269 lines
20 KiB
XML
269 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.859.33634" ID-GBIF-Dataset="1ebfdc7c-50af-4535-bee7-2fb58e3f2b81" ID-PMC="PMC6616056" ID-Pensoft-Pub="1313-2970-859-131" ID-PubMed="31327926" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-859-131" ModsDocOrigin="ZooKeys 859" ModsDocTitle="Call a spade a spade: taxonomy and distribution of Pelobates, with description of a new Balkan endemic" checkinTime="1562137562851" checkinUser="pensoft" docAuthor="Dufresnes, Christophe, Strachinis, Ilias, Tzoras, Elias, Litvinchuk, Spartak N. & Denoel, Mathieu" docDate="2019" docId="4866AFB610CE9977A9841AF1DC13C8C9" docLanguage="en" docName="ZooKeys 859: 131-158" docOrigin="ZooKeys 859" docSource="http://dx.doi.org/10.3897/zookeys.859.33634" docTitle="Pelobates varaldii Pasteur & Bons 1959" docType="treatment" docVersion="5" lastPageNumber="137" masterDocId="FFE13C3F7A123710D3023035D907FF8A" masterDocTitle="Call a spade a spade: taxonomy and distribution of Pelobates, with description of a new Balkan endemic" masterLastPageNumber="158" masterPageNumber="131" pageNumber="134" updateTime="1668167462434" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Call a spade a spade: taxonomy and distribution of Pelobates, with description of a new Balkan endemic</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dufresnes, Christophe</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Strachinis, Ilias</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tzoras, Elias</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Litvinchuk, Spartak N.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Denoel, Mathieu</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>859</mods:number>
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<mods:extent unit="page">
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<mods:start>131</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.859.33634</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.859.33634</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-859-131</mods:identifier>
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<treatment ID-GBIF-Taxon="158498312" LSID="urn:lsid:plazi:treatment:4866AFB610CE9977A9841AF1DC13C8C9" httpUri="http://treatment.plazi.org/id/4866AFB610CE9977A9841AF1DC13C8C9" lastPageId="6" lastPageNumber="137" pageId="3" pageNumber="134">
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<subSubSection pageId="3" pageNumber="134" type="nomenclature">
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<paragraph pageId="3" pageNumber="134">
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<taxonomicName authority="Pasteur & Bons, 1959" authorityName="Pasteur & Bons" authorityYear="1959" class="Amphibia" family="Pelobatidae" genus="Pelobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelobates varaldii" order="Anura" pageId="3" pageNumber="134" phylum="Chordata" rank="species" species="varaldii">Pelobates varaldii Pasteur & Bons, 1959</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="137" pageId="4" pageNumber="135" type="diagnosis">
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<paragraph pageId="4" pageNumber="135">
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<pageBreakToken pageId="4" pageNumber="135" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="137" pageId="4" pageNumber="135">
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A smaller version of
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<taxonomicName lsidName="P. cultripes" pageId="4" pageNumber="135" rank="species" species="cultripes">P. cultripes</taxonomicName>
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(Fig. 2) differing by a few phenotypic features. Unlike
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<taxonomicName lsidName="P. cultripes" pageId="4" pageNumber="135" rank="species" species="cultripes">P. cultripes</taxonomicName>
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, the black coloration of the spades is often concentrated on the edges (
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<bibRefCitation author="Pasteur, G" journalOrPublisher="Serie Zoologique 17, Rabat" pageId="23" pageNumber="154" title="Les batraciens du Maroc. Travaux de l'Institut Scientifique Cherifien." year="1959">Pasteur and Bons 1959</bibRefCitation>
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;
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<bibRefCitation author="Busack, SD" journalOrPublisher="Copeia" pageId="19" pageNumber="150" pagination="107 - 112" title="Pelobatesvaraldii (Anura: Pelobatidae): A morphologically conservative species." url="https://doi.org/10.2307/1444797" volume="1985" year="1985">Busack et al. 1985</bibRefCitation>
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). The cranial braincase is high and narrow in
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<taxonomicName lsidName="P. varaldii" pageId="4" pageNumber="135" rank="species" species="varaldii">P. varaldii</taxonomicName>
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, while it is low and wide in
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<taxonomicName lsidName="P. cultripes" pageId="4" pageNumber="135" rank="species" species="cultripes">P. cultripes</taxonomicName>
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(
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<bibRefCitation author="Pasteur, G" journalOrPublisher="Serie Zoologique 17, Rabat" pageId="23" pageNumber="154" title="Les batraciens du Maroc. Travaux de l'Institut Scientifique Cherifien." year="1959">Pasteur and Bons 1959</bibRefCitation>
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;
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<bibRefCitation pageId="4" pageNumber="135">
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<normalizedToken originalValue="Roček">Rocek</normalizedToken>
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1981
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</bibRefCitation>
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). The background coloration can be yellow, gray, and brown, with dark reticulate patches, and the dorsal surface is abundantly covered by orange dots, most pronounced on the eyelids (usually absent in
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<taxonomicName lsidName="P. cultripes" pageId="4" pageNumber="135" rank="species" species="cultripes">P. cultripes</taxonomicName>
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;
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<bibRefCitation author="Pasteur, G" journalOrPublisher="Serie Zoologique 17, Rabat" pageId="23" pageNumber="154" title="Les batraciens du Maroc. Travaux de l'Institut Scientifique Cherifien." year="1959">Pasteur and Bons 1959</bibRefCitation>
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;
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<bibRefCitation author="Beukema, W" journalOrPublisher="Zootaxa" pageId="18" pageNumber="149" pagination="1 - 60" title="Review of the systematics, distribution, biogeography and natural history of Moroccan amphibians." url="https://doi.org/10.11646/zootaxa.3661.1.1" volume="3661" year="2013">Beukema et al. 2013</bibRefCitation>
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; Fig. 3). Males are usually smaller than females (Fig. 2). Average SVL = 53 mm (range: 36-66 mm) for females (n = 4 populations) and 51 mm
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<pageBreakToken pageId="5" pageNumber="136" start="start">(</pageBreakToken>
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33-65 mm) for males (n = 4 populations) (Suppl. material 1, Table S1; Fig. 2). The karyotype includes six large and seven small pairs of two-armed chromosomes. Large centromeric C-bands appears in the pairs 1, 2, 4, 9, and 12; pericentric bands in the short arms of pair 1 and long arm of pair 8; telomeric bands in the long arms of pairs 1, 2, and 11; the short arm of pair 7 is almost heterochromatic (
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<bibRefCitation author="Herrero, P" journalOrPublisher="Acta Zoologica Cracoviensia" pageId="21" pageNumber="152" pagination="505 - 508" title="Cytotaxonomic studies on Iberian and Moroccan Pelobates (Anura: Pelobatidae)." volume="31" year="1988">Herrero and Talavera 1988</bibRefCitation>
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). The nuclear DNA content averages 7.3 pg (
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<bibRefCitation author="Litvinchuk, SN" journalOrPublisher="Organisms Diversity and Evolution," pageId="21" pageNumber="152" pagination="433 - 451" title="Phylogeographic patterns of genetic diversity in the common spadefoot toad, Pelobatesfuscus (Anura: Pelobatidae), reveals evolutionary history, postglacial range expansion and secondary contact." url="https://doi.org/10.1007/s13127-013-0127-5" volume="13" year="2013">Litvinchuk et al. 2013</bibRefCitation>
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). As shown
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<pageBreakToken pageId="6" pageNumber="137" start="start">in</pageBreakToken>
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Table 1,
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<taxonomicName lsidName="P. varaldii" pageId="6" pageNumber="137" rank="species" species="varaldii">P. varaldii</taxonomicName>
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differs from
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<taxonomicName lsidName="P. cultripes" pageId="6" pageNumber="137" rank="species" species="cultripes">P. cultripes</taxonomicName>
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by 6.0% at mtDNA and 0.40% at nuclear DNA (
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<bibRefCitation author="Dufresnes, C" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="20" pageNumber="151" url="https://doi.org/10.1111/mec.15133" year="2019 b">Dufresnes et al. 2019b</bibRefCitation>
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).
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</paragraph>
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<caption pageId="6" pageNumber="137">
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<paragraph pageId="6" pageNumber="137">
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Figure 2. Between-population variation of average size (
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<normalizedToken originalValue="snout–vent">snout-vent</normalizedToken>
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length - SVL) for each
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<taxonomicName class="Amphibia" family="Pelobatidae" genus="Pelobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelobates" order="Anura" pageId="6" pageNumber="137" phylum="Chordata" rank="genus">Pelobates</taxonomicName>
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species, measured separately for females (pink) and males (blue). This compiles average size-data from 82 populations, representing at least 6,004 individuals (Suppl. material 1, Table S1). For
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<taxonomicName lsidName="P. balcanicus" pageId="6" pageNumber="137" rank="species" species="balcanicus">P. balcanicus</taxonomicName>
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, it only includes populations from the nominal
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<taxonomicName lsidName="P. b." pageId="6" pageNumber="137" rank="subSpecies" species="b." subSpecies="balcanicus">P. b. balcanicus</taxonomicName>
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. For
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<taxonomicName lsidName="P. syriacus" pageId="6" pageNumber="137" rank="species" species="syriacus">P. syriacus</taxonomicName>
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, it only includes populations from
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<taxonomicName lsidName="P. s." pageId="6" pageNumber="137" rank="subSpecies" species="s." subSpecies="boettgeri">P. s. boettgeri</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="137">
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<paragraph pageId="6" pageNumber="137">
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Figure 3. Color variation in
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<taxonomicName class="Amphibia" family="Pelobatidae" genus="Pelobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelobates cultripes" order="Anura" pageId="6" pageNumber="137" phylum="Chordata" rank="species" species="cultripes">Pelobates cultripes</taxonomicName>
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,
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<taxonomicName lsidName="P. varaldii" pageId="6" pageNumber="137" rank="species" species="varaldii">P. varaldii</taxonomicName>
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,
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<taxonomicName lsidName="P. fuscus" pageId="6" pageNumber="137" rank="species" species="fuscus">P. fuscus</taxonomicName>
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and
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<taxonomicName lsidName="P. vespertinus" pageId="6" pageNumber="137" rank="species" species="vespertinus">P. vespertinus</taxonomicName>
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. Photo credits and origins as follows a CD (
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<normalizedToken originalValue="Hérault">Herault</normalizedToken>
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, France) b, c CD (Algarve, Portugal) d A Sanchez Vialas (Spain) e G Martinez (Kenitra, Morocco)
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<normalizedToken originalValue="f–h">f-h</normalizedToken>
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A Sanchez Vialas (Tanger, Morocco) i, j N Suriadna (Ukraine) k CD (Wojewodztwo podkarpackie, Poland) l A
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<normalizedToken originalValue="Nöllert">Noellert</normalizedToken>
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(Burgenland, Austria)
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<normalizedToken originalValue="m–p">m-p</normalizedToken>
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N Suriadna (Ukraine).
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="137">
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<paragraph pageId="6" pageNumber="137">
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Table 1. Pairwise % of genetic differences between
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<taxonomicName class="Amphibia" family="Pelobatidae" genus="Pelobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelobates" order="Anura" pageId="6" pageNumber="137" phylum="Chordata" rank="genus">Pelobates</taxonomicName>
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taxa (from the data of
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<bibRefCitation author="Dufresnes, C" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="20" pageNumber="151" url="https://doi.org/10.1111/mec.15133" year="2019 b">Dufresnes et al. 2019b</bibRefCitation>
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). The estimates below diagonal correspond to mitochondrial DNA (cyt-b + 16S, 1.2 kb); the estimates above diagonal correspond to nuclear DNA (63.5 kb of RAD tags).
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="137">
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<table pageId="6" pageNumber="137">
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. cultripes" pageId="6" pageNumber="137" rank="species" species="cultripes">P. cultripes</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. varaldii" pageId="6" pageNumber="137" rank="species" species="varaldii">P. varaldii</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. fuscus" pageId="6" pageNumber="137" rank="species" species="fuscus">P. fuscus</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. vespertinus" pageId="6" pageNumber="137" rank="species" species="vespertinus">P. vespertinus</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. s." pageId="6" pageNumber="137" rank="subSpecies" species="s." subSpecies="syriacus">P. s. syriacus</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. s." pageId="6" pageNumber="137" rank="subSpecies" species="s." subSpecies="boettgeri">P. s. boettgeri</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. b." pageId="6" pageNumber="137" rank="subSpecies" species="b." subSpecies="balcanicus">P. b. balcanicus</taxonomicName>
