treatments-xml/data/03/B7/F6/03B7F615FFA0FFFEFF0AFAABF041FDFC.xml

<document id="E44CA681E9CB1EA8D193169EE321D89F" ID-DOI="10.11646/phytotaxa.677.1.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="14522214" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734563225261" checkinUser="felipe" docAuthor="Silva, Marcos José Da &amp; Sodré, Rodolfo Carneiro" docDate="2024" docId="03B7F615FFA0FFFEFF0AFAABF041FDFC" docLanguage="en" docName="phytotaxa.677.1.087-094.pdf" docOrigin="Phytotaxa 677 (1)" docSource="https://doi.org/10.11646/phytotaxa.677.1.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Croton espinhacensis M. J. Silva &amp; Sodre 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="92" masterDocId="FF8E8E6DFFA1FFFBFF82FFDFF414FFB9" masterDocTitle="A new species of Croton (Euphorbiaceae) from the Espinhaço Mountain Range, Brazil" masterLastPageNumber="94" masterPageNumber="87" pageNumber="88" updateTime="1734567951438" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="76E5DACAF83CA7692474A00B6151CE32">A new species of Croton (Euphorbiaceae) from the Espinhaço Mountain Range, Brazil</mods:title>
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<mods:namePart id="7D64B5250F722ECAAD624662E8FD6525">Silva, Marcos José Da</mods:namePart>
<mods:affiliation id="7D23EBFDAC83379D3EFED87B109E0138">Laboratório de Morfologia e Taxonomia Vegetal, Instituto de Ciências Biológicas, Departamento de Botânica, Universidade Federal de Goiás, CP 131, 74001 - 970, Goiânia, GO, Brazil</mods:affiliation>
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<mods:namePart id="DA40FD2755CAA0CC6539C73837C060B9">Sodré, Rodolfo Carneiro</mods:namePart>
<mods:affiliation id="2A323FA3D2DC7D7F065CC26B0B6661CC">Programa de Pós-graduação em Ciências Biológicas-Botânica, Departamento de Botânica, Instituto de Biociências, Universidade Estadual Paulista “ Júlio de Mesquita Filho ”, CP 510, 18618 - 970, Botucatu, SP, Brazil</mods:affiliation>
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<paragraph id="8BA14703FFA0FFFAFF0AFAABF70AFA37" blockId="1.[136,798,1396,1423]" box="[136,798,1396,1423]" pageId="1" pageNumber="88">
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<taxonomicName id="4C1E3C80FFA0FFFAFF0AFAABF675FA37" authority="M.J. Silva &amp; Sodre" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[136,609,1396,1423]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="espinhacensis" status="sp. nov.">
<emphasis id="B96A9B11FFA0FFFAFF0AFAABF594FA36" bold="true" box="[136,384,1396,1423]" italics="true" pageId="1" pageNumber="88">Croton espinhacensis</emphasis>
M.J. Silva &amp; Sodré
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, 
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<emphasis id="B96A9B11FFA0FFFAFDEFFAAAF6A8FA37" box="[621,700,1397,1422]" italics="true" pageId="1" pageNumber="88">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
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)
</heading>
</paragraph>
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<paragraph id="8BA14703FFA0FFFAFF0AFA1FF128FA40" blockId="1.[136,1452,1472,1818]" pageId="1" pageNumber="88">
<emphasis id="B96A9B11FFA0FFFAFF0AFA1FF4AEFA6C" bold="true" box="[136,186,1472,1493]" pageId="1" pageNumber="88">Type</emphasis>
:— 
<materialsCitation id="3B764D5EFFA0FFFAFF55FA1FF12CFA40" collectingDate="2016-11-30" collectionCode="A, C, G, J, B, S, E, CEN, UFG, UB" collectorName="Sevilha &amp; Pereira-Silva &amp; Pereira &amp; de Noronha" country="Brazil" county="Fruta de Leite" elevation="892" latitude="-16.211945" location="Ponto" longLatPrecision="21" longitude="-42.481113" municipality="Sitio" pageId="1" pageNumber="88" specimenCode="CEN 103263" specimenCount="2" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="F3090793FFA0FFFAFF55FA1FF527FA6C" box="[215,307,1472,1493]" name="Brazil" pageId="1" pageNumber="88">BRAZIL</collectingCountry>
. 
<collectingRegion id="49DA89E1FFA0FFFAFEBCFA1FF5D0FA6F" box="[318,452,1472,1494]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="88">Minas Gerais</collectingRegion>
: 
<collectingCounty id="62C03F8FFFA0FFFAFE4DFA1FF649FA6F" box="[463,605,1472,1494]" pageId="1" pageNumber="88">Fruta de Leite</collectingCounty>
, 
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2, 
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1, 
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, 
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, 
<date id="FFA061C3FFA0FFFAFBE7FA1FF137FA6F" box="[1125,1315,1472,1494]" pageId="1" pageNumber="88" value="2016-11-30">
<collectingDate id="EFE4982BFFA0FFFAFBE7FA1FF137FA6F" box="[1125,1315,1472,1494]" pageId="1" pageNumber="88" value="2016-11-30">30 November 2016</collectingDate>
</date>
(fl., fr.), 
<emphasis id="B96A9B11FFA0FFFAFA03FA1FF734FA40" italics="true" pageId="1" pageNumber="88">
<collectionCode id="ED0FDFC6FFA0FFFAFA03FA1FF187FA6C" box="[1409,1427,1472,1493]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="88" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA0FFFAFA14FA1FF1BCFA6C" box="[1430,1448,1472,1493]" country="Denmark" name="University of Copenhagen" pageId="1" pageNumber="88" type="Herbarium">C</collectionCode>
. 
<collectorName id="26EB22D5FFA0FFFAFF3FFA3BF512FA40" box="[189,262,1508,1529]" pageId="1" pageNumber="88">Sevilha</collectorName>
, 
<collectionCode id="ED0FDFC6FFA0FFFAFE92FA3BF530FA40" box="[272,292,1508,1529]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="1" pageNumber="88" type="Herbarium">G</collectionCode>
. 
<collectorName id="26EB22D5FFA0FFFAFEAFFA3BF5A4FA40" box="[301,432,1508,1529]" pageId="1" pageNumber="88">Pereira-Silva</collectorName>
, 
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. 
<collectionCode id="ED0FDFC6FFA0FFFAFE49FA3BF5C8FA40" box="[459,476,1508,1529]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="88" type="Herbarium">B</collectionCode>
. 
<collectorName id="26EB22D5FFA0FFFAFE67FA3BF639FA40" box="[485,557,1508,1529]" pageId="1" pageNumber="88">Pereira</collectorName>
&amp; 
<collectionCode id="ED0FDFC6FFA0FFFAFDCEFA3BF64EFA40" box="[588,602,1508,1529]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="1" pageNumber="88" type="Herbarium">S</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA0FFFAFDDFFA3BF67BFA40" box="[605,623,1508,1529]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="88" type="Herbarium">E</collectionCode>
. 
<collectorName id="26EB22D5FFA0FFFAFDFBFA3BF6FEFA40" box="[633,746,1508,1529]" pageId="1" pageNumber="88">de Noronha</collectorName>
5890
</emphasis>
(
<typeStatus id="54A5F9A1FFA0FFFAFCAFFA3BF792FA43" box="[813,902,1508,1530]" pageId="1" pageNumber="88" type="holotype">holotype</typeStatus>
: 
<specimenCode id="DBB8EF78FFA0FFFAFC0DFA3BF001FA43" box="[911,1045,1508,1530]" collectionCode="CEN" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="1" pageNumber="88" type="Herbarium">CEN 103263</specimenCode>
!; 
<typeStatus id="54A5F9A1FFA0FFFAFBA3FA3BF067FA43" box="[1057,1139,1508,1530]" pageId="1" pageNumber="88" type="isotype">isotypes</typeStatus>
: 
<collectionCode id="ED0FDFC6FFA0FFFAFBFFFA3BF0A0FA40" box="[1149,1204,1508,1529]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="1" pageNumber="88" type="Herbarium">CEN</collectionCode>
!, 
<collectionCode id="ED0FDFC6FFA0FFFAFB43FA3BF0ECFA40" box="[1217,1272,1508,1529]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12703" lsid="urn:lsid:biocol.org:col:12703" name="Universidade Federal de Goiás" pageId="1" pageNumber="88" type="Herbarium">UFG</collectionCode>
!, 
<collectionCode id="ED0FDFC6FFA0FFFAFA87FA3BF13AFA40" box="[1285,1326,1508,1529]" country="France" name="Laboratoire de Biostratigraphie" pageId="1" pageNumber="88">UB</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3041488FFA0FFFAFF0AF9D8F553F8A3" pageId="1" pageNumber="88" type="diagnosis">
<paragraph id="8BA14703FFA0FFFAFF0AF9D8F553F8A3" blockId="1.[136,1452,1472,1818]" pageId="1" pageNumber="88">
<emphasis id="B96A9B11FFA0FFFAFF0AF9D8F4FFF9A4" bold="true" box="[136,235,1543,1565]" pageId="1" pageNumber="88">Diagnosis</emphasis>
:—This new species has indumented anthers, an unusual feature in the genus 
<taxonomicName id="4C1E3C80FFA0FFFAFC4FF9D7F004F9A4" authorityName="Linnaeus (1753:" authorityYear="1004" box="[973,1040,1544,1565]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="genus">
<emphasis id="B96A9B11FFA0FFFAFC4FF9D7F004F9A4" box="[973,1040,1544,1565]" italics="true" pageId="1" pageNumber="88">Croton</emphasis>
</taxonomicName>
. It ressembles 
<taxonomicName id="4C1E3C80FFA0FFFAFB1DF9D7F172F9A4" authority="Carn." authorityName="Carn." box="[1183,1382,1544,1565]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="harleyi">
<emphasis id="B96A9B11FFA0FFFAFB1DF9D7F138F9A4" box="[1183,1324,1544,1565]" italics="true" pageId="1" pageNumber="88">Croton harleyi</emphasis>
Carn.
</taxonomicName>
-Torres &amp; Cordeiro (
<emphasis id="B96A9B11FFA0FFFAFEBAF9F3F55FF9F8" box="[312,331,1580,1601]" italics="true" pageId="1" pageNumber="88">in</emphasis>
<bibRefCitation id="EF8F3AF2FFA0FFFAFED2F9F3F690F9F8" author="Carneiro-Torres, D. S. &amp; Cordeiro, I. &amp; Giulietti, A. M. &amp; Berry, P. E. &amp; Riina, R." box="[336,644,1580,1602]" pageId="1" pageNumber="88" pagination="122 - 132" refId="ref4494" refString="Carneiro-Torres, D. S., Cordeiro, I., Giulietti, A. M., Berry, P. E. &amp; Riina, R. (2011) Three new species of Croton (Euphorbiaceae s. s.) from the Brazilian Caatinga. Brittonia 63: 122 - 132. https: // doi. org / 10.1007 / s 12228 - 010 - 9145 - 1" type="journal article" year="2011">
Carneiro-Torres 
<emphasis id="B96A9B11FFA0FFFAFE6CF9F3F60AF9F8" box="[494,542,1580,1601]" italics="true" pageId="1" pageNumber="88">et al.</emphasis>
2011: 128
</bibRefCitation>
), from the Northeastern region of 
<collectingCountry id="F3090793FFA0FFFAFC45F9F3F010F9FB" box="[967,1028,1580,1602]" name="Brazil" pageId="1" pageNumber="88">Brazil</collectingCountry>
, by the shrubby habit, stems densely distally branched with leaves grouped at the apex of branches, bisexual thyrses with unisexual cymes and flowers of both sexes continuous, and staminate flowers with 11 stamens. However, 
<taxonomicName id="4C1E3C80FFA0FFFAFD2AF9ABF75EF930" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[680,842,1652,1673]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA0FFFAFD2AF9ABF75EF930" box="[680,842,1652,1673]" italics="true" pageId="1" pageNumber="88">C. espinhacensis</emphasis>
</taxonomicName>
differs especially in having leaf blades that are chartaceaeous, ovate-lanceolate, ovate-oblong or cordate, with stipitate patelliform nectary glands at the leaf base (
<emphasis id="B96A9B11FFA0FFFAFB1EF947F0A2F914" box="[1180,1206,1688,1709]" italics="true" pageId="1" pageNumber="88">vs.</emphasis>
membranaceous, elliptic blades, and with cylindric nectary glands at the leaf base in 
<taxonomicName id="4C1E3C80FFA0FFFAFC8BF963F77EF968" box="[777,874,1724,1745]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="harleyi">
<emphasis id="B96A9B11FFA0FFFAFC8BF963F77EF968" box="[777,874,1724,1745]" italics="true" pageId="1" pageNumber="88">C. harleyi</emphasis>
</taxonomicName>
), thyrses with 1(–2) pistillate flowers at the base (
<emphasis id="B96A9B11FFA0FFFAFADAF963F167F968" box="[1368,1395,1724,1745]" italics="true" pageId="1" pageNumber="88">vs.</emphasis>
2–4), anthers indumeted (
<emphasis id="B96A9B11FFA0FFFAFEFFF93FF58CF94C" box="[381,408,1760,1781]" italics="true" pageId="1" pageNumber="88">vs.</emphasis>
glabrous), pistillate flowers pedicelate with plane sepals (
<emphasis id="B96A9B11FFA0FFFAFC4BF93FF7C9F94C" box="[969,989,1760,1781]" italics="true" pageId="1" pageNumber="88">vs</emphasis>
. sessile with revolute sepals), and seeds oblong (
<emphasis id="B96A9B11FFA0FFFAFF47F8DBF4F4F8A0" box="[197,224,1796,1817]" italics="true" pageId="1" pageNumber="88">vs.</emphasis>
ellipsoid).
</paragraph>
</subSubSection>
<subSubSection id="C3041488FFA0FFF9FF0AF897F763FC69" lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88" type="description">
<paragraph id="8BA14703FFA0FFF9FF0AF897F763FC69" blockId="1.[136,1452,1864,2071]" lastBlockId="2.[136,1452,158,2093]" lastPageId="2" lastPageNumber="89" pageId="1" pageNumber="88">
<emphasis id="B96A9B11FFA0FFFAFF0AF897F506F8DB" bold="true" box="[136,274,1864,1890]" pageId="1" pageNumber="88">Description</emphasis>
:—Subshrub or shrub 
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tall, erect, monoecious, densely branched distally, adult branches darkish and glabrous and young branches ferrugineous or green-brownish, with stellate or mutiradiate trichomes, the first ones sessile, with 7–10 rays; the second ones multiradiate in 2 or 3 whorls, with stipe measuring 
<quantity id="4CE6EAE6FFA0FFFAFB60F84FF17EF812" box="[1250,1386,1936,1963]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="88" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
long, and rays up to 
<quantity id="4CE6EAE6FFA0FFFAFEA9F86BF595F876" box="[299,385,1972,1999]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="88" unit="mm" value="0.5">0.5 mm</quantity>
long; latex scarce, hyaline; stipules 0.9–1 × 
<quantity id="4CE6EAE6FFA0FFFAFCF3F86BF7EDF876" box="[881,1017,1972,1999]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="88" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
, lanceolate or narrowly triangular with 4–8 ovoid and sessile nectary glands at the base, covered with stellate trichomes on both surfaces, the trichomes more densed externally. leaves entire, spirally alternate along the stems; petioles 
<quantity id="4CE6EAE6FFA0FFFAFC50F823F05BF7AE" box="[978,1103,2044,2071]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="14.0" metricValueMin="5.0" pageId="1" pageNumber="88" unit="cm" value="0.95" valueMax="1.4" valueMin="0.5">0.5–1.4 cm</quantity>
long; blades 1.4–3.9 × 
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, ovate, ovate-lanceolate, ovate-oblong or cordate, with obtuse or slightly cordate base with 2 shortly stipitate patelliform glands 
<quantity id="4CE6EAE6FFA3FFF9FED9FF1DF5F6FF64" box="[347,482,194,221]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="89" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
diam. on the abaxial surface, acute or shortly acuminate apex, entire or slightly serrate or serrulate margin, with sessile colleters in the sinuses, membranaceous, discolorous, adaxial surface light green and adaxial surface green-ferrugineous, the first ones with stellate trichomes with 6–10 rays, the second ones multiradiate in 2 or 3 whorls, with stipe 
<quantity id="4CE6EAE6FFA3FFF9FE4EFEF1F642FEF0" box="[460,598,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="89" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
long, and rays up to 
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long, both, tomentose to the touch; venation brochidodromous, with 4–6 pairs of secondary veins, primary and secondary veins prominent on the abaxial surface and impressed on adaxial surface. Inflorescences bisexual, 
<quantity id="4CE6EAE6FFA3FFF9FC96FEA9F79BFE28" box="[788,911,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="2" pageNumber="89" unit="cm" value="1.2" valueMax="1.5" valueMin="0.9">0.9–1.5 cm</quantity>
long, terminal with 1(–2) pistillate flowers at the base axis, and 9 or 18 staminate flowers continuous along the inflorescence axis; bracts of staminate flowers 2.3–2.4 × 
<quantity id="4CE6EAE6FFA3FFF9FF0AFE61F504FE60" box="[136,272,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="89" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
, lanceolate glabrous internally and pin-cushion externally; bracts of pistillate flowers 4–4.1 × 
<quantity id="4CE6EAE6FFA3FFF9FA9DFE61F1B3FE60" box="[1311,1447,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="89" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
, lanceolate similar to the staminates ones. Staminate flowers 
<quantity id="4CE6EAE6FFA3FFF9FCB6FE3DF786FE44" box="[820,914,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="2" pageNumber="89" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long, yellowish, actinomorphic; pedicels 
<quantity id="4CE6EAE6FFA3FFF9FAEEFE3DF4A0FD98" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="2" pageNumber="89" unit="mm" value="4.05" valueMax="4.1" valueMin="4.0">4–4.1 mm</quantity>
long; calyx 5-lobed, lobes 2.1–2.2 × 
<quantity id="4CE6EAE6FFA3FFF9FDE3FDD9F6F8FD98" box="[609,748,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="2" pageNumber="89" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
, equal, oval, ovate-triangular, with acute apex, slightly united at the base, ventrally glabrous, dorsally stellate; petals 2.4–2.8 × 
<quantity id="4CE6EAE6FFA3FFF9FCE2FDF5F7C0FDFC" box="[864,980,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="2" pageNumber="89" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">1–1.1 mm</quantity>
, elliptic, ovate-elliptic or elliptic-obovate, with a obtuse apex, villous externally with stellate trichomes along the midrib, ciliate on the margins, the longest cilia at the base; stamens 11, 
<quantity id="4CE6EAE6FFA3FFF9FE23FDADF63EFD34" box="[417,554,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="2" pageNumber="89" unit="mm" value="3.75" valueMax="3.8" valueMin="3.7">3.7–3.8 mm</quantity>
long, with a sparsely pubescent indumentum of hispid trichomes on filaments and anthers, anthers oblong with 
<quantity id="4CE6EAE6FFA3FFF9FD88FD49F680FD08" box="[522,660,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="2" pageNumber="89" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
long; disk with 5 obovoid segments, yellowish with margin slightly crenulate; receptacle villous. Pistillate flowers 
<quantity id="4CE6EAE6FFA3FFF9FD1FFD65F732FD6C" box="[669,806,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="2" pageNumber="89" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.2–5.5 mm</quantity>
long; pedicels 
<quantity id="4CE6EAE6FFA3FFF9FC54FD65F05EFD6C" box="[982,1098,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="2" pageNumber="89" unit="mm" value="3.05" valueMax="3.1" valueMin="3.0">3–3.1 mm</quantity>
long; sepals 5, subequal 4.8–5 × 
<quantity id="4CE6EAE6FFA3FFF9FF22FD01F53EFD40" box="[160,298,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="2" pageNumber="89" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6–1.8 mm</quantity>
, lanceolate or narrowly triangular, with acute apex, margin entire, plane, eglandular with multiradiate trichomes on both surfaces, the trichomes more densed externally, free, valvate prefloration; petals 5, glanduliform; ovary 2–2.1 × 
<quantity id="4CE6EAE6FFA3FFF9FEABFCF9F5BBFCF8" box="[297,431,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="2" pageNumber="89" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 mm</quantity>
globose, yellowish-ferrugineous with tomentose indumentum of stellate trichomes; stigmas 3, deeply bifid, 
<quantity id="4CE6EAE6FFA3FFF9FEBCFC95F5DDFCDC" box="[318,457,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="2" pageNumber="89" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6−3.9 mm</quantity>
long, covered with stellate trichomes; disk 5-segmented, receptacle glabrous. Capsules 4.9–5 × 
<quantity id="4CE6EAE6FFA3FFF9FF67FCB1F54DFC30" box="[229,345,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="2" pageNumber="89" unit="mm" value="3.95" valueMax="4.0" valueMin="3.9">3.9–4 mm</quantity>
, oblongoid, ferrugineous, the trichomes multiradiate, sepals accrescent, similar to the length of the fruit, pedicels 
<quantity id="4CE6EAE6FFA3FFF9FEAEFC4DF5A0FC14" box="[300,436,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="2" pageNumber="89" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
long; seeds 4–4.1 × 
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, oblong, grayish, with discretely dark spots; caruncle hatshaped, sessile, yellow-whitish, visible on both surface of the seed.
</paragraph>
</subSubSection>
<subSubSection id="C3041488FFA3FFF9FF3FFC05F12EFB11" pageId="2" pageNumber="89" type="distribution">
<paragraph id="8BA14703FFA3FFF9FF3FFC05F179FB85" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="89">
<emphasis id="B96A9B11FFA3FFF9FF3FFC05F5F7FC4D" bold="true" box="[189,483,986,1012]" pageId="2" pageNumber="89">Distribution and habitat</emphasis>
:—The species is so far known from the Espinhaço Mountain Range, in the Brazilian states of 
<collectingRegion id="49DA89E1FFA3FFF9FF6EFC21F539FBA1" box="[236,301,1022,1048]" country="Brazil" name="Bahia" pageId="2" pageNumber="89">Bahia</collectingRegion>
and 
<collectingRegion id="49DA89E1FFA3FFF9FEE6FC21F5EEFBA1" box="[356,506,1022,1048]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="89">Minas Gerais</collectingRegion>
(
<figureCitation id="13255B86FFA3FFF9FD88FC21F65BFBA1" box="[522,591,1022,1048]" captionStart="FIGURE 2" captionStartId="4.[136,234,1047,1069]" captionTargetBox="[240,1334,203,1018]" captionTargetId="figure-20@4.[222,1365,190,1023]" captionTargetPageId="4" captionText="FIGURE 2. Distribution map of Croton espinhacencis M.J. Silva &amp; Sodré. a map of Brazil with the states of Minas Gerais and Bahia highlighted. b Expanded view of the Minas Gerais and Bahia states, showing their limits and the occurrence points of the new species. Abbreviations for states and federation unit: AL = Alagoas, BA = Bahia, DF = Distrito Federal, ES = Espírito Santo, GO = Goiás, MA = Maranhão, MG = Minas Gerais, MS= Mato Grosso do Sul, MT= Mato Grosso, PI = Piauí; RJ = Rio de Janeiro; SE = Sergipe, SP = São Paulo, and TO = Tocantins." figureDoi="http://doi.org/10.5281/zenodo.14522220" httpUri="https://zenodo.org/record/14522220/files/figure.png" pageId="2" pageNumber="89">Fig. 2</figureCitation>
), where it grows in “cerrado rupestre” and “cerrado denso” in flat or hillside areas between 828 and 
<quantity id="4CE6EAE6FFA3FFF9FE0FFBFDF5F6FB84" box="[397,482,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.391" pageId="2" pageNumber="89" unit="m" value="1391.0">1391 m</quantity>
of altitude. It prefers sandy and litholics soils, as well as plinthosols or latossols.
</paragraph>
<paragraph id="8BA14703FFA3FFF9FF3FFB99F7F7FBD8" blockId="2.[136,1452,158,2093]" box="[189,995,1094,1121]" pageId="2" pageNumber="89">
<emphasis id="B96A9B11FFA3FFF9FF3FFB99F52DFBD9" bold="true" box="[189,313,1094,1120]" pageId="2" pageNumber="89">Phenology</emphasis>
: 
<emphasis id="B96A9B11FFA3FFF9FEC0FB98F54FFBD9" box="[322,347,1095,1120]" italics="true" pageId="2" pageNumber="89">—</emphasis>
Collected with flowers and fruits from November to May.
</paragraph>
<paragraph id="8BA14703FFA3FFF9FF3FFBB5F12EFB11" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="89">
<emphasis id="B96A9B11FFA3FFF9FF3FFBB5F62CFB3D" bold="true" box="[189,568,1130,1156]" pageId="2" pageNumber="89">Preliminary conservation status</emphasis>
:—Species with an estimated EOO and AOO of 
<quantity id="4CE6EAE6FFA3FFF9FBCCFBB5F0FDFB3D" box="[1102,1257,1130,1157]" metricMagnitude="6" metricUnit="m" metricValue="7.825856" pageId="2" pageNumber="89" unit="km" value="7825.856">7,825.856 km</quantity>
<superScript id="7C6BEA4BFFA3FFF9FB6AFBB5F0E4FBC1" attach="left" box="[1256,1264,1130,1144]" fontSize="6" pageId="2" pageNumber="89">2</superScript>
and 
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, respectively. Therefore, it is tentatively classified here as Vulnerable (EN) Criteria B1 subcriteria b(i, iv, v).
</paragraph>
</subSubSection>
<subSubSection id="C3041488FFA3FFFEFF3FFB6DF041FDFC" lastPageId="5" lastPageNumber="92" pageId="2" pageNumber="89" type="etymology">
<paragraph id="8BA14703FFA3FFF9FF3FFB6DF674FB49" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="89">
<emphasis id="B96A9B11FFA3FFF9FF3FFB6DF52AFB75" bold="true" box="[189,318,1202,1228]" pageId="2" pageNumber="89">Etymology</emphasis>
: 
<emphasis id="B96A9B11FFA3FFF9FEC4FB6CF54BFB75" box="[326,351,1203,1228]" italics="true" pageId="2" pageNumber="89">—</emphasis>
The specific epithet 
<taxonomicName id="4C1E3C80FFA3FFF9FDC7FB6DF6FAFB75" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[581,750,1202,1228]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="espinhacensis">
“ 
<emphasis id="B96A9B11FFA3FFF9FDD0FB6DF6FAFB75" box="[594,750,1202,1228]" italics="true" pageId="2" pageNumber="89">espinhacensis</emphasis>
</taxonomicName>
” alludes to the region of the Espinhaço Range, state of 
<collectingRegion id="49DA89E1FFA3FFF9FAE7FB6DF4C5FB49" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="89">Minas Gerais</collectingRegion>
and 
<collectingRegion id="49DA89E1FFA3FFF9FE8AFB09F558FB49" box="[264,332,1238,1264]" country="Brazil" name="Bahia" pageId="2" pageNumber="89">Bahia</collectingRegion>
, where the species lives.
</paragraph>
<paragraph id="8BA14703FFA3FFF9FF3FFB25F6D1F8ED" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="89">
<materialsCitation id="3B764D5EFFA3FFF9FF3FFB25F66EFA71" collectingDate="1994-03-16" collectingDateMax="2020-02-14" collectingDateMin="1994-03-16" collectionCode="V, C, ESA, SPF, R, S, MCCA, M, G, ALCB, BHCB, A, CEN" collectorName="Souza &amp; Fonseca &amp; Monte Azul &amp; Parque Estadual Caminho dos Gerais &amp; Oliveira &amp; Areiao &amp; Ponto &amp; Servilha" country="Brazil" county="Mortugaba" elevation="1000" latitude="-15.441944" location="Fruta de Leite" longLatPrecision="21" longitude="-42.481113" municipality="Jacaraci" pageId="2" pageNumber="89" specimenCount="2" stateProvince="Bahia">
<emphasis id="B96A9B11FFA3FFF9FF3FFB25F62DFAAD" bold="true" box="[189,569,1274,1300]" pageId="2" pageNumber="89">
<typeStatus id="54A5F9A1FFA3FFF9FF3FFB25F62DFAAD" box="[189,569,1274,1300]" isOtherTypeStatus="true" pageId="2" pageNumber="89">Additional specimens examined</typeStatus>
</emphasis>
(
<emphasis id="B96A9B11FFA3FFF9FDC9FB25F6D5FAAD" bold="true" box="[587,705,1274,1300]" pageId="2" pageNumber="89">
<typeStatus id="54A5F9A1FFA3FFF9FDC9FB25F6D5FAAD" box="[587,705,1274,1300]" pageId="2" pageNumber="89" type="paratype">paratypes</typeStatus>
</emphasis>
):— 
<collectingCountry id="F3090793FFA3FFF9FD69FB25F743FAAD" box="[747,855,1274,1300]" name="Brazil" pageId="2" pageNumber="89">BRAZIL</collectingCountry>
. 
<collectingRegion id="49DA89E1FFA3FFF9FCE0FB25F7BCFAAD" box="[866,936,1274,1300]" country="Brazil" name="Bahia" pageId="2" pageNumber="89">Bahia</collectingRegion>
: 
<collectingCounty id="62C03F8FFFA3FFF9FC31FB25F025FAAD" box="[947,1073,1274,1300]" pageId="2" pageNumber="89">Mortugaba</collectingCounty>
, ca. 
<quantity id="4CE6EAE6FFA3FFF9FBE7FB25F08BFAAD" box="[1125,1183,1274,1300]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="89" unit="km" value="8.0">8 km</quantity>
da cidade em direção a 
<collectingMunicipality id="6BC5DD79FFA3FFF9FF0AFAC1F4F6FA81" box="[136,226,1310,1336]" pageId="2" pageNumber="89">Jacaraci</collectingMunicipality>
(“Jacarau” on the label), 
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, 
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, 
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<elevation id="0033A030FFA3FFF9FC97FAC1F779FA80" box="[789,877,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="89" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FCFBFAC1F031FA81" box="[889,1061,1310,1336]" pageId="2" pageNumber="89" value="1994-03-16">
<collectingDate id="EFE4982BFFA3FFF9FCFBFAC1F031FA81" box="[889,1061,1310,1336]" pageId="2" pageNumber="89" value="1994-03-16">16 March 1994</collectingDate>
</date>
(fl.), 
<emphasis id="B96A9B11FFA3FFF9FBE4FAC0F14BFA81" box="[1126,1375,1310,1337]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FBE4FAC0F06DFA80" box="[1126,1145,1311,1337]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="89" type="Museum">V</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FBFFFAC0F085FA80" box="[1149,1169,1311,1337]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="89" type="Herbarium">C</collectionCode>
. 
<collectorName id="26EB22D5FFA3FFF9FB1EFAC0F0CBFA80" box="[1180,1247,1311,1337]" pageId="2" pageNumber="89">Souza</collectorName>
et al. 5527
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FAECFAC1F1B3FA81" box="[1390,1447,1310,1336]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14319" lsid="urn:lsid:biocol.org:col:14319" name="Universidade de São Paulo" pageId="2" pageNumber="89" type="Herbarium">ESA</collectionCode>
, 
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). 
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: 
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, 
<location id="8EC111D8FFA3FFF9FD96FA9DF731FAE4" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FD96FA9DF731FAE4" box="[532,805,1346,1373]" country="Brazil" county="Mortugaba" latitude="-15.441944" longLatPrecision="21" longitude="-42.481113" municipality="Jacaraci" name="Vale do Rio Peixe Bravo" pageId="2" pageNumber="89" stateProvince="Bahia">Vale do Rio Peixe Bravo</location>
, 
<date id="FFA061C3FFA3FFF9FCADFA9DF7E7FAE5" box="[815,1011,1346,1372]" pageId="2" pageNumber="89" value="2020-02-14">
<collectingDate id="EFE4982BFFA3FFF9FCADFA9DF7E7FAE5" box="[815,1011,1346,1372]" pageId="2" pageNumber="89" value="2020-02-14">14 February 2020</collectingDate>
</date>
(fl.), 
<emphasis id="B96A9B11FFA3FFF9FBADFA9CF12EFAE5" box="[1071,1338,1346,1373]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FBADFA9CF054FAE5" box="[1071,1088,1347,1372]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="89">R</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FBC7FA9CF04CFAE4" box="[1093,1112,1347,1373]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="89" type="Herbarium">S</collectionCode>
. 
<collectorName id="26EB22D5FFA3FFF9FBE0FA9CF0D5FAE5" box="[1122,1217,1347,1372]" pageId="2" pageNumber="89">Fonseca</collectorName>
et al. 1329
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FAC5FA9DF1B4FAE5" box="[1351,1440,1346,1372]" pageId="2" pageNumber="89">MCCA</collectionCode>
). 
<collectorName id="26EB22D5FFA3FFF9FF0AFAB8F504FA39" box="[136,272,1382,1409]" pageId="2" pageNumber="89">Monte Azul</collectorName>
, 
<collectorName id="26EB22D5FFA3FFF9FE9EFAB8F6D4FA39" box="[284,704,1382,1409]" pageId="2" pageNumber="89">Parque Estadual Caminho dos Gerais</collectorName>
, 
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, 
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, 
<quantity id="4CE6EAE6FFA3FFF9FC69FAB9F057FA38" box="[1003,1091,1382,1409]" metricMagnitude="3" metricUnit="m" metricValue="1.0459999999999998" pageId="2" pageNumber="89" unit="m" value="1046.0">
<elevation id="0033A030FFA3FFF9FC69FAB9F057FA38" box="[1003,1091,1382,1409]" metricMagnitude="3" metricUnit="m" metricValue="1.0459999999999998" pageId="2" pageNumber="89" unit="m" value="1046.0">1046 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FBCDFAB9F102FA39" box="[1103,1302,1382,1408]" pageId="2" pageNumber="89" value="2017-02-16">
<collectingDate id="EFE4982BFFA3FFF9FBCDFAB9F102FA39" box="[1103,1302,1382,1408]" pageId="2" pageNumber="89" value="2017-02-16">16 February 2017</collectingDate>
</date>
(fl.), 
<emphasis id="B96A9B11FFA3FFF9FAD5FAB8F542FA1D" italics="true" pageId="2" pageNumber="89">
D. 
<collectionCode id="ED0FDFC6FFA3FFF9FAF3FAB8F19AFA38" box="[1393,1422,1383,1409]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="89" type="Herbarium">M</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FA10FAB8F1BCFA38" box="[1426,1448,1383,1409]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="2" pageNumber="89" type="Herbarium">G</collectionCode>
. 
<collectorName id="26EB22D5FFA3FFF9FF0AFA55F4F1FA1D" box="[136,229,1418,1444]" pageId="2" pageNumber="89">Oliveira</collectorName>
et al. 124
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FEE4FA55F5A1FA1D" box="[358,437,1418,1444]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15066" lsid="urn:lsid:biocol.org:col:15066" name="Universidade Federal da Bahia, Campus Universitário de Ondina" pageId="2" pageNumber="89" type="Herbarium">ALCB</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FE42FA55F605FA1D" box="[448,529,1418,1444]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15456" lsid="urn:lsid:biocol.org:col:15456" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="89" type="Herbarium">BHCB</collectionCode>
). 
<location id="8EC111D8FFA3FFF9FDA6FA55F713FA1D" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FDA6FA55F713FA1D" box="[548,775,1418,1444]" country="Brazil" county="Mortugaba" latitude="-15.441944" longLatPrecision="21" longitude="-42.481113" municipality="Jacaraci" name="Rio Pardo de Minas" pageId="2" pageNumber="89" stateProvince="Bahia">Rio Pardo de Minas</location>
, 
<collectorName id="26EB22D5FFA3FFF9FC93FA55F774FA1D" box="[785,864,1418,1444]" pageId="2" pageNumber="89">Areião</collectorName>
, 
<collectorName id="26EB22D5FFA3FFF9FCEEFA54F7B9FA1C" box="[876,941,1419,1445]" pageId="2" pageNumber="89">Ponto</collectorName>
5, 
<geoCoordinate id="EE2A21C4FFA3FFF9FC53FA55F04CFA1C" box="[977,1112,1418,1445]" degrees="15" direction="south" minutes="26" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="31" value="-15.441944">15°26’31”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FBE6FA55F0E0FA1C" box="[1124,1268,1418,1445]" degrees="42" direction="west" minutes="28" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="52" value="-42.481113">42°28’52”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FB7DFA55F15CFA1C" box="[1279,1352,1418,1445]" metricMagnitude="2" metricUnit="m" metricValue="9.02" pageId="2" pageNumber="89" unit="m" value="902.0">
<elevation id="0033A030FFA3FFF9FB7DFA55F15CFA1C" box="[1279,1352,1418,1445]" metricMagnitude="2" metricUnit="m" metricValue="9.02" pageId="2" pageNumber="89" unit="m" value="902.0">902 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FAD7FA55F4D4FA70" pageId="2" pageNumber="89" value="2005-05-23">
<collectingDate id="EFE4982BFFA3FFF9FAD7FA55F4D4FA70" pageId="2" pageNumber="89" value="2005-05-23">23 May 2005</collectingDate>
</date>
(fr), 
<emphasis id="B96A9B11FFA3FFF9FE83FA70F636FA71" box="[257,546,1454,1481]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FE83FA70F502FA70" box="[257,278,1455,1481]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="89" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FE98FA70F53BFA70" box="[282,303,1455,1481]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="89" type="Herbarium">C</collectionCode>
. 
<collectorName id="26EB22D5FFA3FFF9FEBFFA71F58EFA71" box="[317,410,1454,1480]" pageId="2" pageNumber="89">Servilha</collectorName>
et al. 4305
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FDB6FA71F666FA71" box="[564,626,1454,1480]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="2" pageNumber="89" type="Herbarium">CEN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B764D5EFFA3FFF9FD0BFA71F168FA55" collectingDate="2006-04-21" collectionCode="S, ESA, HUEFS" country="Brazil" elevation="868" latitude="-15.831111" location="Rio Pardo de Minas" longLatPrecision="21" longitude="-42.428333" municipality="Novo Horizonte" pageId="2" pageNumber="89" specimenCount="1">
estrada de chão sentido entroncamento 
<collectingMunicipality id="6BC5DD79FFA3FFF9FBD0FA70F11FFA71" box="[1106,1291,1454,1481]" pageId="2" pageNumber="89">Novo Horizonte</collectingMunicipality>
/ 
<location id="8EC111D8FFA3FFF9FA91FA71F4C5FA55" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FA91FA71F4C5FA55" country="Brazil" latitude="-15.831111" longLatPrecision="21" longitude="-42.428333" municipality="Novo Horizonte" name="Rio Pardo de Minas" pageId="2" pageNumber="89">Rio Pardo de Minas</location>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FF5EFA0DF577FA54" box="[220,355,1490,1517]" degrees="15" direction="south" minutes="49" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="52" value="-15.831111">15°49’52”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FEEDFA0DF5EAFA54" box="[367,510,1490,1517]" degrees="42" direction="west" minutes="25" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="42" value="-42.428333">42°25’42”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FD89FA0DF640FA54" box="[523,596,1490,1517]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="2" pageNumber="89" unit="m" value="868.0">
<elevation id="0033A030FFA3FFF9FD89FA0DF640FA54" box="[523,596,1490,1517]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="2" pageNumber="89" unit="m" value="868.0">868 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FDE2FA0DF6E8FA55" box="[608,764,1490,1516]" pageId="2" pageNumber="89" value="2006-04-21">
<collectingDate id="EFE4982BFFA3FFF9FDE2FA0DF6E8FA55" box="[608,764,1490,1516]" pageId="2" pageNumber="89" value="2006-04-21">21 April 2006</collectingDate>
</date>
(fl., fr.), 
<emphasis id="B96A9B11FFA3FFF9FCE6FA0CF0D6FA55" box="[868,1218,1490,1517]" italics="true" pageId="2" pageNumber="89">
D. 
<collectionCode id="ED0FDFC6FFA3FFF9FCFDFA0CF784FA54" box="[895,912,1491,1517]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="89" type="Herbarium">S</collectionCode>
. Carneiro-Torres et al. 695
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FB50FA0DF11FFA55" box="[1234,1291,1490,1516]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14319" lsid="urn:lsid:biocol.org:col:14319" name="Universidade de São Paulo" pageId="2" pageNumber="89" type="Herbarium">ESA</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FA94FA0DF160FA55" box="[1302,1396,1490,1516]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15506" lsid="urn:lsid:biocol.org:col:15506" name="Universidade Estadual de Feira de Santana" pageId="2" pageNumber="89" type="Herbarium">HUEFS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B764D5EFFA3FFF9FA0AFA0DF146F9A9" collectingDate="2008-12-10" collectionCode="A, C, CEN" country="Brazil" elevation="876" latitude="-15.912499" location="Vereda Funda" longLatPrecision="21" longitude="-42.475555" pageId="2" pageNumber="89" specimenCount="1">
<emphasis id="B96A9B11FFA3FFF9FA0AFA0DF1B0FA55" box="[1416,1444,1490,1516]" italics="true" pageId="2" pageNumber="89">ib.</emphasis>
, 
<location id="8EC111D8FFA3FFF9FF0AFA29F532F9A9" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FF0AFA29F532F9A9" box="[136,294,1526,1552]" country="Brazil" latitude="-15.912499" longLatPrecision="21" longitude="-42.475555" name="Vereda Funda" pageId="2" pageNumber="89">Vereda Funda</location>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FEB3FA29F5ADF9A8" box="[305,441,1526,1553]" degrees="15" direction="south" minutes="54" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="45" value="-15.912499">15°54’45”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FE46FA29F640F9A9" box="[452,596,1526,1552]" degrees="42" direction="west" minutes="28" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="32" value="-42.475555">42°28’32”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FDDDFA29F6BCF9A8" box="[607,680,1526,1553]" metricMagnitude="2" metricUnit="m" metricValue="8.76" pageId="2" pageNumber="89" unit="m" value="876.0">
<elevation id="0033A030FFA3FFF9FDDDFA29F6BCF9A8" box="[607,680,1526,1553]" metricMagnitude="2" metricUnit="m" metricValue="8.76" pageId="2" pageNumber="89" unit="m" value="876.0">876 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FD36FA29F79EF9A9" box="[692,906,1526,1552]" pageId="2" pageNumber="89" value="2008-12-10">
<collectingDate id="EFE4982BFFA3FFF9FD36FA29F79EF9A9" box="[692,906,1526,1552]" pageId="2" pageNumber="89" value="2008-12-10">10 December 2008</collectingDate>
</date>
(fl., fr), 
<emphasis id="B96A9B11FFA3FFF9FC69FA28F0E9F9A9" box="[1003,1277,1526,1553]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FC69FA28F7EBF9A8" box="[1003,1023,1527,1553]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="89" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FB89FA28F035F9A8" box="[1035,1057,1527,1553]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="89" type="Herbarium">C</collectionCode>
. Sevilha et al. 4951
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FA8EFA29F15EF9A9" box="[1292,1354,1526,1552]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="2" pageNumber="89" type="Herbarium">CEN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B764D5EFFA3FFF9FADCFA28F115F9E1" collectingDate="2012-03-21" collectionCode="S, W, J, A, M, N, HRCB, UPCB" country="Brazil" elevation="846" location="Corrego da Velha" municipality="Parque Estadual da Serra Nova" pageId="2" pageNumber="89" specimenCount="1">
<collectingMunicipality id="6BC5DD79FFA3FFF9FADCFA28F58AF98C" pageId="2" pageNumber="89">Parque Estadual da Serra Nova</collectingMunicipality>
, trilha para o escorregador no 
<location id="8EC111D8FFA3FFF9FC88F9C5F7CCF98D" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FC88F9C5F7CCF98D" box="[778,984,1562,1588]" country="Brazil" municipality="Parque Estadual da Serra Nova" name="Corrego da Velha" pageId="2" pageNumber="89">Córrego da Velha</location>
, 15°36’53.9–57.3” 
<collectionCode id="ED0FDFC6FFA3FFF9FB36F9C5F0D6F98C" box="[1204,1218,1562,1589]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="89" type="Herbarium">S</collectionCode>
, 42°44’5.6–20.6” 
<collectionCode id="ED0FDFC6FFA3FFF9FA09F9C5F1B3F98C" box="[1419,1447,1562,1589]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="2" pageNumber="89" type="Herbarium">W</collectionCode>
, 
<quantity id="4CE6EAE6FFA3FFF9FF0AF9E1F51EF9E0" box="[136,266,1598,1625]" metricMagnitude="2" metricUnit="m" metricValue="8.459999999999999" metricValueMax="8.58" metricValueMin="8.34" pageId="2" pageNumber="89" unit="m" value="846.0" valueMax="858.0" valueMin="834.0">
<elevation id="0033A030FFA3FFF9FF0AF9E1F51EF9E0" box="[136,266,1598,1625]" metricMagnitude="2" metricUnit="m" metricValue="8.459999999999999" metricValueMax="8.58" metricValueMin="8.34" pageId="2" pageNumber="89" unit="m" value="846.0" valueMax="858.0" valueMin="834.0">834–858 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FE94F9E1F5D7F9E1" box="[278,451,1598,1624]" pageId="2" pageNumber="89" value="2012-03-21">
<collectingDate id="EFE4982BFFA3FFF9FE94F9E1F5D7F9E1" box="[278,451,1598,1624]" pageId="2" pageNumber="89" value="2012-03-21">21 March 2012</collectingDate>
</date>
(fl., fr.), 
<emphasis id="B96A9B11FFA3FFF9FDA9F9E0F02BF9E1" box="[555,1087,1598,1625]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FDA9F9E0F62FF9E0" box="[555,571,1599,1625]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="89" type="Herbarium">J</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FDBDF9E0F647F9E0" box="[575,595,1599,1625]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="89" type="Herbarium">A</collectionCode>
. Lombardi, 
<collectionCode id="ED0FDFC6FFA3FFF9FD59F9E0F6E2F9E1" box="[731,758,1599,1624]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="89" type="Herbarium">M</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FD7BF9E0F704F9E1" box="[761,784,1599,1624]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="2" pageNumber="89" type="Herbarium">N</collectionCode>
. Saka &amp; D. 
<collectionCode id="ED0FDFC6FFA3FFF9FC10F9E0F7B2F9E0" box="[914,934,1599,1625]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="89" type="Herbarium">A</collectionCode>
. Araújo 9029
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FBCCF9E1F08AF9E1" box="[1102,1182,1598,1624]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15505" lsid="urn:lsid:biocol.org:col:15505" name="Universidade Estadual Paulista" pageId="2" pageNumber="89" type="Herbarium">HRCB</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FB29F9E1F0EDF9E1" box="[1195,1273,1598,1624]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15175" lsid="urn:lsid:biocol.org:col:15175" name="Universidade Federal do Paraná" pageId="2" pageNumber="89" type="Herbarium">UPCB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B764D5EFFA3FFF9FA8FF9E1F6BDF951" collectingDate="1994-03-17" collectingDateMax="2016-12-01" collectingDateMin="1994-03-17" collectionCode="J, M, BHCB, A, C, CEN, SPF, K, R" country="Brazil" county="Gerais" elevation="1391" latitude="-15.378056" location="Rubelita" longLatPrecision="21" longitude="-42.698334" municipality="Costa" pageId="2" pageNumber="89" specimenCode="CFCR 15128" specimenCount="1">
<emphasis id="B96A9B11FFA3FFF9FA8FF9E1F137F9E1" box="[1293,1315,1598,1624]" italics="true" pageId="2" pageNumber="89">ib</emphasis>
., trilha para o 
<collectingCounty id="62C03F8FFFA3FFF9FF1CF9BDF4FEF9C5" box="[158,234,1634,1660]" pageId="2" pageNumber="89">Gerais</collectingCounty>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FF74F9BDF568F9C4" box="[246,380,1634,1661]" degrees="15" direction="south" minutes="39" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="40" value="-15.661111">15°39’40”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FE08F9BDF60EF9C4" box="[394,538,1634,1661]" degrees="42" direction="west" minutes="45" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="57" value="-42.765835">42°45’57”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FDA4F9BDF66AF9C4" box="[550,638,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.391" pageId="2" pageNumber="89" unit="m" value="1391.0">
<elevation id="0033A030FFA3FFF9FDA4F9BDF66AF9C4" box="[550,638,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.391" pageId="2" pageNumber="89" unit="m" value="1391.0">1391 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FD08F9BDF72CF9C5" box="[650,824,1634,1660]" pageId="2" pageNumber="89" value="2012-03-22">
<collectingDate id="EFE4982BFFA3FFF9FD08F9BDF72CF9C5" box="[650,824,1634,1660]" pageId="2" pageNumber="89" value="2012-03-22">22 March 2012</collectingDate>
</date>
(fl.), 
<emphasis id="B96A9B11FFA3FFF9FCFBF9BCF062F9C5" box="[889,1142,1634,1661]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FCFBF9BCF79CF9C5" box="[889,904,1635,1660]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="89" type="Herbarium">J</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FC0EF9BCF7BCF9C5" box="[908,936,1635,1660]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="89" type="Herbarium">M</collectionCode>
. 
<collectingMunicipality id="6BC5DD79FFA3FFF9FC31F9BCF7E0F9C4" box="[947,1012,1635,1661]" pageId="2" pageNumber="89">Costa</collectingMunicipality>
et al. 1016
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FB04F9BDF0CCF9C5" box="[1158,1240,1634,1660]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15456" lsid="urn:lsid:biocol.org:col:15456" name="Universidade Federal de Minas Gerais" pageId="2" pageNumber="89" type="Herbarium">BHCB</collectionCode>
). 
<location id="8EC111D8FFA3FFF9FB6EF9BDF15AF9C5" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FB6EF9BDF15AF9C5" box="[1260,1358,1634,1660]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="Rubelita" pageId="2" pageNumber="89">Rubelita</location>
, 
<location id="8EC111D8FFA3FFF9FADBF9BDF19AF9C5" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FADBF9BDF19AF9C5" box="[1369,1422,1634,1660]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="Sitio" pageId="2" pageNumber="89">Sítio</location>
2, 
<location id="8EC111D8FFA3FFF9FF0AF958F4DDF918" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FF0AF958F4DDF918" box="[136,201,1671,1697]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="Ponto" pageId="2" pageNumber="89">Ponto</location>
<quantity id="4CE6EAE6FFA3FFF9FF50F959F52EF918" box="[210,314,1670,1697]" metricMagnitude="3" metricUnit="m" metricValue="3.828" pageId="2" pageNumber="89" unit="m" value="3828.0">
<elevation id="0033A030FFA3FFF9FF50F959F52EF918" box="[210,314,1670,1697]" metricMagnitude="3" metricUnit="m" metricValue="3.828" pageId="2" pageNumber="89" unit="m" value="3828.0">3, 828 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FEC5F959F60BF919" box="[327,543,1670,1696]" pageId="2" pageNumber="89" value="2016-12-01">
<collectingDate id="EFE4982BFFA3FFF9FEC5F959F60BF919" box="[327,543,1670,1696]" pageId="2" pageNumber="89" value="2016-12-01">01 December 2016</collectingDate>
</date>
(fl., fr), 
<emphasis id="B96A9B11FFA3FFF9FD01F958F786F919" box="[643,914,1670,1697]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FD01F958F68CF918" box="[643,664,1671,1697]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="89" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FD1EF958F6A5F918" box="[668,689,1671,1697]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="89" type="Herbarium">C</collectionCode>
. 
<location id="8EC111D8FFA3FFF9FD3FF959F71BF919" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FD3FF959F71BF919" box="[701,783,1670,1696]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="Sevilha" pageId="2" pageNumber="89">Sevilha</location>
et al. 5961
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FC21F959F7F5F919" box="[931,993,1670,1696]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="2" pageNumber="89" type="Herbarium">CEN</collectionCode>
). 
<location id="8EC111D8FFA3FFF9FC76F959F119F919" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FC76F959F119F919" box="[1012,1293,1670,1696]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="Santo Antonio do Retiro" pageId="2" pageNumber="89">Santo Antônio do Retiro</location>
, ca. 
<quantity id="4CE6EAE6FFA3FFF9FAC3F959F19DF919" box="[1345,1417,1670,1696]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="89" unit="km" value="11.0">11 km</quantity>
da cidade em direção a 
<location id="8EC111D8FFA3FFF9FEECF974F5E0F97D" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FEECF974F5E0F97D" box="[366,500,1706,1733]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="Mato Verde" pageId="2" pageNumber="89">Mato Verde</location>
, ca. 
<quantity id="4CE6EAE6FFA3FFF9FDA4F975F64BF97D" box="[550,607,1706,1732]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="89" unit="km" value="2.0">2 km</quantity>
do 
<location id="8EC111D8FFA3FFF9FD0BF975F739F97D" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FD0BF975F739F97D" box="[649,813,1706,1732]" country="Brazil" county="Gerais" latitude="-15.378056" longLatPrecision="21" longitude="-42.698334" municipality="Costa" name="rio Mandacaia" pageId="2" pageNumber="89">rio Mandaçaia</location>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FCB5F975F7AAF97C" box="[823,958,1706,1733]" degrees="15" direction="south" minutes="22" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="41" value="-15.378056">15°22’41”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FC48F975F04EF97C" box="[970,1114,1706,1733]" degrees="42" direction="west" minutes="41" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="54" value="-42.698334">42°41’54”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FBE7F975F0A8F97C" box="[1125,1212,1706,1733]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="89" unit="m" value="1000.0">
<elevation id="0033A030FFA3FFF9FBE7F975F0A8F97C" box="[1125,1212,1706,1733]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="89" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FB45F975F167F97D" box="[1223,1395,1706,1732]" pageId="2" pageNumber="89" value="1994-03-17">
<collectingDate id="EFE4982BFFA3FFF9FB45F975F167F97D" box="[1223,1395,1706,1732]" pageId="2" pageNumber="89" value="1994-03-17">17 March 1994</collectingDate>
</date>
(fl.), 
<emphasis id="B96A9B11FFA3FFF9FF0AF910F60EF951" box="[136,538,1742,1769]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FF0AF910F488F950" box="[136,156,1743,1769]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="89" type="Herbarium">C</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FF22F910F4A9F950" box="[160,189,1743,1769]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="89" type="Herbarium">M</collectionCode>
. Sakuragui et al. 
<specimenCode id="DBB8EF78FFA3FFF9FE06F910F60EF951" box="[388,538,1742,1769]" collectionCode="CFCR" pageId="2" pageNumber="89">CFCR 15128</specimenCode>
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FDA8F911F64FF951" box="[554,603,1742,1768]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15730" lsid="urn:lsid:biocol.org:col:15730" name="Universidade de São Paulo" pageId="2" pageNumber="89" type="Herbarium">SPF</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FDE5F910F66AF950" box="[615,638,1743,1769]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="2" pageNumber="89" type="Herbarium">K</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FD08F911F6B4F951" box="[650,672,1742,1768]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="89">R</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B764D5EFFA3FFF9FD34F911F6D5F8ED" collectingDate="2017-03-15" collectingDateMax="2022-04-04" collectingDateMin="2017-03-15" collectionCode="A, C, CEN, UFG, M, RB, SPF, HUEFS" collectorName="da Cidade" country="Brazil" county="Sucuriu" elevation="986" latitude="-15.801945" location="Parque Estadual de Serra Nova" longLatPrecision="21" longitude="-42.768612" municipality="Serranopolis de Minas" pageId="2" pageNumber="89" specimenCount="1">
<collectingCounty id="62C03F8FFFA3FFF9FD34F911F704F951" box="[694,784,1742,1768]" pageId="2" pageNumber="89">Sucuriu</collectingCounty>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FC9EF911F7B7F950" box="[796,931,1742,1769]" degrees="15" direction="south" minutes="20" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="18" value="-15.338333">15°20’18”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FC32F911F02BF950" box="[944,1087,1742,1769]" degrees="42" direction="west" minutes="45" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="09" value="-42.7525">42°45’09”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FBCEF911F082F950" box="[1100,1174,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="9.86" pageId="2" pageNumber="89" unit="m" value="986.0">
<elevation id="0033A030FFA3FFF9FBCEF911F082F950" box="[1100,1174,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="9.86" pageId="2" pageNumber="89" unit="m" value="986.0">986 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FB21F911F144F951" box="[1187,1360,1742,1769]" pageId="2" pageNumber="89" value="2017-03-15">
<collectingDate id="EFE4982BFFA3FFF9FB21F911F144F951" box="[1187,1360,1742,1769]" pageId="2" pageNumber="89" value="2017-03-15">15 March 2017</collectingDate>
</date>
(fl., fr), 
<emphasis id="B96A9B11FFA3FFF9FF0AF92CF583F8B5" box="[136,407,1778,1805]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FF0AF92CF489F8B4" box="[136,157,1779,1805]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="89" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED0FDFC6FFA3FFF9FF23F92CF4A2F8B4" box="[161,182,1779,1805]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="89" type="Herbarium">C</collectionCode>
. Sevilha et al. 6370
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FE2AF92DF5F0F8B5" box="[424,484,1778,1804]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="2" pageNumber="89" type="Herbarium">CEN</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FE73F92DF63AF8B5" box="[497,558,1778,1804]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12703" lsid="urn:lsid:biocol.org:col:12703" name="Universidade Federal de Goiás" pageId="2" pageNumber="89" type="Herbarium">UFG</collectionCode>
). 
<collectingMunicipality id="6BC5DD79FFA3FFF9FDC0F92DF750F8B5" box="[578,836,1778,1804]" pageId="2" pageNumber="89">Serranópolis de Minas</collectingMunicipality>
, 
<location id="8EC111D8FFA3FFF9FCD2F92CF0A7F8B4" LSID="urn:lsid:plazi:treatment:03B7F615FFA0FFFEFF0AFAABF041FDFC:8EC111D8FFA3FFF9FCD2F92CF0A7F8B4" box="[848,1203,1778,1805]" country="Brazil" county="Sucuriu" latitude="-15.801945" longLatPrecision="21" longitude="-42.768612" municipality="Serranopolis de Minas" name="Parque Estadual de Serra Nova" pageId="2" pageNumber="89">Parque Estadual de Serra Nova</location>
e Talhado, estrada de acesso aos Gerais Santana, cerca de 
<quantity id="4CE6EAE6FFA3FFF9FDA7F8C9F64BF889" box="[549,607,1814,1840]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="89" unit="km" value="1.0">1 km</quantity>
a nordeste 
<collectorName id="26EB22D5FFA3FFF9FD60F8C9F740F889" box="[738,852,1814,1840]" pageId="2" pageNumber="89">da Cidade</collectorName>
das Pedras, 
<geoCoordinate id="EE2A21C4FFA3FFF9FC61F8C9F048F888" box="[995,1116,1814,1841]" degrees="15" direction="south" minutes="48" orientation="latitude" pageId="2" pageNumber="89" precision="15" seconds="7" value="-15.801945">15°48’7”S</geoCoordinate>
, 
<geoCoordinate id="EE2A21C4FFA3FFF9FBEBF8C9F0FFF889" box="[1129,1259,1814,1840]" degrees="42" direction="west" minutes="46" orientation="longitude" pageId="2" pageNumber="89" precision="15" seconds="7" value="-42.768612">42°46’7”W</geoCoordinate>
, 
<quantity id="4CE6EAE6FFA3FFF9FB75F8C9F155F888" box="[1271,1345,1814,1841]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="89" unit="m" value="950.0">
<elevation id="0033A030FFA3FFF9FB75F8C9F155F888" box="[1271,1345,1814,1841]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="89" unit="m" value="950.0">950 m</elevation>
</quantity>
, 
<date id="FFA061C3FFA3FFF9FACCF8C9F4D4F8ED" pageId="2" pageNumber="89" value="2022-04-04">
<collectingDate id="EFE4982BFFA3FFF9FACCF8C9F4D4F8ED" pageId="2" pageNumber="89" value="2022-04-04">04 April 2022</collectingDate>
</date>
(fl.), 
<emphasis id="B96A9B11FFA3FFF9FE82F8E4F5C9F8ED" box="[256,477,1850,1876]" italics="true" pageId="2" pageNumber="89">
<collectionCode id="ED0FDFC6FFA3FFF9FE82F8E4F50FF8ED" box="[256,283,1851,1876]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="89" type="Herbarium">M</collectionCode>
. Verdi et al. 8820
</emphasis>
(
<collectionCode id="ED0FDFC6FFA3FFF9FE6FF8E5F602F8ED" box="[493,534,1850,1876]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="2" pageNumber="89" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FDA3F8E5F644F8ED" box="[545,592,1850,1876]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15730" lsid="urn:lsid:biocol.org:col:15730" name="Universidade de São Paulo" pageId="2" pageNumber="89" type="Herbarium">SPF</collectionCode>
, 
<collectionCode id="ED0FDFC6FFA3FFF9FDD9F8E5F6ADF8ED" box="[603,697,1850,1876]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15506" lsid="urn:lsid:biocol.org:col:15506" name="Universidade Estadual de Feira de Santana" pageId="2" pageNumber="89" type="Herbarium">HUEFS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA14703FFA3FFF9FF3FF881F0BBF794" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="89">
<emphasis id="B96A9B11FFA3FFF9FF3FF881F514F8C1" bold="true" box="[189,256,1886,1912]" pageId="2" pageNumber="89">Notes</emphasis>
:— 
<taxonomicName id="4C1E3C80FFA3FFF9FEA6F880F60DF8C1" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[292,537,1886,1913]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA3FFF9FEA6F880F60DF8C1" box="[292,537,1886,1913]" italics="true" pageId="2" pageNumber="89">Croton espinhacensis</emphasis>
</taxonomicName>
is placed here in 
<taxonomicName id="4C1E3C80FFA3FFF9FD70F880F0F6F8C1" authority="subsect. Geiseleria" box="[754,1250,1886,1913]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="subSection" section="Geiseleria" subSection="Geiseleria">
<emphasis id="B96A9B11FFA3FFF9FD70F880F754F8C0" box="[754,832,1887,1913]" italics="true" pageId="2" pageNumber="89">Croton</emphasis>
sect. 
<emphasis id="B96A9B11FFA3FFF9FC0BF881F7EAF8C1" box="[905,1022,1886,1912]" italics="true" pageId="2" pageNumber="89">Geiseleria</emphasis>
subsect. 
<emphasis id="B96A9B11FFA3FFF9FBEFF881F0F6F8C1" box="[1133,1250,1886,1912]" italics="true" pageId="2" pageNumber="89">Geiseleria</emphasis>
</taxonomicName>
. This decision is based on the combination of the following character states: shrubby or subshrubby habit, alternate leaves no more than 2 to 3 times as long as wide, with sessile or stipitate nectary glands at the apex of the petiole or at the base of the blade and/or in the sinuses of the leaf margin, thyrses without a distinct gap between staminate and pistillate flowers, pistillate flowers with free sepals, glandular or eglandular equal or slightly unequal, developed or glandulifom petals, bifid styles with 6 terminal branch tips, and fruits with an irregularly fimbriate columella apex.
</paragraph>
<caption id="DF61178BFFA2FFF8FF0AF890F7E5F7A0" ID-DOI="http://doi.org/10.5281/zenodo.14522218" ID-Zenodo-Dep="14522218" httpUri="https://zenodo.org/record/14522218/files/figure.png" pageId="3" pageNumber="90" startId="3.[136,234,1871,1893]" targetBox="[189,1365,205,1831]" targetPageId="3" targetType="figure">
<paragraph id="8BA14703FFA2FFF8FF0AF890F7E5F7A0" blockId="3.[136,1452,1871,2073]" pageId="3" pageNumber="90">
<emphasis id="B96A9B11FFA2FFF8FF0AF890F517F8DD" bold="true" box="[136,259,1871,1893]" pageId="3" pageNumber="90">FIGURE 1.</emphasis>
<taxonomicName id="4C1E3C80FFA2FFF8FE8AF88FF689F8DC" authority="M. J. Silva &amp; Sodre." authorityName="M. J. Silva &amp; Sodre." box="[264,669,1871,1893]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="90" phylum="Tracheophyta" rank="species" species="espinhacencis">
<emphasis id="B96A9B11FFA2FFF8FE8AF88FF5CCF8DD" box="[264,472,1871,1893]" italics="true" pageId="3" pageNumber="90">Croton espinhacencis</emphasis>
M.J. Silva &amp; Sodré.
</taxonomicName>
<emphasis id="B96A9B11FFA2FFF8FD20F88FF6BAF8DC" bold="true" box="[674,686,1872,1893]" pageId="3" pageNumber="90">a</emphasis>
fertile branch. 
<emphasis id="B96A9B11FFA2FFF8FCC2F88FF759F8DC" bold="true" box="[832,845,1872,1893]" pageId="3" pageNumber="90">b</emphasis>
leaf blade. 
<emphasis id="B96A9B11FFA2FFF8FC3EF88FF7D3F8DC" bold="true" box="[956,967,1872,1893]" pageId="3" pageNumber="90">c</emphasis>
detail of the glands. 
<emphasis id="B96A9B11FFA2FFF8FB12F88FF089F8DC" bold="true" box="[1168,1181,1872,1893]" pageId="3" pageNumber="90">d</emphasis>
detail of the the leaf margin showing one nectary gland and two colleters. 
<emphasis id="B96A9B11FFA2FFF8FDC8F8ABF641F830" bold="true" box="[586,597,1908,1929]" pageId="3" pageNumber="90">e</emphasis>
. stipule. 
<emphasis id="B96A9B11FFA2FFF8FD2DF8ACF6A3F830" bold="true" box="[687,695,1907,1929]" pageId="3" pageNumber="90">f</emphasis>
detail of inflorescence, showing flowers of both sexes and buds. 
<emphasis id="B96A9B11FFA2FFF8FAB8F8ABF152F830" bold="true" box="[1338,1350,1908,1929]" pageId="3" pageNumber="90">g</emphasis>
bract of a staminate flower. 
<emphasis id="B96A9B11FFA2FFF8FEB1F847F554F814" bold="true" box="[307,320,1944,1965]" pageId="3" pageNumber="90">h</emphasis>
staminate flower. 
<emphasis id="B96A9B11FFA2FFF8FE73F848F5ECF814" bold="true" box="[497,504,1943,1965]" pageId="3" pageNumber="90">i</emphasis>
ventral surface of a staminate petal 
<emphasis id="B96A9B11FFA2FFF8FCD0F848F74EF814" bold="true" box="[850,858,1943,1965]" pageId="3" pageNumber="90">j</emphasis>
dorsal surface of a staminate petal. 
<emphasis id="B96A9B11FFA2FFF8FB30F847F0ABF814" bold="true" box="[1202,1215,1944,1965]" pageId="3" pageNumber="90">k</emphasis>
stamen. 
<emphasis id="B96A9B11FFA2FFF8FA97F847F108F814" bold="true" box="[1301,1308,1944,1965]" pageId="3" pageNumber="90">l</emphasis>
lateral view of anther, note the hispid trichomes on dorsal surface. 
<emphasis id="B96A9B11FFA2FFF8FDFAF863F698F868" bold="true" box="[632,652,1980,2001]" pageId="3" pageNumber="90">m</emphasis>
bract of a pistillate flower. 
<emphasis id="B96A9B11FFA2FFF8FC14F863F7B7F868" bold="true" box="[918,931,1980,2001]" pageId="3" pageNumber="90">n</emphasis>
pistillate flower. 
<emphasis id="B96A9B11FFA2FFF8FBCEF863F04CF868" bold="true" box="[1100,1112,1980,2001]" pageId="3" pageNumber="90">o</emphasis>
. dorsal surface of a pistillate sepal. 
<emphasis id="B96A9B11FFA2FFF8FF0AF83FF481F84C" bold="true" box="[136,149,2016,2037]" pageId="3" pageNumber="90">p</emphasis>
detail of a pistillate flower with ovary removed to show the 5-segmented nectary disck and petals. 
<emphasis id="B96A9B11FFA2FFF8FBD8F83FF073F84C" bold="true" box="[1114,1127,2016,2037]" pageId="3" pageNumber="90">q</emphasis>
capsule with accrescent sepals. 
<emphasis id="B96A9B11FFA2FFF8FA23F83FF1B8F84C" bold="true" box="[1441,1452,2016,2037]" pageId="3" pageNumber="90">r</emphasis>
detail of the fruitg calyx. 
<emphasis id="B96A9B11FFA2FFF8FEFEF7DBF591F7A0" bold="true" box="[380,389,2052,2073]" pageId="3" pageNumber="90">s</emphasis>
seed ventral view. 
<emphasis id="B96A9B11FFA2FFF8FDBCF7DBF652F7A0" bold="true" box="[574,582,2052,2073]" pageId="3" pageNumber="90">t</emphasis>
seed dorsal view. Drawn from the holotype.
</paragraph>
</caption>
<caption id="DF61178BFFA5FFFFFF0AFBC8F584FB04" ID-DOI="http://doi.org/10.5281/zenodo.14522220" ID-Zenodo-Dep="14522220" httpUri="https://zenodo.org/record/14522220/files/figure.png" pageId="4" pageNumber="91" startId="4.[136,234,1047,1069]" targetBox="[240,1334,203,1018]" targetPageId="4" targetType="figure">
<paragraph id="8BA14703FFA5FFFFFF0AFBC8F584FB04" blockId="4.[136,1452,1047,1213]" pageId="4" pageNumber="91">
<emphasis id="B96A9B11FFA5FFFFFF0AFBC8F510FB95" bold="true" box="[136,260,1047,1069]" pageId="4" pageNumber="91">FIGURE 2.</emphasis>
Distribution map of 
<taxonomicName id="4C1E3C80FFA5FFFFFE53FBC8F778FB94" authority="M. J. Silva &amp; Sodre." authorityName="M. J. Silva &amp; Sodre." box="[465,876,1047,1069]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="espinhacencis">
<emphasis id="B96A9B11FFA5FFFFFE53FBC8F6B7FB95" box="[465,675,1047,1068]" italics="true" pageId="4" pageNumber="91">Croton espinhacencis</emphasis>
M.J. Silva &amp; Sodré.
</taxonomicName>
<emphasis id="B96A9B11FFA5FFFFFCF1FBC8F76BFB95" bold="true" box="[883,895,1047,1068]" pageId="4" pageNumber="91">a</emphasis>
map of Brazil with the states of Minas Gerais and Bahia highlighted. 
<emphasis id="B96A9B11FFA5FFFFFE81FBE4F504FBE9" bold="true" box="[259,272,1083,1104]" pageId="4" pageNumber="91">b</emphasis>
Expanded view of the Minas Gerais and Bahia states, showing their limits and the occurrence points of the new species. Abbreviations for states and federation unit: AL = Alagoas, BA = Bahia, DF = Distrito Federal, ES = Espírito Santo, GO = Goiás, MA = Maranhão, MG = Minas Gerais, MS= Mato Grosso do Sul, MT= Mato Grosso, PI = Piauí; RJ = Rio de Janeiro; SE = Sergipe, SP = São Paulo, and TO = Tocantins.
</paragraph>
</caption>
<paragraph id="8BA14703FFA5FFFFFF3FFB33F5D1FA00" blockId="4.[136,1452,1259,1466]" pageId="4" pageNumber="91">
Among the species of this section present in 
<collectingCountry id="F3090793FFA5FFFFFD4EFB34F707FABC" box="[716,787,1259,1285]" name="Brazil" pageId="4" pageNumber="91">Brazil</collectingCountry>
, 
<taxonomicName id="4C1E3C80FFA5FFFFFCA3FB33F002FABC" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[801,1046,1259,1286]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA5FFFFFCA3FB33F002FABC" box="[801,1046,1259,1286]" italics="true" pageId="4" pageNumber="91">Croton espinhacensis</emphasis>
</taxonomicName>
ressembles 
<taxonomicName id="4C1E3C80FFA5FFFFFB26FB33F4E4FA93" authority="(Carneiro et al. 2011)" baseAuthorityName="Carneiro" baseAuthorityYear="2011" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="harleyi">
<emphasis id="B96A9B11FFA5FFFFFB26FB33F103FABC" box="[1188,1303,1259,1286]" italics="true" pageId="4" pageNumber="91">C. harleyi</emphasis>
(Carneiro 
<emphasis id="B96A9B11FFA5FFFFFA15FB33F4B1FA90" italics="true" pageId="4" pageNumber="91">et al.</emphasis>
2011)
</taxonomicName>
, a species from the Caatinga vegetation of the 
<collectingRegion id="49DA89E1FFA5FFFFFC84FAD0F792FA93" box="[774,902,1295,1322]" country="Brazil" name="Bahia" pageId="4" pageNumber="91">Bahia state</collectingRegion>
. Both species share the shrubby habit, the stems densely branched from the upper third, the leaves commonly grouped in terminations of the branches, leaf blades small up to 5 × 
<quantity id="4CE6EAE6FFA5FFFFFEC4FA88F580FACB" box="[326,404,1367,1394]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="91" unit="cm" value="2.5">2.5 cm</quantity>
with nectaries on the base and margin, the thyrses up to 
<quantity id="4CE6EAE6FFA5FFFFFBB0FA87F078FACB" box="[1074,1132,1368,1394]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="91" unit="cm" value="2.0">2 cm</quantity>
long with pistillate flowers continuous to the staminate flowers, the latter with 11 stamens. However, the characters listed in 
<tableCitation id="C69C72B8FFA5FFFFFB30FAA4F117FA2F" box="[1202,1283,1403,1430]" captionStart="TABLE 1" captionStartId="4.[136,219,1515,1536]" captionText="TABLE 1. Differential characters between Croton espinhacensis and C. harleyi." pageId="4" pageNumber="91">Table 1</tableCitation>
certainly serve to differentiate both species.
</paragraph>
<caption id="DF61178BFFA5FFFFFF0AFA34F785F9B9" box="[136,913,1515,1537]" pageId="4" pageNumber="91" startId="4.[136,219,1515,1536]" targetType="table">
<paragraph id="8BA14703FFA5FFFFFF0AFA34F785F9B9" blockId="4.[136,913,1515,1537]" box="[136,913,1515,1537]" pageId="4" pageNumber="91">
<emphasis id="B96A9B11FFA5FFFFFF0AFA34F4FAF9B9" bold="true" box="[136,238,1515,1536]" pageId="4" pageNumber="91">TABLE 1</emphasis>
. Differential characters between 
<taxonomicName id="4C1E3C80FFA5FFFFFDAFFA34F6EFF9B9" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[557,763,1515,1536]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA5FFFFFDAFFA34F6EFF9B9" box="[557,763,1515,1536]" italics="true" pageId="4" pageNumber="91">Croton espinhacensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1E3C80FFA5FFFFFCA8FA34F799F9B9" box="[810,909,1515,1536]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="harleyi">C. harleyi</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA14703FFA5FFFFFF12F9CDF0C8F7AF" pageId="4" pageNumber="91">
<table id="F91EB5A3FFA50004FF12F9CDF198F7AE" box="[144,1420,1554,2071]" gridcols="3" gridrows="13" pageId="4" pageNumber="91">
<tr id="352E4541FFA50004FF12F9CDF198F99C" box="[144,1420,1554,1573]" gridrow="0" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F9CDF5E6F99C" box="[144,498,1554,1573]" gridcol="0" gridrow="0" pageId="4" pageNumber="91">Characters</th>
<th id="76FF2C3DFFA50004FD9CF9CDF7E1F99C" box="[542,1013,1554,1573]" gridcol="1" gridrow="0" pageId="4" pageNumber="91">
<taxonomicName id="4C1E3C80FFA5FFFFFD9CF9CDF6CFF99C" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[542,731,1554,1573]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA5FFFFFD9CF9CDF6CFF99C" bold="true" box="[542,731,1554,1573]" italics="true" pageId="4" pageNumber="91">Croton espinhacensis</emphasis>
</taxonomicName>
</th>
<th id="76FF2C3DFFA50004FB89F9CDF198F99C" box="[1035,1420,1554,1573]" gridcol="2" gridrow="0" pageId="4" pageNumber="91">
<taxonomicName id="4C1E3C80FFA5FFFFFB89F9CDF076F99C" authority="Stipules" authorityName="Stipules" box="[1035,1122,1554,1573]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="91" phylum="Tracheophyta" rank="species" species="harleyi">
<emphasis id="B96A9B11FFA5FFFFFB89F9CDF076F99C" bold="true" box="[1035,1122,1554,1573]" italics="true" pageId="4" pageNumber="91">C. harleyi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="352E4541FFA50004FF12F9E7F198F9F2" box="[144,1420,1592,1611]" gridrow="1" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F9E7F5E6F9F2" box="[144,498,1592,1611]" gridcol="0" gridrow="1" pageId="4" pageNumber="91">Stipules</th>
<td id="76FF2C3DFFA50004FD9CF9E7F7E1F9F2" box="[542,1013,1592,1611]" gridcol="1" gridrow="1" pageId="4" pageNumber="91">0.9–1 mm long, lanceolate or narrowly triangular</td>
<td id="76FF2C3DFFA50004FB89F9E7F198F9F2" box="[1035,1420,1592,1611]" gridcol="2" gridrow="1" pageId="4" pageNumber="91">2–3 mm long, linear</td>
</tr>
<tr id="352E4541FFA50004FF12F982F198F9C8" box="[144,1420,1629,1649]" gridrow="2" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F982F5E6F9C8" box="[144,498,1629,1649]" gridcol="0" gridrow="2" pageId="4" pageNumber="91">Leaf blade</th>
<td id="76FF2C3DFFA50004FD9CF982F7E1F9C8" box="[542,1013,1629,1649]" gridcol="1" gridrow="2" pageId="4" pageNumber="91">Ovate-lanceolate, ovate-oblong or cordate, chartaceous</td>
<td id="76FF2C3DFFA50004FB89F982F198F9C8" box="[1035,1420,1629,1649]" gridcol="2" gridrow="2" pageId="4" pageNumber="91">Elliptic or sometimes ovate, membranaceous</td>
</tr>
<tr id="352E4541FFA50004FF12F95CF198F92F" box="[144,1420,1667,1686]" gridrow="3" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F95CF5E6F92F" box="[144,498,1667,1686]" gridcol="0" gridrow="3" pageId="4" pageNumber="91">Trichomes of the leaves</th>
<td id="76FF2C3DFFA50004FD9CF95CF7E1F92F" box="[542,1013,1667,1686]" gridcol="1" gridrow="3" pageId="4" pageNumber="91">Stellate and multiradiate</td>
<td id="76FF2C3DFFA50004FB89F95CF198F92F" box="[1035,1420,1667,1686]" gridcol="2" gridrow="3" pageId="4" pageNumber="91">Stellate porrect and glandular sessile globose</td>
</tr>
<tr id="352E4541FFA50004FF12F977F198F905" box="[144,1420,1704,1724]" gridrow="4" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F977F5E6F905" box="[144,498,1704,1724]" gridcol="0" gridrow="4" pageId="4" pageNumber="91">Nectary glands at the base of leaf blade</th>
<td id="76FF2C3DFFA50004FD9CF977F7E1F905" box="[542,1013,1704,1724]" gridcol="1" gridrow="4" pageId="4" pageNumber="91">Stipitate patelliform</td>
<td id="76FF2C3DFFA50004FB89F977F198F905" box="[1035,1420,1704,1724]" gridcol="2" gridrow="4" pageId="4" pageNumber="91">Cylindrical</td>
</tr>
<tr id="352E4541FFA50004FF12F911F198F958" box="[144,1420,1742,1761]" gridrow="5" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F911F5E6F958" box="[144,498,1742,1761]" gridcol="0" gridrow="5" pageId="4" pageNumber="91">Colleters in the sinuses of the leaf margin</th>
<td id="76FF2C3DFFA50004FD9CF911F7E1F958" box="[542,1013,1742,1761]" gridcol="1" gridrow="5" pageId="4" pageNumber="91">Present</td>
<td id="76FF2C3DFFA50004FB89F911F198F958" box="[1035,1420,1742,1761]" gridcol="2" gridrow="5" pageId="4" pageNumber="91">Absent</td>
</tr>
<tr id="352E4541FFA50004FF12F92CF198F8BE" box="[144,1420,1779,1799]" gridrow="6" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F92CF5E6F8BE" box="[144,498,1779,1799]" gridcol="0" gridrow="6" pageId="4" pageNumber="91">Number of pistillate flowers per thyrses</th>
<td id="76FF2C3DFFA50004FD9CF92CF7E1F8BE" box="[542,1013,1779,1799]" gridcol="1" gridrow="6" pageId="4" pageNumber="91">1(–2)</td>
<td id="76FF2C3DFFA50004FB89F92CF198F8BE" box="[1035,1420,1779,1799]" gridcol="2" gridrow="6" pageId="4" pageNumber="91">2–4</td>
</tr>
<tr id="352E4541FFA50004FF12F8C6F198F895" box="[144,1420,1817,1836]" gridrow="7" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F8C6F5E6F895" box="[144,498,1817,1836]" gridcol="0" gridrow="7" pageId="4" pageNumber="91">Staminate bracts</th>
<td id="76FF2C3DFFA50004FD9CF8C6F7E1F895" box="[542,1013,1817,1836]" gridcol="1" gridrow="7" pageId="4" pageNumber="91">Lanceolate, 2.3–2.4 mm long</td>
<td id="76FF2C3DFFA50004FB89F8C6F198F895" box="[1035,1420,1817,1836]" gridcol="2" gridrow="7" pageId="4" pageNumber="91">Linear, 3–4 mm long</td>
</tr>
<tr id="352E4541FFA50004FF12F8E1F198F8EB" box="[144,1420,1854,1874]" gridrow="8" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F8E1F5E6F8EB" box="[144,498,1854,1874]" gridcol="0" gridrow="8" pageId="4" pageNumber="91">Staminate flowers</th>
<td id="76FF2C3DFFA50004FD9CF8E1F7E1F8EB" box="[542,1013,1854,1874]" gridcol="1" gridrow="8" pageId="4" pageNumber="91">7–8 mm long, anthers pubescent</td>
<td id="76FF2C3DFFA50004FB89F8E1F198F8EB" box="[1035,1420,1854,1874]" gridcol="2" gridrow="8" pageId="4" pageNumber="91">5–6 mm long, anthers glabrous</td>
</tr>
<tr id="352E4541FFA50004FF12F8BBF198F8CE" box="[144,1420,1892,1911]" gridrow="9" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F8BBF5E6F8CE" box="[144,498,1892,1911]" gridcol="0" gridrow="9" pageId="4" pageNumber="91">Pistillate flowers</th>
<td id="76FF2C3DFFA50004FD9CF8BBF7E1F8CE" box="[542,1013,1892,1911]" gridcol="1" gridrow="9" pageId="4" pageNumber="91">Pedicellate</td>
<td id="76FF2C3DFFA50004FB89F8BBF198F8CE" box="[1035,1420,1892,1911]" gridcol="2" gridrow="9" pageId="4" pageNumber="91">Sessile</td>
</tr>
<tr id="352E4541FFA50004FF12F856F198F878" box="[144,1420,1929,1985]" gridrow="10" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F856F5E6F878" box="[144,498,1929,1985]" gridcol="0" gridrow="10" pageId="4" pageNumber="91">Pistillate sepals</th>
<td id="76FF2C3DFFA50004FD9CF856F7E1F878" box="[542,1013,1929,1985]" gridcol="1" gridrow="10" pageId="4" pageNumber="91">Subequal, lanceolate or narrowly triangular with margin flat, indumented on both surfaces</td>
<td id="76FF2C3DFFA50004FB89F856F198F878" box="[1035,1420,1929,1985]" gridcol="2" gridrow="10" pageId="4" pageNumber="91">Equal, elliptic or ovate with margin revolute, indumented on abaxial surface</td>
</tr>
<tr id="352E4541FFA50004FF12F807F198F855" box="[144,1420,2008,2028]" gridrow="11" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F807F5E6F855" box="[144,498,2008,2028]" gridcol="0" gridrow="11" pageId="4" pageNumber="91">Capsules</th>
<td id="76FF2C3DFFA50004FD9CF807F7E1F855" box="[542,1013,2008,2028]" gridcol="1" gridrow="11" pageId="4" pageNumber="91">4.9–5 × 3.9–4 mm, oblongoid</td>
<td id="76FF2C3DFFA50004FB89F807F198F855" box="[1035,1420,2008,2028]" gridcol="2" gridrow="11" pageId="4" pageNumber="91">5–6 × 5–6 mm, globose</td>
</tr>
<tr id="352E4541FFA50004FF12F7DCF198F7AE" box="[144,1420,2051,2071]" gridrow="12" pageId="4" pageNumber="91">
<th id="76FF2C3DFFA50004FF12F7DCF5E6F7AE" box="[144,498,2051,2071]" gridcol="0" gridrow="12" pageId="4" pageNumber="91">Seeds</th>
<td id="76FF2C3DFFA50004FD9CF7DCF7E1F7AE" box="[542,1013,2051,2071]" gridcol="1" gridrow="12" pageId="4" pageNumber="91">4–4.1 × 1.9–2 mm, oblongoid</td>
<td id="76FF2C3DFFA50004FB89F7DCF198F7AE" box="[1035,1420,2051,2071]" gridcol="2" gridrow="12" pageId="4" pageNumber="91">4–5 × 3–4 mm, ellipsoid</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA14703FFA4FFFEFF3FFF40F717FE61" blockId="5.[136,1452,158,581]" pageId="5" pageNumber="92">
<taxonomicName id="4C1E3C80FFA4FFFEFF3FFF40F5BBFF01" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[189,431,158,185]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA4FFFEFF3FFF40F5BBFF01" box="[189,431,158,185]" italics="true" pageId="5" pageNumber="92">Croton espinhacensis</emphasis>
</taxonomicName>
has been confused in herborized collections (see, e.g., 
<emphasis id="B96A9B11FFA4FFFEFB9CFF40F179FF01" box="[1054,1389,158,185]" italics="true" pageId="5" pageNumber="92">C.M. Sakuragui CFCR 15128</emphasis>
from SPF, K and R; 
<emphasis id="B96A9B11FFA4FFFEFEBBFF1CF63CFF65" box="[313,552,194,221]" italics="true" pageId="5" pageNumber="92">D.M.G. Oliveira 124</emphasis>
from ALCB, BHCB; and 
<emphasis id="B96A9B11FFA4FFFEFCD9FF1CF048FF65" box="[859,1116,194,221]" italics="true" pageId="5" pageNumber="92">V.C. Souza et al. 5527</emphasis>
from ESA and SPF) with 
<taxonomicName id="4C1E3C80FFA4FFFEFA10FF1CF4FBFEB9" authorityName="Muller Argoviensis" authorityYear="1873" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="species" species="atrorufus">
<emphasis id="B96A9B11FFA4FFFEFA10FF1CF4FBFEB9" italics="true" pageId="5" pageNumber="92">C. atrorufus</emphasis>
</taxonomicName>
, a species of 
<taxonomicName id="4C1E3C80FFA4FFFEFE13FF38F5F4FE9D" authority="(Klotzsch 1841: 193) Pax (1890: 39" authorityName="Pax" authorityYear="1858" baseAuthorityName="Klotzsch" baseAuthorityPageNumber="193" baseAuthorityYear="1841" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="subSection" section="Barhamia" subSection="Medea">
<emphasis id="B96A9B11FFA4FFFEFE13FF38F5CBFEB8" box="[401,479,231,257]" italics="true" pageId="5" pageNumber="92">Croton</emphasis>
sect. 
<emphasis id="B96A9B11FFA4FFFEFDA7FF39F680FEB9" box="[549,660,230,256]" italics="true" pageId="5" pageNumber="92">Barhamia</emphasis>
(
<bibRefCitation id="EF8F3AF2FFA4FFFEFD27FF39F79DFEB9" author="Klotzsch, J. F." box="[677,905,230,257]" pageId="5" pageNumber="92" pagination="99 - 106" refId="ref5111" refString="Klotzsch, J. F. (1853) Euphorbiaceae. In: Seemann, B. C. (Ed.) The botany of the voyage of H. M. S. Herald. Reeve &amp; Co., London, pp. 99 - 106." type="book chapter" year="1853">Klotzsch 1853: 104</bibRefCitation>
) 
<bibRefCitation id="EF8F3AF2FFA4FFFEFC18FF39F068FEB9" author="Baillon, H. E." box="[922,1148,230,257]" pageId="5" pageNumber="92" refId="ref4426" refString="Baillon, H. E. (1858) Etude generale du groupe des Euphorbiacees. Librairie de Victor Masson, Paris, 684 pp." type="book" year="1858">Baillon (1858: 367)</bibRefCitation>
subsect. 
<emphasis id="B96A9B11FFA4FFFEFB6BFF39F121FEB9" box="[1257,1333,230,256]" italics="true" pageId="5" pageNumber="92">Medea</emphasis>
(
<bibRefCitation id="EF8F3AF2FFA4FFFEFAC4FF39F4EEFE9D" author="Klotzsch, J. F." pageId="5" pageNumber="92" pagination="175 - 204" refId="ref5033" refString="Klotzsch, J. F. (1841) Neue und weniger gekannte sudamerikanische Euphorbiaceen-Gattungen. Archiv fur Naturgeschichte 7: 175 - 204." type="journal article" year="1841">Klotzsch 1841: 193</bibRefCitation>
) 
<bibRefCitation id="EF8F3AF2FFA4FFFEFE8BFED4F5BFFE9D" author="Pax, F." box="[265,427,266,293]" pageId="5" pageNumber="92" pagination="1 - 119" refId="ref5476" refString="Pax, F. (1890) Euphorbiaceae. In: Engler, A. &amp; Prantl, K. (Eds.) Die naturlichen Pflanzenfamilien 3 (5). Engelmann, Leipzig, pp. 1 - 119." type="book chapter" year="1890">Pax (1890: 39</bibRefCitation>
: 367
</taxonomicName>
) that also can be foud in de Espinhaço Montain Range. In fact, both species sahres the same habit, colors of the branches and leaves, in some case, tha shape of the leaves, and estaminate disc 5-segmented. However, 
<taxonomicName id="4C1E3C80FFA4FFFEFF79FE8CF597FED5" authorityName="Muller Argoviensis" authorityYear="1873" box="[251,387,338,365]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="species" species="atrorufus">
<emphasis id="B96A9B11FFA4FFFEFF79FE8CF597FED5" box="[251,387,338,365]" italics="true" pageId="5" pageNumber="92">C. atrorufus</emphasis>
</taxonomicName>
has stipules laciniate-glandulose (
<emphasis id="B96A9B11FFA4FFFEFC86FE8CF70FFED5" box="[772,795,339,364]" italics="true" pageId="5" pageNumber="92">vs</emphasis>
. entire in 
<taxonomicName id="4C1E3C80FFA4FFFEFC0EFE8CF05CFED5" authorityName="M. J. Silva &amp; Sodre" authorityYear="2024" box="[908,1096,338,365]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="species" species="espinhacensis">
<emphasis id="B96A9B11FFA4FFFEFC0EFE8CF05CFED5" box="[908,1096,338,365]" italics="true" pageId="5" pageNumber="92">C. espinhacensis</emphasis>
</taxonomicName>
), leaves without acropetiolar or basilaminar glands (
<emphasis id="B96A9B11FFA4FFFEFEE9FEA8F596FE29" box="[363,386,375,400]" italics="true" pageId="5" pageNumber="92">vs</emphasis>
. with a pair of basilaminar glands), pistillate sepals with margin laciniate-glandulose (
<emphasis id="B96A9B11FFA4FFFEFAC5FEA8F14AFE29" box="[1351,1374,375,400]" italics="true" pageId="5" pageNumber="92">vs</emphasis>
. entire and without glands), anthers glabrous (
<emphasis id="B96A9B11FFA4FFFEFDBFFE44F640FE0D" box="[573,596,411,436]" italics="true" pageId="5" pageNumber="92">vs</emphasis>
. anthers indumented), styles 4 or 5-fids (
<emphasis id="B96A9B11FFA4FFFEFBA3FE44F02CFE0D" box="[1057,1080,411,436]" italics="true" pageId="5" pageNumber="92">vs</emphasis>
. 2-fids), as well as seeds broadly elliptical with a rugose testa (
<emphasis id="B96A9B11FFA4FFFEFE51FE60F5FEFE61" box="[467,490,447,472]" italics="true" pageId="5" pageNumber="92">vs</emphasis>
. oblongoid and smooth).
</paragraph>
<paragraph id="8BA14703FFA4FFFEFF3FFE3DF041FDFC" blockId="5.[136,1452,158,581]" pageId="5" pageNumber="92">
With the description of this new species, 
<taxonomicName id="4C1E3C80FFA4FFFEFD0DFE3CF06EFE45" authority="Sodre &amp; Secco (2023: 134)" authorityName="Sodre &amp; Secco" authorityPageNumber="134" authorityYear="2023" box="[655,1146,482,509]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="species" species="carajasensis">
<emphasis id="B96A9B11FFA4FFFEFD0DFE3CF729FE45" box="[655,829,483,509]" italics="true" pageId="5" pageNumber="92">C. carajasensis</emphasis>
<bibRefCitation id="EF8F3AF2FFA4FFFEFCC6FE3DF06EFE45" author="Sodre, R. C. &amp; Secco, R. S." box="[836,1146,482,508]" pageId="5" pageNumber="92" pagination="133 - 141" refId="ref6446" refString="Sodre, R. C. &amp; Secco, R. S. (2023) Croton carajasensis, a new species from Serra dos Carajas, Para, Brazilian Amazon. Phytotaxa 630: 133 - 141. https: // doi. org / 10.11646 / phytotaxa. 630.2.4" type="journal article" year="2023">Sodré &amp; Secco (2023: 134)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C1E3C80FFA4FFFEFB30FE3CF53EFD99" authority="M. J. Silva &amp; Sodre" authorityName="M. J. Silva &amp; Sodre" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="species" species="piriquetifolius">
<emphasis id="B96A9B11FFA4FFFEFB30FE3CF166FE45" box="[1202,1394,482,509]" italics="true" pageId="5" pageNumber="92">C. piriquetifolius</emphasis>
M.J. Silva &amp; Sodré
</taxonomicName>
(
<emphasis id="B96A9B11FFA4FFFEFEBEFDD8F546FD99" box="[316,338,519,544]" italics="true" pageId="5" pageNumber="92">in</emphasis>
<bibRefCitation id="EF8F3AF2FFA4FFFEFED9FDD9F641FD99" author="Silva, M. J. &amp; Sodre, R. C. &amp; Alonso, A. A. &amp; Santos, I. S." box="[347,597,518,544]" pageId="5" pageNumber="92" pagination="143 - 156" refId="ref6108" refString="Silva, M. J., Sodre, R. C., Alonso, A. A. &amp; Santos, I. S. (2024) A new species of Croton sect. Geiseleria (Euphorbiaceae) from Brazil, supported by macro- and micromorphological data. Phytotaxa 646: 143 - 156. https: // doi. org / 10.11646 / phytotaxa. 646.2.4" type="journal article" year="2024">
Silva 
<emphasis id="B96A9B11FFA4FFFEFE1FFDD8F5C3FD99" box="[413,471,518,544]" italics="true" pageId="5" pageNumber="92">et al.</emphasis>
2024: 144
</bibRefCitation>
), 
<taxonomicName id="4C1E3C80FFA4FFFEFDE9FDD8F767FD99" box="[619,883,518,545]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="92" phylum="Tracheophyta" rank="section" section="Geiseleria">
<emphasis id="B96A9B11FFA4FFFEFDE9FDD8F6ADFD98" box="[619,697,519,545]" italics="true" pageId="5" pageNumber="92">Croton</emphasis>
sect. 
<emphasis id="B96A9B11FFA4FFFEFD7CFDD9F767FD99" box="[766,883,518,544]" italics="true" pageId="5" pageNumber="92">Geiseleria</emphasis>
</taxonomicName>
is now represented in 
<collectingCountry id="F3090793FFA4FFFEFBFAFDD9F0A8FD99" box="[1144,1212,518,544]" name="Brazil" pageId="5" pageNumber="92">Brazil</collectingCountry>
by 57 species, 39 of which are endemic, which reinforces that 
<collectingCountry id="F3090793FFA4FFFEFDDFFDF5F6B5FDFD" box="[605,673,554,580]" name="Brazil" pageId="5" pageNumber="92">Brazil</collectingCountry>
is the center of diversity of this group.
</paragraph>
</subSubSection>
</treatment>
</document>