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<document id="3175F397005300A49E0ED65C57FFE278" ID-DOI="10.11646/zootaxa.5523.4.5" ID-GBIF-Dataset="c8fd3aa9-6203-4d1b-ae7b-367bf1d498b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13949669" ID-ZooBank="999559E0-4E12-45DC-AED9-88374AA3E3D1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729236181491" checkinUser="plazi" docAuthor="Kuwahara, Gregory K., Marshall, Stephen A. &amp; Yau, Tiffany" docDate="2024" docId="03A1976C12693A7773CA6CEAFA3DF2B6" docLanguage="en" docName="Zootaxa.5523.4.5.pdf" docOrigin="Zootaxa 5523 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helicosina Kuwahara &amp; Marshall &amp; Yau 2024" docType="key" docVersion="8" lastPageNumber="464" masterDocId="FF98EF14126C3A72735D6E59FFA1F664" masterDocTitle="Helicosina, a new genus of Neotropical Limosininae (Diptera: Sphaeroceridae)" masterLastPageNumber="471" masterPageNumber="459" pageNumber="464" updateTime="1729684655948" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7B0EB6F3F2A9256C3CE68AA4EE886218" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C5D0E0D1C0A76E26A27A0B998923DE4F">
<mods:title id="353FFE308C67CDC3D8AE5A0147741EB7">Helicosina, a new genus of Neotropical Limosininae (Diptera: Sphaeroceridae)</mods:title>
</mods:titleInfo>
<mods:name id="B5A6B0740982B9FAC7956DE37795342E" type="personal">
<mods:role id="8DEDED04AC2492BAE3F5F74A274E98DC">
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<mods:namePart id="17BD81302E63563CA1085ED92D4C92D7">Kuwahara, Gregory K.</mods:namePart>
<mods:nameIdentifier id="8F544E9AC2CA92A57F4E0FF85DDF3D2B" type="email">gkuwahar@uoguelph.ca</mods:nameIdentifier>
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<mods:namePart id="871E771082CC5DCD989743F518D95C1A">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="2443D77C82EDD350EAEE0E6E137EEBD0" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="3392254A4CED2C1326473E83D8E19670">Yau, Tiffany</mods:namePart>
<mods:nameIdentifier id="6AE693D46674D4F624EFAB55BE4B0D4D" type="email">tyau@alumni.uoguelph.ca</mods:nameIdentifier>
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<mods:title id="DC6B08DE1F06B89FDC20FA202AF7FCF7">Zootaxa</mods:title>
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<mods:part id="E4223F9BDE74EF4FF3295D060752CE6C">
<mods:date id="ACB15C612BD473435333E228729C000A">2024</mods:date>
<mods:detail id="103A44B7A74E030350F550FDC8207F31" type="pubDate">
<mods:number id="759436D9EE8A42646B995502009B76F8">2024-10-17</mods:number>
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<mods:detail id="B24DB0BC26178C4ADBDB9F82CD6D7B8F" type="volume">
<mods:number id="B0C9FA4B46CA738975A57179D763F544">5523</mods:number>
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<mods:number id="4CFCEDBCC9813069E757221BDED99DDB">4</mods:number>
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<mods:url id="133A3E4CF7123DAA392A5747773A2C72">http://dx.doi.org/10.11646/zootaxa.5523.4.5</mods:url>
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<mods:classification id="FE78FCF70CB45E829F8EC3ECDAE885F4">journal article</mods:classification>
<mods:identifier id="6C04BEB67D3875FF341AAF105475EC64" type="DOI">10.11646/zootaxa.5523.4.5</mods:identifier>
<mods:identifier id="EF36B0FD248EA9E6E21964EA3AD635EA" type="GBIF-Dataset">c8fd3aa9-6203-4d1b-ae7b-367bf1d498b6</mods:identifier>
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<treatment id="03A1976C12693A7773CA6CEAFA3DF2B6" ID-DOI="http://doi.org/10.5281/zenodo.13974001" ID-GBIF-Taxon="244134450" ID-Zenodo-Dep="13974001" LSID="urn:lsid:plazi:treatment:03A1976C12693A7773CA6CEAFA3DF2B6" httpUri="http://treatment.plazi.org/id/03A1976C12693A7773CA6CEAFA3DF2B6" lastPageNumber="464" pageId="5" pageNumber="464">
<subSubSection id="C31275F112693A7773CA6CEAFA35F571" pageId="5" pageNumber="464" type="nomenclature">
<paragraph id="8BB7267A12693A7773CA6CEAFDBEF4A9" blockId="5.[151,543,690,717]" box="[151,543,690,717]" pageId="5" pageNumber="464">
<heading id="D0FF911612693A7773CA6CEAFDBEF4A9" bold="true" box="[151,543,690,717]" fontSize="11" level="1" pageId="5" pageNumber="464" reason="1">
<emphasis id="B97CFA6812693A7773CA6CEAFDBEF4A9" bold="true" box="[151,543,690,717]" pageId="5" pageNumber="464">
Key to the species of
<taxonomicName id="4C085DF912693A7772C06CEBFDBEF4A9" authorityName="Kuwahara &amp; Marshall &amp; Yau" authorityYear="2024" box="[413,543,690,717]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B97CFA6812693A7772C06CEBFDBEF4A9" bold="true" box="[413,543,690,717]" italics="true" pageId="5" pageNumber="464">Helicosina</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB7267A12693A7773CA6CA2FA35F571" blockId="5.[151,1428,763,789]" box="[151,1428,763,789]" pageId="5" pageNumber="464">This key is based almost entirely on male characters, as the females are almost indistinguishable morphologically.</paragraph>
</subSubSection>
<subSubSection id="C31275F112693A7773CA6D19FA3DF2B6" pageId="5" pageNumber="464" type="key">
<keyStep id="30FC3EDF12693A7773CA6D19FA3DF583" pageId="5" pageNumber="464">
<paragraph id="8BB7267A12693A7773CA6D19FA3DF5EB" blockId="5.[151,1437,831,1235]" pageId="5" pageNumber="464">
<keyLead id="30F9854F12693A7773CA6D19FA3DF5EB" pageId="5" pageNumber="464">
1 Body length ≤
<quantity id="4CF08B9F12693A7772326D19FE1AF531" box="[367,443,832,853]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="464" unit="mm" value="1.8">1.8 mm</quantity>
. Male fore and hind femora slightly thickened (length 1.9–2.0 × and 4.4–5.6 × width, respectively) (
<figureCitation id="13333AFF12693A7776286D19FEA2F516" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="5" pageNumber="464">Fig. 2A</figureCitation>
). Surstylus straight along anterior margin (
<figureCitation id="13333AFF12693A7771FA6D04FD54F516" box="[679,757,861,883]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="5" pageNumber="464">Fig. 4C</figureCitation>
). Posterior opening of epandrium roughly triangular, widest ventrally (
<figureCitation id="13333AFF12693A7773B46D23FE95F5EB" box="[233,308,890,912]" captionStart="FIGURE 4" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-29@6.[184,1404,181,1884]" captionTargetPageId="6" captionText="FIGURE 4. Helicosina crena sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral. B, aedeagus and associated structures, left dorsolateral. C, S5–8 and postabdomen, lateroventral. D, epandrium and surstyli, posterior. Abbreviations: bp— basiphallus, cer—cercus, dp—distiphallus, ea—ejaculatory apodeme, epa—epandrium, hyp—hypandrium, pg—postgonite, phal—phallapodeme, sap—subanal plate, ss—surstylus. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982429" httpUri="https://zenodo.org/record/13982429/files/figure.png" pageId="5" pageNumber="464">Fig. 4D</figureCitation>
)..........................................................................
<taxonomicName id="4C085DF912693A7777E66D22FAAEF5F4" authorityName="Kuwahara &amp; Marshall &amp; Yau" authorityYear="2024" box="[1211,1295,891,912]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="species" species="crena" status="sp. nov.">
<emphasis id="B97CFA6812693A7777E66D22FAAEF5F4" box="[1211,1295,891,912]" italics="true" pageId="5" pageNumber="464">H. crena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F471312693A7776496D22FAFEF5F4" box="[1300,1375,891,912]" pageId="5" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
[CRI]
</keyLead>
</paragraph>
<paragraph id="8BB7267A12693A7773CA6DC1FA3DF583" blockId="5.[151,1437,831,1235]" pageId="5" pageNumber="464">
<keyLead id="30F9854F12693A7773CA6DC1FA3DF583" pageId="5" pageNumber="464">
- Body length ≥
<quantity id="4CF08B9F12693A7772286DC1FE63F5C9" box="[373,450,920,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="464" unit="mm" value="1.9">1.9 mm</quantity>
. Male fore and hind femora distinctly thickened (length 2.9–3.2 × and 3.1–3.7 × width, respectively) (
<figureCitation id="13333AFF12693A7773B46DECFEFFF5AE" box="[233,350,949,971]" captionStart="FIGURE 2" captionStartId="3.[152,255,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-73@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Habitus and head photos of Helicosina species.A, Helicosina crena sp. nov. paratype habitus, lateral; B, Helicosina crena sp. nov. paratype head, anterolateral; C, Helicosina sica sp. nov. paratype habitus, lateral; D, Helicosina sica sp. nov. paratype head, anterolateral; E, Helicosina valga sp. nov. paratype habitus, lateral; F, Helicosina valga sp. nov. paratype head, anterolateral. Scale bars indicate 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.13982425" httpUri="https://zenodo.org/record/13982425/files/figure.png" pageId="5" pageNumber="464">Figs 2C, 2E</figureCitation>
). Surstylus sinuate, curved along anterior margin (
<figureCitation id="13333AFF12693A7770606DECFC2AF5AF" box="[829,907,949,971]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Figs 5C</figureCitation>
,
<figureCitation id="13333AFF12693A7770C86DECFC15F5AF" box="[917,948,949,971]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">6C</figureCitation>
). Posterior opening of epandrium ovoid or circular, widest around the middle (
<figureCitation id="13333AFF12693A7772BF6D8BFD90F58C" box="[482,561,978,1000]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Figs 5D</figureCitation>
,
<figureCitation id="13333AFF12693A7771616D8BFDFBF58C" box="[572,602,978,1000]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">6D</figureCitation>
)................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FC3EDF12693A7773CA6DA9FA3DF2B6" pageId="5" pageNumber="464">
<paragraph id="8BB7267A12693A7773CA6DA9FA3CF239" blockId="5.[151,1437,831,1235]" pageId="5" pageNumber="464">
<keyLead id="30F9854F12693A7773CA6DA9FA3CF239" pageId="5" pageNumber="464">
2 Eye height 3.1–3.2 × genal height; face only slightly protruding. Male S5 longer, width 2.4–2.7 × length (
<figureCitation id="13333AFF12693A7777BB6DA9FA92F261" box="[1254,1331,1007,1030]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5A</figureCitation>
). Posterior opening of epandrium almost circular (height 0.9–1.0 × width) (
<figureCitation id="13333AFF12693A7770106A54FC3BF247" box="[845,922,1037,1059]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5D</figureCitation>
). Surstylus bulging basally, apical half gently sinuate (
<figureCitation id="13333AFF12693A7773B46A73FE92F224" box="[233,307,1066,1088]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5C</figureCitation>
). Postgonite parallel-sided basally, truncate apically (
<figureCitation id="13333AFF12693A7770706A73FCD9F224" box="[813,888,1066,1088]" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5B</figureCitation>
). Distiphallus with a lateral row of thickened setulae (
<figureCitation id="13333AFF12693A7776286A73FF5EF239" captionStart="FIGURE 5" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[179,1412,186,1915]" captionTargetId="figure-19@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Helicosina sica sp. nov. male paratype terminalia. A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982433" httpUri="https://zenodo.org/record/13982433/files/figure.png" pageId="5" pageNumber="464">Fig. 5B</figureCitation>
)...............................................................................
<taxonomicName id="4C085DF912693A77779F6A11FAA6F239" authorityName="Kuwahara &amp; Marshall &amp; Yau" authorityYear="2024" box="[1218,1287,1096,1117]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="species" species="sica" status="sp. nov.">
<emphasis id="B97CFA6812693A77779F6A11FAA6F239" box="[1218,1287,1096,1117]" italics="true" pageId="5" pageNumber="464">H. sica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F471312693A7776516A11FAF6F239" box="[1292,1367,1096,1117]" pageId="5" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
[ECU]
</keyLead>
</paragraph>
<paragraph id="8BB7267A12693A7773CA6A3CFA3DF2B6" blockId="5.[151,1437,831,1235]" pageId="5" pageNumber="464">
<keyLead id="30F9854F12693A7773CA6A3CFA3DF2B6" pageId="5" pageNumber="464">
- Eye height 3.6–3.7 × genal height; face distinctly protruding. Male S5 shorter, width 4.2–4.5 × length (
<figureCitation id="13333AFF12693A7777BE6A3CFA92F21F" box="[1251,1331,1125,1147]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6A</figureCitation>
). Posterior opening of epandrium roughly ovoid (height 1.3–1.4 × width) (
<figureCitation id="13333AFF12693A7770166ADBFC38F2FC" box="[843,921,1154,1176]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6D</figureCitation>
). Surstylus roughly parallel-sided basally, apical half strongly bent at about a right angle (
<figureCitation id="13333AFF12693A7771636AF9FD2BF2D2" box="[574,650,1184,1206]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6C</figureCitation>
). Postgonite broadened basally, triangular apically (
<figureCitation id="13333AFF12693A7777216AF9FB69F2D2" box="[1148,1224,1184,1206]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6B</figureCitation>
). Distiphallus densely microtomentose dorsally but without longer setulae (
<figureCitation id="13333AFF12693A7771806AE4FC87F2B7" box="[733,806,1213,1235]" captionStart="FIGURE 6" captionStartId="8.[152,255,1943,1967]" captionTargetBox="[181,1406,186,1915]" captionTargetId="figure-29@8.[176,1411,181,1919]" captionTargetPageId="8" captionText="FIGURE 6. Helicosina valga sp. nov. male paratype terminalia.A, S5–8 and postabdomen, ventral; B, aedeagus and associated structures, left lateral; C, epandrium and surstylus, lateral; D. epandrium and surstyli, posterior. Scale bars indicate 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.13982435" httpUri="https://zenodo.org/record/13982435/files/figure.png" pageId="5" pageNumber="464">Fig. 6B</figureCitation>
)............................
<taxonomicName id="4C085DF912693A7777DF6AE7FB75F2B6" authorityName="Kuwahara &amp; Marshall &amp; Yau" authorityYear="2024" box="[1154,1236,1213,1235]" class="Insecta" family="Sphaeroceridae" genus="Helicosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="464" phylum="Arthropoda" rank="species" species="valga" status="sp. nov.">
<emphasis id="B97CFA6812693A7777DF6AE7FB75F2B6" box="[1154,1236,1213,1235]" italics="true" pageId="5" pageNumber="464">H. valga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F471312693A7777866AE4FA87F2B6" box="[1243,1318,1213,1234]" pageId="5" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
[CRI, PAN]
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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