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<taxonomicName id="6A894D5F1F1B9560FDCAFB5DDF9DFAFD" ID-CoL="MFM2" authority="Morawitz, 1887" authorityName="F. Morawitz" authorityYear="1887" box="[589,999,1276,1303]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FDCAFB5DDF5EFAFC" bold="true" box="[589,804,1276,1303]" italics="true" pageId="89" pageNumber="90">Bombus keriensis</emphasis>
<bibRefCitation id="C9184B2D1F1B9560FCACFB5DDF9DFAFD" author="Morawitz F. F." box="[811,999,1276,1302]" pageId="89" pageNumber="90" pagination="195 - 229" refId="ref68420" refString="Morawitz F. F. 1887. Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva 20 (1886): 195 - 229." type="journal article" year="1887">Morawitz, 1887</bibRefCitation>
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<subSubSection id="E59365571F1B9560FDDCFABEDFA2FAD2" box="[603,984,1311,1337]" pageId="89" pageNumber="90" type="description">
<paragraph id="AD3636DC1F1B9560FDDCFABEDFA2FAD2" blockId="89.[589,999,1276,1337]" box="[603,984,1311,1337]" pageId="89" pageNumber="90">
<figureCitation id="35B22A591F1B9560FDDCFABEDECFFAD2" box="[603,693,1311,1337]" captionStart="Figs 14‒16" captionStartId="25.[189,241,1956,1982]" captionTargetBox="[288,1321,265,1901]" captionTargetId="figure-11@25.[262,1325,265,1917]" captionTargetPageId="25" captionText="Figs 14‒16. Maps of sequenced samples as in Figs 12–13. 14. The tanguticus-group and the lapidarius- group. 15. The sichelii-group. 16. The keriensis-group." figureDoi="http://doi.org/10.5281/zenodo.4064340" httpUri="https://zenodo.org/record/4064340/files/figure.png" pageId="89" pageNumber="90">Figs 16</figureCitation>
, 
<figureCitation id="35B22A591F1B9560FD46FABEDF51FAD2" box="[705,811,1311,1337]" captionStart="Figs 139–180" captionStartId="42.[189,241,1497,1523]" captionTargetBox="[369,1221,268,1431]" captionTargetId="figure-11@42.[366,1221,265,1467]" captionTargetPageId="42" captionText="Figs 139–180. Simplified diagrams for the colour patterns of the hair on the dorsum for the species from the integrative analysis. The dorsum is divided into regions, each of which shows only the predominant or most apparent colour for that region using a simplified colour palette, with olive indicating a mixture of black and yellow hair, and grey indicating a mixture of black and white hair. The keriensis-group. 139. Queen, China-Sichuan. 140. Worker, China-Sichuan. 141. Queen, China-Sichuan. 142. Queen, China-Sichuan. 143. Worker, China-Xizang. 144. Queen, India-Kashmir. 145. Male, China-Qinghai. 146. Male, India-Kashmir. 147. Male, China-Gansu. 148. Queen, Armenia. 149. Queen, Iran. 150. Male, Georgia. 151. Worker, China-Qinghai. 152. Male, China-Qinghai. 153. Queen, Mongolia. 154. Worker, Mongolia. 155. Male, Mongolia. 156. Male, Mongolia. 157. Queen, China-Sichuan. 158. Worker, China-Qinghai. 159. Worker, China-Qinghai. 160. Male, China-Qinghai. 161. Male, China-Qinghai. 162. Male, China-Qinghai. 163. Queen, Pakistan. 164. Queen, Pakistan. 165. Queen, Pakistan. 166. Queen, Pakistan. 167. Queen, Afghanistan. 168. Queen, Pakistan. 169. Queen, Pakistan. 170. Queen, India-Kashmir. 171. Male, India-Kashmir. 172. Male, India-Kashmir. 173. Queen, China- Xinjiang. 174. Queen, Kyrgyzstan. 175. Queen, Kyrgyzstan. 176. Queen, Mongolia. 177. Queen, India- Kashmir. 178. Male, Kazakhstan. 179. Male, China-Xinjiang. 180. Male, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.4064358" httpUri="https://zenodo.org/record/4064358/files/figure.png" pageId="89" pageNumber="90">163–172</figureCitation>
, 
<figureCitation id="35B22A591F1B9560FCB0FABEDF1CFAD2" box="[823,870,1311,1337]" captionStart="Figs 199‒204" captionStartId="45.[189,241,1852,1878]" captionTargetBox="[230,1342,827,1806]" captionTargetId="figure-159@45.[194,1393,771,1810]" captionTargetPageId="45" captionText="Figs 199‒204. Morphology of the male genitalia for species of the subgenus Melanobombus von Dalla Torre, 1880 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 199. Bombus alagesianus Reinig, 1930 stat. rev., Georgia. 200. B. tibeticus sp. nov., China-Qinghai. 201. B. incertoides Vogt, 1911 stat. rev., Mongolia. 202. B. qilianensis sp. nov., China-Qinghai. 203. B. keriensis Morawitz, 1887, India-Kashmir. 204. B. separandus Vogt, 1909 stat. rev., Kyrgyzstan. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4064364" httpUri="https://zenodo.org/record/4064364/files/figure.png" pageId="89" pageNumber="90">203</figureCitation>
, 
<figureCitation id="35B22A591F1B9560FCF6FABEDFDAFAD2" box="[881,928,1311,1337]" captionStart="Figs 209–210" captionStartId="93.[189,241,1884,1910]" captionTargetBox="[190,1374,318,1788]" captionTargetId="figure-11@93.[189,1398,318,1792]" captionTargetPageId="93" captionText="Figs 209–210. Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau. 209. Bombus keriensis Morawitz, 1887 from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405). 210. B. separandus Vogt, 1909 stat. rev. from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="89" pageNumber="90">209</figureCitation>
, 
<figureCitation id="35B22A591F1B9560FC2BFABEDFA2FAD2" box="[940,984,1311,1337]" captionStart="Figs 211–212" captionStartId="100.[189,241,1742,1768]" captionTargetBox="[319,1269,265,1703]" captionTargetId="figure-10@100.[319,1269,265,1703]" captionTargetPageId="100" captionText="Figs 211–212. Maps of sequenced samples showing the most frequent colour patterns locally. 211. The sichelii-complex, consisting of the single polytypic species B. sichelii Radoszkowski, 1859 (inset: B. sichelii worker, China-Neimenggu, photo PW). 212. The keriensis-complex, consisting of B. alagesianus Reinig, 1930 stat. rev. (blue spots), B. tibeticus sp. nov. (dark green spots), B. incertoides Vogt, 1911 stat. rev. (light green spots), B. qilianensis sp. nov. (yellow spots), B. keriensis Morawitz, 1887 (orange spots), and B. separandus Vogt, 1909 stat. rev. (red spots) (inset: B.qilianensis sp. nov. queen, China-Sichuan, photo PW). Maps projected onto a sphere in ArcGIS using World_Shaded_Relief basemap © 2014 ESRI and showing boundaries between countries as recognised by the UN." figureDoi="http://doi.org/10.5281/zenodo.4064372" httpUri="https://zenodo.org/record/4064372/files/figure.png" pageId="89" pageNumber="90">212</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1B9560FF3AFAC0DFD0FA0E" pageId="89" pageNumber="90" type="reference_group">
<paragraph id="AD3636DC1F1B9560FF3AFAC0DEEBFA90" blockId="89.[189,1014,1377,1509]" box="[189,657,1377,1403]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="8D9911F21F1B9560FF3AFAC0DEEBFA90" box="[189,657,1377,1403]" pageId="89" pageNumber="90">
<treatmentCitation id="2C2810CD1F1B9560FF3AFAC0DEF6FA90" author="Morawitz F. F." box="[189,652,1377,1403]" page="199" pageId="89" pageNumber="90" year="1887">
<taxonomicName id="6A894D5F1F1B9560FF3AFAC0DEF6FA90" ID-CoL="MFM2" authority="Morawitz, 1887: 199" authorityName="F. Morawitz" authorityPageNumber="199" authorityYear="1887" box="[189,652,1377,1403]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FF3AFAC0DDF1FA90" box="[189,395,1377,1403]" italics="true" pageId="89" pageNumber="90">Bombus keriensis</emphasis>
<bibRefCitation id="C9184B2D1F1B9560FE15FAC0DEF6FA90" author="Morawitz F. F." box="[402,652,1377,1403]" pageId="89" pageNumber="90" pagination="195 - 229" refId="ref68420" refString="Morawitz F. F. 1887. Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva 20 (1886): 195 - 229." type="journal article" year="1887">Morawitz, 1887: 199</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1B9560FF3AFA25DF8CFA75" blockId="89.[189,1014,1377,1509]" box="[189,1014,1412,1438]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="8D9911F21F1B9560FF3AFA25DF8CFA75" box="[189,1014,1412,1438]" pageId="89" pageNumber="90">
<treatmentCitation id="2C2810CD1F1B9560FF3AFA25DF88FA75" author="Skorikov A. S." box="[189,1010,1412,1438]" page="127" pageId="89" pageNumber="90" year="1914">
<taxonomicName id="6A894D5F1F1B9560FF3AFA25DF88FA75" ID-CoL="7K98T" authority="Skorikov, 1914 a: 127" authorityName="Skorikov" authorityPageNumber="127" authorityYear="1914" box="[189,1010,1412,1438]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="subSpecies" species="separandus" subSpecies="meridialis">
<emphasis id="9FFDEACE1F1B9560FF3AFA25DE66FA75" box="[189,540,1412,1438]" italics="true" pageId="89" pageNumber="90">Lapidariobombus separandus</emphasis>
subsp. 
<emphasis id="9FFDEACE1F1B9560FDF2FA25DE96FA75" box="[629,748,1412,1438]" italics="true" pageId="89" pageNumber="90">meridialis</emphasis>
<bibRefCitation id="C9184B2D1F1B9560FD74FA25DF88FA75" author="Skorikov A. S." box="[755,1010,1412,1438]" pageId="89" pageNumber="90" pagination="119 - 129" refId="ref70701" refString="Skorikov A. S. 1914 a. Les formes nouvelles des bourdons (Hymenoptera, Bombidae). VI. Russkoe entomologicheskoe Obozrenie 14: 119 - 129." type="journal article" year="1914">Skorikov, 1914a: 127</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1B9560FF3AFA06DECAFA2A" blockId="89.[189,1014,1377,1509]" box="[189,688,1447,1474]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="8D9911F21F1B9560FF3AFA06DECAFA2A" box="[189,688,1447,1474]" pageId="89" pageNumber="90">
<taxonomicName id="6A894D5F1F1B9560FF3AFA06DDE6FA2A" ID-CoL="MFX2" authorityName="Wang" authorityYear="1982" box="[189,412,1447,1473]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="trilineatus">
<emphasis id="9FFDEACE1F1B9560FF3AFA06DDE6FA2A" box="[189,412,1447,1473]" italics="true" pageId="89" pageNumber="90">Bombus trilineatus</emphasis>
</taxonomicName>
S.-F. 
<treatmentCitation id="2C2810CD1F1B9560FE67FA09DED7FA2A" author="Wang S. - F." box="[480,685,1447,1474]" page="441" pageId="89" pageNumber="90" year="1982">
<bibRefCitation id="C9184B2D1F1B9560FE67FA09DED7FA2A" author="Wang S. - F." box="[480,685,1447,1474]" pageId="89" pageNumber="90" pagination="427 - 447" refId="ref72195" refString="Wang S. - F. 1982. Hymenoptera: Apidae - Bombus. In: Qinghai-Tibet Plateau Comprehensive Scientific Investigation Team of the Chinese Academy of Sciences (ed.) Insects of Xizang: 427 - 447. Science Press, Beijing." type="book chapter" year="1982">Wang, 1982: 441</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1B9560FF3AFA6BDFD0FA0E" blockId="89.[189,1014,1377,1509]" box="[189,938,1482,1509]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="8D9911F21F1B9560FF3AFA6BDFD0FA0E" box="[189,938,1482,1509]" pageId="89" pageNumber="90">
<treatmentCitation id="2C2810CD1F1B9560FF3AFA6BDFDCFA0E" author="Tkalcu B." box="[189,934,1482,1509]" page="57" pageId="89" pageNumber="90" year="1989">
<taxonomicName id="6A894D5F1F1B9560FF3AFA6BDFDCFA0E" ID-CoL="5KT75" authority="Tkalcu, 1989: 57" authorityName="Tkalcu" authorityPageNumber="57" authorityYear="1989" box="[189,934,1482,1509]" class="Insecta" family="Apidae" genus="Pyrobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="subSpecies" species="keriensis" subSpecies="karakorumensis">
<emphasis id="9FFDEACE1F1B9560FF3AFA6BDDC5FA0F" box="[189,447,1482,1508]" italics="true" pageId="89" pageNumber="90">Pyrobombus keriensis</emphasis>
subsp. 
<emphasis id="9FFDEACE1F1B9560FD9EFA6BDEAEFA0F" box="[537,724,1482,1508]" italics="true" pageId="89" pageNumber="90">karakorumensis</emphasis>
<bibRefCitation id="C9184B2D1F1B9560FD5CFA6BDFDCFA0E" author="Tkalcu B." box="[731,934,1482,1509]" pageId="89" pageNumber="90" pagination="39 - 60" refId="ref71707" refString="Tkalcu B. 1989. Neue Taxa asiatischer Hummeln (Hymenoptera, Apoidea). Acta entomologica bohemoslovaca 86: 39 - 60." type="journal article" year="1989">Tkalců, 1989: 57</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1B9563FF3AF9ADDF96F93D" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="description">
<paragraph id="AD3636DC1F1B9560FF3AF9ADDDD0F9A2" blockId="89.[189,1398,1548,1609]" pageId="89" pageNumber="90">
The taxonomic limits of the species 
<taxonomicName id="6A894D5F1F1B9560FD06F9ACDF74F9CD" box="[641,782,1548,1574]" class="Insecta" family="Apidae" genus="Pyrobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FD06F9ACDF74F9CD" box="[641,782,1548,1574]" italics="true" pageId="89" pageNumber="90">B. keriensis</emphasis>
</taxonomicName>
have been a particular long-standing or persistent taxonomic problem.
</paragraph>
<paragraph id="AD3636DC1F1B9560FF3AF9D0DFB5F93A" blockId="89.[189,1399,1648,1745]" pageId="89" pageNumber="90">
<bibRefCitation id="C9184B2D1F1B9560FF3AF9D0DD0EF961" author="Vogt O." box="[189,372,1648,1675]" pageId="89" pageNumber="90" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt (1909: 41</bibRefCitation>
, 62) treated 
<taxonomicName id="6A894D5F1F1B9560FD8AF9D0DEE2F961" box="[525,664,1648,1674]" class="Insecta" family="Apidae" genus="Pyrobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FD8AF9D0DEE2F961" box="[525,664,1648,1674]" italics="true" pageId="89" pageNumber="90">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1B9560FD25F9D0DE98F960" box="[674,738,1649,1675]" pageId="89" pageNumber="90" sensu="stricto">s. str.</taxonomicNameLabel>
as a species separate from his taxon 
<taxonomicName id="6A894D5F1F1B9560FB2FF9D1D955F961" baseAuthorityName="Vogt" baseAuthorityYear="1909" box="[1192,1327,1648,1674]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1B9560FB2FF9D1D955F961" box="[1192,1327,1648,1674]" italics="true" pageId="89" pageNumber="90">separandus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="C9184B2D1F1B9560FAC7F9D0DD4DF945" author="Vogt O." pageId="89" pageNumber="90" pagination="31 - 74" refId="ref72005" refString="Vogt O. 1911. Studien ʾ ber das Artproblem. 2. Mitteilung. Uber das Variieren der Hummeln. 2. Teil. (Schluss). Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1911: 31 - 74." type="journal article" year="1911">Vogt (1911: 58)</bibRefCitation>
characterised and distinguished 
<taxonomicName id="6A894D5F1F1B9560FD31F935DF44F946" box="[694,830,1683,1709]" class="Insecta" family="Apidae" genus="Pyrobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FD31F935DF44F946" box="[694,830,1683,1709]" italics="true" pageId="89" pageNumber="90">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1B9560FCC1F935DFF9F945" box="[838,899,1684,1710]" pageId="89" pageNumber="90" sensu="stricto">s. str.</taxonomicNameLabel>
from 
<taxonomicName id="6A894D5F1F1B9560FC4CF935D808F946" baseAuthorityName="Vogt" baseAuthorityYear="1909" box="[971,1138,1683,1709]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1B9560FC4CF935D808F946" box="[971,1138,1683,1709]" italics="true" pageId="89" pageNumber="90">B. separandus</emphasis>
</taxonomicName>
by having from a few yellow hairs on the face to having the hair of the face entirely pale.
</paragraph>
<paragraph id="AD3636DC1F1B9560FF3AF959DE38F80D" blockId="89.[189,1399,1784,2022]" pageId="89" pageNumber="90">
<bibRefCitation id="C9184B2D1F1B9560FF3AF959DD04F8F9" author="Skorikov A. S." box="[189,382,1784,1810]" pageId="89" pageNumber="90" pagination="175 - 247" refId="ref70808" refString="Skorikov A. S. 1931. Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: Lindholm V. A. (ed.) Abhandlungen der Pamir-Expedition 1928: 175 - 247. Academy of Sciences of the USSR, Leningrad." type="journal article" year="1931">Skorikov (1931)</bibRefCitation>
listed 
<taxonomicName id="6A894D5F1F1B9560FE4BF958DE2FF8F9" box="[460,597,1784,1810]" class="Insecta" family="Apidae" genus="Pyrobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FE4BF958DE2FF8F9" box="[460,597,1784,1810]" italics="true" pageId="89" pageNumber="90">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1B9560FDDBF958DEE0F8F8" box="[604,666,1785,1811]" pageId="89" pageNumber="90" sensu="stricto">s. str.</taxonomicNameLabel>
and 
<taxonomicName id="6A894D5F1F1B9560FD55F958DF00F8F9" baseAuthorityName="Vogt" baseAuthorityYear="1909" box="[722,890,1784,1810]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1B9560FD55F958DF00F8F9" box="[722,890,1784,1810]" italics="true" pageId="89" pageNumber="90">B. separandus</emphasis>
</taxonomicName>
as two species but discussed them together under a joint heading, “ 
<taxonomicName id="6A894D5F1F1B9560FE5DF8BADFACF8DE" authority="(F. Mor.) 1886" baseAuthorityName="F. Mor." baseAuthorityYear="1886" box="[474,982,1819,1846]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FE5DF8BADF67F8DE" box="[474,797,1819,1845]" italics="true" pageId="89" pageNumber="90">Lapidariobombus keriensis</emphasis>
(F. Mor.) 1886
</taxonomicName>
und 
<taxonomicName id="6A894D5F1F1B9560FB92F8BADD2BF8B2" authority="(Vogt) 1909" baseAuthorityName="Vogt" baseAuthorityYear="1909" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1B9560FB92F8BAD90CF8DE" box="[1045,1398,1819,1845]" italics="true" pageId="89" pageNumber="90">Lapidariobombus separandus</emphasis>
(Vogt) 1909
</taxonomicName>
”. While he mentioned the difference in face colour, he described 
<taxonomicName id="6A894D5F1F1B9560FBEAF89ED882F8B2" box="[1133,1272,1855,1881]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1B9560FBEAF89ED882F8B2" box="[1133,1272,1855,1881]" italics="true" pageId="89" pageNumber="90">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1B9560FA85F89ED938F8B2" box="[1282,1346,1855,1881]" pageId="89" pageNumber="90" sensu="stricto">s. str.</taxonomicNameLabel>
and 
<taxonomicName id="6A894D5F1F1B9560FF3AF8C2DD1FF897" baseAuthorityName="Vogt" baseAuthorityYear="1909" box="[189,357,1890,1916]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1B9560FF3AF8C2DD1FF897" box="[189,357,1890,1916]" italics="true" pageId="89" pageNumber="90">B. separandus</emphasis>
</taxonomicName>
as hardly distinguishable (his 
<taxonomicName id="6A894D5F1F1B9560FD4CF8C3D95BF897" authority="Skorikov, 1931" authorityName="Skorikov" authorityYear="1931" box="[715,1313,1890,1916]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="anargumentosus">
<emphasis id="9FFDEACE1F1B9560FD4CF8C3D81EF897" box="[715,1124,1890,1916]" italics="true" pageId="89" pageNumber="90">Lapidariobombus anargumentosus</emphasis>
<bibRefCitation id="C9184B2D1F1B9560FBEBF8C3D95BF897" author="Skorikov A. S." box="[1132,1313,1890,1916]" pageId="89" pageNumber="90" pagination="175 - 247" refId="ref70808" refString="Skorikov A. S. 1931. Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: Lindholm V. A. (ed.) Abhandlungen der Pamir-Expedition 1928: 175 - 247. Academy of Sciences of the USSR, Leningrad." type="journal article" year="1931">Skorikov, 1931</bibRefCitation>
</taxonomicName>
, on the same page is recognised here from the pattern of punctures on the ocello-ocular area of the head of the 
<typeStatus id="7232887E1F1B9560FF3AF808DD5EF828" box="[189,292,1961,1987]" pageId="89" pageNumber="90" type="syntype">syntypes</typeStatus>
in the ZIN as conspecific with 
<taxonomicName id="6A894D5F1F1B9560FD0BF808D86DF829" authority="Vogt, 1911" authorityName="Vogt" authorityYear="1911" box="[652,1047,1960,1987]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="biroi" status="syn. nov." subGenus="Pyrobombus">
<emphasis id="9FFDEACE1F1B9560FD0BF808DEDFF829" box="[652,677,1961,1986]" italics="true" pageId="89" pageNumber="90">B.</emphasis>
(
<emphasis id="9FFDEACE1F1B9560FD32F809DF32F829" box="[693,840,1960,1986]" italics="true" pageId="89" pageNumber="90">Pyrobombus</emphasis>
) 
<emphasis id="9FFDEACE1F1B9560FCDEF809DFEBF829" box="[857,913,1960,1986]" italics="true" pageId="89" pageNumber="90">biroi</emphasis>
<bibRefCitation id="C9184B2D1F1B9560FC10F808D86DF829" author="Vogt O." box="[919,1047,1960,1987]" pageId="89" pageNumber="90" pagination="31 - 74" refId="ref72005" refString="Vogt O. 1911. Studien ʾ ber das Artproblem. 2. Mitteilung. Uber das Variieren der Hummeln. 2. Teil. (Schluss). Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1911: 31 - 74." type="journal article" year="1911">Vogt, 1911</bibRefCitation>
</taxonomicName>
, 
<taxonomicNameLabel id="84CE57B51F1B9560FBA4F808D8F7F828" box="[1059,1165,1961,1987]" pageId="89" pageNumber="90">syn. nov.</taxonomicNameLabel>
, and is not a species of the subgenus 
<taxonomicName id="6A894D5F1F1B9560FEFBF86ADE4BF80E" authorityName="von Dalla Torre" authorityYear="1880" box="[380,561,1995,2021]" class="Insecta" family="Apidae" genus="Melanobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="9FFDEACE1F1B9560FEFBF86ADE4BF80E" box="[380,561,1995,2021]" italics="true" pageId="89" pageNumber="90">Melanobombus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="AD3636DC1F189563FF3AFEA8DDB4FE81" blockId="90.[189,1398,265,362]" pageId="90" pageNumber="91">
<bibRefCitation id="C9184B2D1F189563FF3AFEA8DD1EFECF" author="Reinig W. F." box="[189,356,265,292]" pageId="90" pageNumber="91" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig (1935)</bibRefCitation>
included the taxon 
<taxonomicName id="6A894D5F1F189563FDD1FEA8DEC4FEC8" box="[598,702,265,291]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FDD1FEA8DEC4FEC8" box="[598,702,265,291]" italics="true" pageId="90" pageNumber="91">keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FD4FFEABDF73FECF" box="[712,777,266,292]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
as a part of a broad species 
<taxonomicName id="6A894D5F1F189563FBEEFEABD88EFEC8" box="[1129,1268,265,291]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="lato" species="keriensis">
<emphasis id="9FFDEACE1F189563FBEEFEABD88EFEC8" box="[1129,1268,265,291]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FB78FEABD93BFEC8" box="[1279,1345,265,292]" pageId="90" pageNumber="91" sensu="lato">s. lat.</taxonomicNameLabel>
that also included the taxa 
<emphasis id="9FFDEACE1F189563FE48FE8DDFCCFEAD" box="[463,950,300,326]" italics="true" pageId="90" pageNumber="91">
<taxonomicName id="6A894D5F1F189563FE48FE8DDE2CFEAD" baseAuthorityName="Vogt" baseAuthorityYear="1909" box="[463,598,300,326]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">separandus</taxonomicName>
<taxonomicName id="6A894D5F1F189563FDE6FE8CDF7BFEAD" authorityName="Vogt" authorityYear="1911" baseAuthorityName="Vogt" baseAuthorityYear="1911" box="[609,769,300,326]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="incertoides">+ incertoides</taxonomicName>
<taxonomicName id="6A894D5F1F189563FC8BFE8CDFCCFEAD" authorityName="Reinig" authorityYear="1930" box="[780,950,300,326]" class="Insecta" family="Apidae" genus="Lapidariobombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="alagesianus">+ alagesianus</taxonomicName>
</emphasis>
. This interpretation was followed by 
<bibRefCitation id="C9184B2D1F189563FF3AFEEEDD0CFE81" author="Williams P. H." box="[189,374,335,362]" pageId="90" pageNumber="91" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams (1991</bibRefCitation>
, 
<bibRefCitation id="C9184B2D1F189563FE05FEF1DDBBFE81" author="Williams P. H." box="[386,449,336,362]" pageId="90" pageNumber="91" pagination="79 - 152" refId="ref72435" refString="Williams P. H. 1998. An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology) 67: 79 - 152. Available and updated from www. nhm. ac. uk / bombus / [accessed 2019]." type="journal article" year="1998">1998</bibRefCitation>
).
</paragraph>
<paragraph id="AD3636DC1F189563FF3AFE30DE2BFE1A" blockId="90.[189,1399,401,498]" pageId="90" pageNumber="91">
<bibRefCitation id="C9184B2D1F189563FF3AFE30DD2DFE40" author="Wang S. - F." box="[189,343,401,427]" pageId="90" pageNumber="91" pagination="427 - 447" refId="ref72195" refString="Wang S. - F. 1982. Hymenoptera: Apidae - Bombus. In: Qinghai-Tibet Plateau Comprehensive Scientific Investigation Team of the Chinese Academy of Sciences (ed.) Insects of Xizang: 427 - 447. Science Press, Beijing." type="book chapter" year="1982">Wang (1982)</bibRefCitation>
described her taxon 
<taxonomicName id="6A894D5F1F189563FDD6FE30DEB0FE40" authorityName="Wang" authorityYear="1982" box="[593,714,401,427]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="trilineatus">
<emphasis id="9FFDEACE1F189563FDD6FE30DEB0FE40" box="[593,714,401,427]" italics="true" pageId="90" pageNumber="91">trilineatus</emphasis>
</taxonomicName>
without reference to 
<taxonomicName id="6A894D5F1F189563FC4DFE33D82EFE40" box="[970,1108,401,427]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F189563FC4DFE33D82EFE40" box="[970,1108,401,427]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
, comparing it instead to the more distantly related “ 
<taxonomicName id="6A894D5F1F189563FD85FE14DF9BFE25" authority="Richards" authorityName="Richards" box="[514,993,436,463]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="variety" species="occulatus" variety="haemorrhous">
<emphasis id="9FFDEACE1F189563FD85FE14DEEEFE25" box="[514,660,436,462]" italics="true" pageId="90" pageNumber="91">B. occulatus</emphasis>
var. 
<emphasis id="9FFDEACE1F189563FD56FE15DF17FE25" box="[721,877,436,462]" italics="true" pageId="90" pageNumber="91">haemorrhous</emphasis>
Richards
</taxonomicName>
” [sic] (= 
<taxonomicName id="6A894D5F1F189563FBD4FE14D894FE25" authorityName="Smith" authorityYear="1852" box="[1107,1262,436,462]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="9FFDEACE1F189563FBD4FE14D894FE25" box="[1107,1262,436,462]" italics="true" pageId="90" pageNumber="91">B. simillimus</emphasis>
</taxonomicName>
) for colour pattern and details of morphology.
</paragraph>
<paragraph id="AD3636DC1F189563FF3AFDB8DE8DFD77" blockId="90.[189,1399,537,669]" pageId="90" pageNumber="91">
Our PTP analysis (
<figureCitation id="35B22A591F189563FE27FDB8DD8DFDD8" box="[416,503,537,563]" captionStart="Fig" captionStartId="21.[189,232,1953,1979]" captionTargetBox="[233,1316,282,1913]" captionTargetId="figure-12@21.[233,1355,265,1914]" captionTargetPageId="21" captionText="Fig. 10. October 2019 (Fig. 8) Bayesian PTP analysis of MrBayes tree of unique COI barcodes (UHF- PTP). Symbols as in Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.4064332" httpUri="https://zenodo.org/record/4064332/files/figure.png" pageId="90" pageNumber="91">Fig. 10</figureCitation>
) of coalescents in the COI gene within the 
<taxonomicName id="6A894D5F1F189563FB80FDB8D815FDD8" box="[1031,1135,537,563]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F189563FB80FDB8D815FDD8" box="[1031,1135,537,563]" italics="true" pageId="90" pageNumber="91">keriensis</emphasis>
</taxonomicName>
-complex supports six species including 
<taxonomicName id="6A894D5F1F189563FE17FD9CDE62FDBD" box="[400,536,572,598]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FE17FD9CDE62FDBD" box="[400,536,572,598]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FDA7FD9DDE1AFDBD" box="[544,608,572,598]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
, corroborated by differences in morphology. These species are also supported by the absence of a positive divergence-with-distance relationship among them (
<figureCitation id="35B22A591F189563FB64FDFED942FD92" box="[1251,1336,607,633]" captionStart="Figs 17–20" captionStartId="26.[189,241,1773,1799]" captionTargetBox="[191,1390,661,1666]" captionTargetId="figure-140@26.[189,1398,653,1671]" captionTargetPageId="26" captionText="Figs 17–20. Plots of (y-axis) pairwise proportion of genetic divergence between COI barcode region sequences against (x-axis) pairwise Great Circle geographical distance in km between sample sites with linear trend lines (black). 17. Genetic divergences within the candidate species for the sichelii-complex identified using Bayesian UF-PTP (Mantel r = 0.52). 18. Genetic divergences among the candidate species for the sichelii-complex identified using Bayesian UF-PTP (Mantel r = 0.21). 19. Genetic divergences within the candidate species for the keriensis-complex identified using Bayesian UF-PTP (Mantel r = 0.16). 20. Genetic divergences among the candidate species for the keriensis-complex identified using Bayesian UF-PTP (relationship not positive). For details of the measurements see the" figureDoi="http://doi.org/10.5281/zenodo.4064342" httpUri="https://zenodo.org/record/4064342/files/figure.png" pageId="90" pageNumber="91">Fig. 20</figureCitation>
) (see Divergence and geographical distance, page 12).
</paragraph>
<paragraph id="AD3636DC1F189563FF3AFD65D91EFADE" blockId="90.[189,1399,708,1333]" pageId="90" pageNumber="91">
From morphology, after examining&gt;1000 specimens of the 
<taxonomicName id="6A894D5F1F189563FC03FD65DF96FD35" box="[900,1004,708,734]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F189563FC03FD65DF96FD35" box="[900,1004,708,734]" italics="true" pageId="90" pageNumber="91">keriensis</emphasis>
</taxonomicName>
-group from across Asia and after comparing the distribution of states of several characters as part of this study (PW), Vogt’s character state of at least a few yellow hairs on the face does appear to remain one of the most consistently diagnostic character states for 
<taxonomicName id="6A894D5F1F189563FD9BFC8FDED9FCAC" box="[540,675,813,839]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FD9BFC8FDED9FCAC" box="[540,675,813,839]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FD2EFC8FDE9FFCA3" box="[681,741,814,840]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
(by comparison with COI sequences), at least for queens in most parts of the species’ range (but see the comments on 
<taxonomicName id="6A894D5F1F189563FC1BFCF0D83CFC81" baseAuthorityName="Vogt" baseAuthorityYear="1909" box="[924,1094,848,874]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F189563FC1BFCF0D83CFC81" box="[924,1094,848,874]" italics="true" pageId="90" pageNumber="91">B. separandus</emphasis>
</taxonomicName>
regarding the Mongolian taxon 
<taxonomicName id="6A894D5F1F189563FE80FCD5DD20FC65" authorityName="Skorikov" authorityYear="1910" box="[263,346,884,910]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="9FFDEACE1F189563FE80FCD5DD20FC65" box="[263,346,884,910]" italics="true" pageId="90" pageNumber="91">kozlovi</emphasis>
</taxonomicName>
). Several aspects of the colour pattern vary strongly within the species: (1) 
<taxonomicName id="6A894D5F1F189563FB6BFCD5D90CFC65" box="[1260,1398,884,910]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FB6BFCD5D90CFC65" box="[1260,1398,884,910]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FF3AFC36DC84FC5A" box="[189,254,919,945]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
includes both yellow-banded and white-banded colour patterns; (2) the extent of pale hair on the face varies; (3) the extent of the black hair between the wing bases varies; (4) the extent of the pale hair on the side and ventral area (leg bases) of the thorax varies; (5) the extent of the pale hair posteriorly on T3 varies (the ‘ciliation’ described by authors); (6) the contrast of the pale posterior fringes on T4–5 varies. Many of these characters might appear to show a continuum of variation that is also continuous with the variation within 
<taxonomicName id="6A894D5F1F189563FD4DFBE9D83BFB8A" authority="(Williams 1991)" baseAuthorityName="Williams" baseAuthorityYear="1991" box="[714,1089,1095,1121]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F189563FD4DFBE9DF0EFB8A" box="[714,884,1095,1121]" italics="true" pageId="90" pageNumber="91">B. separandus</emphasis>
(
<bibRefCitation id="C9184B2D1F189563FC02FBE6D842FB8A" author="Williams P. H." box="[901,1080,1095,1121]" pageId="90" pageNumber="91" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams 1991</bibRefCitation>
)
</taxonomicName>
, which might then appear consistent with the two taxa being parts of the same species (
<bibRefCitation id="C9184B2D1F189563FC26FBCBD840FB6E" author="Reinig W. F." box="[929,1082,1130,1157]" pageId="90" pageNumber="91" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig 1935</bibRefCitation>
). However, evidence from COI barcodes shows that the two gene-coalescent groups differ most consistently in the presence of pale hairs on the face of 
<taxonomicName id="6A894D5F1F189563FE2EFB10DE48FB21" box="[425,562,1200,1226]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FE2EFB10DE48FB21" box="[425,562,1200,1226]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FDBCFB10DE01FB20" box="[571,635,1201,1227]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
, supporting Vogt’s characterisation of the species, and providing integrated morphological support for its status as a separate species. The overlapping colour variation of 
<taxonomicName id="6A894D5F1F189563FF3AFB59DD1FFAFA" baseAuthorityName="Williams" baseAuthorityYear="1991" box="[189,357,1271,1297]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F189563FF3AFB59DD1FFAFA" box="[189,357,1271,1297]" italics="true" pageId="90" pageNumber="91">B. separandus</emphasis>
</taxonomicName>
but only in 
<collectingCountry id="D59E764C1F189563FE71FB56DE12FAFA" box="[502,616,1271,1297]" name="Mongolia" pageId="90" pageNumber="91">Mongolia</collectingCountry>
(where 
<taxonomicName id="6A894D5F1F189563FD4EFB59DF29FAFA" box="[713,851,1271,1297]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FD4EFB59DF29FAFA" box="[713,851,1271,1297]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FCDCFB56DFE3FAFA" box="[859,921,1271,1297]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
does not occur) does not negate the fact that 
<taxonomicName id="6A894D5F1F189563FF77FABADD03FADF" box="[240,377,1306,1332]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FF77FABADD03FADF" box="[240,377,1306,1332]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FE07FABADDC4FADE" box="[384,446,1307,1333]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
is recognisable by morphological characters throughout its geographical range.
</paragraph>
<paragraph id="AD3636DC1F189563FF3AFAFADF96F93D" blockId="90.[189,1399,1371,1750]" pageId="90" pageNumber="91">
There are some yellow-banded queens from the Pir Panjal and Great Himalaya ranges (part of 
<taxonomicName id="6A894D5F1F189563FB77FAFDD90CFA9E" box="[1264,1398,1371,1397]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FB77FAFDD90CFA9E" box="[1264,1398,1371,1397]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FF3AFADEDC81FA72" box="[189,251,1407,1433]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
according our COI coalescent results, 
<figureCitation id="35B22A591F189563FD45FADEDF6CFA72" box="[706,790,1407,1433]" captionStart="Fig" captionStartId="21.[189,232,1953,1979]" captionTargetBox="[233,1316,282,1913]" captionTargetId="figure-12@21.[233,1355,265,1914]" captionTargetPageId="21" captionText="Fig. 10. October 2019 (Fig. 8) Bayesian PTP analysis of MrBayes tree of unique COI barcodes (UHF- PTP). Symbols as in Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.4064332" httpUri="https://zenodo.org/record/4064332/files/figure.png" pageId="90" pageNumber="91">Fig. 10</figureCitation>
) that have very few pale hairs on the face, although these specimens still have a few yellow hairs on the leg bases and have a few black hairs in the pale bands of the thoracic dorsum. Many of the available specimens have failed repeated attempts at sequencing, so the species’ diagnosis remains tentative. Particularly influential in the interpretation of these individuals is one sequenced yellow-banded queen with very few pale hairs on the face, on the underside of the thorax, and on T3, from the Pir Panjal range (PW: ML405, 
<figureCitation id="35B22A591F189563FCF2F98EDFA4F9A2" box="[885,990,1583,1609]" captionStart="Figs 139–180" captionStartId="42.[189,241,1497,1523]" captionTargetBox="[369,1221,268,1431]" captionTargetId="figure-11@42.[366,1221,265,1467]" captionTargetPageId="42" captionText="Figs 139–180. Simplified diagrams for the colour patterns of the hair on the dorsum for the species from the integrative analysis. The dorsum is divided into regions, each of which shows only the predominant or most apparent colour for that region using a simplified colour palette, with olive indicating a mixture of black and yellow hair, and grey indicating a mixture of black and white hair. The keriensis-group. 139. Queen, China-Sichuan. 140. Worker, China-Sichuan. 141. Queen, China-Sichuan. 142. Queen, China-Sichuan. 143. Worker, China-Xizang. 144. Queen, India-Kashmir. 145. Male, China-Qinghai. 146. Male, India-Kashmir. 147. Male, China-Gansu. 148. Queen, Armenia. 149. Queen, Iran. 150. Male, Georgia. 151. Worker, China-Qinghai. 152. Male, China-Qinghai. 153. Queen, Mongolia. 154. Worker, Mongolia. 155. Male, Mongolia. 156. Male, Mongolia. 157. Queen, China-Sichuan. 158. Worker, China-Qinghai. 159. Worker, China-Qinghai. 160. Male, China-Qinghai. 161. Male, China-Qinghai. 162. Male, China-Qinghai. 163. Queen, Pakistan. 164. Queen, Pakistan. 165. Queen, Pakistan. 166. Queen, Pakistan. 167. Queen, Afghanistan. 168. Queen, Pakistan. 169. Queen, Pakistan. 170. Queen, India-Kashmir. 171. Male, India-Kashmir. 172. Male, India-Kashmir. 173. Queen, China- Xinjiang. 174. Queen, Kyrgyzstan. 175. Queen, Kyrgyzstan. 176. Queen, Mongolia. 177. Queen, India- Kashmir. 178. Male, Kazakhstan. 179. Male, China-Xinjiang. 180. Male, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.4064358" httpUri="https://zenodo.org/record/4064358/files/figure.png" pageId="90" pageNumber="91">Figs 170</figureCitation>
, 
<figureCitation id="35B22A591F189563FC6DF98ED862F9A2" box="[1002,1048,1583,1609]" captionStart="Figs 209–210" captionStartId="93.[189,241,1884,1910]" captionTargetBox="[190,1374,318,1788]" captionTargetId="figure-11@93.[189,1398,318,1792]" captionTargetPageId="93" captionText="Figs 209–210. Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau. 209. Bombus keriensis Morawitz, 1887 from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405). 210. B. separandus Vogt, 1909 stat. rev. from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="90" pageNumber="91">209</figureCitation>
) that groups with 
<taxonomicName id="6A894D5F1F189563FB69F991D90CF9A2" box="[1262,1398,1583,1609]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F189563FB69F991D90CF9A2" box="[1262,1398,1583,1609]" italics="true" pageId="90" pageNumber="91">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F189563FF3AF9F3DC83F987" box="[189,249,1618,1644]" pageId="90" pageNumber="91" sensu="stricto">s. str.</taxonomicNameLabel>
(
<figureCitation id="35B22A591F189563FE8FF9F3DD21F987" box="[264,347,1618,1644]" captionStart="Fig" captionStartId="21.[189,232,1953,1979]" captionTargetBox="[233,1316,282,1913]" captionTargetId="figure-12@21.[233,1355,265,1914]" captionTargetPageId="21" captionText="Fig. 10. October 2019 (Fig. 8) Bayesian PTP analysis of MrBayes tree of unique COI barcodes (UHF- PTP). Symbols as in Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.4064332" httpUri="https://zenodo.org/record/4064332/files/figure.png" pageId="90" pageNumber="91">Fig. 10</figureCitation>
) rather than with 
<taxonomicName id="6A894D5F1F189563FDAFF9F2DEB4F987" baseAuthorityName="Williams" baseAuthorityYear="1991" box="[552,718,1618,1644]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F189563FDAFF9F2DEB4F987" box="[552,718,1618,1644]" italics="true" pageId="90" pageNumber="91">B. separandus</emphasis>
</taxonomicName>
(diagnostic base positions 267T, 286T, 301T, 306C, 339C, 363T, 423C, 498T, 529A, 540C, 541T, 556T). This is one of the specimens that most closely resembles some yellow-banded 
<taxonomicName id="6A894D5F1F189563FE30F938DE25F959" baseAuthorityName="Williams" baseAuthorityYear="1991" box="[439,607,1688,1714]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F189563FE30F938DE25F959" box="[439,607,1688,1714]" italics="true" pageId="90" pageNumber="91">B. separandus</emphasis>
</taxonomicName>
also from Kashmir (
<figureCitation id="35B22A591F189563FCD6F939DFC0F959" box="[849,954,1688,1714]" captionStart="Figs 139–180" captionStartId="42.[189,241,1497,1523]" captionTargetBox="[369,1221,268,1431]" captionTargetId="figure-11@42.[366,1221,265,1467]" captionTargetPageId="42" captionText="Figs 139–180. Simplified diagrams for the colour patterns of the hair on the dorsum for the species from the integrative analysis. The dorsum is divided into regions, each of which shows only the predominant or most apparent colour for that region using a simplified colour palette, with olive indicating a mixture of black and yellow hair, and grey indicating a mixture of black and white hair. The keriensis-group. 139. Queen, China-Sichuan. 140. Worker, China-Sichuan. 141. Queen, China-Sichuan. 142. Queen, China-Sichuan. 143. Worker, China-Xizang. 144. Queen, India-Kashmir. 145. Male, China-Qinghai. 146. Male, India-Kashmir. 147. Male, China-Gansu. 148. Queen, Armenia. 149. Queen, Iran. 150. Male, Georgia. 151. Worker, China-Qinghai. 152. Male, China-Qinghai. 153. Queen, Mongolia. 154. Worker, Mongolia. 155. Male, Mongolia. 156. Male, Mongolia. 157. Queen, China-Sichuan. 158. Worker, China-Qinghai. 159. Worker, China-Qinghai. 160. Male, China-Qinghai. 161. Male, China-Qinghai. 162. Male, China-Qinghai. 163. Queen, Pakistan. 164. Queen, Pakistan. 165. Queen, Pakistan. 166. Queen, Pakistan. 167. Queen, Afghanistan. 168. Queen, Pakistan. 169. Queen, Pakistan. 170. Queen, India-Kashmir. 171. Male, India-Kashmir. 172. Male, India-Kashmir. 173. Queen, China- Xinjiang. 174. Queen, Kyrgyzstan. 175. Queen, Kyrgyzstan. 176. Queen, Mongolia. 177. Queen, India- Kashmir. 178. Male, Kazakhstan. 179. Male, China-Xinjiang. 180. Male, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.4064358" httpUri="https://zenodo.org/record/4064358/files/figure.png" pageId="90" pageNumber="91">Figs 177</figureCitation>
, 
<figureCitation id="35B22A591F189563FC40F939DF8FF959" box="[967,1013,1688,1714]" captionStart="Figs 209–210" captionStartId="93.[189,241,1884,1910]" captionTargetBox="[190,1374,318,1788]" captionTargetId="figure-11@93.[189,1398,318,1792]" captionTargetPageId="93" captionText="Figs 209–210. Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau. 209. Bombus keriensis Morawitz, 1887 from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405). 210. B. separandus Vogt, 1909 stat. rev. from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="90" pageNumber="91">210</figureCitation>
) (the difference in the breadth of the black band between the wings is not diagnostic for these species).
</paragraph>
</subSubSection>
<subSubSection id="E59365571F189563FF3AF95CDE53F838" pageId="90" pageNumber="91" type="diagnosis">
<paragraph id="AD3636DC1F189563FF3AF95CDD4CF8FC" blockId="90.[189,310,1789,1815]" box="[189,310,1789,1815]" pageId="90" pageNumber="91">
<heading id="F67E81B01F189563FF3AF95CDD4CF8FC" bold="true" box="[189,310,1789,1815]" fontSize="11" level="3" pageId="90" pageNumber="91" reason="3">
<emphasis id="9FFDEACE1F189563FF3AF95CDD4CF8FC" bold="true" box="[189,310,1789,1815]" pageId="90" pageNumber="91">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F189563FF6DF88DDD2BF8AD" blockId="90.[189,1399,1836,2003]" box="[234,337,1836,1862]" pageId="90" pageNumber="91">
<emphasis id="9FFDEACE1F189563FF6DF88DDD2BF8AD" bold="true" box="[234,337,1836,1862]" pageId="90" pageNumber="91">Females</emphasis>
</paragraph>
<paragraph id="AD3636DC1F189563FF3AF8EEDE53F838" blockId="90.[189,1399,1836,2003]" pageId="90" pageNumber="91">
Queens medium-sized body length 
<quantity id="6A719B391F189563FDDCF8EEDEA7F882" box="[603,733,1871,1897]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="90" pageNumber="91" unit="mm" value="19.0" valueMax="21.0" valueMin="17.0">17–21 mm</quantity>
, workers 
<quantity id="6A719B391F189563FCCAF8EEDFBAF882" box="[845,960,1871,1897]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="90" pageNumber="91" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">9–14 mm</quantity>
. Can be distinguished in Central Asia by the combination of hair of the face usually with some pale hairs, the leg bases usually with some pale hairs, T2 posteriorly entirely pale without black hairs, with T3 laterally usually with some pale hairs (cf. 
<taxonomicName id="6A894D5F1F189563FF6BF818DD18F839" authorityName="Radoszkowski" authorityYear="1860" box="[236,354,1976,2002]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="sichelii">
<emphasis id="9FFDEACE1F189563FF6BF818DD18F839" box="[236,354,1976,2002]" italics="true" pageId="90" pageNumber="91">B. sichelii</emphasis>
</taxonomicName>
, 
<taxonomicName id="6A894D5F1F189563FEF7F818DE62F839" baseAuthorityName="Williams" baseAuthorityYear="1991" box="[368,536,1976,2002]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F189563FEF7F818DE62F839" box="[368,536,1976,2002]" italics="true" pageId="90" pageNumber="91">B. separandus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E59365571F199562FF6DFEA8DF06FE1D" pageId="91" pageNumber="92" type="description">
<paragraph id="AD3636DC1F199562FF6DFEA8DD4FFEC8" blockId="91.[189,1399,265,502]" box="[234,309,265,291]" pageId="91" pageNumber="92">
<emphasis id="9FFDEACE1F199562FF6DFEA8DD4FFEC8" bold="true" box="[234,309,265,291]" pageId="91" pageNumber="92">Males</emphasis>
</paragraph>
<paragraph id="AD3636DC1F199562FF3AFE8DDF06FE1D" blockId="91.[189,1399,265,502]" pageId="91" pageNumber="92">
Body length 
<quantity id="6A719B391F199562FED2FE8ADDADFEAD" box="[341,471,299,326]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="91" pageNumber="92" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11–15 mm</quantity>
. Can be distinguished reliably at present only by their COI sequence, although in the western Himalaya they may be distinguished by yellow hair between the wing bases and on T3 posterio-laterally. Genitalia (
<figureCitation id="35B22A591F199562FDCDFED3DECFFE67" box="[586,693,370,396]" captionStart="Figs 199‒204" captionStartId="45.[189,241,1852,1878]" captionTargetBox="[230,1342,827,1806]" captionTargetId="figure-159@45.[194,1393,771,1810]" captionTargetPageId="45" captionText="Figs 199‒204. Morphology of the male genitalia for species of the subgenus Melanobombus von Dalla Torre, 1880 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 199. Bombus alagesianus Reinig, 1930 stat. rev., Georgia. 200. B. tibeticus sp. nov., China-Qinghai. 201. B. incertoides Vogt, 1911 stat. rev., Mongolia. 202. B. qilianensis sp. nov., China-Qinghai. 203. B. keriensis Morawitz, 1887, India-Kashmir. 204. B. separandus Vogt, 1909 stat. rev., Kyrgyzstan. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4064364" httpUri="https://zenodo.org/record/4064364/files/figure.png" pageId="91" pageNumber="92">Fig. 203</figureCitation>
) with the gonostylus shorter than broad, its inner basal projection reduced to a short stub (cf. 
<emphasis id="9FFDEACE1F199562FDFAFE34DEADFE44" box="[637,727,405,431]" italics="true" pageId="91" pageNumber="92">rufipes-</emphasis>
group, 
<emphasis id="9FFDEACE1F199562FCADFE34DFF6FE44" box="[810,908,405,431]" italics="true" pageId="91" pageNumber="92">festivus-</emphasis>
group, 
<taxonomicName id="6A894D5F1F199562FC58FE34D802FE44" authorityName="Smith" authorityYear="1852" box="[991,1144,405,431]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="rufofasciatus">
<emphasis id="9FFDEACE1F199562FC58FE34D802FE44" box="[991,1144,405,431]" italics="true" pageId="91" pageNumber="92">rufofasciatus</emphasis>
</taxonomicName>
-group); volsella with the inner distal corner broadly produced but without a narrow hook (cf. 
<emphasis id="9FFDEACE1F199562FB9CFE18D80FFE38" box="[1051,1141,441,467]" italics="true" pageId="91" pageNumber="92">rufipes-</emphasis>
group, 
<emphasis id="9FFDEACE1F199562FB4EFE18D951FE38" box="[1225,1323,441,467]" italics="true" pageId="91" pageNumber="92">festivus-</emphasis>
group, 
<taxonomicName id="6A894D5F1F199562FF3AFE7DDD2CFE1D" authorityName="Smith" authorityYear="1852" box="[189,342,476,502]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="rufofasciatus">
<emphasis id="9FFDEACE1F199562FF3AFE7DDD2CFE1D" box="[189,342,476,502]" italics="true" pageId="91" pageNumber="92">rufofasciatus</emphasis>
</taxonomicName>
-group); eye unenlarged relative to female eye.
</paragraph>
</subSubSection>
<subSubSection id="E59365571F199562FF3AFDBCDE1AF907" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="AD3636DC1F199562FF3AFDBCDDD0FDDC" blockId="91.[189,426,541,567]" box="[189,426,541,567]" pageId="91" pageNumber="92">
<heading id="F67E81B01F199562FF3AFDBCDDD0FDDC" bold="true" box="[189,426,541,567]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="9FFDEACE1F199562FF3AFDBCDDD0FDDC" bold="true" box="[189,426,541,567]" pageId="91" pageNumber="92">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F199562FF6DFDEDDD86FD8E" blockId="91.[189,1398,587,685]" box="[234,508,587,614]" pageId="91" pageNumber="92">
<emphasis id="9FFDEACE1F199562FF6DFDEDDD86FD8E" bold="true" box="[234,508,587,614]" pageId="91" pageNumber="92">
<typeStatus id="7232887E1F199562FF6DFDEDDD1CFD8D" box="[234,358,588,614]" pageId="91" pageNumber="92" type="lectotype">Lectotype</typeStatus>
designation
</emphasis>
</paragraph>
<paragraph id="AD3636DC1F199562FF3AFDCEDD4AFD46" blockId="91.[189,1398,587,685]" pageId="91" pageNumber="92">
<materialsCitation id="1DE13C811F199562FF3AFDCEDD4AFD46" ID-GBIF-Occurrence="2867606329" collectionCode="ZIN" collectorName="Przewalsky" country="China" elevation="2743" location="Keria Mts" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<collectingCountry id="D59E764C1F199562FF3AFDCEDD62FD62" box="[189,280,623,649]" name="China" pageId="91" pageNumber="92">CHINA</collectingCountry>
• 
<specimenCount id="BB8FFD551F199562FEAAFDCEDD39FD62" box="[301,323,623,649]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
(queen), 
<typeStatus id="7232887E1F199562FE29FDCEDE60FD62" box="[430,538,623,649]" pageId="91" pageNumber="92" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="6A894D5F1F199562FDBBFDCEDFB3FD62" authority="Morawitz, 1887" authorityName="F. Morawitz" authorityYear="1887" box="[572,969,623,649]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F199562FDBBFDCEDF72FD62" box="[572,776,623,649]" italics="true" pageId="91" pageNumber="92">Bombus keriensis</emphasis>
<bibRefCitation id="C9184B2D1F199562FC8AFDCEDFB3FD62" author="Morawitz F. F." box="[781,969,623,649]" pageId="91" pageNumber="92" pagination="195 - 229" refId="ref68420" refString="Morawitz F. F. 1887. Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva 20 (1886): 195 - 229." type="journal article" year="1887">Morawitz, 1887</bibRefCitation>
</taxonomicName>
; 
<location id="A85660071F199562FC54FDCED831FD62" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FC54FDCED831FD62" box="[979,1099,623,649]" country="China" name="Keria Mts" pageId="91" pageNumber="92">Keria Mts</location>
; 
<elevation id="26A4D1EF1F199562FBD2FDCED89EFD62" box="[1109,1252,623,649]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="91" pageNumber="92" unit="ft" value="9000.0">
<quantity id="6A719B391F199562FBD2FDCED8DCFD62" box="[1109,1190,623,649]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="91" pageNumber="92" unit="ft" value="9000.0">9000 ft</quantity>
a.s.l.
</elevation>
; 
<collectorName id="007C530A1F199562FB68FDCED90CFD62" box="[1263,1398,623,649]" pageId="91" pageNumber="92">Przewalsky</collectorName>
leg.; 
<collectionCode id="CB98AE191F199562FF7FFD32DD57FD46" box="[248,301,659,685]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="91" pageNumber="92" type="Museum">ZIN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="AD3636DC1F199562FF3AFD72DF1EFBC0" blockId="91.[189,1399,723,1067]" pageId="91" pageNumber="92">
Morawitz’s original description of several females of 
<taxonomicName id="6A894D5F1F199562FCCDFD75DFACFD06" authorityName="F. Morawitz" authorityYear="1887" box="[842,982,723,749]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F199562FCCDFD75DFACFD06" box="[842,982,723,749]" italics="true" pageId="91" pageNumber="92">B. keriensis</emphasis>
</taxonomicName>
gives the type locality as “Keria” [= Keriya/Yutian, east of Hotan /Hetian / Khotan], presumably not in the low-lying town but higher in the neighbouring Karakax 
<collectingRegion id="6F4DF83E1F199562FE7BFCBBDE4DFCDF" box="[508,567,794,820]" country="Myanmar" name="Shan" pageId="91" pageNumber="92">Shan</collectingRegion>
(of the western Kunlun 
<collectingRegion id="6F4DF83E1F199562FCD8FCBBDFE0FCDF" box="[863,922,794,820]" country="Myanmar" name="Shan" pageId="91" pageNumber="92">Shan</collectingRegion>
[shan = mountains], southern 
<collectingRegion id="6F4DF83E1F199562FA8EFCBBD909FCDF" box="[1289,1395,794,820]" country="China" name="Xinjiang Uygur" pageId="91" pageNumber="92">Xinjiang</collectingRegion>
, and probably near the village of Polu / Pulu/ Pulucun [cun = village], 
<geoCoordinate id="C8BD501B1F199562FB8CFC9CD8E3FCB3" box="[1035,1177,829,856]" degrees="36.1850" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="36.185">36.1850° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FB2EFC9CD94EFCB3" box="[1193,1332,829,856]" degrees="81.4827" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="81.4827">81.4827° E</geoCoordinate>
. This village was visited by N. Prshewalsky while searching for a pass southwards into Ladakh during his fourth (and final) Central Asian expedition in the summer of 1885 (
<bibRefCitation id="C9184B2D1F199562FC6CFC22D8E3FC75" author="Rayfield D." box="[1003,1177,899,926]" pageId="91" pageNumber="92" refId="ref69618" refString="Rayfield D. 1976. The Dream of Lhasa. The Life of Nikolay Przhevalsky (1839 - 88), Explorer of Central Asia. Elek Books, London." type="book" year="1976">Rayfield 1976</bibRefCitation>
). If the “9000” on the 
<typeStatus id="7232887E1F199562FF6DFC06DD3FFC2A" box="[234,325,935,961]" pageId="91" pageNumber="92" type="syntype">syntype</typeStatus>
specimen label (below) were an elevation in feet then this is substantially higher than the elevation of Keriya (ca 
<quantity id="6A719B391F199562FE50FC6BDE36FC0E" box="[471,588,970,997]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="91" pageNumber="92" unit="ft" value="4500.0">4500 feet</quantity>
), so a collecting site near Pulu is highly likely. These mountains are very arid, so likely sites with a reliable water supply and food plants every summer sufficient to support bumblebee colonies are restricted (
<bibRefCitation id="C9184B2D1F199562FDD1FBB1DF2DFBC0" author="Williams P. H. &amp; Bystriakova N. &amp; Huang J. - X. &amp; Miao Z. - Y. &amp; An J. - D." box="[598,855,1040,1067]" pageId="91" pageNumber="92" pagination="164 - 181" refId="ref73123" refString="Williams P. H., Bystriakova N., Huang J. - X., Miao Z. - Y. &amp; An J. - D. 2015 a. Bumblebees, climate and glaciers across the Tibetan plateau (Apidae: Bombus Latreille). Systematics and Biodiversity 13: 164 - 181. https: // doi. org / 10.1080 / 14772000.2014.982228" type="journal article" year="2015">
Williams 
<emphasis id="9FFDEACE1F199562FD4EFBB0DF7EFBC1" box="[713,772,1040,1066]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
2015a
</bibRefCitation>
).
</paragraph>
<paragraph id="AD3636DC1F199562FF3AFBF3D86DFA88" blockId="91.[189,1399,1106,1379]" pageId="91" pageNumber="92">
The 
<collectionCode id="CB98AE191F199562FF74FBF3DD5EFB87" box="[243,292,1106,1132]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="91" pageNumber="92" type="Museum">ZIN</collectionCode>
collection contains a queen that agrees with the original description and carries seven labels: (1)gold disc;(2)handwritten white label“Keria / Gebirg./ 9000”reverse“2/10/[illegible]”;(3)handwritten white label “Przewalsky”; (4) handwritten white label “ 
<taxonomicName id="6A894D5F1F199562FC21FB39D87FFB59" authorityName="Latreille" authorityYear="1802" box="[934,1029,1176,1202]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="9FFDEACE1F199562FC21FB39D87FFB59" box="[934,1029,1176,1202]" italics="true" pageId="91" pageNumber="92">Bombus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="6A894D5F1F199562FB93FB39D806FB59" authorityName="F. Morawitz" authorityYear="1887" box="[1044,1148,1176,1202]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F199562FB93FB39D806FB59" box="[1044,1148,1176,1202]" italics="true" pageId="91" pageNumber="92">keriensis</emphasis>
</taxonomicName>
<specimenCount id="BB8FFD551F199562FB16FB38D8D2FB58" box="[1169,1192,1177,1203]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
. / F. Morawitz.”; (5) printed white label in Cyrillic “[K. 
<emphasis id="9FFDEACE1F199562FD0CFB1DDF63FB3E" box="[651,793,1212,1237]" italics="true" pageId="91" pageNumber="92">F. Morawitz</emphasis>
]”; (6) red label partly printed in black “ 
<emphasis id="9FFDEACE1F199562FB70FB1DDD66FB13" italics="true" pageId="91" pageNumber="92">
<typeStatus id="7232887E1F199562FB70FB1DD90CFB3E" box="[1271,1398,1212,1237]" pageId="91" pageNumber="92" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="6A894D5F1F199562FF3AFB7FDD66FB13" authorityName="Latreille" authorityYear="1802" box="[189,284,1246,1272]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">Bombus</taxonomicName>
</emphasis>
/ 
<taxonomicName id="6A894D5F1F199562FEACFB7FDE4CFB13" authority="F. Morawitz" authorityName="F. Morawitz" authorityYear="1887" box="[299,566,1246,1273]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F199562FEACFB7FDDE9FB13" box="[299,403,1246,1272]" italics="true" pageId="91" pageNumber="92">keriensis</emphasis>
F. Morawitz
</taxonomicName>
/ design. Podbolotsk.” (unpublished); (7) red label printed in black “ 
<emphasis id="9FFDEACE1F199562FF4DFAA0DD04FAF0" bold="true" box="[202,382,1281,1307]" pageId="91" pageNumber="92">
<typeStatus id="7232887E1F199562FF4DFAA0DD04FAF0" box="[202,382,1281,1307]" pageId="91" pageNumber="92" type="lectotype">LECTOTYPE</typeStatus>
</emphasis>
<specimenCount id="BB8FFD551F199562FE08FAA3DDDFFAF7" box="[399,421,1282,1308]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
/ 
<taxonomicName id="6A894D5F1F199562FE33FAA3DE69FAF7" authorityName="Latreille" authorityYear="1802" box="[436,531,1282,1308]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="9FFDEACE1F199562FE33FAA3DE69FAF7" box="[436,531,1282,1308]" italics="true" pageId="91" pageNumber="92">Bombus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="6A894D5F1F199562FDA5FAA0DEF4FAF7" authorityName="F. Morawitz" authorityYear="1887" box="[546,654,1281,1308]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F199562FDA5FAA0DEF4FAF7" bold="true" box="[546,654,1281,1308]" italics="true" pageId="91" pageNumber="92">keriensis</emphasis>
</taxonomicName>
/ 
<bibRefCitation id="C9184B2D1F199562FD1AFAA3DF18FAF7" author="Morawitz F. F." box="[669,866,1282,1308]" pageId="91" pageNumber="92" pagination="195 - 229" refId="ref68420" refString="Morawitz F. F. 1887. Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva 20 (1886): 195 - 229." type="journal article" year="1887">Morawitz, 1887</bibRefCitation>
/ det. 
<collectingCountry id="D59E764C1F199562FC2AFAA3DFA8FAF7" box="[941,978,1282,1308]" name="Philippines" pageId="91" pageNumber="92">PH</collectingCountry>
Williams 2017”. This specimen, which is complete, is recognised as one of Morawitz’s 
<typeStatus id="7232887E1F199562FCC3FA84DFD1FAD4" box="[836,939,1317,1343]" pageId="91" pageNumber="92" type="syntype">syntypes</typeStatus>
and is designated here as the 
<typeStatus id="7232887E1F199562FA8CFA84D90DFAD4" box="[1291,1399,1317,1343]" pageId="91" pageNumber="92" type="lectotype">lectotype</typeStatus>
in order to reduce uncertainty in the identity and application of the name.
</paragraph>
<paragraph id="AD3636DC1F199562FF3AFA28DE27FA4F" blockId="91.[189,605,1417,1444]" box="[189,605,1417,1444]" pageId="91" pageNumber="92">
<emphasis id="9FFDEACE1F199562FF3AFA28DDC8FA4F" bold="true" box="[189,434,1417,1444]" pageId="91" pageNumber="92">Material sequenced</emphasis>
(
<specimenCount id="BB8FFD551F199562FE44FA2BDE2FFA4F" box="[451,597,1417,1444]" pageId="91" pageNumber="92" type="generic">7 specimens</specimenCount>
)
</paragraph>
<paragraph id="AD3636DC1F199562FF3AFA19DE1AF907" blockId="91.[189,1399,1464,1772]" pageId="91" pageNumber="92">
<materialsCitation id="1DE13C811F199562FF3AFA19DF60FA1D" ID-GBIF-Occurrence="2867606525" collectingDate="1985-08" collectionCode="PW" collectorName="P. Williams" country="India" latitude="34.0548" location="Mt Apharwat" longLatPrecision="6" longitude="74.3856" municipality="Gulmarg" pageId="91" pageNumber="92" specimenCode="ML405" specimenCount="1" specimenCount-female="1" stateProvince="Kashmir">
<collectingCountry id="D59E764C1F199562FF3AFA19DD74FA39" box="[189,270,1464,1490]" name="India" pageId="91" pageNumber="92">INDIA</collectingCountry>
– 
<emphasis id="9FFDEACE1F199562FEBFFA19DDDDFA39" bold="true" box="[312,423,1464,1490]" pageId="91" pageNumber="92">
<collectingRegion id="6F4DF83E1F199562FEBFFA19DDDDFA39" box="[312,423,1464,1490]" pageId="91" pageNumber="92">Kashmir</collectingRegion>
</emphasis>
• 
<specimenCount id="BB8FFD551F199562FE4CFA19DD87FA38" box="[459,509,1464,1491]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(queen); 
<collectingMunicipality id="4D52ACA61F199562FDFEFA19DE9EFA39" box="[633,740,1464,1490]" pageId="91" pageNumber="92">Gulmarg</collectingMunicipality>
, 
<location id="A85660071F199562FD71FA18DFE1FA39" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FD71FA18DFE1FA39" box="[758,923,1464,1491]" country="India" latitude="34.0548" longLatPrecision="6" longitude="74.3856" municipality="Gulmarg" name="Mt Apharwat" pageId="91" pageNumber="92" stateProvince="Kashmir">Mt Apharwat</location>
; 
<geoCoordinate id="C8BD501B1F199562FC2BFA19D847FA38" box="[940,1085,1464,1491]" degrees="34.0548" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="34.0548">34.0548° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FBC8FA19D8A6FA38" box="[1103,1244,1464,1491]" degrees="74.3856" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="74.3856">74.3856° E</geoCoordinate>
; 
<date id="D937101C1F199562FB6BFA19D908FA38" box="[1260,1394,1464,1491]" pageId="91" pageNumber="92" value="1985-08">
<collectingDate id="C973E9F41F199562FB6BFA19D908FA38" box="[1260,1394,1464,1491]" pageId="91" pageNumber="92" value="1985-08">Aug. 1985</collectingDate>
</date>
; 
<collectorName id="007C530A1F199562FF3AFA7DDD3BFA1E" box="[189,321,1499,1526]" pageId="91" pageNumber="92">P. Williams</collectorName>
leg.; BOLD seq: 6877G02; 
<collectionCode id="CB98AE191F199562FD0EFA7DDEC3FA1D" box="[649,697,1500,1526]" country="Thailand" name="Paleontological Collections" pageId="91" pageNumber="92">PW</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FD45FA7ADF60FA1D" box="[706,794,1499,1526]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="91" pageNumber="92">ML405</specimenCode>
</materialsCitation>
• 
<materialsCitation id="1DE13C811F199562FCB6FA7ADF0DF9F2" ID-GBIF-Occurrence="2867606516" collectingDate="2009-07-21" collectionCode="RR" country="India" latitude="34.0212" location="Mt Apharwat" longLatPrecision="6" longitude="74.3208" municipality="Gulmarg" pageId="91" pageNumber="92" specimenCode="ML204" specimenCount="1" specimenCount-female="1" stateProvince="Kashmir">
<specimenCount id="BB8FFD551F199562FCB6FA7ADF26FA1D" box="[817,860,1499,1526]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(queen); 
<collectingMunicipality id="4D52ACA61F199562FC4DFA7AD84FFA1E" box="[970,1077,1499,1525]" pageId="91" pageNumber="92">Gulmarg</collectingMunicipality>
, 
<location id="A85660071F199562FBC7FA7DD8A4FA1E" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FBC7FA7DD8A4FA1E" box="[1088,1246,1499,1526]" country="India" latitude="34.0212" longLatPrecision="6" longitude="74.3208" municipality="Gulmarg" name="Mt Apharwat" pageId="91" pageNumber="92" stateProvince="Kashmir">Mt Apharwat</location>
; 
<geoCoordinate id="C8BD501B1F199562FB6EFA7AD908FA1D" box="[1257,1394,1499,1526]" degrees="34.0212" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="34.0212">34.0212° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FF3AFA5EDD38F9F2" box="[189,322,1535,1561]" degrees="74.3208" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="74.3208">74.3208° E</geoCoordinate>
; 
<date id="D937101C1F199562FECAFA5EDDA6F9F2" box="[333,476,1534,1561]" pageId="91" pageNumber="92" value="2009-07-21">
<collectingDate id="C973E9F41F199562FECAFA5EDDA6F9F2" box="[333,476,1534,1561]" pageId="91" pageNumber="92" value="2009-07-21">21 Jul. 2009</collectingDate>
</date>
; BOLD seq: 1552D08; 
<collectionCode id="CB98AE191F199562FD6DFA5EDF6CF9F2" box="[746,790,1535,1561]" pageId="91" pageNumber="92">RR</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FC99FA5EDF0DF9F2" box="[798,887,1535,1561]" collectionCode="RR" pageId="91" pageNumber="92">ML204</specimenCode>
</materialsCitation>
• 
<materialsCitation id="1DE13C811F199562FC0CFA5EDFF8F9D7" ID-GBIF-Occurrence="2867606494" collectingDate="2007-07-30" collectionCode="RR" country="India" latitude="33.4746" location="Padum" longLatPrecision="6" longitude="76.8871" municipality="Zanskar" pageId="91" pageNumber="92" specimenCode="ML203" specimenCount="1" specimenCount-female="1" stateProvince="Kashmir">
<specimenCount id="BB8FFD551F199562FC0CFA5EDFCFF9F2" box="[907,949,1535,1561]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(queen); 
<collectingMunicipality id="4D52ACA61F199562FBA6FA5FD8FBF9F3" box="[1057,1153,1534,1560]" pageId="91" pageNumber="92">Zanskar</collectingMunicipality>
, 
<location id="A85660071F199562FB0CFA5FD89AF9F3" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FB0CFA5FD89AF9F3" box="[1163,1248,1534,1560]" country="India" latitude="33.4746" longLatPrecision="6" longitude="76.8871" municipality="Zanskar" name="Padum" pageId="91" pageNumber="92" stateProvince="Kashmir">Padum</location>
; 
<geoCoordinate id="C8BD501B1F199562FB6DFA5FD908F9F2" box="[1258,1394,1534,1561]" degrees="33.4746" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="33.4746">33.4746° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FF3AF983DD3EF9D7" box="[189,324,1570,1596]" degrees="76.8871" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="76.8871">76.8871° E</geoCoordinate>
; 
<date id="D937101C1F199562FEC8F983DD9BF9D7" box="[335,481,1570,1596]" pageId="91" pageNumber="92" value="2007-07-30">
<collectingDate id="C973E9F41F199562FEC8F983DD9BF9D7" box="[335,481,1570,1596]" pageId="91" pageNumber="92" value="2007-07-30">30 Jul. 2007</collectingDate>
</date>
; BOLD seq: 1552D11; 
<collectionCode id="CB98AE191F199562FD73F983DF5AF9D7" box="[756,800,1570,1596]" pageId="91" pageNumber="92">RR</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FCADF983DFF8F9D7" box="[810,898,1570,1596]" collectionCode="RR" pageId="91" pageNumber="92">ML203</specimenCode>
</materialsCitation>
• 
<materialsCitation id="1DE13C811F199562FC1EF983D869F9B4" ID-GBIF-Occurrence="2867606572" collectingDate="2011-07-31" collectionCode="RR" collectorName="R. Raina" country="India" latitude="33.4746" location="Padum" longLatPrecision="6" longitude="76.8871" municipality="Zanskar" pageId="91" pageNumber="92" specimenCode="ML243" specimenCount="1" specimenCount-female="1" stateProvince="Kashmir">
<specimenCount id="BB8FFD551F199562FC1EF983DFBFF9D7" box="[921,965,1570,1596]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(worker); same collection data as for preceding; 
<date id="D937101C1F199562FEB8F9E4DDAAF9B4" box="[319,464,1605,1631]" pageId="91" pageNumber="92" value="2011-07-31">
<collectingDate id="C973E9F41F199562FEB8F9E4DDAAF9B4" box="[319,464,1605,1631]" pageId="91" pageNumber="92" value="2011-07-31">31 Jul. 2011</collectingDate>
</date>
; 
<collectorName id="007C530A1F199562FE5CF9E4DE3AF9B4" box="[475,576,1605,1631]" pageId="91" pageNumber="92">R. Raina</collectorName>
leg.; BOLD seq: 1555C12; 
<collectionCode id="CB98AE191F199562FC01F9E4DFC8F9B4" box="[902,946,1605,1631]" pageId="91" pageNumber="92">RR</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FC3CF9E4D869F9B4" box="[955,1043,1605,1631]" collectionCode="RR" pageId="91" pageNumber="92">ML243</specimenCode>
</materialsCitation>
• 
<materialsCitation id="1DE13C811F199562FBADF9E4D8C5F969" ID-GBIF-Occurrence="2867606395" collectingDate="2012-07-23" collectionCode="RR" collectorName="R. Raina" country="India" latitude="34.539" location="Razdaan" longLatPrecision="6" longitude="74.6377" pageId="91" pageNumber="92" specimenCode="ML244" specimenCount="1" specimenCount-female="1" stateProvince="Kashmir">
<specimenCount id="BB8FFD551F199562FBADF9E4D82FF9B4" box="[1066,1109,1605,1631]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(worker); 
<location id="A85660071F199562FB57F9E4D94FF9B4" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FB57F9E4D94FF9B4" box="[1232,1333,1605,1631]" country="India" latitude="34.539" longLatPrecision="6" longitude="74.6377" name="Razdaan" pageId="91" pageNumber="92" stateProvince="Kashmir">Razdaan</location>
pass; 
<geoCoordinate id="C8BD501B1F199562FF3AF9C9DD3DF968" box="[189,327,1640,1667]" degrees="34.5390" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="34.539">34.5390° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FED5F9C9DDA2F968" box="[338,472,1640,1667]" degrees="74.6377" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="74.6377">74.6377° E</geoCoordinate>
; 
<date id="D937101C1F199562FE63F9C9DE0CF969" box="[484,630,1640,1666]" pageId="91" pageNumber="92" value="2012-07-23">
<collectingDate id="C973E9F41F199562FE63F9C9DE0CF969" box="[484,630,1640,1666]" pageId="91" pageNumber="92" value="2012-07-23">23 Jul. 2012</collectingDate>
</date>
; 
<collectorName id="007C530A1F199562FD05F9C8DE9DF969" box="[642,743,1640,1667]" pageId="91" pageNumber="92">R. Raina</collectorName>
leg.; BOLD seq: 1555D01; 
<collectionCode id="CB98AE191F199562FBB7F9C8D826F968" box="[1072,1116,1641,1667]" pageId="91" pageNumber="92">RR</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FBE1F9C9D8C5F969" box="[1126,1215,1640,1666]" collectionCode="RR" pageId="91" pageNumber="92">ML244</specimenCode>
</materialsCitation>
• 
<materialsCitation id="1DE13C811F199562FB51F9C9DDDCF922" ID-GBIF-Occurrence="2867606464" collectingDate="2012-07-24" collectionCode="RR" collectorName="R. Raina" country="India" latitude="34.6464" location="Dawar" longLatPrecision="6" longitude="74.7171" pageId="91" pageNumber="92" specimenCode="ML305" specimenCount="1" specimenCount-female="1" stateProvince="Himachal Pradesh">
<specimenCount id="BB8FFD551F199562FB51F9C9D97BF968" box="[1238,1281,1640,1667]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(worker); 
<location id="A85660071F199562FF3AF92DDD75F94D" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FF3AF92DDD75F94D" box="[189,271,1676,1702]" country="India" latitude="34.6464" longLatPrecision="6" longitude="74.7171" name="Dawar" pageId="91" pageNumber="92" stateProvince="Himachal Pradesh">Dawar</location>
; 
<geoCoordinate id="C8BD501B1F199562FE9AF92ADDD0F94D" box="[285,426,1675,1702]" degrees="34.6464" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5" value="34.6464">34.6464° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FE3FF92ADE3BF94D" box="[440,577,1675,1702]" degrees="74.7171" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="5" value="74.7171">74.7171° E</geoCoordinate>
; 
<date id="D937101C1F199562FDD7F92ADE92F94E" box="[592,744,1675,1701]" pageId="91" pageNumber="92" value="2012-07-24">
<collectingDate id="C973E9F41F199562FDD7F92ADE92F94E" box="[592,744,1675,1701]" pageId="91" pageNumber="92" value="2012-07-24">24 Jul. 2012</collectingDate>
</date>
; 
<collectorName id="007C530A1F199562FD70F92DDF25F94E" box="[759,863,1675,1702]" pageId="91" pageNumber="92">R. Raina</collectorName>
leg.; BOLD seq: 1555H03; 
<collectionCode id="CB98AE191F199562FB30F92DD899F94D" box="[1207,1251,1676,1702]" pageId="91" pageNumber="92">RR</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FB77F92AD936F94D" box="[1264,1356,1675,1702]" collectionCode="RR" pageId="91" pageNumber="92">ML305</specimenCode>
. — 
<emphasis id="9FFDEACE1F199562FF3AF90FDDDCF922" bold="true" box="[189,422,1710,1737]" pageId="91" pageNumber="92">
<collectingRegion id="6F4DF83E1F199562FF3AF90FDDDCF922" box="[189,422,1710,1737]" country="India" name="Himachal Pradesh" pageId="91" pageNumber="92">Himachal Pradesh</collectingRegion>
</emphasis>
</materialsCitation>
• 
<materialsCitation id="1DE13C811F199562FE38F90EDE26F907" ID-GBIF-Occurrence="2867606336" collectingDate="1986-09-08" collectionCode="PW" collectorName="A. Hutchings" country="India" latitude="31.6" location="Saichu Nulla" longLatPrecision="5555" longitude="78.267" pageId="91" pageNumber="92" specimenCode="ML159" specimenCount="1" specimenCount-female="1" stateProvince="Kashmir">
<specimenCount id="BB8FFD551F199562FE38F90EDD96F922" box="[447,492,1711,1737]" pageId="91" pageNumber="92" type="female">1 ♀</specimenCount>
(worker); 
<location id="A85660071F199562FDEDF90FDF7DF923" LSID="urn:lsid:plazi:treatment:252087CA1F1B9565FDCAFB5DD92EFEBA:A85660071F199562FDEDF90FDF7DF923" box="[618,775,1710,1736]" country="India" latitude="31.6" longLatPrecision="5555" longitude="78.267" name="Saichu Nulla" pageId="91" pageNumber="92" stateProvince="Kashmir">Saichu Nulla</location>
; 
<geoCoordinate id="C8BD501B1F199562FC94F90FDF08F922" box="[787,882,1710,1737]" degrees="31.6" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="5555" value="31.6">31.6° N</geoCoordinate>
, 
<geoCoordinate id="C8BD501B1F199562FCF9F90FDF8DF922" box="[894,1015,1710,1737]" degrees="78.267" direction="east" orientation="longitude" pageId="91" pageNumber="92" precision="55" value="78.267">78.267° E</geoCoordinate>
; 
<date id="D937101C1F199562FB83F90ED8EEF923" box="[1028,1172,1710,1737]" pageId="91" pageNumber="92" value="1986-09-08">
<collectingDate id="C973E9F41F199562FB83F90ED8EEF923" box="[1028,1172,1710,1737]" pageId="91" pageNumber="92" value="1986-09-08">8 Sep. 1986</collectingDate>
</date>
; 
<collectorName id="007C530A1F199562FB19F90ED946F923" box="[1182,1340,1710,1737]" pageId="91" pageNumber="92">A. Hutchings</collectorName>
leg.; BOLD seq: 6877B05; 
<collectionCode id="CB98AE191F199562FE41F973DD8CF907" box="[454,502,1746,1772]" country="Thailand" name="Paleontological Collections" pageId="91" pageNumber="92">PW</collectionCode>
: 
<specimenCode id="FD2F9EA71F199562FD87F970DE26F907" box="[512,604,1745,1772]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="91" pageNumber="92">ML159</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E59365571F199562FF3AF8B2DE4BF849" pageId="91" pageNumber="92" type="distribution">
<paragraph id="AD3636DC1F199562FF3AF8B2DDD7F8C6" blockId="91.[189,429,1811,1837]" box="[189,429,1811,1837]" pageId="91" pageNumber="92">
<heading id="F67E81B01F199562FF3AF8B2DDD7F8C6" bold="true" box="[189,429,1811,1837]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="9FFDEACE1F199562FF3AF8B2DDD7F8C6" bold="true" box="[189,429,1811,1837]" pageId="91" pageNumber="92">Global distribution</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F199562FF3AF8E3DE4BF849" blockId="91.[189,1399,1857,1955]" pageId="91" pageNumber="92">
(West Qinghai-Tibetan-Plateau species) 
<emphasis id="9FFDEACE1F199562FD19F8E0DF62F8B0" bold="true" box="[670,792,1857,1883]" pageId="91" pageNumber="92">Himalaya</emphasis>
: 
<collectingCountry id="D59E764C1F199562FCAEF8E3DF80F8B7" box="[809,1018,1858,1884]" name="Afghanistan" pageId="91" pageNumber="92">AFGHANISTAN</collectingCountry>
, 
<collectingCountry id="D59E764C1F199562FB8EF8E3D8E2F8B7" box="[1033,1176,1858,1884]" name="Pakistan" pageId="91" pageNumber="92">PAKISTAN</collectingCountry>
, 
<collectingCountry id="D59E764C1F199562FB21F8E3D884F8B7" box="[1190,1278,1858,1884]" name="India" pageId="91" pageNumber="92">INDIA</collectingCountry>
: Kashmir, 
<collectingRegion id="6F4DF83E1F199562FF3AF8C4DDEDF894" box="[189,407,1893,1919]" country="India" name="Himachal Pradesh" pageId="91" pageNumber="92">Himachal Pradesh</collectingRegion>
. – 
<emphasis id="9FFDEACE1F199562FE30F8C4DE51F894" bold="true" box="[439,555,1893,1919]" pageId="91" pageNumber="92">East Asia</emphasis>
: 
<collectingCountry id="D59E764C1F199562FDBCF8C4DEE7F894" box="[571,669,1893,1919]" name="China" pageId="91" pageNumber="92">CHINA</collectingCountry>
: 
<collectingRegion id="6F4DF83E1F199562FD20F8C4DF7AF894" box="[679,768,1893,1919]" country="China" name="Xizang" pageId="91" pageNumber="92">Xizang</collectingRegion>
. (IOZ, NHMUK, PW, RR, ZIN, ZSM.) The species is usually not common (
<figureCitation id="35B22A591F199562FE3AF829DE5EF849" box="[445,548,1928,1954]" captionStart="Figs 211–212" captionStartId="100.[189,241,1742,1768]" captionTargetBox="[319,1269,265,1703]" captionTargetId="figure-10@100.[319,1269,265,1703]" captionTargetPageId="100" captionText="Figs 211–212. Maps of sequenced samples showing the most frequent colour patterns locally. 211. The sichelii-complex, consisting of the single polytypic species B. sichelii Radoszkowski, 1859 (inset: B. sichelii worker, China-Neimenggu, photo PW). 212. The keriensis-complex, consisting of B. alagesianus Reinig, 1930 stat. rev. (blue spots), B. tibeticus sp. nov. (dark green spots), B. incertoides Vogt, 1911 stat. rev. (light green spots), B. qilianensis sp. nov. (yellow spots), B. keriensis Morawitz, 1887 (orange spots), and B. separandus Vogt, 1909 stat. rev. (red spots) (inset: B.qilianensis sp. nov. queen, China-Sichuan, photo PW). Maps projected onto a sphere in ArcGIS using World_Shaded_Relief basemap © 2014 ESRI and showing boundaries between countries as recognised by the UN." figureDoi="http://doi.org/10.5281/zenodo.4064372" httpUri="https://zenodo.org/record/4064372/files/figure.png" pageId="91" pageNumber="92">Fig. 212</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1E9565FF3AFEA9D92EFEBA" box="[189,1364,264,337]" pageId="92" pageNumber="93" type="biology_ecology">
<paragraph id="AD3636DC1F1E9565FF3AFEA9DD45FEC9" blockId="92.[189,319,264,290]" box="[189,319,264,290]" pageId="92" pageNumber="93">
<heading id="F67E81B01F1E9565FF3AFEA9DD45FEC9" bold="true" box="[189,319,264,290]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="9FFDEACE1F1E9565FF3AFEA9DD45FEC9" bold="true" box="[189,319,264,290]" pageId="92" pageNumber="93">Behaviour</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFE96D92EFEBA" blockId="92.[189,1364,311,337]" box="[189,1364,311,337]" pageId="92" pageNumber="93">
Food-plant generalists (
<bibRefCitation id="C9184B2D1F1E9565FE55FE96DEFEFEBA" author="Williams P. H." box="[466,644,311,337]" pageId="92" pageNumber="93" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams 1991</bibRefCitation>
). Male mate-searching patrolling behaviour (
<bibRefCitation id="C9184B2D1F1E9565FB13FE96D93DFEBA" author="Williams P. H." box="[1172,1351,311,337]" pageId="92" pageNumber="93" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>