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</td>
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. b." pageId="6" pageNumber="137" rank="subSpecies" species="b." subSpecies="chloeae">P. b. chloeae</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. cultripes" pageId="6" pageNumber="137" rank="species" species="cultripes">P. cultripes</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. varaldii" pageId="6" pageNumber="137" rank="species" species="varaldii">P. varaldii</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. fuscus" pageId="6" pageNumber="137" rank="species" species="fuscus">P. fuscus</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. vespertinus" pageId="6" pageNumber="137" rank="species" species="vespertinus">P. vespertinus</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. s." pageId="6" pageNumber="137" rank="subSpecies" species="s." subSpecies="syriacus">P. s. syriacus</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. s." pageId="6" pageNumber="137" rank="subSpecies" species="s." subSpecies="boettgeri">P. s. boettgeri</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. b." pageId="6" pageNumber="137" rank="subSpecies" species="b." subSpecies="balcanicus">P. b. balcanicus</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="137">
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<td colspan="1" pageId="6" pageNumber="137" rowspan="1">
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<taxonomicName lsidName="P. b." pageId="6" pageNumber="137" rank="subSpecies" species="b." subSpecies="chloeae">P. b. chloeae</taxonomicName>
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="137" type="taxonomy">
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<paragraph pageId="6" pageNumber="137">Taxonomy.</paragraph>
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<paragraph pageId="6" pageNumber="137">
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The nomen
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<taxonomicName class="Amphibia" family="Pelobatidae" genus="Pelobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelobates varaldii" order="Anura" pageId="6" pageNumber="137" phylum="Chordata" rank="species" species="varaldii">Pelobates varaldii</taxonomicName>
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Pasteur & Bons, 1959 is the only one ever proposed for the Moroccan populations of spadefoot toads; holotype: MNHN-RA-1959.1; type locality: Merja Samora, Morocco. The ancient split of
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<taxonomicName lsidName="P. varaldii" pageId="6" pageNumber="137" rank="species" species="varaldii">P. varaldii</taxonomicName>
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, dating back to the Messinian Salinity Crisis (5.3 My), supports its specific distinction from
|
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<taxonomicName lsidName="P. cultripes" pageId="6" pageNumber="137" rank="species" species="cultripes">P. cultripes</taxonomicName>
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(
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<bibRefCitation author="Busack, SD" journalOrPublisher="Copeia" pageId="19" pageNumber="150" pagination="107 - 112" title="Pelobatesvaraldii (Anura: Pelobatidae): A morphologically conservative species." url="https://doi.org/10.2307/1444797" volume="1985" year="1985">Busack et al. 1985</bibRefCitation>
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||
;
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||
<bibRefCitation author="Crottini, A" journalOrPublisher="Molecular Ecology" pageId="19" pageNumber="150" pagination="2734 - 2754" title="Fossorial but widespread: the phylogeography of the common spadefoot toad (Pelobatesfuscus), and the role of the Po Valley as a major source of genetic variability." url="https://doi.org/10.1111/j.1365-294X.2007.03274.x" volume="16" year="2007">Crottini et al. 2007</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="137" type="distribution">
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<paragraph pageId="6" pageNumber="137">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="137">
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It is endemic to north-western Morocco (0-350 m elevation a.s.l.), found along the Atlantic coast, from the south of Tanger to Oualidia, where it is rare (
|
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<bibRefCitation author="de Pous, P" journalOrPublisher="Basic and Applied Herpetology" pageId="19" pageNumber="150" pagination="57 - 71" title="Distribution review, habitat suitability and conservation of the endangered and endemic Moroccan spadefoot toad (Pelobatesvaraldii)." url="https://doi.org/10.11160/bah.11015" volume="26" year="2012">de Pous et al. 2012</bibRefCitation>
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;
|
||
<bibRefCitation author="Beukema, W" journalOrPublisher="Zootaxa" pageId="18" pageNumber="149" pagination="1 - 60" title="Review of the systematics, distribution, biogeography and natural history of Moroccan amphibians." url="https://doi.org/10.11646/zootaxa.3661.1.1" volume="3661" year="2013">Beukema et al. 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Frost, D" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="20" pageNumber="151" url="http://research.amnh.org/herpetology/amphibia/index.html" year="2019">Frost 2019</bibRefCitation>
|
||
). IUCN status: Endangered (
|
||
<bibRefCitation author="Salvador, A" journalOrPublisher="Historia Naturalis Ranarum Nostratium in Qua Omnes earum Proprietates Praesertim quae ad Generationem Ipsarum Pertinent, Fusius Enarrantur / Die natuerliche Historie der Froesche hiesigen Landes, Albrecht von Haller, Nuernberg" pageId="23" pageNumber="154" url="https://doi.org/10.2305/IUCN.UK.2004.RLTS.T58054A11723994.en" year="2009">Salvador et al. 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="137" type="diversity">
|
||
<paragraph pageId="6" pageNumber="137">Diversity.</paragraph>
|
||
<paragraph pageId="6" pageNumber="137">
|
||
To our knowledge,
|
||
<taxonomicName lsidName="P. varaldii" pageId="6" pageNumber="137" rank="species" species="varaldii">P. varaldii</taxonomicName>
|
||
has not been the focus of any phylogeographic or population genetic work.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |