211 lines
21 KiB
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211 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.805.28757" ID-GBIF-Dataset="c0cb883f-3c6c-4d0b-a280-04157f2ad247" ID-PMC="PMC6299064" ID-Pensoft-Pub="1313-2970-805-45" ID-PubMed="30584392" ID-ZBK="5EF6D879D86F49279B12C0BC91DC6FF2" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-805-45" ModsDocOrigin="ZooKeys 805" ModsDocTitle="A new genus and species of Brontinae from Borneo (Coleoptera, Silvanidae)" checkinTime="1545315517247" checkinUser="pensoft" docAuthor="Yoshida, Takahiro & Hirowatari, Toshiya" docDate="2018" docId="B68C81A5A35C9034A095ACF9617E3691" docLanguage="en" docName="ZooKeys 805: 45-57" docOrigin="ZooKeys 805" docSource="http://dx.doi.org/10.3897/zookeys.805.28757" docTitle="Borneophanus Yoshida & Hirowatari, 2018, gen. n." docType="treatment" docUuid="E5B80A06-C636-4CFF-AA37-A3A86EB7EB49" docUuidSource="ZooBank" docVersion="5" lastPageNumber="49" masterDocId="FFA49D22D31FFF9FFFBDEF54FFFCFFEE" masterDocTitle="A new genus and species of Brontinae from Borneo (Coleoptera, Silvanidae)" masterLastPageNumber="57" masterPageNumber="45" pageNumber="47" updateTime="1668166574091" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and species of Brontinae from Borneo (Coleoptera, Silvanidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoshida, Takahiro</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hirowatari, Toshiya</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>805</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>57</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.805.28757</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.805.28757</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-805-45</mods:identifier>
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<mods:identifier type="ZBK">5EF6D879D86F49279B12C0BC91DC6FF2</mods:identifier>
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<mods:identifier type="ZooBank">5EF6D879D86F49279B12C0BC91DC6FF2</mods:identifier>
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<treatment ID-GBIF-Taxon="154126816" LSID="urn:lsid:zoobank.org:act:E5B80A06-C636-4CFF-AA37-A3A86EB7EB49" httpUri="http://treatment.plazi.org/id/B68C81A5A35C9034A095ACF9617E3691" lastPageId="4" lastPageNumber="49" pageId="2" pageNumber="47">
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<subSubSection pageId="2" pageNumber="47" type="nomenclature">
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<paragraph pageId="2" pageNumber="47">
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<taxonomicName LSID="http://zoobank.org/E5B80A06-C636-4CFF-AA37-A3A86EB7EB49" class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Borneophanus</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="47">gen. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="47" type="type species">
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<paragraph pageId="2" pageNumber="47">Type species.</paragraph>
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<paragraph pageId="2" pageNumber="47">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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Yoshida & Hirowatari, sp. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="47" type="diagnosis">
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<paragraph pageId="2" pageNumber="47">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="47">
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Among telephanine genera, this new genus shares the following character states with
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<taxonomicName class="Insecta" family="Silvanidae" genus="Telephanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telephanus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Telephanus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Silvanidae" genus="Indophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indophanus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Indophanus</taxonomicName>
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Pal, 1982: apical maxillary palpomere securiform; apical labial palpomere securiform; scutellary striole absent (some species of Malagasy
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<taxonomicName class="Insecta" family="Silvanidae" genus="Telephanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telephanus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Telephanus</taxonomicName>
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have a scutellary striole). This new genus differs from these genera by the combination of the following character states: distinct pair of longitudinal frontal lines present (absent in
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<taxonomicName class="Insecta" family="Silvanidae" genus="Telephanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telephanus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Telephanus</taxonomicName>
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); scutellar shield with a transverse carina and excavate posteriorly (flat in
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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); antennomere IV normal (markedly long, approximately twice as long as combined length of II and III in
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<taxonomicName class="Insecta" family="Silvanidae" genus="Indophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indophanus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Indophanus</taxonomicName>
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) (
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<bibRefCitation author="Thomas, MC" journalOrPublisher="Insecta Mundi" pageId="12" pageNumber="57" pagination="1 - 14" title="A new genus of telephanine Silvanidae (Coleoptera: Cucujoidea), with a diagnosis of the tribe and key to genera." url="http://digitalcommons.unl.edu/insectamundi/576" volume="0048" year="2008">Thomas and Nearns 2008</bibRefCitation>
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;
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<bibRefCitation author="Sen Gupta, T" journalOrPublisher="Zoological Survey of India, Calcutta" pageId="11" pageNumber="56" title="Fauna of India and the Adjacent Countries. Calvicornia [sic]: Coleoptera. Family Silvanidae." year="1996">Sen Gupta and Pal 1996</bibRefCitation>
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). In addition, this new genus possesses the following characteristic morphology: the asymmetric shaped antennomere X (somewhat asymmetric in
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<taxonomicName class="Insecta" family="Silvanidae" genus="Bolianus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bolianus" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">Bolianus</taxonomicName>
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); the sharply protruding elytral apices; the non-folded internal sac; the partly coiled flagellum; and the very long and rolled up spermathecal duct.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47" type="description">
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<paragraph pageId="2" pageNumber="47">Description.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
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Body densely covered with pubescence. Head (Figs 1-3) with indistinct frontoclypeal suture, with distinct pair of longitudinal frontal lines; eyes somewhat small, hemispherical; temples moderate in size; labrum (Fig. 3B) rectangular; antenna (Fig. 3A) long, with long scape; antennomere X asymmetrically enlarged; mandible (Fig. 3C, D) tridentate, with a ventral tooth and an inner lateral tooth near apex, ventrally with asperities near the inner tooth, with fine, random punctation, dorsolaterally with medium length to long setae, dorsally with a mycangium near base; maxilla (Fig. 3E) with lacinia and galea; galea divided into distigalea and basigalea; maxillary palp 4-segmented; apical palpomere securiform, dorsally and largely with digitiform sensilla on middle of apical palpomere; labium (Fig. 3F) divided from ligula; prementum widened distally; articulation of palps tight; labial palp 3-segmented; palpomere 2 extended outwards strongly and broadly, with large distal area; palpomere 3 securiform, strongly widened distally; mentum widened proximally, widest near base. Pronotum (Figs 1A, 2) without teeth on lateral margins, with rounded anterior angles, densely covered with setae of various lengths, without microsculpture. Thoracic ventrites (Fig. 1B) with narrowly separated pro- and mesocoxal cavities, with somewhat widely separated metacoxal cavities. Scutellar shield (Figs 1A, 2) with a transverse carina, excavate posteriorly. Legs (Figs 1, 4A, B) somewhat thin; tarsomeres 2 and 3 lobed, not bilobed. Abdominal ventrites (Fig. 4A) with somewhat wide intercoxal process, without sexual dimorphism. Elytra (Figs 1A, 2) long, without scutellary striole; lateral margins very narrowly explanate; apices somewhat elongate and narrowly rounded, with short setae on interstices. Male tergite and sternite VIII square and not divided (Fig. 4D); spiculum gastrale (Fig. 4E) thin and Y-shaped, with branches covered with membrane; median lobe (Fig. 4H) without setae; internal sac (Fig. 4H) not folded, with partly coiled flagellum, exposed around apex of median lobe; parameres (Fig. 4F, G) flat. Female with elongate gonostyli (Fig. 6B); gonocoxite
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<pageBreakToken pageId="3" pageNumber="48" start="start">(</pageBreakToken>
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Fig. 6B) with setae of various lengths; spermathecal duct (Figs 6A, 7A) connected to basal portion of bursa copulatrix, very long, rolled up and forming large ellipsoid.
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</paragraph>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 1. Habitus of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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gen. et sp. n., holotype, male. A dorsal B ventral aspect.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 2. Close-up image of head, pronotum, scutellar shield and basis of elytra of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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gen. et sp. n., holotype, male.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 3. Antenna and mouth parts of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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gen. et sp. n., paratype, male. A Right antenna B labrum, dorsal view C, D right mandible, ventral view (C) and dorsal view (D) E right maxilla, dorsal view F labium, ventral view. An arrow indicates digitiform sensilla.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 4. Tarsomeres and abdominal parts including the male genital structures of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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gen. et sp. n., holotype. A, B Tarsomeres, dorsal view (A) and lateral view (B) C abdominal ventries D eighth abdominal segments, ventral view E spiculum gastrale, ventral view F tegmen, ventral view G paramere, dorsal view H median lobe and internal sac, ventral view.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 5. SEM images of apical palpomere of left maxillae (
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) and a right maxilla (D), dorsal view, with digitiform sensilla. A
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus dentatus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="dentatus">Psammoecus dentatus</taxonomicName>
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B
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<taxonomicName class="Insecta" family="Silvanidae" genus="Telephanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telephanus paradoxus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="paradoxus">Telephanus paradoxus</taxonomicName>
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C
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<taxonomicName class="Insecta" family="Silvanidae" genus="Australohyliota" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Australohyliota mcleayi" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="mcleayi">Australohyliota mcleayi</taxonomicName>
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D
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<taxonomicName class="Insecta" family="Cucujidae" genus="Cucujus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucujus bicolor" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">Cucujus bicolor</taxonomicName>
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. Circles indicate digitiform sensilla.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 6. Female genital structures of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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gen. et sp. n. A female genital structures B gonocoxites and gonostyli C, D spermatheca, lateral view (C) and basal view (D). Scale bar provided for
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="48">
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<paragraph pageId="3" pageNumber="48">
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Figure 7. Female genital structures of telephanine species. A
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<taxonomicName class="Insecta" family="Silvanidae" genus="Borneophanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Borneophanus spinosus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="spinosus">Borneophanus spinosus</taxonomicName>
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gen. et sp. n. B
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus dentatus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="dentatus">Psammoecus dentatus</taxonomicName>
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C
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<taxonomicName class="Insecta" family="Silvanidae" genus="Cryptamorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptamorpha desjardinsi" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="desjardinsi">Cryptamorpha desjardinsi</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="distribution">
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<paragraph pageId="3" pageNumber="48">Distribution.</paragraph>
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<paragraph pageId="3" pageNumber="48">Malaysia (Sabah and Sarawak states).</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="48" type="etymology">
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<paragraph pageId="3" pageNumber="48">Etymology.</paragraph>
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<paragraph pageId="3" pageNumber="48">The new genus name is composed of two words, the locality, Borneo where this new genus was collected, and the Greek phanos meaning bright.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48" type="remarks">
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<paragraph pageId="3" pageNumber="48">Remarks.</paragraph>
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<paragraph pageId="3" pageNumber="48">
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We found sensilla, called digitiform sensilla, each located in a groove on the dorsum of the apical maxillary palpomere.
