treatments-xml/data/03/99/87/03998788FF94041CCB3BFA373CF4FE62.xml

<document id="D36625560F12970973AA6B121F6BC441" ID-DOI="10.6620/ZS.2018.57-27" ID-ISSN="1810-522X" ID-PMC="PMC6517715" ID-PubMed="31966267" ID-Zenodo-Dep="13314991" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721916104468" checkinUser="felipe" docAuthor="Cai, Yixiong &amp; Ng, Peter Kee Lin" docDate="2018" docId="03998788FF94041CCB3BFA373CF4FE62" docLanguage="en" docName="ZoolStud.57.27.1-33.pdf" docOrigin="Zoological Studies 57 (27)" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Caridina spinicrus Cai &amp; Ng, 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="FFA0FFF0FF9C0410CA0AFFEB3F02FFF5" masterDocTitle="Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="9" updateTime="1723578273083" updateUser="ExternalLinkService">
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<mods:title id="A56EB1B39D2C290B0B3435406122D743">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="1B21DCC3C707281FBE3D729414B6E838">Cai, Yixiong</mods:namePart>
<mods:affiliation id="9D946E0169D460358B562D7BB051EFF9">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<mods:namePart id="F8BA3119301C1FD736FE4518DB58F2F0">Ng, Peter Kee Lin</mods:namePart>
<mods:affiliation id="2097D2E9D185FE0AC58CA9FD2C850F03">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of &amp; National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<mods:title id="8412F3474D9AB97774E84DFEF8FED708">Zoological Studies</mods:title>
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<mods:date id="6DC48A3C406EBD2D2A4038E56F35851D">2018</mods:date>
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<treatment id="03998788FF94041CCB3BFA373CF4FE62" ID-DOI="http://doi.org/10.5281/zenodo.12826764" ID-Zenodo-Dep="12826764" LSID="urn:lsid:zoobank.org:act:EF868C18-D01C-4920-A9C4-51DFACB8D42C" httpUri="http://treatment.plazi.org/id/03998788FF94041CCB3BFA373CF4FE62" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
<subSubSection id="C32A6515FF940418CB3BFA373D81FA03" box="[305,643,1500,1526]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B8F369EFF940418CB3BFA373D81FA03" blockId="8.[183,766,1500,1610]" box="[305,643,1500,1526]" pageId="8" pageNumber="9">
<heading id="D0C781F2FF940418CB3BFA373D81FA03" bold="true" box="[305,643,1500,1526]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="9" reason="2">
<emphasis id="B944EA8CFF940418CB3BFA373D81FA03" bold="true" box="[305,643,1500,1526]" pageId="8" pageNumber="9">
<taxonomicName id="4C304D1DFF940418CB3BFA373D18FA03" ID-CoL="8VKFL" authorityName="Cai &amp; Ng" authorityYear="2018" box="[305,538,1500,1526]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF940418CB3BFA373D18FA03" bold="true" box="[305,538,1500,1526]" italics="true" pageId="8" pageNumber="9">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF940418C82BFA363D81FA03" box="[545,643,1501,1526]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32A6515FF94041CCB94FA163CF4FE62" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
<paragraph id="8B8F369EFF940418CB94FA163D15F9E3" blockId="8.[183,766,1500,1610]" box="[414,535,1533,1558]" pageId="8" pageNumber="9">
(
<figureCitation id="130B2A1BFF940418CBAFFA163D12F9E3" box="[421,528,1533,1558]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[136,170,1898,1918]" captionStartId-1="10.[174,208,1898,1918]" captionStartId-2="11.[136,170,1898,1918]" captionTargetBox-0="[252,1337,267,1858]" captionTargetBox-1="[301,1322,254,1855]" captionTargetBox-2="[204,1333,269,1863]" captionTargetId-0="figure-6@9.[196,1354,249,1870]" captionTargetId-1="figure-58@10.[275,1351,249,1870]" captionTargetId-2="figure-58@11.[195,1354,249,1870]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." captionText-1="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." captionText-2="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi-0="http://doi.org/10.5281/zenodo.13315007" figureDoi-1="http://doi.org/10.5281/zenodo.13315009" figureDoi-2="http://doi.org/10.5281/zenodo.13315011" httpUri-0="https://zenodo.org/record/13315007/files/figure.png" httpUri-1="https://zenodo.org/record/13315009/files/figure.png" httpUri-2="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Figs. 7-9</figureCitation>
)
</paragraph>
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<uri id="FFA13A9CFF940418CABDF9F33D37F9BF" pageId="8" pageNumber="9">
urn:lsid:zoobank.org:act: 
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<paragraph id="8B8F369EFF940418CAE0F99B3D4DF8DC" blockId="8.[174,776,1648,1961]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0F99B3EEFF97C" box="[234,493,1648,1673]" italics="true" pageId="8" pageNumber="9">Material examined</emphasis>
: 
<materialsCitation id="3B583CC3FF940418C80FF99B3FF2F8FC" collectingDate="1993-10-30" collectionCode="IZCAS" collectorName="Y. Cai" country="China" location="Goulou Cave" pageId="8" pageNumber="9" specimenCode="DE 5031" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="548B883CFF940418C80FF99B3D7DF97C" box="[517,639,1648,1673]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
: male, cl 
<quantity id="4CC89B7BFF940418CAA4F97B3E11F95C" box="[174,275,1680,1705]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="9" unit="mm" value="5.5">5.5 mm</quantity>
, 
<collectionCode id="ED21AE5BFF940418CB28F97B3E76F95F" box="[290,372,1680,1706]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="9">IZCAS</collectionCode>
<specimenCode id="DB969EE5FF940418CB8AF97B3EF7F95C" box="[384,501,1680,1705]" collectionCode="IZCAS" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="9">DE 5031</specimenCode>
, artificial pond inside 
<location id="8EEF6045FF940418CAA4F95B3E4CF93F" LSID="urn:lsid:plazi:treatment:03998788FF94041CCB3BFA373CF4FE62:8EEF6045FF940418CAA4F95B3E4CF93F" box="[174,334,1712,1738]" country="China" name="Goulou Cave" pageId="8" pageNumber="9" stateProvince="Guangxi">Goulou Cave</location>
in 
<taxonomicName id="4C304D1DFF940418CB7DF95B3D19F93F" authority="County" authorityName="County" box="[375,539,1712,1738]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beiliu">Beiliu County</taxonomicName>
, 
<collectingRegion id="49F4F87CFF940418C822F95B3C01F93C" box="[552,771,1712,1738]" country="China" name="Guangxi" pageId="8" pageNumber="9">Guangxi Province</collectingRegion>
, southern 
<collectingCountry id="F327760EFF940418CB3CF93B3E87F91C" box="[310,389,1744,1770]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
, coll. 
<collectorName id="26C55348FF940418CBECF93B3D38F91C" box="[486,570,1744,1770]" pageId="8" pageNumber="9">Y. Cai</collectorName>
et al., 
<date id="FF8E105EFF940418C8A0F93B3FF2F8FC" pageId="8" pageNumber="9" value="1993-10-30">
<collectingDate id="EFCAE9B6FF940418C8A0F93B3FF2F8FC" pageId="8" pageNumber="9" value="1993-10-30">30 Oct 1993</collectingDate>
</date>
</materialsCitation>
; 
<typeStatus id="548B883CFF940418CAF5F91B3E81F8FC" box="[255,387,1776,1801]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D36FD17FF940418CB98F91B3EF9F8FC" box="[402,507,1776,1801]" count="2" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
, cl 5.5-6.0 mm, ZRC 2018.0482, same data as 
<typeStatus id="548B883CFF940418CBE8F8FB3D49F8DC" box="[482,587,1808,1833]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8F369EFF940418CAE0F8DB3A14FE83" blockId="8.[174,776,1648,1961]" lastBlockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
<emphasis id="B944EA8CFF940418CAE0F8DB3E83F8BC" box="[234,385,1840,1865]" italics="true" pageId="8" pageNumber="9">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF940418C814F8DB3D87F8BC" box="[542,645,1840,1865]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7A</figureCitation>
) straight, reaching anterior half of second segment of antennular peduncle, dorsally with 17 teeth, of which 7 on carapace behind posterior margin of orbital margin, occupying 0.14 carapace length, armed ventrally with only 1 small tooth. Antennal spine sharp, completely fused with suborbital angle. Pterygostomian angle rounded.
</paragraph>
<paragraph id="8B8F369EFF940418C985FE963B34FE03" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
Telson (
<figureCitation id="130B2A1BFF940418CE0DFE963B73FE63" box="[1031,1137,381,406]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Fig. 9B</figureCitation>
) with posteromedian projection, posterior margin rounded, lateral pair of spines longer than intermediate pairs, sub-lateral pair being shortest.
</paragraph>
<paragraph id="8B8F369EFF940418C985FE163CBCFD43" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
Eyes well developed (
<figureCitation id="130B2A1BFF940418CEA3FE163A0FFDE3" box="[1193,1293,509,534]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7A</figureCitation>
). Antennular peduncle 0.50 times as long as carapace. Stylocerite reaching 0.7 basal segment length of antennular peduncle. Scaphocerite (
<figureCitation id="130B2A1BFF940418CF49FDB63AA1FD83" box="[1347,1443,605,630]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8H</figureCitation>
) 3.0 times as long as wide, outer margin slightly concave.
</paragraph>
<paragraph id="8B8F369EFF940418C985FD563CDDFB43" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF940418CF1DFD563A6CFD23" box="[1303,1390,701,726]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7B</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF940418C950FD163CB8FCE2" box="[858,954,765,791]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7C</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with setae and teeth. Maxilla (
<figureCitation id="130B2A1BFF940418CF76FCD63C75FC83" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7D</figureCitation>
) with slender palp, upper endite subdivided, scaphognathite tapering with numerous long setae posteriorly. Palp of first maxilliped (
<figureCitation id="130B2A1BFF940418CF73FC763C77FC23" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7E</figureCitation>
) ending as triangular projection. Endopod of second maxilliped (
<figureCitation id="130B2A1BFF940418CE4DFC363BA7FC03" box="[1095,1189,989,1014]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7E</figureCitation>
) with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF940418CF42FC163AA6FBE2" box="[1352,1444,1021,1047]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="8" pageNumber="9">Fig. 7G</figureCitation>
) reaching beyond distal end of scaphocerite, ending in single terminal claw; exopod reaching 0.5 times penultimate segment length, ultimate segment as long as pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<paragraph id="8B8F369EFF940418C985FB563AA9F842" blockId="8.[850,1452,253,1975]" pageId="8" pageNumber="9">
First pereiopod (
<figureCitation id="130B2A1BFF940418CE53FB563BDAFB23" box="[1113,1240,1213,1238]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8A, B</figureCitation>
) reaching end of basal segment of antennular peduncle, merus 2.7 times as long as broad; carpus concave anteriorly, 1.7 times as long as high, chela 2.5 times as long as broad, fingers 1.3 times as long as palm. Second pereiopod (
<figureCitation id="130B2A1BFF940418CE47FAB63BACFA82" box="[1101,1198,1373,1399]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8C</figureCitation>
) reaching to end of antennular peduncle, carpus 5.2 times as long as high, chela slightly shorter than carpus, 2.7 times as long as broad, fingers 1.7 times as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF940418CEA2FA363A0BFA03" box="[1192,1289,1501,1526]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="8" pageNumber="9">Fig. 8D</figureCitation>
) long, stout, reaching beyond distal end of scaphocerite. Merus 4.0 times as long as broad; carpus 2.4 times as long as broad, ventral surface armed with a row of 16 closely packed strong spines. Propodus strongly curved inwards, 9.0 times as long as broad, 4.0 times as long as dactylus (claw included), ventral surface armed with row of 23 strong spines, similar to those of at carpus, but more widely lined anteriorly; dactylus terminating as a claw, with 5 accessory spines on flexor margin. Fourth pereiopod (
<figureCitation id="130B2A1BFF940418C9EAF8D63B8BF8A3" box="[992,1161,1853,1879]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Fig. 9C, D, E</figureCitation>
) reaching to distal end of antennular peduncle, similar to third pereiopod in form, with strong spines on ventral surfaces of carpus and propodus. Fifth pereiopod (
<figureCitation id="130B2A1BFF940418CF23F8763AA6F842" box="[1321,1444,1949,1975]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="8" pageNumber="9">Fig. 9F, G</figureCitation>
)
</paragraph>
<footnote id="E82B2A90FF940418CE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">
<paragraph id="8B8F369EFF940418CE96F8363AA9F81A" blockId="8.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF950419CADDFC2C3FF1FC2E" blockId="9.[215,243,967,987]" box="[215,243,967,987]" pageId="9" pageNumber="10">(B)</paragraph>
<caption id="DF4F6616FF950419CA82F8813B74F843" ID-DOI="http://doi.org/10.5281/zenodo.13315007" ID-Zenodo-Dep="13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,170,1898,1918]" targetBox="[252,1337,267,1858]" targetPageId="9" targetType="figure">
<paragraph id="8B8F369EFF950419CA82F8813B74F843" blockId="9.[136,1415,1898,1974]" pageId="9" pageNumber="10">
<emphasis id="B944EA8CFF950419CA82F8813FC5F88B" bold="true" box="[136,199,1898,1918]" pageId="9" pageNumber="10">Fig. 7.</emphasis>
<taxonomicName id="4C304D1DFF950419CADDF8813E80F88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[215,386,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF950419CADDF8813E80F88B" box="[215,386,1898,1918]" italics="true" pageId="9" pageNumber="10">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF950419CB83F8813ED4F88B" box="[393,470,1898,1918]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
(A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482).
</paragraph>
</caption>
<footnote id="E82B2A90FF950419CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="9" pageNumber="10">
<paragraph id="8B8F369EFF950419CA82F8363E9AF81A" blockId="9.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="9" pageNumber="10">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF96041ACAA4F8813D18F843" ID-DOI="http://doi.org/10.5281/zenodo.13315009" ID-Zenodo-Dep="13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="10" pageNumber="11" startId="10.[174,208,1898,1918]" targetBox="[301,1322,254,1855]" targetPageId="10" targetType="figure">
<paragraph id="8B8F369EFF96041ACAA4F8813D18F843" blockId="10.[174,1453,1898,1974]" pageId="10" pageNumber="11">
<emphasis id="B944EA8CFF96041ACAA4F8813FE9F88B" bold="true" box="[174,235,1898,1918]" pageId="10" pageNumber="11">Fig. 8.</emphasis>
<taxonomicName id="4C304D1DFF96041ACAFDF8813EA3F88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[247,417,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF96041ACAFDF8813EA3F88B" box="[247,417,1898,1918]" italics="true" pageId="10" pageNumber="11">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF96041ACBADF8813EF0F88B" box="[423,498,1898,1918]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482).
</paragraph>
</caption>
<footnote id="E82B2A90FF96041ACE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="10" pageNumber="11">
<paragraph id="8B8F369EFF96041ACE96F8363AA9F81A" blockId="10.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="10" pageNumber="11">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF97041BCA82F8813FFEF843" ID-DOI="http://doi.org/10.5281/zenodo.13315011" ID-Zenodo-Dep="13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="11" pageNumber="12" startId="11.[136,170,1898,1918]" targetBox="[204,1333,269,1863]" targetPageId="11" targetType="figure">
<paragraph id="8B8F369EFF97041BCA82F8813FFEF843" blockId="11.[136,1414,1898,1974]" pageId="11" pageNumber="12">
<emphasis id="B944EA8CFF97041BCA82F8813FC5F88B" bold="true" box="[136,199,1898,1918]" pageId="11" pageNumber="12">Fig. 9.</emphasis>
<taxonomicName id="4C304D1DFF97041BCADFF8813E7DF88B" authorityName="Cai &amp; Ng" authorityYear="2018" box="[213,383,1898,1918]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF97041BCADFF8813E7DF88B" box="[213,383,1898,1918]" italics="true" pageId="11" pageNumber="12">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF97041BCB8DF8813ED0F88B" box="[391,466,1898,1918]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482).
</paragraph>
</caption>
<footnote id="E82B2A90FF97041BCA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="11" pageNumber="12">
<paragraph id="8B8F369EFF97041BCA82F8363E9AF81A" blockId="11.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="11" pageNumber="12">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF90041CCAA4FF163E0AFE43" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">reaching distal end of antennular peduncle; merus 4.0 times as long as broad; carpus 4.0 times as long as broad; propodus slightly curved inwards, 10 times as long as broad, 3.8 times as long as dactylus; dactylus with 50 spinules on flexor margin.</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FE563DA8FCE3" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
Endopod of male first pleopod (
<figureCitation id="130B2A1BFF90041CC893FE563DFCFE23" box="[665,766,445,470]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8E</figureCitation>
) reaching to 0.67 endopod length, subrectangular, distal end rounded, inner margin slightly curved backwards, 3.0 times as long as wide; appendix interna situated at posterior 0.2 endopod length, extending to end of endopod. Appendix masculina of male second pleopod (
<figureCitation id="130B2A1BFF90041CCBFFFD963D7AFD62" box="[501,632,637,663]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8F, G</figureCitation>
) narrowing down distally, with 2 rows of spines at inner surface, reaching to 0.75 of endopod length; appendix interna of male second pleopod small, reaching to 0.4 appendix masculina length.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FCF63C07FCC3" blockId="12.[174,775,253,1974]" box="[234,773,797,822]" pageId="12" pageNumber="13">
Uropodal diaeresis (
<figureCitation id="130B2A1BFF90041CCBD2FCF63D32FCC3" box="[472,560,797,822]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="12" pageNumber="13">Fig. 7H</figureCitation>
) with 13 spinules.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FCD63E8FFC23" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CCAE0FCD63E4AFCA3" box="[234,328,829,854]" italics="true" pageId="12" pageNumber="13">Habitat</emphasis>
: 
<taxonomicName id="4C304D1DFF90041CCB55FCD63D52FCA3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[351,592,829,854]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCB55FCD63D52FCA3" box="[351,592,829,854]" italics="true" pageId="12" pageNumber="13">Caridina spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CC854FCD63DCAFCA3" box="[606,712,829,854]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
was collected together with 
<taxonomicName id="4C304D1DFF90041CCBE9FCB63D4DFC83" authorityName="Cai &amp; Ng" authorityYear="2018" box="[483,591,861,886]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCBE9FCB63D4DFC83" box="[483,591,861,886]" italics="true" pageId="12" pageNumber="13">C. beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CC851FCB63DC0FC83" box="[603,706,861,886]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
from an artificial concrete pond inside Goulou Cave in 
<taxonomicName id="4C304D1DFF90041CCADEFC763E98FC43" authority="County." authorityName="County." box="[212,410,925,951]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu">Beiliu County.</taxonomicName>
The water comes from a subterranean river.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FC363D08FBA3" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CCAE0FC363E64FC03" box="[234,358,989,1014]" italics="true" pageId="12" pageNumber="13">Etymology</emphasis>
: 
<taxonomicName id="4C304D1DFF90041CCB7CFC363D48FC03" authorityName="Cai &amp; Ng" authorityYear="2018" box="[374,586,989,1014]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CCB7CFC363D48FC03" box="[374,586,989,1014]" italics="true" pageId="12" pageNumber="13">Caridina spinicrus</emphasis>
</taxonomicName>
is named for its distinctive spiny third and fourth pereiopods, from the Latin for spiny and legs (crura). The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0FBB63E56F983" blockId="12.[174,775,253,1974]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CCAE0FBB63E54FB83" box="[234,342,1117,1142]" italics="true" pageId="12" pageNumber="13">Remarks</emphasis>
: With regard to the spiny third and fourth pereiopods, only one species, 
<taxonomicName id="4C304D1DFF90041CC891FB963EE8FB43" authority="Gurney, 1984" authorityName="Gurney" authorityYear="1984" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crurispinata">
<emphasis id="B944EA8CFF90041CC891FB963E3BFB43" italics="true" pageId="12" pageNumber="13">Caridina crurispinata</emphasis>
<bibRefCitation id="EFA14B6FFF90041CCB49FB763EE8FB43" author="Gurney AR" box="[323,490,1181,1207]" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
</taxonomicName>
, described from a cave in 
<collectingCountry id="F327760EFF90041CCAC7FB563E64FB23" box="[205,358,1213,1238]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
, shares such features. Compared with 
<taxonomicName id="4C304D1DFF90041CCAE0FB363EA2FB03" authorityName="Gurney" authorityYear="1984" box="[234,416,1245,1270]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crurispinata">
<emphasis id="B944EA8CFF90041CCAE0FB363EA2FB03" box="[234,416,1245,1270]" italics="true" pageId="12" pageNumber="13">C. crurispinata</emphasis>
</taxonomicName>
, the spines on the third and fourth pereiopods in 
<taxonomicName id="4C304D1DFF90041CCBC9FB163D5CFAE3" authorityName="Cai &amp; Ng" authorityYear="2018" box="[451,606,1277,1302]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCBC9FB163D5CFAE3" box="[451,606,1277,1302]" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CC861FB163DD0FAE3" box="[619,722,1277,1302]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
are more prominent (
<figureCitation id="130B2A1BFF90041CCB75FAF63EDCFAC3" box="[383,478,1309,1334]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8D</figureCitation>
; 
<figureCitation id="130B2A1BFF90041CCBE6FAF63D8DFAC3" box="[492,655,1309,1335]" captionStart="Fig" captionStartId="11.[136,170,1898,1918]" captionTargetBox="[204,1333,269,1863]" captionTargetId="figure-58@11.[195,1354,249,1870]" captionTargetPageId="11" captionText="Fig. 9. Caridina spinicrus sp. nov. (A) telson, (B) distal portion of telson, (C, D) fourth pereiopod, (E) dactylus of fourth pereiopod, (F) fifth pereiopod, (G) dactylus of fifth pereiopod. Scale bars: A, C, D, F = 1.0 mm; B, E, G = 0.2 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315011" httpUri="https://zenodo.org/record/13315011/files/figure.png" pageId="12" pageNumber="13">Fig. 9C, D, E</figureCitation>
) (vs. less prominent; cf. 
<bibRefCitation id="EFA14B6FFF90041CCB6BFAD63D09FAA3" author="Gurney AR" box="[353,523,1341,1367]" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
: fig. 6C). Apart from this, 
<taxonomicName id="4C304D1DFF90041CCAE4FAB63E85FA83" authorityName="Cai &amp; Ng" authorityYear="2018" box="[238,391,1373,1398]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CCAE4FAB63E85FA83" box="[238,391,1373,1398]" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C304D1DFF90041CC8E1FAB63E3AFA63" authorityName="Gurney" authorityYear="1984" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crurispinata">
<emphasis id="B944EA8CFF90041CC8E1FAB63E3AFA63" italics="true" pageId="12" pageNumber="13">C. crurispinata</emphasis>
</taxonomicName>
by the straight rostrum which reaches to the anterior half of the second segment of antennular peduncle (
<figureCitation id="130B2A1BFF90041CCBDFFA563D39FA23" box="[469,571,1469,1494]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
) (vs. gradually curving upwards and reaching beyond the antennal scale; cf. 
<bibRefCitation id="EFA14B6FFF90041CCB2BFA163EC4F9E3" author="Gurney AR" box="[289,454,1533,1559]" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
: fig. 4B); and the presence of an appendix interna on the endopod of the male first pleopod (
<figureCitation id="130B2A1BFF90041CCB68F9D63EC1F9A3" box="[354,451,1597,1622]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8E</figureCitation>
) (vs. absent; cf. 
<bibRefCitation id="EFA14B6FFF90041CC8AEF9D63FECF983" author="Gurney AR" pageId="12" pageNumber="13" pagination="567 - 590" refId="ref15183" refString="Gurney AR. 1984. Freshwater shrimps genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with description of four new species. J Nat Hist 18: 567 - 590, figures 1 - 14." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
: fig. 7B).
</paragraph>
<paragraph id="8B8F369EFF90041CCAE0F9963BB7FEA3" blockId="12.[174,775,253,1974]" lastBlockId="12.[850,1451,253,407]" pageId="12" pageNumber="13">
Compared with other Chinese species, 
<taxonomicName id="4C304D1DFF90041CC8E1F9963E15F943" authorityName="Cai &amp; Ng" authorityYear="2018" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CC8E1F9963E15F943" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
also superficially resembles the sympatric 
<taxonomicName id="4C304D1DFF90041CCAA4F9563E12F923" authorityName="Cai &amp; Ng" authorityYear="2018" box="[174,272,1725,1750]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="beiliu" status="sp. nov.">
<emphasis id="B944EA8CFF90041CCAA4F9563E12F923" box="[174,272,1725,1750]" italics="true" pageId="12" pageNumber="13">C. beiliu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF90041CCB13F9563E75F923" box="[281,375,1725,1750]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
However, 
<taxonomicName id="4C304D1DFF90041CCBF2F9563D87F923" authorityName="Cai &amp; Ng" authorityYear="2018" box="[504,645,1725,1750]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spinicrus">
<emphasis id="B944EA8CFF90041CCBF2F9563D87F923" box="[504,645,1725,1750]" italics="true" pageId="12" pageNumber="13">C. spinicrus</emphasis>
</taxonomicName>
can easily be separated by the rostrum not reaching beyond the end of the second segment of the antennular peduncle (
<figureCitation id="130B2A1BFF90041CCB3CF8F63E9AF8C3" box="[310,408,1821,1846]" captionStart="Fig" captionStartId="9.[136,170,1898,1918]" captionTargetBox="[252,1337,267,1858]" captionTargetId="figure-6@9.[196,1354,249,1870]" captionTargetPageId="9" captionText="Fig. 7. Caridina spinicrus sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 2.0 mm; B, C, H = 0.5 mm; D-G = 1.0 mm. (A, male, cl 5.5 mm, holotype, IZCAS DE 5031; B-H, male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315007" httpUri="https://zenodo.org/record/13315007/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
) (vs. beyond the end of the second segment of the antennular peduncle; 
<figureCitation id="130B2A1BFF90041CC8D0F8D63FE1F883" captionStart="Fig" captionStartId="13.[136,170,1898,1918]" captionTargetBox="[244,1338,260,1856]" captionTargetId="figure-58@13.[206,1344,249,1873]" captionTargetPageId="13" captionText="Fig. 10. Caridina beiliu sp. nov. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped, (G) third maxilliped, (H) uropodal diaeresis. Scale bars: A = 1.5 mm, B-G = 1.0 mm, H = 0.5 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483)." figureDoi="http://doi.org/10.5281/zenodo.13315013" httpUri="https://zenodo.org/record/13315013/files/figure.png" pageId="12" pageNumber="13">Fig. 10A</figureCitation>
); the more slender first pereiopod having the dactylus 2.5 times as long as broad (
<figureCitation id="130B2A1BFF90041CC874F8963DFDF863" box="[638,767,1917,1942]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8A, B</figureCitation>
) (vs. 1.5 times; 
<figureCitation id="130B2A1BFF90041CCB51F8763EC3F843" box="[347,449,1949,1974]" captionStart="Fig" captionStartId="14.[174,208,1925,1945]" captionTargetBox="[205,1322,249,1864]" captionTargetId="figure-6@14.[202,1347,249,1871]" captionTargetPageId="14" captionText="Fig. 11. Caridina beiliu sp. nov. (A) first pereiopod, (B) second pereiopod, (C) third pereiopod, (D) dactylus of third pereiopod, (E) fifth pereiopod, (F) dactylus of fifth pereiopod. Scale bars: A-C, E = 1.0 mm; D, F = 0.2 mm. (male, cl 6.2 mm, paratype, ZRC 2018.0483). © 2018 Academia Sinica, Taiwan" figureDoi="http://doi.org/10.5281/zenodo.13315015" httpUri="https://zenodo.org/record/13315015/files/figure.png" pageId="12" pageNumber="13">Fig. 11A</figureCitation>
); and the male first pleopod having an appendix interna that reaches to the distal end of the endopod (
<figureCitation id="130B2A1BFF90041CCE95FEF63BF9FEC3" box="[1183,1275,285,310]" captionStart="Fig" captionStartId="10.[174,208,1898,1918]" captionTargetBox="[301,1322,254,1855]" captionTargetId="figure-58@10.[275,1351,249,1870]" captionTargetPageId="10" captionText="Fig. 8. Caridina spinicrus sp. nov. (A, B) first pereiopod, (C) second pereiopod, (D) third pereiopod, (E) male first pleopod, (F) appendix masculina and appendix interna of male second pleopod, (G) male second pleopod. Scale bars: A-D, H = 1.0 mm; E-G = 0.5 mm. (male, cl 6.0 mm, paratype, ZRC 2018.0482)." figureDoi="http://doi.org/10.5281/zenodo.13315009" httpUri="https://zenodo.org/record/13315009/files/figure.png" pageId="12" pageNumber="13">Fig. 8E</figureCitation>
) (vs. distinctly reaching beyond; 
<figureCitation id="130B2A1BFF90041CCE2CFED63B7BFEA3" box="[1062,1145,317,342]" captionStart="Fig" captionStartId="15.[136,170,1898,1918]" captionTargetBox="[142,1414,261,1859]" captionTargetId="figure-6@15.[114,1423,249,1899]" captionTargetPageId="15" captionText="Fig. 12. Caridina beiliu sp. nov. (A) telson, (B) distal portion of telson, (C) scaphocerite, (D, E, F) endopod of male first pleopod, (G) appendix masculina and appendix interna of male second pleopod. Scale bars: A, C = 1.0 mm, B, D-F = 0.2 mm. (A-D, G. male, cl 6.2 mm, paratype, ZRC 2018.0483; E. F., male, cl 4.8 mm, paratype, RMNH.CRUS.D.57291)." figureDoi="http://doi.org/10.5281/zenodo.13315017" httpUri="https://zenodo.org/record/13315017/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
D-F).
</paragraph>
<paragraph id="8B8F369EFF90041CC985FEB63CF4FE62" blockId="12.[850,1451,253,407]" pageId="12" pageNumber="13">
<emphasis id="B944EA8CFF90041CC985FEB63B28FE83" box="[911,1066,349,374]" italics="true" pageId="12" pageNumber="13">Distribution</emphasis>
: 
<collectingRegion id="49F4F87CFF90041CCE4BFEB63A31FE83" box="[1089,1331,349,375]" country="China" name="Guangxi" pageId="12" pageNumber="13">Guangxi Province</collectingRegion>
(Goulou Cave), 
<collectingCountry id="F327760EFF90041CC9A2FE963CF0FE62" box="[936,1010,381,407]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>