The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species Author Ho, - Z. text Zootaxa 2019 2019-10-18 4687 1 1 174 journal article 25190 10.11646/zootaxa.4687.1.1 de00ca6c-2843-4ae1-a39c-a29603947612 1175-5326 3508098 CE73264D-C234-4B82-A634-CAD6254C5957 Atyphella carolinae Olivier 1911b Atyphella carolinae Olivier 1911b: 173 . Wittmer 1958: 67 . Blair 1940: 134 . Ballantyne & Lambkin 2000: 15 . Luciola (Luciola) carolinae (Olivier) . McDermott 1966: 100 . Ballantyne & Buck 1979: 121 . Magnalata carolinae (Olivier) . Ballantyne & Lambkin 2009: 68 , figs 225, 226, 233, 234, 237. Type . Male. REPUBLIC OF PALAU . ( NHML ) . FIGURES 180−187 . Atyphella abdominalis comb. nov. male (USNM). 180, 181 habitus: 180 dorsal; 181 ventral; 182, 184 abdomen: 182 ventral; 184 dorsal; 183 pronotum dorsal; 185−187 aedeagus in aedeagal sheath: 185 ventral; 186 dorsal; 187 right lateral. Diagnosis. Repeated with slight modifications from Ballantyne & Lambkin 2009: 68 . Males exist in three colour variations: 1. Dorsal surface of elytra and pronotum entirely brown to dark brown ( Ballantyne & Lambkin 2009 fig. 225), or pronotum may be mottled brownish orange ( Ballantyne & Lambkin 2009 fig. 226). 2. Pronotum, MS and MN orange with extensive darker brown markings, elytra dark brown. 3. Pronotum MS and MN orange with no darker markings; elytra dark brown. Colour pattern 1 similar to Luciola hypocrita (Olivier) , which has a black dorsal surface, very shiny elytra (elytral surface dull in carolinae ), no flat hypomeral areas (posterior half flat in carolinae ), a pale ventral surface, no apparent LO in V7 , paired LOs in V6 (full LOs in both these ventrites in carolinae ), and posterolateral eye excavations (no eye excavations in carolinae ). Distinguished from Atyphella aphrogeneia Ballantyne et Buck 1979 by the variations in pronotal colouration, the pronotal width/humeral width ratio, and the apical elytral modifications in Aty. aphrogeneia (the apical area of both suture and lateral margin as well as the elytral apex are swollen).