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<document id="63FADDAD4292C53F3EF83F38C4C14942" ID-DOI="10.5252/zoosystema2024v46a25" ID-ISSN="1638-9387" ID-Zenodo-Dep="13907152" ID-ZooBank="D4AC1F89-AC34-43C4-9761-3F2015A02265" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728463297646" checkinUser="plazi" docAuthor="Segniagbeto, Gabriel Hoinsoudé, Mark-Oliver, Annemarie Ohler, Luca, Mark-Oliver Rödel, Alain, Luca Luiselli &amp; Dubois, Alain" docDate="2024" docId="03F187A4FFA1FFF7AB5CFB29FE6DF826" docLanguage="en" docName="zoosystema.2024.46.25.pdf" docOrigin="Zoosystema 46 (25)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a25.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Kassina fusca SchiOtz 1967" docType="treatment" docVersion="2" lastPageNumber="650" masterDocId="FFC8FFDCFFB4FFE2AA73FFBDFFEFFFCF" masterDocTitle="Amphibians of Togo: taxonomy, distribution and conservation status" masterLastPageNumber="670" masterPageNumber="631" pageNumber="650" updateTime="1728463900544" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3F36ADD56C226552E93EAAAF9FEA7797">Amphibians of Togo: taxonomy, distribution and conservation status</mods:title>
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<mods:namePart id="B132628CA21F4EFEFBE1F2441B0B1B33">Segniagbeto, Gabriel Hoinsoudé</mods:namePart>
<mods:affiliation id="B7E7F45138AEFFFDBE91051908C85A22">Laboratory of Ecology and Ecotoxicology, Faculty of Sciences, University of Lomé, BP 6057 Lomé, Togo, BP 1515, Lomé (Togo) and Institute for Development, Ecology, Conservation and Cooperation, Rome (Italy) gsegniagbeto @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="57EB07AFAEB91781B1E5CFD75C164387">Mark-Oliver, Annemarie Ohler</mods:namePart>
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<mods:namePart id="E7D52B820C2A84898AC10E70168B31AE">Luca, Mark-Oliver Rödel</mods:namePart>
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<mods:namePart id="9BA9F92D88FA109407B440B2DD942022">Alain, Luca Luiselli</mods:namePart>
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<mods:namePart id="0EC437CBC687036D67A325E7EEC95D7C">Dubois, Alain</mods:namePart>
<mods:affiliation id="C379B9B670553B8B0DF9695BA619553D">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE-PSL, Université des Antilles, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr</mods:affiliation>
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<heading id="D0AF81DEFFA1FFF7AB5CFB29FDB3FB61" box="[303,604,1170,1198]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="650" reason="2">
<taxonomicName id="4C584D31FFA1FFF7AB5CFB29FDB3FB61" ID-CoL="3R3JM" authority="SchiOtz, 1967" authorityName="SchiOtz" authorityYear="1967" box="[303,604,1170,1198]" class="Amphibia" family="Hyperoliidae" genus="Kassina" kingdom="Animalia" order="Anura" pageId="21" pageNumber="650" phylum="Chordata" rank="species" species="fusca">
<emphasis id="B92CEAA0FFA1FFF7AB5CFB29FE2FFB62" bold="true" box="[303,448,1171,1198]" italics="true" pageId="21" pageNumber="650">Kassina fusca</emphasis>
<bibRefCitation id="EFC94B43FFA1FFF7ABB4FB2FFDB3FB61" author="SCHIOTZ A." box="[455,604,1170,1198]" pageId="21" pageNumber="650" pagination="1 - 346" refId="ref38284" refString="SCHIOTZ A. 1967. - The treefrogs (Rhacophoridae) of West Africa. Spolia zoologica Musei haunienses 25: 1 - 346." type="journal article" year="1967">SchiØtz, 1967</bibRefCitation>
</taxonomicName>
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(
<figureCitation id="13632A37FFA1FFF7ABE0FB09FE18FB01" box="[403,503,1204,1230]" captionStart="FIG" captionStartId="22.[132,143,1609,1627]" captionTargetBox="[132,1455,215,1565]" captionTargetId="figure-258@22.[698,1267,754,1296]" captionTargetPageId="22" captionText="FIG. 10. — Representatives of anuran species from Togo in life: A, B, Hyperolius torrentis Schiøtz, 1967: A, (Kalaré); B, (Diguengué); C, Hyperolius sylvaticus Schiøtz, 1967 (Diguengué);D, Kassina fusca Schiøtz, 1967 (Bounako); E, Kassina schioetzi RÖdel, Grafe, Rudolf &amp; Ernst, 2002 (Tsévié); F, Kassina senegalensis (Duméril &amp; Bibron, 1841) (Tsévié)." figureDoi="http://doi.org/10.5281/zenodo.13907172" httpUri="https://zenodo.org/record/13907172/files/figure.png" pageId="21" pageNumber="650">Fig. 10D</figureCitation>
)
</paragraph>
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. —
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<collectingCountry id="F34F7622FFA1FFF7AB1FFB4AFE72FAC0" box="[364,413,1271,1295]" name="Togo" pageId="21" pageNumber="650">Togo</collectingCountry>
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; Kanté; MNHN-RA-1998.2012.
</paragraph>
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. —
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<collectingCountry id="F34F7622FFA1FFF7ABEFFA8CFDCBFA87" box="[412,548,1328,1353]" name="Burkina Faso" pageId="21" pageNumber="650">Burkina Faso</collectingCountry>
</emphasis>
<specimenCount id="9D5EFD3BFFA1FFF7A84DFA8DFD87FA84" box="[574,616,1328,1355]" count="1" pageId="21" pageNumber="650" type="male">1 ♂</specimenCount>
; Lay; MNHN- RA-1998.2302 •
<specimenCount id="9D5EFD3BFFA1FFF7AB5CFAF6FEBCFAA9" box="[303,339,1355,1382]" count="1" pageId="21" pageNumber="650" type="female">1 ♀</specimenCount>
; origin unknown; MNHN-RA-1998.2301.
</paragraph>
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. — Small to medium sized frog (SVL
<quantity id="4CA09B57FFA1FFF7A8BEFA3EFF0DFA7A" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.75" metricValueMin="3.15" pageId="21" pageNumber="650" unit="mm" value="34.5" valueMax="37.5" valueMin="31.5">31.5- 37.5 mm</quantity>
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, 62.0 mm
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), with a fairly robust and stocky body. Snout oval or rounded. Head as wide as long (HW 26-37% SVL; HL 26-34% SVL). Tympanum distinct (TYD 4-5% SVL).Tibia short (TL 30-39% SVL). Webbing absent, phalanges all free. Skin smooth on back, flank with glandular warts; very granular chest and belly.
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. — Dorsal pattern with large brown, dark, light or white spots of irregular shape. The dorsal surfaces of the forearm, thigh and tibia have the same pattern as the back. The belly, chest and throat are white.
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. — In males, the throat is black, with welldeveloped glandular disc on subgular vocal sac.
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. — This species is mainly a Sudanian savannah species (
<bibRefCitation id="EFC94B43FFA1FFF7AB47F8F0FE56F8AA" author="HUGHES B." box="[308,441,1869,1893]" pageId="21" pageNumber="650" pagination="29 - 38" refId="ref34770" refString="HUGHES B. 1988. - Herpetology in Ghana (West Africa). British Herpetological Society Bulletin 25: 29 - 38." type="journal article" year="1988">Hughes 1988</bibRefCitation>
;
<bibRefCitation id="EFC94B43FFA1FFF7ABB6F8F0FD95F8AA" author="BOHME W. &amp; MEINIG H. &amp; RODEL M. - O." box="[453,634,1869,1893]" pageId="21" pageNumber="650" pagination="7 - 26" refId="ref32943" refString="BOHME W., MEINIG H. &amp; RODEL M. - O. 1996. - New records of amphibians and reptiles from Burkina ̿ aso and Mali. British Herpetological Society Bulletin 55: 7 - 26." type="journal article" year="1996">
Böhme
<emphasis id="B92CEAA0FFA1FFF7A861F8F3FDAEF8AA" box="[530,577,1869,1893]" italics="true" pageId="21" pageNumber="650">et al.</emphasis>
1996
</bibRefCitation>
;
<bibRefCitation id="EFC94B43FFA1FFF7A8F5F8F0FD15F8AA" author="RODEL M. - O." box="[646,762,1869,1893]" pageId="21" pageNumber="650" refId="ref37146" refString="RODEL M. - O. 2000. - Herpetofauna of West Africa. Vol 1. Amphibians of theWest African savanna. ̿ rankfurt am Main, Chimaira, 335 p." type="book" year="2000">Rödel 2000</bibRefCitation>
). Its distribution in
<collectingCountry id="F34F7622FFA1FFF7AB40F8D5FE8AF84F" box="[307,357,1896,1920]" name="Togo" pageId="21" pageNumber="650">Togo</collectingCountry>
is limited to ecological zones I and II, similar to the species distribution in neighbouring
<collectingCountry id="F34F7622FFA1FFF7A822F83FFD64F855" box="[593,651,1922,1946]" name="Benin" pageId="21" pageNumber="650">Benin</collectingCountry>
(
<bibRefCitation id="EFC94B43FFA1FFF7A8E8F83EFF57F87B" author="NAGO S. G. A. &amp; GRELL O. &amp; SINSIN B. &amp; RODEL M. - O." pageId="21" pageNumber="650" pagination="93 - 108" refId="ref36329" refString="NAGO S. G. A., GRELL O., SINSIN B. &amp; RODEL M. - O. 2006. - The amphibian fauna of the Pendjari National Park and surroundings, northern Benin. Salamandra 42: 93 - 108." type="journal article" year="2006">
Nago
<emphasis id="B92CEAA0FFA1FFF7A8ABF83EFCE7F855" box="[728,776,1922,1946]" italics="true" pageId="21" pageNumber="650">et al.</emphasis>
2006
</bibRefCitation>
). The species was reported in
<collectingCountry id="F34F7622FFA1FFF7AB93F820FDFDF87A" box="[480,530,1949,1973]" name="Togo" pageId="21" pageNumber="650">Togo</collectingCountry>
by
<bibRefCitation id="EFC94B43FFA1FFF7A844F820FD20F87A" author="BOURGAT R." box="[567,719,1949,1973]" pageId="21" pageNumber="650" pagination="597 - 624" refId="ref33022" refString="BOURGAT R. 1979. - Trematodes d'amphibiens du Togo. Bulletin du Museum national d'Histoire naturelle (4 e serie) 3: 597 - 624." type="journal article" year="1979">Bourgat (1979)</bibRefCitation>
from Kanté. We observed this species in Bounaka and the northern Fazao Malfakassa National Park.
</paragraph>
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</treatment>
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