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<document id="E4BEFA62E7104827FF1843F42D768796" ID-CLB-Dataset="20773" ID-DOI="10.1111/syen.12522" ID-GBIF-Dataset="564eaad8-6245-4f42-9a88-42b97bf30807" ID-Zenodo-Dep="5975346" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1644003985773" checkinUser="tatiana" docAuthor="Boudinot, Brendon E., Borowiec, Marek L. &amp; Prebus, Matthew M." docDate="2022" docId="03B987A9436FFF9CFCA4FA32FAAED8CA" docLanguage="en" docName="systEnt.47.113-151.pdf.imf" docOrigin="Systematic Entomology 47" docTitle="Sussudio Boudinot &amp; Borowiec &amp; Prebus 2022, gen.nov." docType="treatment" docUuid="BFF515E7-9966-443F-8A34-555A4B41906B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="143" masterDocId="FF80FFD14372FF82FFF5FF95FF99D837" masterDocTitle="Phylogeny, evolution, and classification of the ant genus Lasius, the tribe Lasiini and the subfamily Formicinae (Hymenoptera: Formicidae)" masterLastPageNumber="151" masterPageNumber="113" pageNumber="142" updateTime="1730916979833" updateUser="juliana">
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<mods:title id="C774C8015AA8ABE2C22BFE3CCCB21AF5">Phylogeny, evolution, and classification of the ant genus Lasius, the tribe Lasiini and the subfamily Formicinae (Hymenoptera: Formicidae)</mods:title>
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<mods:name id="654D87FCE7FCE4A8621B2038FB42FFE6" type="personal">
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<mods:namePart id="BC256852A38BEE72A34FFCB3E3969AD8">Boudinot, Brendon E.</mods:namePart>
<mods:affiliation id="34FEE7458AEDE9B7104D7D762D861661">Department of Entomology and Nematology, University of California, Davis CA, U. S. A. &amp; Friedrich-Schiller-Universität Jena, Institut für Spezielle Zoologie, Jena, Germany</mods:affiliation>
</mods:name>
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<mods:namePart id="E5108E3F576AC5201E059AE215FA361D">Borowiec, Marek L.</mods:namePart>
<mods:affiliation id="B24228024B4A1CBB0554DE61AE1E349B">Department of Entomology and Nematology, University of California, Davis CA, U. S. A. &amp; Department of Plant Pathology, Entomology and Nematology, University of Idaho, Moscow ID, U. S. A. &amp; Institute for Bioinformatics and Evolutionary Studies, University of Idaho, Moscow ID, U. S. A.</mods:affiliation>
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<mods:namePart id="7749F1EB6D8B55EA01C96EAE0559A527">Prebus, Matthew M.</mods:namePart>
<mods:affiliation id="327D3B3E066E8F40464C46655E0D6DD9">Department of Entomology and Nematology, University of California, Davis CA, U. S. A. &amp; School of Life Sciences, Arizona State University, Tempe AZ, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="BB6243F835D47B2C747911BFCA7CC33E" type="email">mprebus@gmail.com</mods:nameIdentifier>
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<mods:title id="9B5BD7DD0AD95F8A840DE0BE7962A124">Systematic Entomology</mods:title>
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<mods:part id="2293B3284C2C67033508B2C1D46E0FEA">
<mods:date id="5015E4884D428E4FDA464450D1F734A8">2022</mods:date>
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<mods:number id="3E6818B4F189E6B255B5083C40902C77">2022-01-10</mods:number>
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<treatment id="03B987A9436FFF9CFCA4FA32FAAED8CA" ID-DOI="http://doi.org/10.5281/zenodo.6303238" ID-GBIF-Taxon="193278222" ID-Zenodo-Dep="6303238" LSID="urn:lsid:zoobank.org:act:BFF515E7-9966-443F-8A34-555A4B41906B" httpUri="http://treatment.plazi.org/id/03B987A9436FFF9CFCA4FA32FAAED8CA" lastPageId="30" lastPageNumber="143" pageId="29" pageNumber="142">
<subSubSection id="C30A6534436FFF9FFCA4FA32FBCFDD8A" box="[849,1110,1447,1469]" pageId="29" pageNumber="142" type="nomenclature">
<paragraph id="8BAF36BF436FFF9FFCA4FA32FBCFDD8A" blockId="29.[830,1437,276,1618]" box="[849,1110,1447,1469]" pageId="29" pageNumber="142">
<heading id="D0E781D3436FFF9FFCA4FA32FBCBDD8A" box="[849,1106,1447,1469]" pageId="29" pageNumber="142">
<emphasis id="B964EAAD436FFF9FFCA4FA32FBCFDD8A" bold="true" box="[849,1110,1447,1469]" pageId="29" pageNumber="142">
Genus †
<taxonomicName id="4C104D3C436FFF9FFC51FA32FC64DD8B" authority="Boudinot &amp; Borowiec &amp; Prebus, 2022" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[932,1021,1447,1468]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="142" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B964EAAD436FFF9FFC51FA32FC64DD8B" bold="true" box="[932,1021,1447,1468]" italics="true" pageId="29" pageNumber="142">Sussudio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757D6436FFF9FFBF6FA32FBCFDD8A" box="[1027,1110,1447,1469]" pageId="29" pageNumber="142" rank="genus">gen.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6534436FFF9FFCA4FA50FC2DDDCF" pageId="29" pageNumber="142" type="type_taxon">
<paragraph id="8BAF36BF436FFF9FFCA4FA50FC2DDDCF" blockId="29.[830,1437,276,1618]" pageId="29" pageNumber="142">
<emphasis id="B964EAAD436FFF9FFCA4FA50FBDFDDED" box="[849,1094,1477,1498]" italics="true" pageId="29" pageNumber="142">Type and included species</emphasis>
. †
<taxonomicName id="4C104D3C436FFF9FFBA8FA50FADCDDED" authority="Wheeler, 1915" authorityName="Wheeler" authorityYear="1915" box="[1117,1349,1477,1498]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="142" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B964EAAD436FFF9FFBA8FA50FB2ADDED" box="[1117,1203,1477,1498]" italics="true" pageId="29" pageNumber="142">P. boreus</emphasis>
<bibRefCitation id="EF814B4E436FFF9FFB42FA50FADCDDED" author="Wheeler, W. M." box="[1207,1349,1477,1498]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="29" pageNumber="142" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
</taxonomicName>
. Original designation.
</paragraph>
</subSubSection>
<subSubSection id="C30A6534436FFF9CFCA4F994FD76DA72" lastPageId="30" lastPageNumber="143" pageId="29" pageNumber="142" type="description">
<paragraph id="8BAF36BF436FFF9FFCA4F994FB66DE03" blockId="29.[830,1437,276,1618]" pageId="29" pageNumber="142">
<emphasis id="B964EAAD436FFF9FFCA4F994FC7DDE21" box="[849,996,1537,1558]" italics="true" pageId="29" pageNumber="142">ZooBank LSID</emphasis>
. http://zoobank.org/urn:lsid:zoobank.org:act: BFF515E7-9966-443F-8A34-555A4B41906B
</paragraph>
<paragraph id="8BAF36BF436FFF9FFCA4F9A8FB89DE65" blockId="29.[830,1437,276,1618]" box="[849,1040,1597,1618]" pageId="29" pageNumber="142">
<emphasis id="B964EAAD436FFF9FFCA4F9A8FB93DE65" box="[849,1034,1597,1618]" italics="true" pageId="29" pageNumber="142">Definition (worker)</emphasis>
.
</paragraph>
<paragraph id="8BAF36BF436FFF9FFCB9F9E8FA92DEA5" blockId="29.[844,1437,1661,1863]" box="[844,1291,1661,1682]" pageId="29" pageNumber="142">
1. With characters of the
<taxonomicName id="4C104D3C436FFF9FFBB7F9E8FB36DEA5" baseAuthorityName="Lepeletier de Saint-Fargeau" box="[1090,1199,1661,1682]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
(Note 1).
</paragraph>
<paragraph id="8BAF36BF436FFF9FFCB9F90EFB7BDEF8" blockId="29.[844,1437,1661,1863]" pageId="29" pageNumber="142">
2.
<emphasis id="B964EAAD436FFF9FFC9CF909FBDFDE86" box="[873,1094,1692,1713]" italics="true" pageId="29" pageNumber="142">Cranium size variable</emphasis>
,
<emphasis id="B964EAAD436FFF9FFBA3F909FB1FDEF8" italics="true" pageId="29" pageNumber="142">width somewhat broader than to about twice that of mesosoma</emphasis>
(Note 2).
</paragraph>
<paragraph id="8BAF36BF436FFF9FFCB9F94DFB5EDEDA" blockId="29.[844,1437,1661,1863]" box="[844,1223,1752,1773]" pageId="29" pageNumber="142">3. Mandible triangular, with 78 teeth.</paragraph>
<paragraph id="8BAF36BF436FFF9FFCB9F963FBC7DF1E" blockId="29.[844,1437,1661,1863]" pageId="29" pageNumber="142">4. Dorsal mandibular groove extending along lateral mandibular margin in dorsal view.</paragraph>
<paragraph id="8BAF36BF436FFF9FFCB9F8A7FA04DF70" blockId="29.[844,1437,1661,1863]" box="[844,1437,1842,1863]" pageId="29" pageNumber="142">5. Palps reduced, not reaching midlength of postgenal bridge.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF50FF39FDF1D8F6" blockId="30.[153,758,172,581]" box="[165,616,172,193]" pageId="30" pageNumber="143">6. Frontal carinae raised above antennal toruli.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF50FF5FFD07D8E8" blockId="30.[153,758,172,581]" box="[165,670,202,223]" pageId="30" pageNumber="143">7. Antennal toruli abutting posterior clypeal margin.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF50FF7DFDA2D8CA" blockId="30.[153,758,172,581]" box="[165,571,232,253]" pageId="30" pageNumber="143">8. Antennomere count unknown (Note 3).</paragraph>
<paragraph id="8BAF36BF436CFF9CFF50FE93FD81D92C" blockId="30.[153,758,172,581]" box="[165,536,262,283]" pageId="30" pageNumber="143">9. Antennomere III longer than broad.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFEB6FD5BD90F" blockId="30.[153,758,172,581]" box="[153,706,291,312]" pageId="30" pageNumber="143">10. Compound eyes situated posterior to head midlength.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFED4FD7CD961" blockId="30.[153,758,172,581]" box="[153,741,321,342]" pageId="30" pageNumber="143">11. Compound eyes not enormous and long axes subparallel.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFECAFD9ED943" blockId="30.[153,758,172,581]" box="[153,519,351,372]" pageId="30" pageNumber="143">12. Ocelli minutely present or absent.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFEE8FD5ED9A5" blockId="30.[153,758,172,581]" box="[153,711,381,402]" pageId="30" pageNumber="143">13. Promesonotum domed, raised well above propodeum.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFE0EFD92D987" blockId="30.[153,758,172,581]" box="[153,523,411,432]" pageId="30" pageNumber="143">14. Metanotal groove well-developed.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFE2CFDD5D9DC" blockId="30.[153,758,172,581]" pageId="30" pageNumber="143">15. Petiolar foramen in profile view clearly raised and margined with conspicuous thickened carina or lip.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFE61FDEDDA10" blockId="30.[153,758,172,581]" pageId="30" pageNumber="143">16. Petiolar node squamiform, tall; dorsoventral height equal to or possibly exceeding dorsum of propodeum.</paragraph>
<paragraph id="8BAF36BF436CFF9CFF6CFDA5FD76DA72" blockId="30.[153,758,172,581]" box="[153,751,560,581]" pageId="30" pageNumber="143">17. Abdominal segment III without raised tergosternal fusion.</paragraph>
</subSubSection>
<subSubSection id="C30A6534436CFF9CFF62FDFFFE7BDC4C" pageId="30" pageNumber="143" type="discussion">
<paragraph id="8BAF36BF436CFF9CFF62FDFFFEC7DAB7" blockId="30.[151,759,618,1863]" box="[151,350,618,640]" pageId="30" pageNumber="143">
<emphasis id="B964EAAD436CFF9CFF62FDFFFECEDAB7" bold="true" box="[151,343,618,640]" pageId="30" pageNumber="143">Notes on definition</emphasis>
:
</paragraph>
<paragraph id="8BAF36BF436CFF9CFF62FD1CFDF7DB59" blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
<emphasis id="B964EAAD436CFF9CFF62FD1CFF7CDAA9" box="[151,229,649,670]" italics="true" pageId="30" pageNumber="143">Note 1.</emphasis>
Several characters were not possible to evaluate. These include the following: Reduction of third tooth from mandibular apex, palp formula, the metapleural gland, ventral length of the petiolar foramen, relative separation of the meso- and metacoxae, presence or absence of setal double-row on ventral tibial surfaces, petiolar apodeme conformation, cross-sectional shape of the petiolar sternum, and presence or absence of the posthelcial sulcus.
</paragraph>
<paragraph id="8BAF36BF436CFF9CFF62FCE2FE12DC16" blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
<emphasis id="B964EAAD436CFF9CFF62FCE2FF47DBBB" box="[151,222,887,908]" italics="true" pageId="30" pageNumber="143">Note 2.</emphasis>
The massive head of †
<taxonomicName id="4C104D3C436CFF9CFE35FCE2FD81DBBB" authorityName="Wheeler" authorityYear="1915" box="[448,536,887,908]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B964EAAD436CFF9CFE35FCE2FD81DBBB" box="[448,536,887,908]" italics="true" pageId="30" pageNumber="143">P. boreus</emphasis>
</taxonomicName>
led
<bibRefCitation id="EF814B4E436CFF9CFDB1FCE2FD45DBBB" author="Wheeler, W. M." box="[580,732,887,908]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="30" pageNumber="143" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler (1915)</bibRefCitation>
to erroneously assume that the specimens were majors of
<taxonomicName id="4C104D3C436CFF9CFD4AFC00FE92DBFF" authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFD4AFC00FE92DBFF" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
</taxonomicName>
; Wheeler did note the posteriorly set eyes of †
<taxonomicName id="4C104D3C436CFF9CFD11FC26FF62DBD1" authorityName="Wheeler" authorityYear="1915" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B964EAAD436CFF9CFD11FC26FF62DBD1" italics="true" pageId="30" pageNumber="143">P. boreus</emphasis>
</taxonomicName>
, one of the distinctions between it and
<taxonomicName id="4C104D3C436CFF9CFD84FC44FD76DBD1" authorityName="Emery" authorityYear="1887" box="[625,751,977,998]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFD84FC44FD76DBD1" box="[625,751,977,998]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
</taxonomicName>
, but vacillated based on the insufficient knowledge of the latter genus at that time.
</paragraph>
<paragraph id="8BAF36BF436CFF9CFF62FBBFFE7BDC4C" blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
<emphasis id="B964EAAD436CFF9CFF62FBBFFF7BDC08" box="[151,226,1066,1087]" italics="true" pageId="30" pageNumber="143">Note 3.</emphasis>
The examined specimens are missing their terminal antennomeres, and no previous descriptions of †
<taxonomicName id="4C104D3C436CFF9CFD6EFBDDFD6CDC6A" authorityName="Wheeler" authorityYear="1915" box="[667,757,1096,1117]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B964EAAD436CFF9CFD6EFBDDFD6CDC6A" box="[667,757,1096,1117]" italics="true" pageId="30" pageNumber="143">P. boreus</emphasis>
</taxonomicName>
include an antennomere count.
</paragraph>
</subSubSection>
<subSubSection id="C30A6534436CFF9CFF5FFB11FDE9DC81" pageId="30" pageNumber="143" type="etymology">
<paragraph id="8BAF36BF436CFF9CFF5FFB11FDE9DC81" blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
<emphasis id="B964EAAD436CFF9CFF5FFB11FE89DCAE" box="[170,272,1156,1177]" italics="true" pageId="30" pageNumber="143">Etymology</emphasis>
. Asimplification of pseudo-
<taxonomicName id="4C104D3C436CFF9CFD9AFB11FD74DCAE" authorityName="Emery" authorityYear="1887" box="[623,749,1156,1177]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFD9AFB11FD74DCAE" box="[623,749,1156,1177]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
</taxonomicName>
inspired by the musician Phil Collins. Masculine.
</paragraph>
</subSubSection>
<subSubSection id="C30A6534436CFF9CFF5FFB55FAAED8CA" pageId="30" pageNumber="143" type="discussion">
<paragraph id="8BAF36BF436CFF9CFF5FFB55FAAED8CA" blockId="30.[151,759,618,1863]" lastBlockId="30.[821,1427,172,253]" pageId="30" pageNumber="143">
<emphasis id="B964EAAD436CFF9CFF5FFB55FE96DCE2" box="[170,271,1216,1237]" italics="true" pageId="30" pageNumber="143">Comments</emphasis>
. †
<taxonomicName id="4C104D3C436CFF9CFEC4FB55FE4FDCE2" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[305,470,1216,1237]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus" status="comb. nov.">
<emphasis id="B964EAAD436CFF9CFEC4FB55FE4FDCE2" box="[305,470,1216,1237]" italics="true" pageId="30" pageNumber="143">Sussudio boreus</emphasis>
</taxonomicName>
<emphasis id="B964EAAD436CFF9CFE13FB2AFDD5DCE2" bold="true" box="[486,588,1215,1237]" pageId="30" pageNumber="143">
<taxonomicNameLabel id="A25757D6436CFF9CFE13FB2AFDD5DCE2" box="[486,588,1215,1237]" pageId="30" pageNumber="143" rank="species">comb.nov.</taxonomicNameLabel>
</emphasis>
was previously placed in the
<taxonomicName id="4C104D3C436CFF9CFEE3FB48FEC1DCC5" authorityName="Ashmead" authorityYear="1905" box="[278,344,1245,1266]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="tribe" tribe="Lasiini">Lasiini</taxonomicName>
(
<bibRefCitation id="EF814B4E436CFF9CFE91FB48FE78DCC5" author="Emery, C." box="[356,481,1245,1266]" journalOrPublisher="Subfam. Formicinae. Genera Insectorum" pageId="30" pageNumber="143" pagination="1 - 302" part="183" refId="ref33484" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." title="Hymenoptera. Fam. Formicidae" type="journal article" year="1925">Emery, 1925</bibRefCitation>
,
<bibRefCitation id="EF814B4E436CFF9CFE1EFB48FD68DCC5" author="Dlussky, G. M. &amp; Fedoseeva, E. B." bookContentInfo="232 pp." box="[491,753,1245,1266]" editor="Ponomarenko, A. G." journalOrPublisher="Moskva: Nauka" pageId="30" pageNumber="143" pagination="70 - 144" refId="ref33007" refString="Dlussky, G. M. &amp; Fedoseeva, E. B. (1988) Origin and early stages of evolution in ants [in Russian]. Pp. 70 - 144 in: Ponomarenko, A. G. (ed.) Cretaceous Biocenotic Crisis and Insect Evolution. [in Russian.]. Moskva: Nauka, 232 pp." title="Origin and early stages of evolution in ants" type="book chapter" volumeTitle="Cretaceous Biocenotic Crisis and Insect Evolution" year="1988">Dlussky &amp; Fedoseeva 1988</bibRefCitation>
,
<bibRefCitation id="EF814B4E436CFF9CFF62FB6EFE80DD27" author="Bolton, B." bookContentInfo="p. 222" box="[151,281,1275,1296]" journalOrPublisher="Harvard University Press, Cambridge" pageId="30" pageNumber="143" refId="ref31447" refString="Bolton, B. (1994) Identification Guide to the Ant Genera of the World, p. 222. Harvard University Press, Cambridge." title="Identification Guide to the Ant Genera of the World" type="book" year="1994">Bolton, 1994</bibRefCitation>
), and has been considered to be a member of the genus
<taxonomicName id="4C104D3C436CFF9CFF20FA8CFECADD19" authorityName="Emery" authorityYear="1887" box="[213,339,1305,1326]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFF20FA8CFECADD19" box="[213,339,1305,1326]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF814B4E436CFF9CFEAAFA8CFE76DD19" author="Wheeler, W. M." box="[351,495,1305,1326]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="30" pageNumber="143" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
;
<bibRefCitation id="EF814B4E436CFF9CFE0DFA8CFD72DD19" author="LaPolla, J. S. &amp; Dlussky, G. M." box="[504,747,1305,1326]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="30" pageNumber="143" pagination="258 - 273" part="112" refId="ref35305" refString="LaPolla, J. S. &amp; Dlussky, G. M. (2010) Review of fossil Prenolepis genus-group species (Hymenoptera: Formicidae). Proceedings of the Entomological Society of Washington, 112, 258 - 273." title="Review of fossil Prenolepis genus-group species (Hymenoptera: Formicidae)" type="journal article" year="2010">LaPolla &amp; Dlussky, 2010</bibRefCitation>
). Based on an examination of a syntype (GZG-BST04646) and a nontype (NHMW1984-31-211) worker, as well as the original description from
<bibRefCitation id="EF814B4E436CFF9CFEB6FAE7FE46DDB0" author="Wheeler, W. M." box="[323,479,1394,1415]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="30" pageNumber="143" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler (1915)</bibRefCitation>
, we reject both placements. Neither specimen can be mistaken due to the unique combination of characters they display, including setation, high petiolar nodes, massive crania, and mesosomal form. †
<taxonomicName id="4C104D3C436CFF9CFDACFA59FD6CDDD6" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[601,757,1484,1505]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B964EAAD436CFF9CFDACFA59FD6CDDD6" box="[601,757,1484,1505]" italics="true" pageId="30" pageNumber="143">Sussudio boreus</emphasis>
</taxonomicName>
does not display the modifications of the third abdominal segment characteristic of the
<taxonomicName id="4C104D3C436CFF9CFE3EF99DFDABDE2A" box="[459,562,1544,1565]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFE3EF99DFDABDE2A" box="[459,562,1544,1565]" italics="true" pageId="30" pageNumber="143">Prenolepis</emphasis>
</taxonomicName>
genus group, also rejecting its original placement in
<taxonomicName id="4C104D3C436CFF9CFE14F9B0FDC6DE0D" authorityName="Emery" authorityYear="1887" box="[481,607,1573,1594]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFE14F9B0FDC6DE0D" box="[481,607,1573,1594]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
</taxonomicName>
(see diagnostic Note 2 above). †
<taxonomicName id="4C104D3C436CFF9CFEC9F9D6FE08DE6F" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[316,401,1603,1624]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFEC9F9D6FE08DE6F" box="[316,401,1603,1624]" italics="true" pageId="30" pageNumber="143">Sussudio</emphasis>
</taxonomicName>
is excluded from the
<taxonomicName id="4C104D3C436CFF9CFD9CF9D6FD32DE6F" authorityName="Ashmead" authorityYear="1905" box="[617,683,1603,1624]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="tribe" tribe="Lasiini">Lasiini</taxonomicName>
overall by two additional characters: (i) The petiolar node, which is completely vertical and very tall dorsoventrally, is as tall as or possibly taller than the propodeum; and (ii) the raised and conspicuously lipped petiolar foramen. No lasiine, extant or extinct, has such a tall petiolar node, nor do any species display the petiolar foramen conformation observed in †
<taxonomicName id="4C104D3C436CFF9CFD86F963FD51DF3C" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[627,712,1782,1803]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFD86F963FD51DF3C" box="[627,712,1782,1803]" italics="true" pageId="30" pageNumber="143">Sussudio</emphasis>
</taxonomicName>
. We cannot confidently place †
<taxonomicName id="4C104D3C436CFF9CFE69F881FE68DF1E" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[412,497,1812,1833]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAD436CFF9CFE69F881FE68DF1E" box="[412,497,1812,1833]" italics="true" pageId="30" pageNumber="143">Sussudio</emphasis>
</taxonomicName>
in any extant tribe, therefore, we consider †
<emphasis id="B964EAAD436CFF9CFEA3F8A7FDA3DF70" box="[342,570,1842,1863]" italics="true" pageId="30" pageNumber="143">
<taxonomicName id="4C104D3C436CFF9CFEA3F8A7FE32DF70" authority="Boudinot &amp; Borowiec &amp; Prebus, 2022" authorityName="Boudinot &amp; Borowiec &amp; Prebus" authorityYear="2022" box="[342,427,1842,1863]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus" status="incertae sedis">Sussudio</taxonomicName>
<taxonomicNameLabel id="A25757D6436CFF9CFE46F8A7FDA3DF70" box="[435,570,1842,1863]" pageId="30" pageNumber="143">incertae sedis</taxonomicNameLabel>
</emphasis>
in the
<taxonomicName id="4C104D3C436CFF9CFD76F8A7FD6ADF70" baseAuthorityName="Lepeletier de Saint-Fargeau" box="[643,755,1842,1863]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
. As with other fossil taxa, which are of uncertain placement in the
<taxonomicName id="4C104D3C436CFF9CFCAFFF5FFC50D8E8" baseAuthorityName="Lepeletier de Saint-Fargeau" box="[858,969,202,223]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
, micro-CT scanning could be used to evaluate the form of the helcium and third abdominal sternum.
</paragraph>
</subSubSection>
</treatment>
</document>