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<bibRefCitation author="Lawrence, JF" journalOrPublisher="CSIRO, Collingwood" pageId="11" pageNumber="56" title="Australian beetles: Morphology, classification and keys." year="2013">
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Lawrence and
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<normalizedToken originalValue="Ślipiński">Ślipinski</normalizedToken>
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(2013)
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</bibRefCitation>
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stated that such sensilla on apical maxillary palpomere may be present in adephagan and polyphagan beetles, but they have never been described in
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<taxonomicName family="Silvanidae" lsidName="" pageId="3" pageNumber="48" rank="family">Silvanidae</taxonomicName>
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. We confirmed these in genera possessing securiform apical maxillary palpomeres (
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<taxonomicName lsidName="" pageId="3" pageNumber="48" rank="tribe" tribe="Telephanini">Telephanini</taxonomicName>
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:
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus dentatus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="dentatus">Psammoecus dentatus</taxonomicName>
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Grouvelle, 1883;
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<taxonomicName class="Insecta" family="Silvanidae" genus="Telephanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telephanus paradoxus" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="paradoxus">Telephanus paradoxus</taxonomicName>
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Reitter, 1874;
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<taxonomicName class="Insecta" family="Silvanidae" genus="Cryptamorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptamorpha desjardinsi" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="desjardinsi">Cryptamorpha desjardinsi</taxonomicName>
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(
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<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
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, 1844),
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<taxonomicName lsidName="" pageId="3" pageNumber="48" rank="tribe" tribe="Brontini">Brontini</taxonomicName>
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:
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<taxonomicName class="Insecta" family="Silvanidae" genus="Australohyliota" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Australohyliota mcleayi" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="mcleayi">Australohyliota mcleayi</taxonomicName>
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(Olliff, 1885),
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<taxonomicName family="Cucujidae" lsidName="" pageId="3" pageNumber="48" rank="family">Cucujidae</taxonomicName>
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:
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<taxonomicName class="Insecta" family="Cucujidae" genus="Cucujus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucujus bicolor" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">Cucujus bicolor</taxonomicName>
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Smith, 1851), although, these sensilla vary in the position and size of the area among the genera (Fig. 5).
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</paragraph>
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<paragraph lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
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The spermathecal duct of this new genus is extremely long, likely correlated to the partly coiled flagellum of the male genitalia. However, among
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<taxonomicName lsidName="" pageId="3" pageNumber="48" rank="tribe" tribe="Telephanini">Telephanini</taxonomicName>
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, female genital structures have not been studied. For comparison, we examined the female genitalia of
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<taxonomicName lsidName="P. dentatus" pageId="3" pageNumber="48" rank="species" species="dentatus">P. dentatus</taxonomicName>
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(Fig. 7B) and
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<taxonomicName lsidName="C. desjardinsi" pageId="3" pageNumber="48" rank="species" species="desjardinsi">C. desjardinsi</taxonomicName>
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(Fig. 7C) and found that their sper
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<pageBreakToken pageId="4" pageNumber="49" start="start">mathecal</pageBreakToken>
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ducts are apparently shorter than in the new genus. Additionally, we found further differences in the female genitalia (e.g., gonostyli; spermatheca), thus, future comprehensive study of telephanine female genitalia seems likely to provide some useful taxonomic characters and/or their phylogenetic insights.
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</paragraph>
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<paragraph pageId="4" pageNumber="49">
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The biology of the new genus is unknown except for the information that one paratype of the type series was collected by beating foliage. This new genus possesses lobed tarsomeres which are shared by most telephanines and seem to be related to the ecology living on dead leaves (
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<bibRefCitation author="Thomas, MC" journalOrPublisher="Coleopterists Bulletin" pageId="11" pageNumber="56" pagination="43 - 55" title="A new species of apterous Telephanus (Coleoptera: Silvanidae) with a discussion of phylogenetic relationships of the Silvanidae." volume="38" year="1984">Thomas 1984</bibRefCitation>
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), thus, they may have ordinary habits of telephanine beetles.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |