193 lines
23 KiB
XML
193 lines
23 KiB
XML
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<mods:title id="7A80206A89C4A0BD5B91029280924221">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<mods:namePart id="0DC07B430F282773A60B58D9C18E9D64">Ho, - Z.</mods:namePart>
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<mods:title id="43C7FF042A657A2F426972457B0823C2">Zootaxa</mods:title>
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<mods:date id="E00D85B34305E652FF42BC9F8F32A315">2019</mods:date>
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<mods:number id="F1D485C9EACD68E660C5181AAB40B732">4687</mods:number>
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<treatment id="6C3DA91C5101186AFF0EF966EE5C1E5B" ID-DOI="http://doi.org/10.5281/zenodo.4688825" ID-GBIF-Taxon="159762448" ID-Zenodo-Dep="4688825" LSID="urn:lsid:plazi:treatment:6C3DA91C5101186AFF0EF966EE5C1E5B" httpUri="http://treatment.plazi.org/id/6C3DA91C5101186AFF0EF966EE5C1E5B" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59">
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<paragraph id="E42B180A5101186CFF0EF966ED191B53" blockId="58.[151,545,1785,1812]" box="[151,545,1785,1812]" pageId="58" pageNumber="59">
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<heading id="BF63AF665101186CFF0EF966ED191B53" bold="true" box="[151,545,1785,1812]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<taxonomicName id="239463895101186CFF0EF966ED191B53" ID-CoL="8NC9Y" authority="Olivier 1911 b" authorityName="Olivier" authorityYear="1911" box="[151,545,1785,1812]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carolinae">
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<emphasis id="D6E0C4185101186CFF0EF966ED191B53" bold="true" box="[151,545,1785,1812]" pageId="58" pageNumber="59">
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<emphasis id="D6E0C4185101186CFF0EF966EE421B53" bold="true" box="[151,378,1785,1812]" italics="true" pageId="58" pageNumber="59">Atyphella carolinae</emphasis>
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<bibRefCitation id="800565FB5101186CFE18F966ED191B53" author="Olivier" box="[385,545,1785,1812]" firstAuthor="Olivier" pageId="58" pageNumber="59" pagination="171 - 175" refId="ref120903" refString="Olivier, E. (1911 b) Le genre Atyphella Oll. et descriptions des especes nouvelle (Col. Lampyridae). Bulletin de la Societe Entomologique de France, 8, 171 - 175. https: // doi. org / 10.5962 / bhl. part. 2971" type="journal article" year="1911" yearSuffix="b">Olivier 1911b</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B815101186CFF0EF8A1EB6B1BD3" pageId="58" pageNumber="59" type="reference_group">
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<paragraph id="E42B180A5101186CFF0EF8A1EBD51B10" blockId="58.[151,1261,1854,1971]" box="[151,1261,1854,1879]" pageId="58" pageNumber="59">
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<taxonomicName id="239463895101186CFF0EF8A1ED101B11" ID-CoL="8NC9Y" authority="Olivier 1911 b: 173" authorityName="Olivier" authorityPageNumber="173" authorityYear="1911" box="[151,552,1854,1879]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carolinae">
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<emphasis id="D6E0C4185101186CFF0EF8A1EE671B11" box="[151,351,1854,1878]" italics="true" pageId="58" pageNumber="59">Atyphella carolinae</emphasis>
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<treatmentCitation id="65353E1B5101186CFEFCF8A1ED101B11" author="Olivier, E." box="[357,552,1854,1879]" page="173" pageId="58" pageNumber="59" year="1911">
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<bibRefCitation id="800565FB5101186CFEFCF8A1ED101B11" author="Olivier" box="[357,552,1854,1879]" firstAuthor="Olivier" pageId="58" pageNumber="59" pagination="171 - 175" refId="ref120903" refString="Olivier, E. (1911 b) Le genre Atyphella Oll. et descriptions des especes nouvelle (Col. Lampyridae). Bulletin de la Societe Entomologique de France, 8, 171 - 175. https: // doi. org / 10.5962 / bhl. part. 2971" type="journal article" year="1911" yearSuffix="b">Olivier 1911b: 173</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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<treatmentCitation id="65353E1B5101186CFDA8F8A1EDDF1B10" author="Wittmer, W." box="[561,743,1854,1879]" page="67" pageId="58" pageNumber="59" year="1958">
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<bibRefCitation id="800565FB5101186CFDA8F8A1EDDF1B10" author="Wittmer" box="[561,743,1854,1879]" firstAuthor="Wittmer" pageId="58" pageNumber="59" pagination="67 - 68" refId="ref122540" refString="Wittmer, W. (1958) Insects of Micronesia Coleoptera: Lampyridae: Cantharidae, Malachiidae, Prionoceridae. Bernice P. Bishop Museum Insects of Micronesia, 16 (2), 67 - 68." type="journal article" year="1958">Wittmer 1958: 67</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="65353E1B5101186CFD68F8A1ECAB1B11" author="Blair, K. G." box="[753,915,1854,1879]" page="134" pageId="58" pageNumber="59" year="1940">
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<bibRefCitation id="800565FB5101186CFD68F8A1ECAB1B11" author="Blair" box="[753,915,1854,1879]" firstAuthor="Blair" pageId="58" pageNumber="59" pagination="131" refId="ref115247" refString="Blair, K. G. (1940) Coleoptera from the Caroline Islands. Occasional Papers of Bernice P. Bishop Museum, Honolulu Hawaii, XVI (6), 131." type="journal article" year="1940">Blair 1940: 134</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="65353E1B5101186CFC04F8A1EBD11B10" author="Ballantyne, L. A. & Lambkin, C." box="[925,1257,1854,1879]" page="15" pageId="58" pageNumber="59" year="2000">
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<bibRefCitation id="800565FB5101186CFC04F8A1EBD11B10" author="Ballantyne & Lambkin" box="[925,1257,1854,1879]" firstAuthor="Ballantyne" pageId="58" pageNumber="59" pagination="15 - 93" refId="ref114393" refString="Ballantyne, L. A. & Lambkin, C. (2000) Lampyridae of Australia (Coleoptera: Lampyridae: Luciolinae: Luciolini). Memoirs of the Queensland Museum, 46 (1), 15 - 93." type="journal article" year="2000">Ballantyne & Lambkin 2000: 15</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="E42B180A5101186CFF0EF8C2EB791B32" blockId="58.[151,1261,1854,1971]" box="[151,1089,1884,1909]" pageId="58" pageNumber="59">
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<treatmentCitationGroup id="C4843F245101186CFF0EF8C2EB791B32" box="[151,1089,1884,1909]" pageId="58" pageNumber="59">
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<taxonomicName id="239463895101186CFF0EF8C2ED2A1B32" ID-CoL="8MYP6" authority="(Olivier)" baseAuthorityName="Olivier" box="[151,530,1884,1909]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carolinae" subGenus="Luciola">
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<emphasis id="D6E0C4185101186CFF0EF8C2EE881B32" box="[151,432,1885,1909]" italics="true" pageId="58" pageNumber="59">Luciola (Luciola) carolinae</emphasis>
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(Olivier)
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</taxonomicName>
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.
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<treatmentCitation id="65353E1B5101186CFD82F8C1EC391B32" author="McDermott, F. A." box="[539,769,1884,1909]" page="100" pageId="58" pageNumber="59" year="1966">
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<bibRefCitation id="800565FB5101186CFD82F8C1EC391B32" author="McDermott" box="[539,769,1884,1909]" firstAuthor="McDermott" pageId="58" pageNumber="59" pagination="1 - 149" refId="ref119380" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.), Coleopterorum Catalogus Supplementa. Pars 9. Editio Secunda. W. Junk, S'Gravenhage, pp. 1 - 149." type="book chapter" year="1966">McDermott 1966: 100</bibRefCitation>
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</treatmentCitation>
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.
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||
<treatmentCitation id="65353E1B5101186CFC92F8C3EB061B32" author="Ballantyne, L. A. & Buck, E." box="[779,1086,1884,1909]" page="121" pageId="58" pageNumber="59" year="1979">
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<bibRefCitation id="800565FB5101186CFC92F8C3EB061B32" author="Ballantyne & Buck" box="[779,1086,1884,1909]" firstAuthor="Ballantyne" pageId="58" pageNumber="59" pagination="117 - 137" refId="ref114081" refString="Ballantyne, L. A. & Buck, E. (1979) Taxonomy and behavior of Luciola (Luciola) aphrogeneia, a new surf firefly from Papua New Guinea. Transactions of the American Entomological Society, 105, 117 - 137." type="journal article" year="1979">Ballantyne & Buck 1979: 121</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="E42B180A5101186CFF0EF8E3EB6B1BD3" blockId="58.[151,1261,1854,1971]" box="[151,1107,1915,1940]" pageId="58" pageNumber="59">
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<treatmentCitationGroup id="C4843F245101186CFF0EF8E3EB6B1BD3" box="[151,1107,1915,1940]" pageId="58" pageNumber="59">
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<taxonomicName id="239463895101186CFF0EF8E3EEF71BD3" ID-CoL="8MYTM" authority="(Olivier)" baseAuthorityName="Olivier" box="[151,463,1915,1940]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="carolinae">
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<emphasis id="D6E0C4185101186CFF0EF8E3EE551BD3" box="[151,365,1916,1940]" italics="true" pageId="58" pageNumber="59">Magnalata carolinae</emphasis>
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(Olivier)
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</taxonomicName>
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.
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<treatmentCitation id="65353E1B5101186CFE41F8E4EC1B1BD3" ID-DOI="http://doi.org/10.5281/zenodo.5324394" ID-Zenodo-Dep="5324394" author="Ballantyne, L. A. & Lambkin, C. L." box="[472,803,1915,1940]" httpUri="http://treatment.plazi.org/id/0394D665BE6FFFF8FF3C543626CEEA42" page="68" pageId="58" pageNumber="59" year="2009">
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<bibRefCitation id="800565FB5101186CFE41F8E4EC1B1BD3" author="Ballantyne & Lambkin" box="[472,803,1915,1940]" firstAuthor="Ballantyne" pageId="58" pageNumber="59" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009: 68</bibRefCitation>
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</treatmentCitation>
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, figs 225, 226, 233, 234, 237.
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B815101186CFF0EF804EDA81BF5" box="[151,656,1946,1971]" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="E42B180A5101186CFF0EF804EDA81BF5" blockId="58.[151,1261,1854,1971]" box="[151,656,1946,1971]" pageId="58" pageNumber="59">
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<materialsCitation id="54FC12575101186CFF0EF804EDB41BF5" ID-GBIF-Occurrence="2427486670" box="[151,652,1946,1971]" collectionCode="NHML" country="Palau" location="REPUBLIC OF PALAU" pageId="58" pageNumber="59" specimenCount="1" typeStatus="holotype">
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<emphasis id="D6E0C4185101186CFF0EF804EFEA1BF4" bold="true" box="[151,210,1947,1971]" pageId="58" pageNumber="59">
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<typeStatus id="3B2FA6A85101186CFF0EF804EFF61BF4" box="[151,206,1947,1971]" pageId="58" pageNumber="59">Type</typeStatus>
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.
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</emphasis>
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Male.
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<emphasis id="D6E0C4185101186CFE83F805ED1F1BF4" bold="true" box="[282,551,1946,1971]" pageId="58" pageNumber="59">
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<collectingCountry id="9C83589A5101186CFE83F805ED1A1BF4" box="[282,546,1946,1971]" name="Palau" pageId="58" pageNumber="59">REPUBLIC OF PALAU</collectingCountry>
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||
.
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</emphasis>
|
||
(
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<collectionCode id="828580CF5101186CFDACF805EDBE1BF5" box="[565,646,1946,1970]" collectionName="Libya, Tripoli, Natural History Museum" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="58" pageNumber="59">NHML</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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</subSubSection>
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<caption id="B0EB48825100186DFF0EF81BEE2C1BA2" ID-DOI="http://doi.org/10.5281/zenodo.3508178" ID-Zenodo-Dep="3508178" httpUri="https://zenodo.org/record/3508178/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,264,1924,1949]" targetBox="[203,1377,166,1905]" targetPageId="59">
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<paragraph id="E42B180A5100186DFF0EF81BEE2C1BA2" blockId="59.[151,1436,1924,2021]" pageId="59" pageNumber="60">
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<emphasis id="D6E0C4185100186DFF0EF81BEE521BDA" bold="true" box="[151,362,1924,1949]" pageId="59" pageNumber="60">FIGURES 180−187</emphasis>
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.
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<taxonomicName id="239463895100186DFEE0F81AED671BDA" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1886" box="[377,607,1925,1949]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="abdominalis" status="comb. nov.">
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<emphasis id="D6E0C4185100186DFEE0F81AED671BDA" box="[377,607,1925,1949]" italics="true" pageId="59" pageNumber="60">Atyphella abdominalis</emphasis>
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||
</taxonomicName>
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||
<emphasis id="D6E0C4185100186DFDFEF81AEDE51BDA" bold="true" box="[615,733,1925,1949]" pageId="59" pageNumber="60">
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||
<taxonomicNameLabel id="CDD379635100186DFDFEF81AEDE51BDA" box="[615,733,1925,1949]" pageId="59" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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male (USNM). 180, 181 habitus: 180 dorsal; 181 ventral; 182, 184 abdomen: 182 ventral; 184 dorsal; 183 pronotum dorsal; 185−187 aedeagus in aedeagal sheath: 185 ventral; 186 dorsal; 187 right lateral.
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</paragraph>
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||
</caption>
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||
<subSubSection id="AC8E4B815107186AFF0EFF07EE5C1E5B" pageId="60" pageNumber="61" type="diagnosis">
|
||
<paragraph id="E42B180A5107186AFF0EFF07EE5C1E5B" blockId="60.[151,1437,152,540]" pageId="60" pageNumber="61">
|
||
<emphasis id="D6E0C4185107186AFF0EFF07EE2A1CF4" bold="true" box="[151,274,152,179]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
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||
Repeated with slight modifications from
|
||
<bibRefCitation id="800565FB5107186AFD75FF06EB581CF4" author="Ballantyne & Lambkin" box="[748,1120,153,179]" firstAuthor="Ballantyne" pageId="60" pageNumber="61" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009: 68</bibRefCitation>
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||
. Males exist in three colour variations: 1. Dorsal surface of elytra and pronotum entirely brown to dark brown (
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||
<bibRefCitation id="800565FB5107186AFBB6FF22EA481C90" author="Ballantyne & Lambkin" box="[1071,1392,189,215]" firstAuthor="Ballantyne" pageId="60" pageNumber="61" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009</bibRefCitation>
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||
fig. 225), or pronotum may be mottled brownish orange (
|
||
<bibRefCitation id="800565FB5107186AFD77FF7EEB0B1CBC" author="Ballantyne & Lambkin" box="[750,1075,225,251]" firstAuthor="Ballantyne" pageId="60" pageNumber="61" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009</bibRefCitation>
|
||
fig. 226). 2. Pronotum, MS and MN orange with extensive darker brown markings, elytra dark brown. 3. Pronotum MS and MN orange with no darker markings; elytra dark brown. Colour pattern 1 similar to
|
||
<taxonomicName id="239463895107186AFCCDFEB6EBBA1D03" authority="(Olivier)" baseAuthorityName="Olivier" box="[852,1154,297,324]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="hypocrita">
|
||
<emphasis id="D6E0C4185107186AFCCDFEB6EB221D04" box="[852,1050,297,323]" italics="true" pageId="60" pageNumber="61">Luciola hypocrita</emphasis>
|
||
(Olivier)
|
||
</taxonomicName>
|
||
, which has a black dorsal surface, very shiny elytra (elytral surface dull in
|
||
<taxonomicName id="239463895107186AFD37FED2EC201D20" box="[686,792,333,359]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="D6E0C4185107186AFD37FED2EC201D20" box="[686,792,333,359]" italics="true" pageId="60" pageNumber="61">carolinae</emphasis>
|
||
</taxonomicName>
|
||
), no flat hypomeral areas (posterior half flat in
|
||
<taxonomicName id="239463895107186AFABBFED2EAB41D20" box="[1314,1420,333,359]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="D6E0C4185107186AFABBFED2EAB41D20" box="[1314,1420,333,359]" italics="true" pageId="60" pageNumber="61">carolinae</emphasis>
|
||
</taxonomicName>
|
||
), a pale ventral surface, no apparent LO in
|
||
<date id="902A3ECA5107186AFDF3FEEEEDB71DCC" box="[618,655,369,395]" pageId="60" pageNumber="61">V7</date>
|
||
, paired LOs in
|
||
<date id="902A3ECA5107186AFCA5FEEEEC661DCC" box="[828,862,369,395]" pageId="60" pageNumber="61">V6</date>
|
||
(full LOs in both these ventrites in
|
||
<taxonomicName id="239463895107186AFB6BFEEEEA641DCC" box="[1266,1372,369,395]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="D6E0C4185107186AFB6BFEEEEA641DCC" box="[1266,1372,369,395]" italics="true" pageId="60" pageNumber="61">carolinae</emphasis>
|
||
</taxonomicName>
|
||
), and posterolateral eye excavations (no eye excavations in
|
||
<taxonomicName id="239463895107186AFD64FE0AEC5F1DE8" box="[765,871,405,431]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="carolinae">
|
||
<emphasis id="D6E0C4185107186AFD64FE0AEC5F1DE8" box="[765,871,405,431]" italics="true" pageId="60" pageNumber="61">carolinae</emphasis>
|
||
</taxonomicName>
|
||
). Distinguished from
|
||
<taxonomicName id="239463895107186AFBFAFE0AEEBD1D94" authority="Ballantyne et Buck 1979" authorityName="Ballantyne et Buck" authorityYear="1979" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="aphrogeneia">
|
||
<emphasis id="D6E0C4185107186AFBFAFE0AEA5C1DE8" box="[1123,1380,405,431]" italics="true" pageId="60" pageNumber="61">Atyphella aphrogeneia</emphasis>
|
||
<bibRefCitation id="800565FB5107186AFAF5FE0AEEBD1D94" author="Ballantyne et Buck" firstAuthor="Ballantyne" pageId="60" pageNumber="61" pagination="117 - 137" refId="ref114081" refString="Ballantyne, L. A. & Buck, E. (1979) Taxonomy and behavior of Luciola (Luciola) aphrogeneia, a new surf firefly from Papua New Guinea. Transactions of the American Entomological Society, 105, 117 - 137." type="journal article" year="1979">
|
||
Ballantyne
|
||
<emphasis id="D6E0C4185107186AFF69FE25EE3C1D94" box="[240,260,442,467]" italics="true" pageId="60" pageNumber="61">et</emphasis>
|
||
Buck 1979
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
by the variations in pronotal colouration, the pronotal width/humeral width ratio, and the apical elytral modifications in
|
||
<taxonomicName id="239463895107186AFE52FE42EDB21DB0" authorityName="Ballantyne et Buck" authorityYear="1979" box="[459,650,477,503]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="aphrogeneia">
|
||
<emphasis id="D6E0C4185107186AFE52FE42EDB21DB0" box="[459,650,477,503]" italics="true" pageId="60" pageNumber="61">Aty. aphrogeneia</emphasis>
|
||
</taxonomicName>
|
||
(the apical area of both suture and lateral margin as well as the elytral apex are swollen).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |