9DB2C6FDC" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B41F4258CEA47389E3E8B4B3205C105E" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
|
|
</taxonomicName>
|
|
with the inclusion of the species. Once the new combination has been published, the nomenclatural act is valid and herein confirmed (see below). As
|
|
<taxonomicName id="EF4C0E55393787E4FA03B3D3430C582D" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
|
|
<emphasis id="44D1F9A6586F5014649516526617C2F6" italics="true" pageId="0" pageNumber="309">Chacoikeontus clavifemur</emphasis>
|
|
</taxonomicName>
|
|
is the type-species of the genus, by implication
|
|
<taxonomicName id="4A35590B2FC355C4D48CFF101EDDBD99" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B823A569FF954DF378207E0BA71B1EF1" italics="true" pageId="0" pageNumber="309">Chacoikeontus</emphasis>
|
|
</taxonomicName>
|
|
was considered a junior subjective synonym of
|
|
<taxonomicName id="B9E9725B2837B250CB9BCA3ECFDBA73F" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="53EB5D86A29D5913A4FBDFF726E4DF75" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9A7C4FA069678BD1EA3BCDA4D304A773" pageId="0" pageNumber="309" type="species composition">
|
|
<paragraph id="412B3F442CEB022E1DE44E15B4BC4E01" pageId="0" pageNumber="309">Species composition.</paragraph>
|
|
<paragraph id="7E33DD58B826C621E8DB83470DC2234B" pageId="0" pageNumber="309">
|
|
<taxonomicName id="11606A3F778523E4208770C79110B14B" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus beni" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="beni">
|
|
<emphasis id="BDCBE4F94B3EC35E471A4B30DF80CF47" italics="true" pageId="0" pageNumber="309">Metasarcus beni</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="3C9AA5B3AC5326A78FD05F5C03D5EAA9" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="4394FD8CF494F03A345FC9316FDD13B5" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus bergmani" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bergmani">
|
|
<emphasis id="0302D4BF967D61C49419DC664DA2EC01" italics="true" pageId="0" pageNumber="309">Metasarcus bergmani</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="BFB8DEB19BD4DCAAEF7F0C338D268092" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="1D476C1DBF73B3CE38B2665D8CA26027" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">
|
|
<emphasis id="704E8C2BE367DB7B6607F9067E1C3FE8" italics="true" pageId="0" pageNumber="309">Metasarcus bolivianus</emphasis>
|
|
</taxonomicName>
|
|
Roewer, 1913;
|
|
<taxonomicName id="E7EB390091D1309B8EA0EC8622C75961" baseAuthorityName="Roewer" baseAuthorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
|
|
<emphasis id="3D33109F6033D12FA19DDEA3187A3C7D" italics="true" pageId="0" pageNumber="309">Metasarcus clavifemur</emphasis>
|
|
</taxonomicName>
|
|
(Roewer, 1929);
|
|
<taxonomicName id="E140AC369373D2F1D2CBB369534ABAA9" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
|
|
<emphasis id="BD02F6E1F4D46F4FD4C42B6F138B2C0B" italics="true" pageId="0" pageNumber="309">Metasarcus fellinii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="AE620EC0582541FFD4A36AB7B59904DA" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="E852977040B23FC7B3647B8554B75A76" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
|
|
<emphasis id="03C86C4C72FBFD2B19099E30E5408746" italics="true" pageId="0" pageNumber="309">Metasarcus kurosawai</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="D749F959EE53155F2B10FBFE7C0E6266" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="87593BF2AD6E58C18394AB64E33B285D" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus limachii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="limachii">
|
|
<emphasis id="74633BD9CB9E1DE162EBDBD71F9AF98E" italics="true" pageId="0" pageNumber="309">Metasarcus limachii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="FE8F5CB7691BC1651C44CE08FD507779" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName id="B216E42752590B297180C2B638172766" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
|
|
<emphasis id="4951CE3EDF9AB887791BB653FCE90644" italics="true" pageId="0" pageNumber="309">Metasarcus trispinosus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="1A3D3F52B996B0F8AADBEB215CEBC5D9" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
and
|
|
<taxonomicName id="C0656C7E8DD08AED45649EE50ABB6432" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
|
|
<emphasis id="E56AAB4C2353F083C75BA5E40B974DE1" italics="true" pageId="0" pageNumber="309">Metasarcus vacafloresae</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="5721F88F58A9054A23D2A98FB98F1269" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |