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<document id="D2B6CE950241E597DDB77D9A83EEAD72" ID-DOI="10.11646/zootaxa.5584.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14812232" ID-ZooBank="726DBD8C-38B1-4F43-8457-62F56BD9130D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738776724264" checkinUser="plazi" docAuthor="Sorokina, Vera S." docDate="2025" docId="03FA87F7FFC4FF0978EF994AFF76FB5B" docLanguage="en" docName="zootaxa.5584.1.1.pdf" docOrigin="Zootaxa 5584 (1)" docSource="https://doi.org/10.11646/zootaxa.5584.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spilogona ciliatocosta" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FFC3FF8FFFCEFF0278789C14FF82FF9A" masterDocTitle="New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="11" updateTime="1738777252473" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0842078D866479419F7B443D4FAAC458">New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species</mods:title>
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<treatment id="03FA87F7FFC4FF0978EF994AFF76FB5B" ID-DOI="http://doi.org/10.5281/zenodo.14812244" ID-Zenodo-Dep="14812244" LSID="urn:lsid:plazi:treatment:03FA87F7FFC4FF0978EF994AFF76FB5B" httpUri="http://treatment.plazi.org/id/03FA87F7FFC4FF0978EF994AFF76FB5B" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="C349656AFFC4FF0878EF994AFDE9FAE2" box="[151,619,1374,1401]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BEC36E1FFC4FF0878EF994AFDE9FAE2" blockId="10.[151,619,1374,1401]" box="[151,619,1374,1401]" pageId="10" pageNumber="11">
<heading id="D0A4818DFFC4FF0878EF994AFDE9FAE2" bold="true" box="[151,619,1374,1401]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C534D62FFC4FF0878EF994AFDE9FAE2" ID-CoL="4Z3TF" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,619,1374,1401]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0878EF994AFDE9FAE2" bold="true" box="[151,619,1374,1401]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878EF994AFE20FAE3" bold="true" box="[151,418,1374,1401]" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
(Schnabl, 1915)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC4FF0878EF99B7FE57F9CE" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BEC36E1FFC4FF0878EF99B7FBB0FA43" blockId="10.[151,1436,1443,1621]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4311CFFFC4FF0878EF99B7FBB0FA43" pageId="10" pageNumber="11">
<taxonomicName id="4C534D62FFC4FF0878EF99B7FDECFA21" authority="Schnabl" authorityName="Schnabl" box="[151,622,1443,1467]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta" subGenus="Spilogona">
<emphasis id="B927EAF3FFC4FF0878EF99B7FE96FA21" box="[151,276,1443,1467]" italics="true" pageId="10" pageNumber="11">Limnophora</emphasis>
(
<emphasis id="B927EAF3FFC4FF08795D99B7FE0EFA21" box="[293,396,1443,1467]" italics="true" pageId="10" pageNumber="11">Spilogona</emphasis>
)
<emphasis id="B927EAF3FFC4FF0879E599B7FD91FA21" box="[413,531,1443,1467]" italics="true" pageId="10" pageNumber="11">ciliatocosta</emphasis>
<materialsCitation id="3B3B3CBCFFC4FF087A6499B7FDECFA21" box="[540,622,1443,1467]" collectingDate="1909-07" collectorName="F. Zaitsev" country="Russia" county="Kara district" location="West Siberia" municipality="Yugorskiy" pageId="10" pageNumber="11" specimenCount="1">Schnabl</materialsCitation>
</taxonomicName>
in
<treatmentCitation id="0AF210F0FFC4FF087AEA99B7FC01FA21" author="Becker, T. &amp; Dziedzicki, H. &amp; Schnabl, J. &amp; Villeneuve, J." box="[658,899,1443,1467]" page="31" pageId="10" pageNumber="11" year="1915">
<bibRefCitation id="EFC24B10FFC4FF087AEA99B7FC01FA21" author="Becker, T. &amp; Dziedzicki, H. &amp; Schnabl, J. &amp; Villeneuve, J." box="[658,899,1443,1467]" pageId="10" pageNumber="11" pagination="1 - 67" refId="ref18056" refString="Becker, T., Dziedzicki, H., Schnabl, J. &amp; Villeneuve, J. (1915) Resultats scientifiques de l'Expedition des freres Kuznecov (Kouznetzov) a l'Oural Arctique en 1909, sous la direction de H. Backlund. Diptera. Zapiski Imperatorskoi Akademii Nauk (VIII), Fiziko-Matematicheskoe Otdeleniye, 28 (7), 1-67." type="journal article" year="1915">
Becker
<emphasis id="B927EAF3FFC4FF087A9A99B0FC95FA21" box="[738,791,1443,1467]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
, 1915: 31
</bibRefCitation>
</treatmentCitation>
. Type-locality: “in der Karskaja Tundra” [
<collectingCountry id="F3447671FFC4FF087D4499B7FA14FA21" box="[1340,1430,1443,1467]" name="Russia" pageId="10" pageNumber="11">RUSSIA</collectingCountry>
,
<location id="8E8C603AFFC4FF0878BF99D6FEC8FA40" LSID="urn:lsid:plazi:treatment:03FA87F7FFC4FF0978EF994AFF76FB5B:8E8C603AFFC4FF0878BF99D6FEC8FA40" box="[199,330,1474,1498]" country="Russia" county="Kara district" municipality="Yugorskiy" name="West Siberia" pageId="10" pageNumber="11">West Siberia</location>
,
<collectingMunicipality id="6B88AC9BFFC4FF08792B99D6FE39FA40" box="[339,443,1474,1498]" pageId="10" pageNumber="11">Yugorskiy</collectingMunicipality>
peninsula,
<collectingCounty id="628D4E6DFFC4FF087A5799D6FD33FA40" box="[559,689,1474,1498]" pageId="10" pageNumber="11">Kara district</collectingCounty>
;
<date id="FFED1021FFC4FF087AC399D6FC93FA40" box="[699,785,1474,1498]" pageId="10" pageNumber="11" value="1909-07">
<collectingDate id="EFA9E9C9FFC4FF087AC399D6FC93FA40" box="[699,785,1474,1498]" pageId="10" pageNumber="11" value="1909-07">vii.1909</collectingDate>
</date>
; collector
<collectorName id="26A65337FFC4FF087B0399D6FC62FA40" box="[891,992,1474,1498]" pageId="10" pageNumber="11">F. Zaitsev</collectorName>
] [seen].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878EF99F4FCDDF98D" blockId="10.[151,1436,1443,1621]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4311CFFFC4FF0878EF99F4FCDDF98D" pageId="10" pageNumber="11">
<treatmentCitation id="0AF210F0FFC4FF0878EF99F4FD9BFA63" author="Huckett, H. C." box="[151,537,1504,1529]" page="18" pageId="10" pageNumber="11" year="1965">
<taxonomicName id="4C534D62FFC4FF0878EF99F4FD9BFA63" ID-CoL="4Z3TQ" authority="Huckett, 1965: 18" authorityName="Huckett" authorityPageNumber="18" authorityYear="1965" box="[151,537,1504,1529]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF0878EF99F4FEDFFA62" box="[151,349,1504,1528]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
<materialsCitation id="3B3B3CBCFFC4FF08791A99F4FD9BFA63" box="[354,537,1504,1529]" collectingDate="1951-07-14" collectorName="J. G. Chillcott" country="Canada" location="Spence Bay" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Northwest Territories">
<bibRefCitation id="EFC24B10FFC4FF08791A99F4FD9BFA63" author="Huckett, H. C." box="[354,537,1504,1529]" pageId="10" pageNumber="11" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 18</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, 190, 201 (key), and 216. Type-locality: “
<location id="8E8C603AFFC4FF087BC399F5FBB4FA63" LSID="urn:lsid:plazi:treatment:03FA87F7FFC4FF0978EF994AFF76FB5B:8E8C603AFFC4FF087BC399F5FBB4FA63" box="[955,1078,1505,1529]" country="Canada" name="Spence Bay" pageId="10" pageNumber="11" stateProvince="Northwest Territories">Spence Bay</location>
,
<collectingRegion id="4997F803FFC4FF087C4799F4FA98FA62" box="[1087,1306,1504,1528]" country="Canada" name="Northwest Territories" pageId="10" pageNumber="11">Northwest Territories</collectingRegion>
, 7.14.51 [i.e.
<date id="FFED1021FFC4FF0878BF99EBFEBCF98D" box="[199,318,1535,1559]" pageId="10" pageNumber="11" value="1951-07-14">
<collectingDate id="EFA9E9C9FFC4FF0878BF99EBFEBCF98D" box="[199,318,1535,1559]" pageId="10" pageNumber="11" value="1951-07-14">14.vii.1951</collectingDate>
</date>
],
<collectorName id="26A65337FFC4FF08792899EBFE58F98D" box="[336,474,1535,1559]" pageId="10" pageNumber="11">J.G. Chillcott</collectorName>
” [
<collectingCountry id="F3447671FFC4FF08799799EBFDDCF98D" box="[495,606,1535,1559]" name="Canada" pageId="10" pageNumber="11">CANADA</collectingCountry>
, NWT] [seen].
<emphasis id="B927EAF3FFC4FF087A8699EBFCDDF98D" bold="true" box="[766,863,1535,1559]" pageId="10" pageNumber="11">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878EF9A0AFE57F9CE" blockId="10.[151,1436,1443,1621]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4311CFFFC4FF0878EF9A0AFE57F9CE" pageId="10" pageNumber="11">
<taxonomicName id="4C534D62FFC4FF0878EF9A0AFDA4F9AC" ID-CoL="4Z3TF" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,550,1566,1590]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0878EF9A0AFEF8F9AC" box="[151,378,1566,1590]" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
(Schnabl, 1915)
</taxonomicName>
;
<materialsCitation id="3B3B3CBCFFC4FF087A489A0AFE52F9CE" collectorName="Sorokina &amp; Thidrikh" location="Wrangel Island" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Chukotka">
<treatmentCitation id="0AF210F0FFC4FF087A489A0AFC8BF9AC" author="Sorokina, V. S." box="[560,777,1566,1590]" page="331" pageId="10" pageNumber="11" year="2012">
<bibRefCitation id="EFC24B10FFC4FF087A489A0AFC8BF9AC" author="Sorokina, V. S." box="[560,777,1566,1590]" pageId="10" pageNumber="11" pagination="328 - 332" refId="ref19142" refString="Sorokina, V. S. (2012 a) The Muscidae (Diptera) of the Russian tundra zones. Report 1. Caucasian Entomological Bulletin, 8 (2), 328-332. [in Russian, with English abstract]" type="journal article" year="2012">Sorokina, 2012a: 331</bibRefCitation>
</treatmentCitation>
(Taymyr Peninsula);
<collectorName id="26A65337FFC4FF087B9D9A0AFBC2F9AC" box="[997,1088,1566,1590]" pageId="10" pageNumber="11">Sorokina</collectorName>
&amp;
<collectorName id="26A65337FFC4FF087C189A0AFB39F9AC" box="[1120,1211,1566,1590]" pageId="10" pageNumber="11">Thidrikh</collectorName>
, 2021: 219 (
<collectingRegion id="4997F803FFC4FF087D409A0AFA1EF9AC" box="[1336,1436,1566,1590]" country="Russia" name="Chukotka" pageId="10" pageNumber="11">Chukotka</collectingRegion>
AO, incl.
<location id="8E8C603AFFC4FF0879539A28FE4BF9CE" LSID="urn:lsid:plazi:treatment:03FA87F7FFC4FF0978EF994AFF76FB5B:8E8C603AFFC4FF0879539A28FE4BF9CE" box="[299,457,1596,1620]" name="Wrangel Island" pageId="10" pageNumber="11" stateProvince="Chukotka">Wrangel Island</location>
)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC4FF0978EF9A97FEDCFCA3" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BEC36E1FFC4FF0978EF9A97FE80FE87" blockId="10.[151,1437,1667,2017]" lastBlockId="11.[151,1437,151,825]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878EF9A97FF58F907" bold="true" box="[151,218,1667,1693]" pageId="10" pageNumber="11">Notes</emphasis>
. The type material of
<taxonomicName id="4C534D62FFC4FF0879B59A97FD59F907" box="[461,731,1667,1693]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0879B59A97FD59F907" box="[461,731,1667,1693]" italics="true" pageId="10" pageNumber="11">Limnophora ciliatocosta</emphasis>
</taxonomicName>
(in
<collectionCode id="ED42AE24FFC4FF087B7E9A97FCC3F907" box="[774,833,1667,1693]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="11">ZISP</collectionCode>
) and
<taxonomicName id="4C534D62FFC4FF087B059A97FBD0F906" authorityName="Huckett" authorityYear="1965" box="[893,1106,1667,1693]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF087B059A97FBD0F906" box="[893,1106,1667,1693]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
</taxonomicName>
(in
<collectionCode id="ED42AE24FFC4FF087C059A97FB3AF907" box="[1149,1208,1667,1693]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="10" pageNumber="11">CNC</collectionCode>
) was examined. The descriptions of both species and the male terminalia are the same. However, the illustrations of the male terminalia of
<taxonomicName id="4C534D62FFC4FF0878CF9ADFFE2DF97F" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[183,431,1739,1765]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0878CF9ADFFE2DF97F" box="[183,431,1739,1765]" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC24B10FFC4FF0879A49ADFFD04F97F" author="Hennig, W." box="[476,646,1738,1765]" pageId="10" pageNumber="11" pagination="233 - 288" refId="ref18511" refString="Hennig, W. (1959 a) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 204. E. Schweizerbart, Stuttgart, pp. 233-288." type="book chapter" year="1959">Hennig (1959a</bibRefCitation>
: Taf. XV, Fig. 299) and of
<taxonomicName id="4C534D62FFC4FF087BCF9ADFFB0DF97E" authorityName="Huckett" authorityYear="1965" box="[951,1167,1739,1765]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF087BCF9ADFFB0DF97E" box="[951,1167,1739,1765]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC24B10FFC4FF087CC49ADFFA1AF97F" author="Huckett, H. C." box="[1212,1432,1738,1765]" pageId="10" pageNumber="11" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett (1965: 352</bibRefCitation>
, Fig. 165) are slightly different in the width of the surstylus. I examined many specimens of Palaearctic
<taxonomicName id="4C534D62FFC4FF087D549AFBFE95F8B7" baseAuthorityName="Schnabl" baseAuthorityYear="1915" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF087D549AFBFE95F8B7" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
</taxonomicName>
and Nearctic
<taxonomicName id="4C534D62FFC4FF0879CF9B07FD12F8B6" authorityName="Huckett" authorityYear="1965" box="[439,656,1811,1837]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF0879CF9B07FD12F8B6" box="[439,656,1811,1837]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
</taxonomicName>
and I have concluded that the width of the surstylus depends on the angle of viewing the terminalia and is variable. The cercal plate of these species is very distinctive because of the different length of right and left cerci (
<bibRefCitation id="EFC24B10FFC4FF087A339B4FFCABF8EF" author="Huckett, H. C." box="[587,809,1882,1909]" pageId="10" pageNumber="11" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 346</bibRefCitation>
, Fig. 96). Hennig in his monograph illustrated the male terminalia (
<bibRefCitation id="EFC24B10FFC4FF0879619B6BFE44F803" author="Hennig, W." box="[281,454,1918,1945]" pageId="10" pageNumber="11" pagination="233 - 288" refId="ref18511" refString="Hennig, W. (1959 a) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 204. E. Schweizerbart, Stuttgart, pp. 233-288." type="book chapter" year="1959">Hennig, 1959a</bibRefCitation>
: Taf. XIII, Fig. 276; 1959b: Taf. XVIII, Fig. 361) of a specimen from Wrangel Island named as “
<taxonomicName id="4C534D62FFC4FF08796C9BB7FDD8F827" authority="Frey" authorityName="Frey" authorityYear="1915" box="[276,602,1955,1981]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macropyga">
<emphasis id="B927EAF3FFC4FF08796C9BB7FD9CF826" box="[276,542,1955,1981]" italics="true" pageId="10" pageNumber="11">Limnophora macropyga</emphasis>
Frey
</taxonomicName>
” (det. A. Stackelberg). These figures correspond well with both
<taxonomicName id="4C534D62FFC4FF087D649BB7FA1EF827" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1308,1436,1955,1981]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF087D649BB7FA1EF827" box="[1308,1436,1955,1981]" italics="true" pageId="10" pageNumber="11">ciliatocosta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFC4FF0878BE9BD3FEA4F87A" authorityName="Huckett" authorityYear="1965" box="[198,294,1991,2016]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF0878BE9BD3FEA4F87A" box="[198,294,1991,2016]" italics="true" pageId="10" pageNumber="11">contigua</emphasis>
</taxonomicName>
. I examined this specimen and found that the terminalia is mounted on a permanent slide preparation, which is why it is slightly compressed, and the surstylus thus appears wider than that of the Hennig’s figure of the type of
<taxonomicName id="4C534D62FFC5FF0979609CAFFDAAFF4F" box="[280,552,187,213]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0979609CAFFDAAFF4F" box="[280,552,187,213]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
. Unfortunately, the terminalia of the
<typeStatus id="54E88843FFC5FF097BBC9CAFFB98FF4F" box="[964,1050,187,213]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF097C479CAFFACDFF4F" box="[1087,1359,187,213]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097C479CAFFACDFF4F" box="[1087,1359,187,213]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
is also mounted on a poorly prepared permanent mount, and it was impossible to describe the form and exact width of the surstylus.
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978BF9D33FEBEFE4B" blockId="11.[151,1437,151,825]" pageId="11" pageNumber="12">
Specimens of both sexes of these species are distinctive, with a completely black body, facial edge projecting beyond level of profrons, mid tibia with
<emphasis id="B927EAF3FFC5FF097A0B9D5FFDFDFEFE" box="[627,639,331,356]" italics="true" pageId="11" pageNumber="12">v</emphasis>
setae and hind femur without
<emphasis id="B927EAF3FFC5FF097B9F9D5FFB83FEFE" box="[999,1025,331,356]" italics="true" pageId="11" pageNumber="12">pv</emphasis>
setae, scutellum with downwardly-directed preapical setulae on upper border of declivities, shining prementum, and costa with costal spinules. I therefore conclude that
<taxonomicName id="4C534D62FFC5FF0979D39D87FCB2FE37" authority="Huckett, 1965" authorityName="Huckett" authorityYear="1965" box="[427,816,402,429]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC5FF0979D39D87FD04FE36" box="[427,646,403,429]" italics="true" pageId="11" pageNumber="12">Spilogona contigua</emphasis>
<bibRefCitation id="EFC24B10FFC5FF097AE89D87FCB2FE37" author="Huckett, H. C." box="[656,816,402,429]" pageId="11" pageNumber="12" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965</bibRefCitation>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="4C534D62FFC5FF097CF19D87FEBAFE4B" authority="Schnabl, 1915" authorityName="Schnabl" authorityYear="1915" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097CF19D87FA1EFE37" box="[1161,1436,403,429]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
Schnabl, 1915
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978BF9DCFFA93FD6B" blockId="11.[151,1437,151,825]" pageId="11" pageNumber="12">
In the
<collectionCode id="ED42AE24FFC5FF09796B9DCFFECEFE6F" box="[275,332,475,501]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="12">ZISP</collectionCode>
, apart from the male
<typeStatus id="54E88843FFC5FF097A3C9DCFFD19FE6F" box="[580,667,475,501]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF097ABD9DCFFC54FE6F" box="[709,982,475,501]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097ABD9DCFFC54FE6F" box="[709,982,475,501]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
there is a female named as
<taxonomicName id="4C534D62FFC5FF097D6C9DCFFE95FD83" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097D6C9DCFFE95FD83" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
. The locality is the same as the male (Karskaja Tundra). However, this specimen does not agree with the male
<typeStatus id="54E88843FFC5FF0978879E37FED4FDA7" box="[255,342,547,573]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF0979049E37FD0EFDA7" box="[380,652,547,573]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0979049E37FD0EFDA7" box="[380,652,547,573]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC24B10FFC5FF097AE29E37FCCAFDA7" author="Hennig, W." box="[666,840,546,573]" pageId="11" pageNumber="12" pagination="233 - 288" refId="ref18511" refString="Hennig, W. (1959 a) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 204. E. Schweizerbart, Stuttgart, pp. 233-288." type="book chapter" year="1959">Hennig (1959a)</bibRefCitation>
also had doubts about the identity of this female. The female
<typeStatus id="54E88843FFC5FF0978939E53FEC0FDFB" box="[235,322,583,609]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF0979109E53FDF5FDFB" box="[360,631,583,609]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0979109E53FDF5FDFB" box="[360,631,583,609]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
does not have the facial edge projecting beyond level of profrons, costa with very strong costal spines, mid tibia without
<emphasis id="B927EAF3FFC5FF097AB89E7FFD4EFD1E" box="[704,716,619,644]" italics="true" pageId="11" pageNumber="12">v</emphasis>
setae and hind femur with several
<emphasis id="B927EAF3FFC5FF097C209E7FFBF0FD1E" box="[1112,1138,619,644]" italics="true" pageId="11" pageNumber="12">pv</emphasis>
setae, proboscis short and thickened, prementum dusted, body with grey dust, not black, thorax grey with three distinct brown longitudinal stripes and with 2+3
<emphasis id="B927EAF3FFC5FF0979F89EA7FE18FD57" box="[384,410,691,717]" italics="true" pageId="11" pageNumber="12">dc</emphasis>
setae, abdomen wet, but with traces of paired brown spots, postgenital plate with hairs. This female is definitely a different species and the specimen runs to
<taxonomicName id="4C534D62FFC5FF097B1E9EC3FA8EFD6B" authority="(Ringdahl, 1926)" baseAuthorityName="Ringdahl" baseAuthorityYear="1926" box="[870,1292,727,753]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tornensis">
<emphasis id="B927EAF3FFC5FF097B1E9EC3FBC1FD6A" box="[870,1091,727,753]" italics="true" pageId="11" pageNumber="12">Spilogona tornensis</emphasis>
(Ringdahl, 1926)
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978BF9EEFFEDCFCA3" blockId="11.[151,1437,151,825]" pageId="11" pageNumber="12">
<taxonomicName id="4C534D62FFC5FF0978BF9EEFFE3DFC8F" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[199,447,763,789]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0978BF9EEFFE3DFC8F" box="[199,447,763,789]" italics="true" pageId="11" pageNumber="12">Spilogona ciliatocosta</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C534D62FFC5FF097AF29EEFFB91FC8F" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[650,1043,762,789]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tundrae">
<emphasis id="B927EAF3FFC5FF097AF29EEFFCDAFC8F" box="[650,856,763,789]" italics="true" pageId="11" pageNumber="12">Spilogona tundrae</emphasis>
(Schnabl, 1915)
</taxonomicName>
. These species can be separated as following couple:
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0978EF9F70FABFFBE3" pageId="11" pageNumber="12" type="key">
<keyStep id="30A72E44FFC5FF0978EF9F70FA1EFB91" pageId="11" pageNumber="12">
<key id="FFC261D0FFC5FF0978EF9F70FA1EFB91" pageId="11" pageNumber="12">
<paragraph id="8BEC36E1FFC5FF0978EF9F70FA1FFC29" blockId="11.[151,1437,867,1036]" pageId="11" pageNumber="12">
<keyLead id="30A295D4FFC5FF0978EF9F70FA1FFC29" pageId="11" pageNumber="12">
1 Larger species (
<quantity id="4CAB9B04FFC5FF0979029F77FE4CFCE3" box="[378,462,867,889]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="11" pageNumber="12" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
), costal setulae weak. Male: fore tibia with
<emphasis id="B927EAF3FFC5FF097B0A9F70FCFCFCE3" box="[882,894,868,889]" italics="true" pageId="11" pageNumber="12">p</emphasis>
seta in middle; fronto-orbital plates separated by a wide frontal vitta, which is as wide as width of outer margins of ocelli or wider; abdomen shining; anal segment swollen, when viewed in profile bulging towards the dorsal region. Female: abdomen blackish.............................
<taxonomicName id="4C534D62FFC5FF097D439F8BFA1FFC29" ID-CoL="4Z427" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1339,1437,926,948]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tundrae">
<emphasis id="B927EAF3FFC5FF097D439F8BFA1FFC29" box="[1339,1437,926,948]" italics="true" pageId="11" pageNumber="12">S. tundrae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978EF9FA8FA1EFB91" blockId="11.[151,1437,867,1036]" pageId="11" pageNumber="12">
<keyLead id="30A295D4FFC5FF0978EF9FA8FA1EFB91" pageId="11" pageNumber="12">
- Smaller species (
<quantity id="4CAB9B04FFC5FF0979FA9FAFFE65FC4B" box="[386,487,955,977]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="11" pageNumber="12" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
), costal setulae coarsened proximad of costal spine. Male: fore tibia without
<emphasis id="B927EAF3FFC5FF097CC19FA8FB47FC4B" box="[1209,1221,956,977]" italics="true" pageId="11" pageNumber="12">p</emphasis>
seta in middle; fronto-orbital plates touching or separated by a narrow frontal vitta; abdomen subshining; anal segment flattish or appressed dorsad, not bulging toward the dorsal region. Female: abdomen bronzy.......................................
<taxonomicName id="4C534D62FFC5FF097D6F9FE3FA1EFB91" ID-CoL="4Z3TF" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1303,1436,1014,1036]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097D6F9FE3FA1EFB91" box="[1303,1436,1014,1036]" italics="true" pageId="11" pageNumber="12">S. ciliatocosta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
<paragraph id="8BEC36E1FFC5FF0978EF982FFABFFBE3" blockId="11.[151,1437,1083,1217]" pageId="11" pageNumber="12">
All of the characters given above for
<taxonomicName id="4C534D62FFC5FF097A20982FFCD0FBCF" ID-CoL="4Z3TF" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[600,850,1083,1109]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097A20982FFCD0FBCF" box="[600,850,1083,1109]" italics="true" pageId="11" pageNumber="12">Spilogona ciliatocosta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFC5FF097BE8982FFBE3FBCF" ID-CoL="4Z427" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[912,1121,1083,1109]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tundrae">
<emphasis id="B927EAF3FFC5FF097BE8982FFBE3FBCF" box="[912,1121,1083,1109]" italics="true" pageId="11" pageNumber="12">Spilogona tundrae</emphasis>
</taxonomicName>
are variable, but are more commonly as stated in the couplet. The male terminalia provides the most reliable identification character.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0978BF9896FF76FB5B" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BEC36E1FFC5FF0978BF9896FF76FB5B" blockId="11.[151,1437,1083,1217]" pageId="11" pageNumber="12">
<emphasis id="B927EAF3FFC5FF0978BF9896FEE3FB06" bold="true" box="[199,353,1154,1180]" pageId="11" pageNumber="12">Distribution.</emphasis>
PALAEARCTIC:
<collectingCountry id="F3447671FFC5FF097A469897FD0BFB07" box="[574,649,1155,1181]" name="Russia" pageId="11" pageNumber="12">Russia</collectingCountry>
(Arctic Siberia and Far East). NEARCTIC: Alaska, Yukon, NWT, Quebec.
</paragraph>
</subSubSection>
</treatment>
</document>
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