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<document id="91FDBF0683D10781742ED7B4DFD78F1E" ID-DOI="10.11646/zootaxa.4808.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3933160" ID-ZooBank="15876EA8-DC4E-456C-82CF-21E1DA2479F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594120931585" checkinUser="plazi" docAuthor="Vu, Ha T., Nguyen, Hung D., Le, Son X., Eguchi, Katsuyuki, Nguyen, Anh D. &amp; Tran, Binh T. T." docDate="2020" docId="BE3287F3FF83D212FF1CF95798A4B1E4" docLanguage="en" docName="zootaxa..4808.3.1.pdf" docOrigin="Zootaxa 4808 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otostigmus Porat 1876" docType="key" docVersion="5" lastPageNumber="435" masterDocId="420BFF8BFFA2D230FF8BFF829D38B016" masterDocTitle="A review and notes on the phylogenetic relationship of the centipede genus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae) from Vietnam" masterLastPageNumber="438" masterPageNumber="401" pageNumber="434" updateTime="1698777941745" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8F543A25044E8DEDDDD1C42D30E4FEAA">A review and notes on the phylogenetic relationship of the centipede genus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae) from Vietnam</mods:title>
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<mods:namePart id="2EC9992FA4414758E32144060E564615">Nguyen, Hung D.</mods:namePart>
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<mods:namePart id="86515889EAD1FEB4DF95B7512A4B57E3">Le, Son X.</mods:namePart>
<mods:affiliation id="361781FD05EEA56CBCEAE52636DFD9F2">Institute of Tropical Ecology, Vietnamese-Russian Tropical Center, 2 Nguyen Van Huyen Str., Cau Giay District, Hanoi, VIETNAM. lesonenv 86 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1648 - 6773</mods:affiliation>
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<mods:namePart id="7B84339F226DC1C6F0F748F5C6B2EC19">Eguchi, Katsuyuki</mods:namePart>
<mods:affiliation id="45359E2A5AF9F75C8724FC8CDF8E1031">Department of Biology, Graduate School of Science, Tokyo Metropolitan University, JAPAN. &amp; antist 2007 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1054 - 1295</mods:affiliation>
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<mods:namePart id="18B39EA8A4989C5C3AA2EC2695DF2BB9">Nguyen, Anh D.</mods:namePart>
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Key to
<taxonomicName id="F19B4D66FF83D211FF7AF9569C42B6F8" ID-CoL="63HN6" ID-ENA="390858" authorityName="Porat" authorityYear="1876" box="[241,378,1748,1774]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="33" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="04EFEAF7FF83D211FF7AF9569C42B6F8" bold="true" box="[241,378,1748,1774]" italics="true" pageId="33" pageNumber="434">Otostigmus</emphasis>
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species in
<collectingCountry id="4E8C7675FF83D211FE75F9569F50B6F8" box="[510,616,1748,1774]" name="Vietnam" pageId="33" pageNumber="434">Vietnam</collectingCountry>
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<paragraph id="362436E5FF83D211FF1CF89B98A4B738" blockId="33.[151,1436,1817,2044]" box="[151,1436,1817,1839]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF89B98A4B738" box="[151,1436,1817,1839]" pageId="33" pageNumber="434">1. Terga with longitudinal keels........................................................................... 2</keyLead>
</paragraph>
<paragraph id="362436E5FF83D211FF1CF8B598A4B75D" blockId="33.[151,1436,1817,2044]" box="[151,1436,1846,1868]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF8B598A4B75D" box="[151,1436,1846,1868]" pageId="33" pageNumber="434">- Terga without longitudinal keels......................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="8D6F2E40FF83D211FF1CF8D698A4B790" pageId="33" pageNumber="434">
<paragraph id="362436E5FF83D211FF1CF8D698A4B77F" blockId="33.[151,1436,1817,2044]" box="[151,1436,1876,1898]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF8D698A4B77F" box="[151,1436,1876,1898]" pageId="33" pageNumber="434">
2. Terga with 3 or fewer longitudinal keels, antennae with 17 segments, teeth 4+4............................
<taxonomicName id="F19B4D66FF83D211FAADF8D698A4B77F" ID-CoL="4B7V6" authorityName="Chamberlin" authorityYear="1913" box="[1318,1436,1876,1897]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="33" pageNumber="434" phylum="Arthropoda" rank="species" species="amballae">
<emphasis id="04EFEAF7FF83D211FAADF8D698A4B77F" box="[1318,1436,1876,1897]" italics="true" pageId="33" pageNumber="434">O. amballae</emphasis>
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</keyLead>
</paragraph>
<paragraph id="362436E5FF83D211FF1CF8F398A4B790" blockId="33.[151,1436,1817,2044]" box="[151,1436,1905,1927]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF8F398A4B790" box="[151,1436,1905,1927]" pageId="33" pageNumber="434">
- Terga with 57 longitudinal keels, antennae with 21 segments, teeth 4(6)+4(6)..............................
<taxonomicName id="F19B4D66FF83D211FAB4F8F398A4B790" ID-CoL="4B7WK" authorityName="Porat" authorityYear="1876" box="[1343,1436,1905,1926]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="33" pageNumber="434" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="04EFEAF7FF83D211FAB4F8F398A4B790" box="[1343,1436,1905,1926]" italics="true" pageId="33" pageNumber="434">O. scaber</emphasis>
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<paragraph id="362436E5FF83D211FF1CF80C98A4B7B5" blockId="33.[151,1436,1817,2044]" box="[151,1436,1934,1956]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF80C98A4B7B5" box="[151,1436,1934,1956]" pageId="33" pageNumber="434">3. Antennae with 1718 antennomeres with more than three glabrous ones.......................................... 4</keyLead>
</paragraph>
<paragraph id="362436E5FF83D211FF1CF82E98A4B7D7" blockId="33.[151,1436,1817,2044]" box="[151,1436,1963,1986]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF82E98A4B7D7" box="[151,1436,1963,1986]" pageId="33" pageNumber="434">- Antennae with 2021 antennomeres with first two ones and 1/31/2 of the third glabrous............................ 5</keyLead>
</paragraph>
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<keyStep id="8D6F2E40FF83D211FF1CF84B98A4B7EA" pageId="33" pageNumber="434">
<paragraph id="362436E5FF83D211FF1CF84B98A4B7C8" blockId="33.[151,1436,1817,2044]" box="[151,1436,1993,2015]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF84B98A4B7C8" box="[151,1436,1993,2015]" pageId="33" pageNumber="434">4. Antennae with 3 antennomeres glabrous................................................................... 7</keyLead>
</paragraph>
<paragraph id="362436E5FF83D211FF1CF86598A4B7EA" blockId="33.[151,1436,1817,2044]" box="[151,1436,2022,2044]" pageId="33" pageNumber="434">
<keyLead id="8D6A95D0FF83D211FF1CF86598A4B7EA" box="[151,1436,2022,2044]" pageId="33" pageNumber="434">
- Antennae with 4 antennomeres glabrous.........................................................
<taxonomicName id="F19B4D66FF83D211FA97F86598A4B7EA" ID-CoL="4B7WH" authorityName="Schileyko" authorityYear="1995" box="[1308,1436,2023,2044]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="33" pageNumber="434" phylum="Arthropoda" rank="species" species="reservatus">
<emphasis id="04EFEAF7FF83D211FA97F86598A4B7EA" box="[1308,1436,2023,2044]" italics="true" pageId="33" pageNumber="434">O. reservatus</emphasis>
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<paragraph id="362436E5FF80D212FF1CFF1B98A4B0B8" blockId="34.[151,1437,153,498]" box="[151,1436,153,175]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFF1B98A4B0B8" box="[151,1436,153,175]" pageId="34" pageNumber="435">
5. Each coxosternal tooth with 6(8)+6(8) small teeth.................................................
<taxonomicName id="F19B4D66FF80D212FAA9FF1898A4B0B8" ID-CoL="4B7VZ" authorityName="Haase" authorityYear="1887" box="[1314,1436,153,175]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="04EFEAF7FF80D212FAA9FF1898A4B0B8" box="[1314,1436,153,175]" italics="true" pageId="34" pageNumber="435">O. multidens</emphasis>
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<paragraph id="362436E5FF80D212FF1CFF3598A4B0DB" blockId="34.[151,1437,153,498]" box="[151,1436,183,205]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFF3598A4B0DB" box="[151,1436,183,205]" pageId="34" pageNumber="435">- Each coxosternal tooth with 3+3 or 4+4 small teeth.......................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="8D6F2E40FF80D212FF1CFF5698A4B154" pageId="34" pageNumber="435">
<paragraph id="362436E5FF80D212FF1CFF5698A5B111" blockId="34.[151,1437,153,498]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFF5698A5B111" pageId="34" pageNumber="435">
6. Penultimate prefemur with one spine, terminal leg with one tarsal spur, coxopleural process short, with 23 apical spines...............................................................................................
<taxonomicName id="F19B4D66FF80D212FAA7FF7098A5B111" ID-CoL="4B7WN" authorityName="Porat" authorityYear="1876" box="[1324,1437,242,263]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="04EFEAF7FF80D212FAA7FF7098A5B111" box="[1324,1437,242,263]" italics="true" pageId="34" pageNumber="435">O. spinosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="362436E5FF80D212FF16FE8D98A4B154" blockId="34.[151,1437,153,498]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF16FE8D98A4B154" pageId="34" pageNumber="435">
- Penultimate prefemur without spine, terminal leg without tarsal spur, coxopleural process long, with 57 apical spines.................................................................................................
<taxonomicName id="F19B4D66FF80D212FABCFEAF98A4B154" ID-CoL="75DNQ" baseAuthorityName="Kohlrausch" baseAuthorityYear="1878" box="[1335,1436,301,322]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="astenus">
<emphasis id="04EFEAF7FF80D212FABCFEAF98A4B154" box="[1335,1436,301,322]" italics="true" pageId="34" pageNumber="435">O. astenus</emphasis>
</taxonomicName>
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<keyStep id="8D6F2E40FF80D212FF1CFEC898A4B18F" pageId="34" pageNumber="435">
<paragraph id="362436E5FF80D212FF1CFEC898A4B16A" blockId="34.[151,1437,153,498]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFEC898A4B16A" pageId="34" pageNumber="435">
7. Coxopleural process with 57 apical spines, 23 lateral spines; terminal leg with many unarranged small spines (more than 26 spines) or some rows of bigger spines............................................................
<taxonomicName id="F19B4D66FF80D212FAA8FEE598A4B16A" ID-CoL="4B7V5" authorityName="Haase" authorityYear="1887" box="[1315,1436,359,380]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="04EFEAF7FF80D212FAA8FEE598A4B16A" box="[1315,1436,359,380]" italics="true" pageId="34" pageNumber="435">O. aculeatus</emphasis>
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<paragraph id="362436E5FF80D212FF1CFE0798A4B18F" blockId="34.[151,1437,153,498]" box="[151,1436,388,410]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFE0798A4B18F" box="[151,1436,388,410]" pageId="34" pageNumber="435">- Coxopleural process with 13 apical spines................................................................ 8</keyLead>
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</keyStep>
<keyStep id="8D6F2E40FF80D212FF1CFE2098A4B1E4" pageId="34" pageNumber="435">
<paragraph id="362436E5FF80D212FF1CFE2098A4B1C2" blockId="34.[151,1437,153,498]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFE2098A4B1C2" pageId="34" pageNumber="435">
8. Coxopleural process with 3 apical spines, without lateral spines. Sterna 220 with complete paramedian sutures......................................................................................................
<taxonomicName id="F19B4D66FF80D212FAB5FE3D98A4B1C2" ID-CoL="4B7WD" authorityName="Karsch" authorityYear="1881" box="[1342,1436,447,468]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="04EFEAF7FF80D212FAB5FE3D98A4B1C2" box="[1342,1436,447,468]" italics="true" pageId="34" pageNumber="435">O. politus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="362436E5FF80D212FF1CFE5F98A4B1E4" blockId="34.[151,1437,153,498]" box="[151,1436,476,498]" pageId="34" pageNumber="435">
<keyLead id="8D6A95D0FF80D212FF1CFE5F98A4B1E4" box="[151,1436,476,498]" pageId="34" pageNumber="435">
- Coxopleural process with one apical spine, without lateral spine. Sterna without paramedian sutures..........
<taxonomicName id="F19B4D66FF80D212FAA2FE5F98A4B1E4" ID-CoL="4B7WX" authorityName="Schileyko" authorityYear="1992" box="[1321,1436,477,498]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="voprosus">
<emphasis id="04EFEAF7FF80D212FAA2FE5F98A4B1E4" box="[1321,1436,477,498]" italics="true" pageId="34" pageNumber="435">O. voprosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>