446 lines
38 KiB
XML
446 lines
38 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5643921" ID-GBIF-Dataset="7399d769-249a-47c9-b099-719ecd66851c" ID-GBIF-Taxon="189980188" ID-Zenodo-Dep="5643921" LSID="urn:lsid:zoobank.org:act:7827D48E-E121-4904-9636-3A46AE42B369" approvedBy="jonas" checkinTime="1635963396380" checkinUser="jonas" docAuthor="Wagner, Philipp, Ihlow, Flora, Hartmann, Timo, Flecks, Morris, Schmitz, Andreas & Böhme, Wolfgang" docDate="2021" docId="023E3900242FFFF163B1F9EC2CC9C9ED" docLanguage="en" docName="BonnzoolBull.70.1.141-171.pdf" docOrigin="Bonn zoological Bulletin 70 (1)" docTitle="Calotes goetzi P. & F. & T. & M. & A. & W. 2021, sp. n." docType="treatment" docUuid="7827D48E-E121-4904-9636-3A46AE42B369" docUuidSource="ZooBank" docVersion="8" lastPageId="25" lastPageNumber="166" masterDocId="FE0741782424FFE86326FFAC2F20CF1D" masterDocTitle="Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae)" masterLastPageNumber="171" masterPageNumber="141" pageId="11" pageNumber="152" updateTime="1636080518133" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wagner, Philipp</mods:namePart>
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<mods:nameIdentifier type="LSID">0575CE45-AB5C-4493-A8AD-DD399E18284A</mods:nameIdentifier>
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<mods:affiliation>Allwetterzoo Münster, Sentruper Strasse 315, D- 48161 Münster, Germany & Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA</mods:affiliation>
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<mods:nameIdentifier type="email">wagner@allwetterzoo.de</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ihlow, Flora</mods:namePart>
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<mods:nameIdentifier type="LSID">10DB565B-DA19-484E-AE50-ED29945658E8</mods:nameIdentifier>
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<mods:affiliation>Museum of Zoology Senckenberg, Dresden, Königsbrücker Landstrasse 159, D- 01109 Dresden, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hartmann, Timo</mods:namePart>
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<mods:nameIdentifier type="LSID">A911B13A-B364-424E-8579-AE23820F1DC1</mods:nameIdentifier>
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<mods:affiliation>ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Flecks, Morris</mods:namePart>
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<mods:nameIdentifier type="LSID">974BF839-926B-4039-9617-135F5CEC2596</mods:nameIdentifier>
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<mods:affiliation>ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schmitz, Andreas</mods:namePart>
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<mods:nameIdentifier type="LSID">5B1146D2-B9A6-48E6-9E4E-EB1C8ED1AE7A</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Geneva, UREC-Herpetology & Ichthyology, Route de Malagnou 1, CH- 1208 Genève, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Böhme, Wolfgang</mods:namePart>
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<mods:nameIdentifier type="LSID">FFAC2972-9F52-404B-BA9C-489C7793FF8D</mods:nameIdentifier>
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<mods:affiliation>ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Bonn zoological Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-05-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>70</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>141</mods:start>
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<mods:end>171</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">https://doi.org/10.20363/BZB-2021.70.1.141</mods:identifier>
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<mods:identifier type="GBIF-Dataset">7399d769-249a-47c9-b099-719ecd66851c</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5643781</mods:identifier>
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<mods:identifier type="ZooBank">4E7B4E14-7682-465D-A2D9-A26EA04AB3FD</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5643921" ID-GBIF-Taxon="189980188" ID-Zenodo-Dep="5643921" LSID="urn:lsid:zoobank.org:act:7827D48E-E121-4904-9636-3A46AE42B369" httpUri="http://treatment.plazi.org/id/023E3900242FFFF163B1F9EC2CC9C9ED" lastPageId="25" lastPageNumber="166" pageId="11" pageNumber="152">
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<subSubSection box="[151,367,1600,1625]" pageId="11" pageNumber="152" type="nomenclature">
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<paragraph blockId="11.[151,722,1600,1685]" box="[151,367,1600,1625]" pageId="11" pageNumber="152">
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<taxonomicName authority="P. & F. & T. & M. & A. & W., 2021" authorityName="P. & F. & T. & M. & A. & W." authorityYear="2021" box="[151,303,1600,1625]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="152" phylum="Chordata" rank="species" species="goetzi" status="sp. n.">
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<emphasis bold="true" box="[151,303,1600,1625]" italics="true" pageId="11" pageNumber="152">Calotes goetzi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[310,367,1601,1624]" pageId="11" pageNumber="152" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="152" type="description">
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<paragraph blockId="11.[151,722,1600,1685]" pageId="11" pageNumber="152">
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(
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<figureCitation box="[159,224,1631,1656]" captionStart="Fig" captionStartId="7.[151,186,1855,1877]" captionTargetBox="[162,1361,258,1801]" captionTargetId="figure-15@7.[160,1390,227,1834]" captionTargetPageId="7" captionText="Fig. 2. The Bayesian consensus tree based on 988 bp of mitochondrial DNA (12S rRNA and COI) shows six distinct lineages within Calotes mystaceus. Node support in terms of Bayesian posterior probabilities is indicated by circles at nodes (nodes with a BPP ≥ 0.90 are white, BPP ≥ 0.95 are grey, BPP ≥ 0.99 are black, values <0.90 are not marked). Outgroup (Calotes versicolor) not shown for clarity. Numbers in parentheses behind taxa refer to localities mapped in Fig. 1." figureDoi="http://doi.org/10.5281/zenodo.5643789" httpUri="https://zenodo.org/record/5643789/files/figure.png" pageId="11" pageNumber="152">Fig. 2</figureCitation>
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, Clade A)
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<emphasis box="[151,722,1666,1685]" italics="true" pageId="11" pageNumber="152">
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<uri box="[151,722,1666,1685]" pageId="11" pageNumber="152">
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urn:lsid:zoobank.org:act:
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<uuid box="[348,722,1666,1685]" pageId="11" pageNumber="152">7827D48E-E121-4904-9636-3A46AE42B369</uuid>
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</uri>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="152" type="materials_examined">
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<paragraph blockId="11.[150,756,1724,1965]" pageId="11" pageNumber="152">
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<materialsCitation ID-GBIF-Occurrence="3397335304" collectionCode="ZFMK" country="Cambodia" latitude="13.699167" location="Kbal Spean within the Phnom Kulen National Park" longLatPrecision="1" longitude="103.99861" municipality="Kbal Spean within the Phnom Kulen National Park" pageId="11" pageNumber="152" specimenCode="ZFMK 92606" specimenCount="1" specimenCount-adult="1" stateProvince="Siem Reap Province" typeStatus="holotype">
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<emphasis bold="true" box="[174,285,1724,1748]" pageId="11" pageNumber="152">
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<typeStatus box="[174,280,1724,1748]" pageId="11" pageNumber="152">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode box="[290,437,1724,1748]" pageId="11" pageNumber="152">
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<collectionCode box="[290,364,1725,1748]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" pageId="11" pageNumber="152">ZFMK</collectionCode>
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92606
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</specimenCode>
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(
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<specimenCount box="[449,564,1724,1748]" pageId="11" pageNumber="152" type="male">adult male</specimenCount>
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,
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<figureCitation box="[574,635,1724,1748]" captionStart="Fig" captionStartId="14.[189,224,1922,1944]" captionTargetBox="[218,1407,227,1908]" captionTargetId="figure-18@14.[218,1407,227,1908]" captionTargetPageId="14" captionText="Fig. 7. Calotes goetzi sp. n. A. Holotype (ZFMK 92606, adult male), general view from above. B. Holotype, lateral view.C. Living holotype from Kbal Spean, Cambodia." figureDoi="http://doi.org/10.5281/zenodo.5643803" httpUri="https://zenodo.org/record/5643803/files/figure.png" pageId="11" pageNumber="152">Fig. 7</figureCitation>
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A-C, Clade
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<collectionCode box="[151,172,1755,1779]" pageId="11" pageNumber="152">A</collectionCode>
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) from
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<collectingCountry box="[240,352,1755,1779]" name="Cambodia" pageId="11" pageNumber="152" value="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion box="[360,577,1755,1779]" country="Cambodia" name="Siem Reap" pageId="11" pageNumber="152">Siem Reap Province</collectingRegion>
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, near
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<collectingMunicipality country="Cambodia" latitude="13.699167" longLatPrecision="1" longitude="103.99861" name="Kbal Spean within the Phnom Kulen National Park" pageId="11" pageNumber="152" stateProvince="Siem Reap Province">Kbal Spean within the Phnom Kulen National Park</collectingMunicipality>
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[
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<geoCoordinate box="[594,749,1786,1810]" degrees="13.699167" direction="north" orientation="latitude" pageId="11" pageNumber="152" precision="1" value="13.699167">13.699167° N</geoCoordinate>
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,
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<geoCoordinate box="[151,310,1817,1841]" degrees="103.998611" direction="east" orientation="longitude" pageId="11" pageNumber="152" precision="1" value="103.99861">103.998611° E</geoCoordinate>
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]
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="11.[150,756,1724,1965]" pageId="11" pageNumber="152">
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<materialsCitation ID-GBIF-Occurrence="3397335313" collectionCode="ZFMK" country="Cambodia" latitude="13.699167" location="Kbal Spean within the Phnom Kulen National Park" longLatPrecision="1" longitude="103.99861" municipality="Kbal Spean within the Phnom Kulen National Park" pageId="11" pageNumber="152" specimenCode="ZFMK 88341, ZFMK 92607" specimenCount="2" specimenCount-adult="2" stateProvince="Siem Reap Province" typeStatus="paratype">
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<emphasis bold="true" box="[174,295,1848,1872]" pageId="11" pageNumber="152">
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<typeStatus box="[174,290,1848,1872]" pageId="11" pageNumber="152">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCode box="[303,452,1848,1872]" pageId="11" pageNumber="152">
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<collectionCode box="[303,377,1849,1872]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" pageId="11" pageNumber="152">ZFMK</collectionCode>
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88341
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</specimenCode>
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(
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<specimenCount box="[468,587,1848,1872]" pageId="11" pageNumber="152" type="male">adult male</specimenCount>
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),
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<specimenCode box="[607,756,1848,1872]" pageId="11" pageNumber="152">
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<collectionCode box="[607,681,1849,1872]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" pageId="11" pageNumber="152">ZFMK</collectionCode>
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92607
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</specimenCode>
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(
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<specimenCount box="[158,294,1879,1903]" pageId="11" pageNumber="152" type="female">adult female</specimenCount>
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) from the same locality as the
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<typeStatus box="[622,719,1879,1903]" pageId="11" pageNumber="152">holotype</typeStatus>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="152" type="diagnosis">
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<paragraph blockId="11.[150,756,1724,1965]" lastBlockId="11.[793,1401,1106,1965]" pageId="11" pageNumber="152">
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<emphasis bold="true" box="[174,291,1910,1934]" pageId="11" pageNumber="152">Diagnosis.</emphasis>
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Alarge species of
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<taxonomicName authorityName="Cuvier" authorityYear="1817" box="[483,564,1910,1934]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="152" phylum="Chordata" rank="genus">
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<emphasis box="[483,564,1910,1934]" italics="true" pageId="11" pageNumber="152">Calotes</emphasis>
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</taxonomicName>
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, with a maximum SVL of
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<quantity box="[236,324,1941,1965]" metricMagnitude="-1" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="152" unit="mm" value="143.0">143 mm</quantity>
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in males and
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<quantity box="[474,562,1941,1965]" metricMagnitude="-1" metricUnit="m" metricValue="1.22" pageId="11" pageNumber="152" unit="mm" value="122.0">122 mm</quantity>
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in females. It can be distinguished from other species of the group by the combination of the following characters: 1) head, body and limbs robust, tail long but not as long as in
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<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1837" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="152" phylum="Chordata" rank="species" species="mystaceus">
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<emphasis italics="true" pageId="11" pageNumber="152">C. mystaceus</emphasis>
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</taxonomicName>
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; 2) body scales mid-sized, homogeneous, keeled, arranged in regular rows; 3) upper dorsolateral scales pointing up- and backwards; 4) body scales arranged in 45–60 rows around midbody; 5) two short and separated spines, surrounded by a ring of scales between the tympanum and the vertebral crest on both sides of the head; 6) vertebral crest continuous from above the tympanum to the hind limbs, composed of erected scales, directed posteriorly, highest slightly in front of the insertion of the front limbs, becoming gradually shorter towards the hind limbs; 7) vertebral scales, including vertebral spines,
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<quantity metricMagnitude="0" metricUnit="m" metricValue="1.1303" metricValueMax="1.3208" metricValueMin="0.9398" pageId="11" pageNumber="152" unit="in" value="44.5" valueMax="52.0" valueMin="37.0">37– 52 in</quantity>
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males and
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<quantity box="[967,1061,1539,1563]" metricMagnitude="0" metricUnit="m" metricValue="1.3081" metricValueMax="1.524" metricValueMin="1.0922" pageId="11" pageNumber="152" unit="in" value="51.5" valueMax="60.0" valueMin="43.0">43–60 in</quantity>
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females; 8) oblique skin fold in front of the fore limbs; 9) head, body, and limbs bluish in males; 19) males with a white stripe from between nostril and orbit along the upper lip and the tympanum to the front limb insertion; 11) three to five distinct dark brown dorsolateral blotches.
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="16" lastPageNumber="157" pageId="11" pageNumber="152" type="description">
|
||
<paragraph blockId="11.[793,1401,1106,1965]" lastBlockId="16.[187,797,239,1965]" lastPageId="16" lastPageNumber="157" pageId="11" pageNumber="152">
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<emphasis bold="true" box="[816,1169,1724,1748]" pageId="11" pageNumber="152">
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||
Description of the
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<typeStatus box="[1064,1164,1724,1748]" pageId="11" pageNumber="152">holotype</typeStatus>
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.
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</emphasis>
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Large male (SVL
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<quantity box="[794,891,1755,1779]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="11" pageNumber="152" unit="mm" value="118.0">118 mm</quantity>
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). Body robust, tail relatively short,
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<quantity box="[1304,1398,1755,1779]" metricMagnitude="-1" metricUnit="m" metricValue="2.36" pageId="11" pageNumber="152" unit="mm" value="236.0">236 mm</quantity>
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||
long. Fore- and hind limbs relatively slender, fourth finger and toe longest. Head distinct from the neck, posterior jaw angles heavily swollen. Tip of the snout blunt, rostral small. Nostril large, in a single scale, separated from the rostral by one elongated scale and from the first two supralabial scales by two rectangular scales. Canthus rostralis distinct, formed by nine scales from the nostril to above the orbit and continuous with supraciliary scales. Lateral sides of the head flat. Supralabial scales 9/9, infralabial scales 10/10. Supralabial scales separated from the orbit by four rows of small scales. Eight scales between the orbit and the tympanum, two of them slightly ridged. Tympanum distinct, two spines, surrounded by a ring of scales, between the tympanum and the vertebral crest. Mental scale small, bordered by two postmental scales which are separated from each other. First pair of the postmental scales in contact with the infralabial scale. Scales on the dorsal part of the chin smooth to feebly keeled, becoming strongly keeled towards the throat. Vertebral crest continuous from above the tympanum to the tail, spines highest above the insertion of the front limbs and gradually decreasing towards the tail. Dorsal and lateral scales strongly keeled, pointing up- and backwards. Ventral scales parallelly keeled. Caudal scales smooth to feebly keeled, directed backwards. Subcaudal scales parallel and strongly keeled.
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</paragraph>
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||
<paragraph blockId="16.[187,797,239,1965]" pageId="16" pageNumber="157">
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<emphasis bold="true" box="[212,397,858,882]" pageId="16" pageNumber="157">Male coloration.</emphasis>
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Head, body and limbs bluish. Awhite stripe, as high as the tympanum, is present from between nostril and orbit along the upper lip and the tympanum to the insertion of the front limb. The stripe is followed by three to five large reddish-brown to dark-brown blotches, with the first above the insertion of the fore limbs and the latest at around midbody or above the insertion of the hindlimbs, sometimes continuing on the tail. Throat coloration darker than the chin and the head coloration. From Thailand, males are known to have bright blue hindlimbs and crest scales. Intermediate males with a blue head and a white stripe, but body coloration brownish-black with indistinct light brown blotches. Non-brilliant coloration in males from Thailand reddish-brown, with a white non-continuous stripe from the orbit to above the hindlimbs. Blotches brownish, darker than the body coloration.
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</paragraph>
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<paragraph blockId="16.[187,797,239,1965]" lastBlockId="16.[831,1438,239,1439]" pageId="16" pageNumber="157">
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<emphasis bold="true" box="[212,326,1384,1408]" pageId="16" pageNumber="157">Variation.</emphasis>
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Bodymeasurements and meristic characters for adult individuals are given in
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<tableCitation box="[566,662,1415,1439]" captionStart="Table 4" captionStartId="11.[151,207,757,778]" captionTargetBox="[181,1368,845,1030]" captionTargetPageId="11" captionText="Table 4. Eigenvalues and percent of explained variance per principal component of mensural and meristic data of males and females as shown in Figs 3–4." httpUri="http://table.plazi.org/id/DEE8D89E242FFFE363B1FD592E87CC38" pageId="16" pageNumber="157" tableUuid="DEE8D89E242FFFE363B1FD592E87CC38">Tables 4</tableCitation>
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. Specimens ranged in size between the smallest adult with a SVL of
|
||
<quantity box="[190,267,1477,1501]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="16" pageNumber="157" unit="mm" value="84.0">84 mm</quantity>
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(MNHN 1884.548) and the largest female with a SVL of
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||
<quantity box="[304,396,1508,1532]" metricMagnitude="-1" metricUnit="m" metricValue="1.26" pageId="16" pageNumber="157" unit="mm" value="126.0">126 mm</quantity>
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(ZFMK 44893) to the largest male measuring
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<quantity box="[307,394,1539,1563]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="16" pageNumber="157" unit="mm" value="145.0">145 mm</quantity>
|
||
(MNHN 1884.546). In general, adult males are larger than females, because of longer SVLs, tail lengths, head lengths, and widths and slightly fewer vertebral scales including the crest scales. Heads of adult males and females are proportionally equal in length, but male heads are wider than those of females, probably because of the swollen jaw angle. Dorsal coloration differs between sexes and both sexes are able to change the coloration. Coloration characters being lighter in females; head and body blue. Lateral stripe from between the nostril and orbit along the upper lips and the tympanum to the first blotch. Sometimes the first two brownish blotches are framed by scales in the same coloration as the lateral stripe. Three to five blotches present, from above the insertion of the fore limbs to about mid-body or even hindlimb insertion. Non-display coloration of females with lateral parts of the body gray and dorsal parts brown, with three to four darker crossbands between the lateral stripes on both sides of the body. Lateral stripe white, from the mental scale along the upper lips and the tympanum to the hindlimbs. Head light blue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="157" type="etymology">
|
||
<paragraph blockId="16.[831,1438,239,1439]" pageId="16" pageNumber="157">
|
||
<emphasis bold="true" box="[854,983,456,480]" pageId="16" pageNumber="157">Etymology.</emphasis>
|
||
The specific epithet is a patronym formed in the genitive singular honoring Dr. Stephan Goetz, Munich, Germany, in recognition of his longtime support of species conservation efforts in Cambodia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="157" type="distribution">
|
||
<paragraph blockId="16.[831,1438,239,1439]" pageId="16" pageNumber="157">
|
||
<emphasis bold="true" box="[854,1001,580,604]" pageId="16" pageNumber="157">Distribution.</emphasis>
|
||
<taxonomicName authority="P. & F. & T. & M. & A. & W., 2021" authorityName="P. & F. & T. & M. & A. & W." authorityYear="2021" box="[1010,1165,580,604]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="goetzi" status="sp. n.">
|
||
<emphasis box="[1010,1165,580,604]" italics="true" pageId="16" pageNumber="157">Calotes goetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1174,1234,580,603]" pageId="16" pageNumber="157" rank="species">sp. n.</taxonomicNameLabel>
|
||
is distributed in a large area of Indochina and inhabits most of the previous range of
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1837" box="[931,1074,642,665]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="mystaceus">
|
||
<emphasis box="[931,1074,642,665]" italics="true" pageId="16" pageNumber="157">C. mystaceus</emphasis>
|
||
</taxonomicName>
|
||
. The new species is documented by museum specimens and photo vouchers from Cambodia, China (Yunnan province), Laos, Myanmar, and Thailand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="157" type="biology_ecology">
|
||
<paragraph blockId="16.[831,1438,239,1439]" pageId="16" pageNumber="157">
|
||
<emphasis bold="true" box="[854,950,766,790]" pageId="16" pageNumber="157">Ecology.</emphasis>
|
||
<taxonomicName authority="P. & F. & T. & M. & A. & W., 2021" authorityName="P. & F. & T. & M. & A. & W." authorityYear="2021" box="[959,1113,765,789]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="goetzi" status="sp. n.">
|
||
<emphasis box="[959,1113,765,789]" italics="true" pageId="16" pageNumber="157">Calotes goetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1124,1183,766,789]" pageId="16" pageNumber="157" rank="species">sp. n.</taxonomicNameLabel>
|
||
is diurnal and semi-arboreal to arboreal and can easily climb on tree trunks at a height of 5–10 meters or above. The species is known from dipterocarp lowland forests, cultural landscape and secondary forest. At its
|
||
<typeStatus box="[1087,1133,890,914]" pageId="16" pageNumber="157">type</typeStatus>
|
||
locality the species was observed in more open habitats with a closed canopy, but also within dense monsoon tropical forests and solitary trees in rather open areas. Here it occurs sympatrically with
|
||
<taxonomicName baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[921,1122,1013,1037]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="versicolor">
|
||
<emphasis box="[921,1122,1013,1037]" italics="true" pageId="16" pageNumber="157">Calotes versicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Hawkeswood TJ & Sommung A" journalOrPublisher="Calodema" pageId="16" pageNumber="157" pagination="1 - 8" part="607" refId="ref16767" refString="Hawkeswood TJ, Sommung A (2018) First record of the Blue Forest Lizard, Calotes mystaceus Dumeril & Bibron, 1837 (Reptilia: Agamidae) from Ubon Ratchathani Province, Thailand, with a review of literature on the biology and distribution of the species in Thailand. Calodema 607: 1 - 8" title="First record of the Blue Forest Lizard, Calotes mystaceus Dumeril & Bibron, 1837 (Reptilia: Agamidae) from Ubon Ratchathani Province, Thailand, with a review of literature on the biology and distribution of the species in Thailand" type="journal article" year="2018">Hawkeswood & Sommung (2018)</bibRefCitation>
|
||
report it from farmland with e.g., rice, coconut, banana, and durian plantations.
|
||
<bibRefCitation author="Chan-Ard T & Parr JWR & Nabhitabhata J" box="[1184,1437,1075,1099]" journalOrPublisher="Oxford University Press, New York" pageId="16" pageNumber="157" refId="ref16112" refString="Chan-Ard T, Parr JWR, Nabhitabhata J (2015) A Field Guide to the Reptiles of Thailand. Oxford University Press, New York" title="A Field Guide to the Reptiles of Thailand" type="book" year="2015">Chan-Ard et al. (2015)</bibRefCitation>
|
||
report the species from a wide range of forest
|
||
<typeStatus box="[1350,1406,1106,1130]" pageId="16" pageNumber="157">types</typeStatus>
|
||
in Thailand, including tree-lined agricultural lands and grasslands. Similar to other
|
||
<taxonomicName box="[1149,1322,1167,1191]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis box="[1149,1230,1167,1191]" italics="true" pageId="16" pageNumber="157">Calotes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="P. & F. & T. & M. & A. & W., 2021" authorityName="P. & F. & T. & M. & A. & W." authorityYear="2021" box="[1337,1437,1168,1191]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="goetzi" status="sp. n.">
|
||
<emphasis box="[1337,1437,1168,1191]" italics="true" pageId="16" pageNumber="157">C. goetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[831,891,1199,1222]" pageId="16" pageNumber="157" rank="species">sp. n.</taxonomicNameLabel>
|
||
feeds on a variety of arthropods.
|
||
<bibRefCitation author="Chan-Ard T & Parr JWR & Nabhitabhata J" journalOrPublisher="Oxford University Press, New York" pageId="16" pageNumber="157" refId="ref16112" refString="Chan-Ard T, Parr JWR, Nabhitabhata J (2015) A Field Guide to the Reptiles of Thailand. Oxford University Press, New York" title="A Field Guide to the Reptiles of Thailand" type="book" year="2015">Chan-Ard et al. (2015)</bibRefCitation>
|
||
report from Thailand, that mature individuals are territorial, chasing away possible intruders. Eggs are buried in soft soil and later guarded by the males.
|
||
<bibRefCitation author="Amber ED & Waengsothorn S & Strine CT" journalOrPublisher="Herpetological Review" pageId="16" pageNumber="157" pagination="640" part="48" refId="ref15924" refString="Amber ED, Waengsothorn S, Strine CT (2017) Calotes mystaceus (Moustached Crested Lizard). Defensive behaviors. Herpetological Review 48: 640" title="Calotes mystaceus (Moustached Crested Lizard). Defensive behaviors" type="journal article" year="2017">Amber et al. (2017)</bibRefCitation>
|
||
recognized a possible ontogenetic shift in defense strategies of
|
||
<taxonomicName authority="P. & F. & T. & M. & A. & W., 2021" authorityName="P. & F. & T. & M. & A. & W." authorityYear="2021" box="[1056,1151,1354,1377]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="goetzi" status="sp. n.">
|
||
<emphasis box="[1056,1151,1354,1377]" italics="true" pageId="16" pageNumber="157">C. goetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1158,1215,1354,1377]" pageId="16" pageNumber="157" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<taxonomicName authorityName="Dumeril & Bibron" authorityYear="1837" box="[1230,1371,1354,1377]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="157" phylum="Chordata" rank="species" species="mystaceus">
|
||
<emphasis box="[1230,1371,1354,1377]" italics="true" pageId="16" pageNumber="157">C. mystaceus</emphasis>
|
||
</taxonomicName>
|
||
at the time of their publication), with older individuals utilizing color change, while juveniles do not.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="166" type="materials_examined">
|
||
<paragraph blockId="25.[150,761,1451,1971]" lastBlockId="25.[790,1399,1297,1971]" pageId="25" pageNumber="166">
|
||
<emphasis bold="true" box="[328,557,1730,1754]" pageId="25" pageNumber="166">
|
||
<taxonomicName authority="P. & F. & T. & M. & A. & W., 2021" authorityName="P. & F. & T. & M. & A. & W." authorityYear="2021" box="[328,484,1730,1754]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="166" phylum="Chordata" rank="species" species="goetzi" status="sp. n.">
|
||
<emphasis bold="true" box="[328,484,1730,1754]" italics="true" pageId="25" pageNumber="166">Calotes goetzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[494,557,1730,1754]" pageId="25" pageNumber="166" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis box="[577,590,1731,1754]" italics="true" pageId="25" pageNumber="166">n</emphasis>
|
||
=
|
||
<specimenCount box="[626,656,1730,1754]" pageId="25" pageNumber="166">71</specimenCount>
|
||
):
|
||
<materialsCitation ID-GBIF-Occurrence="3398324327" collectionCode="BMNH" pageId="25" pageNumber="166" specimenCode="BMNH 1868.4.3.61, BMNH 1868.4.3.63, BMNH 1891.11.26.29, BMNH 1914.4.25.1, BMNH 1929.12.1.11, BMNH 1933.3.10.1, BMNH 1933.3.10.2" specimenCount="1">
|
||
<specimenCode pageId="25" pageNumber="166">BMNH 1868.4.3.61</specimenCode>
|
||
,
|
||
<specimenCode box="[290,505,1761,1785]" pageId="25" pageNumber="166">BMNH 1868.4.3.63</specimenCode>
|
||
,
|
||
<specimenCode box="[514,754,1761,1785]" pageId="25" pageNumber="166">BMNH 1891.11.26.29</specimenCode>
|
||
,
|
||
<specimenCode box="[150,381,1792,1816]" pageId="25" pageNumber="166">BMNH 1914.4.25.1</specimenCode>
|
||
,
|
||
<specimenCode box="[408,652,1792,1816]" pageId="25" pageNumber="166">BMNH 1929.12.1.11</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">BMNH 1933.3.10.1</specimenCode>
|
||
,
|
||
<specimenCode box="[303,531,1823,1847]" pageId="25" pageNumber="166">BMNH 1933.3.10.2</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3398324315" collectionCode="MNHN" pageId="25" pageNumber="166" specimenCode="MNHN 1884.546, MNHN 1884.547, MNHN 1884.548, MNHN 1893.335, MNHN 1893.336" specimenCount="1">
|
||
<specimenCode box="[552,755,1823,1847]" pageId="25" pageNumber="166">MNHN 1884.546</specimenCode>
|
||
,
|
||
<specimenCode box="[151,344,1854,1878]" pageId="25" pageNumber="166">MNHN 1884.547</specimenCode>
|
||
,
|
||
<specimenCode box="[357,550,1854,1878]" pageId="25" pageNumber="166">MNHN 1884.548</specimenCode>
|
||
,
|
||
<specimenCode box="[562,755,1854,1878]" pageId="25" pageNumber="166">MNHN 1893.335</specimenCode>
|
||
,
|
||
<specimenCode box="[153,353,1885,1909]" pageId="25" pageNumber="166">MNHN 1893.336</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3398324325" collectionCode="NME" pageId="25" pageNumber="166" specimenCode="NME R0581/09, NME R0584/09, NME R0585/09, NME R0686/11, NME R0751/12, NME R0783/13, NME R0784/13, NME R0785/13, NME R0786/13, NME R0787/13, NME R0790/14" specimenCount="1">
|
||
<specimenCode box="[372,555,1885,1909]" pageId="25" pageNumber="166">NME R0581/09</specimenCode>
|
||
,
|
||
<specimenCode box="[574,756,1885,1909]" pageId="25" pageNumber="166">NME R0584/09</specimenCode>
|
||
,
|
||
<specimenCode box="[151,338,1916,1940]" pageId="25" pageNumber="166">NME R0585/09</specimenCode>
|
||
,
|
||
<specimenCode box="[360,546,1916,1940]" pageId="25" pageNumber="166">NME R0686/11</specimenCode>
|
||
,
|
||
<specimenCode box="[569,754,1916,1940]" pageId="25" pageNumber="166">NME R0751/12</specimenCode>
|
||
,
|
||
<specimenCode box="[151,324,1947,1971]" pageId="25" pageNumber="166">NME R0783/13</specimenCode>
|
||
,
|
||
<specimenCode box="[334,507,1947,1971]" pageId="25" pageNumber="166">NME R0784/13</specimenCode>
|
||
,
|
||
<specimenCode box="[517,690,1946,1970]" pageId="25" pageNumber="166">NME R0785/13</specimenCode>
|
||
,
|
||
<specimenCode box="[700,898,1297,1970]" pageId="25" pageNumber="166">NME R0786/13</specimenCode>
|
||
,
|
||
<specimenCode box="[918,1101,1297,1321]" pageId="25" pageNumber="166">NME R0787/13</specimenCode>
|
||
,
|
||
<specimenCode box="[1121,1304,1297,1321]" pageId="25" pageNumber="166">NME R0790/14</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3398324303" collectionCode="ZFMK" pageId="25" pageNumber="166" specimenCode="ZFMK 16640, ZFMK 16641, ZFMK 40561, ZFMK 43906, ZFMK 43930, ZFMK 43931, ZFMK 43932, ZFMK 44893, ZFMK 44894, ZFMK 45490, ZFMK 45491, ZFMK 45492, ZFMK 45493, ZFMK 45494, ZFMK 45495, ZFMK 45496, ZFMK 45497, ZFMK 45498, ZFMK 45499, ZFMK 45500, ZFMK 45501, ZFMK 45502, ZFMK 45553, ZFMK 45554, ZFMK 47080, ZFMK 47081, ZFMK 49164, ZFMK 49202, ZFMK 49219, ZFMK 49220, ZFMK 49221, ZFMK 49242, ZFMK 49243, ZFMK 49244, ZFMK 49245, ZFMK 49246, ZFMK 55610, ZFMK 55611, ZFMK 55612, ZFMK 84867, ZFMK 88341, ZFMK 92606, ZFMK 92607" specimenCount="1">
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 16640</specimenCode>
|
||
,
|
||
<specimenCode box="[879,1038,1329,1353]" pageId="25" pageNumber="166">ZFMK 16641</specimenCode>
|
||
,
|
||
<specimenCode box="[1058,1216,1329,1353]" pageId="25" pageNumber="166">ZFMK 40561</specimenCode>
|
||
,
|
||
<specimenCode box="[1236,1394,1329,1353]" pageId="25" pageNumber="166">ZFMK 43906</specimenCode>
|
||
,
|
||
<specimenCode box="[791,949,1362,1386]" pageId="25" pageNumber="166">ZFMK 43930</specimenCode>
|
||
,
|
||
<specimenCode box="[969,1127,1362,1386]" pageId="25" pageNumber="166">ZFMK 43931</specimenCode>
|
||
,
|
||
<specimenCode box="[1146,1305,1362,1386]" pageId="25" pageNumber="166">ZFMK 43932</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 44893</specimenCode>
|
||
,
|
||
<specimenCode box="[879,1038,1395,1419]" pageId="25" pageNumber="166">ZFMK 44894</specimenCode>
|
||
,
|
||
<specimenCode box="[1058,1216,1394,1418]" pageId="25" pageNumber="166">ZFMK 45490</specimenCode>
|
||
,
|
||
<specimenCode box="[1236,1394,1394,1418]" pageId="25" pageNumber="166">ZFMK 45491</specimenCode>
|
||
,
|
||
<specimenCode box="[791,949,1427,1451]" pageId="25" pageNumber="166">ZFMK 45492</specimenCode>
|
||
,
|
||
<specimenCode box="[969,1127,1427,1451]" pageId="25" pageNumber="166">ZFMK 45493</specimenCode>
|
||
,
|
||
<specimenCode box="[1146,1305,1427,1451]" pageId="25" pageNumber="166">ZFMK 45494</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 45495</specimenCode>
|
||
,
|
||
<specimenCode box="[879,1038,1459,1483]" pageId="25" pageNumber="166">ZFMK 45496</specimenCode>
|
||
,
|
||
<specimenCode box="[1058,1216,1459,1483]" pageId="25" pageNumber="166">ZFMK 45497</specimenCode>
|
||
,
|
||
<specimenCode box="[1236,1394,1459,1483]" pageId="25" pageNumber="166">ZFMK 45498</specimenCode>
|
||
,
|
||
<specimenCode box="[791,949,1492,1516]" pageId="25" pageNumber="166">ZFMK 45499</specimenCode>
|
||
,
|
||
<specimenCode box="[969,1127,1492,1516]" pageId="25" pageNumber="166">ZFMK 45500</specimenCode>
|
||
,
|
||
<specimenCode box="[1146,1305,1492,1516]" pageId="25" pageNumber="166">ZFMK 45501</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 45502</specimenCode>
|
||
,
|
||
<specimenCode box="[879,1038,1524,1548]" pageId="25" pageNumber="166">ZFMK 45553</specimenCode>
|
||
,
|
||
<specimenCode box="[1058,1216,1524,1548]" pageId="25" pageNumber="166">ZFMK 45554</specimenCode>
|
||
,
|
||
<specimenCode box="[1236,1394,1525,1549]" pageId="25" pageNumber="166">ZFMK 47080</specimenCode>
|
||
,
|
||
<specimenCode box="[791,949,1557,1581]" pageId="25" pageNumber="166">ZFMK 47081</specimenCode>
|
||
,
|
||
<specimenCode box="[969,1127,1557,1581]" pageId="25" pageNumber="166">ZFMK 49164</specimenCode>
|
||
,
|
||
<specimenCode box="[1146,1305,1557,1581]" pageId="25" pageNumber="166">ZFMK 49202</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 49219</specimenCode>
|
||
,
|
||
<specimenCode box="[879,1038,1589,1613]" pageId="25" pageNumber="166">ZFMK 49220</specimenCode>
|
||
,
|
||
<specimenCode box="[1058,1216,1589,1613]" pageId="25" pageNumber="166">ZFMK 49221</specimenCode>
|
||
,
|
||
<specimenCode box="[1236,1394,1589,1613]" pageId="25" pageNumber="166">ZFMK 49242</specimenCode>
|
||
,
|
||
<specimenCode box="[791,949,1622,1646]" pageId="25" pageNumber="166">ZFMK 49243</specimenCode>
|
||
,
|
||
<specimenCode box="[969,1127,1622,1646]" pageId="25" pageNumber="166">ZFMK 49244</specimenCode>
|
||
,
|
||
<specimenCode box="[1146,1305,1622,1646]" pageId="25" pageNumber="166">ZFMK 49245</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 49246</specimenCode>
|
||
,
|
||
<specimenCode box="[880,1038,1654,1678]" pageId="25" pageNumber="166">ZFMK 55610</specimenCode>
|
||
,
|
||
<specimenCode box="[1058,1216,1654,1678]" pageId="25" pageNumber="166">ZFMK 55611</specimenCode>
|
||
,
|
||
<specimenCode box="[1235,1394,1654,1678]" pageId="25" pageNumber="166">ZFMK 55612</specimenCode>
|
||
,
|
||
<specimenCode box="[791,949,1687,1711]" pageId="25" pageNumber="166">ZFMK 84867</specimenCode>
|
||
,
|
||
<specimenCode box="[969,1127,1687,1711]" pageId="25" pageNumber="166">ZFMK 88341</specimenCode>
|
||
,
|
||
<specimenCode box="[1146,1305,1687,1711]" pageId="25" pageNumber="166">ZFMK 92606</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZFMK 92607</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3398324309" collectionCode="ZMB" pageId="25" pageNumber="166" specimenCode="ZMB 11603A, ZMB 30186, ZMB 30188A, ZMB 30197, ZMB 6034" specimenCount="1">
|
||
<specimenCode box="[871,1022,1719,1743]" pageId="25" pageNumber="166">ZMB 11603A</specimenCode>
|
||
,
|
||
<specimenCode box="[1033,1166,1719,1743]" pageId="25" pageNumber="166">ZMB 30186</specimenCode>
|
||
,
|
||
<specimenCode box="[1177,1330,1719,1743]" pageId="25" pageNumber="166">ZMB 30188A</specimenCode>
|
||
,
|
||
<specimenCode pageId="25" pageNumber="166">ZMB 30197</specimenCode>
|
||
,
|
||
<specimenCode box="[873,998,1752,1776]" pageId="25" pageNumber="166">ZMB 6034</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |