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<document id="1274A742FB76A16E7E42E99C7E36EFEE" ID-DOI="10.11646/zootaxa.5584.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="14812374" ID-ZooBank="8F289D1B-CED1-4506-B31B-467A59B36A10" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738777092588" checkinUser="plazi" docAuthor="Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E. &amp; Bahder, Brian W." docDate="2025" docId="0176194EFFB4C23BFF4948F4FE3B0683" docLanguage="en" docName="zootaxa.5584.1.7.pdf" docOrigin="Zootaxa 5584 (1)" docSource="https://doi.org/10.11646/zootaxa.5584.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agoo keili Bahder &amp; Bartlett 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="119" masterDocId="FD4F6136FFB7C23DFFDE4E14FF95004F" masterDocTitle="Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica" masterLastPageNumber="125" masterPageNumber="113" pageNumber="116" updateTime="1738777566703" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="876DB683E18629452D483384DCBD8CD7">Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica</mods:title>
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<mods:namePart id="140A95B1490D13799A187B40CFE82B75">Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:affiliation id="374F537FBE8CE660072271E9E5818ADD">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica</mods:affiliation>
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<mods:namePart id="7C8D56125E49A33B939B6469B77992B0">Zumbado Echavarria, Marco A.</mods:namePart>
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<mods:namePart id="D4BC17D090A49F4B6132C86E4C6A3EF9">Bartlett, Charles R.</mods:namePart>
<mods:affiliation id="BB408C94929AE7EB5510C028BF2A7094">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA</mods:affiliation>
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<mods:namePart id="C66ADE1FD2F8FF381B583B9398812493">Helmick, Ericka E.</mods:namePart>
<mods:affiliation id="DEFE24E7A52372D0CF1325E73AA06555">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
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<mods:namePart id="2AC5CC4B919A6C5223F152041704A3E9">Bahder, Brian W.</mods:namePart>
<mods:affiliation id="CDB18170B2D4E98BB688841880225028">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
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<mods:date id="0FDAB55217CB9A2957A29C192808D83F">2025</mods:date>
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<mods:number id="BA2648B30A3C005139B6F47D40C4BA7F">2025-02-05</mods:number>
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<treatment id="0176194EFFB4C23BFF4948F4FE3B0683" ID-DOI="http://doi.org/10.5281/zenodo.14812370" ID-Zenodo-Dep="14812370" LSID="urn:lsid:plazi:treatment:0176194EFFB4C23BFF4948F4FE3B0683" httpUri="http://treatment.plazi.org/id/0176194EFFB4C23BFF4948F4FE3B0683" lastPageId="6" lastPageNumber="119" pageId="3" pageNumber="116">
<subSubSection id="C1C5FBD3FFB4C23EFF4948F4FDF106B5" box="[151,612,1759,1786]" pageId="3" pageNumber="116" type="nomenclature">
<paragraph id="8960A858FFB4C23EFF4948F4FDF106B5" blockId="3.[151,612,1759,1822]" box="[151,612,1759,1786]" pageId="3" pageNumber="116">
<heading id="D2281F34FFB4C23EFF4948F4FDF106B5" bold="true" box="[151,612,1759,1786]" fontSize="11" level="1" pageId="3" pageNumber="116" reason="1">
<emphasis id="BBAB744AFFB4C23EFF4948F4FDF106B5" bold="true" box="[151,612,1759,1786]" pageId="3" pageNumber="116">
<taxonomicName id="4EDFD3DBFFB4C23EFF4948F4FE6A06B5" authority="Bahder &amp; Bartlett" authorityName="Bahder &amp; Bartlett" authorityYear="2025" box="[151,511,1759,1786]" class="Insecta" family="Derbidae" genus="Agoo" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="keili" status="sp. nov.">
<emphasis id="BBAB744AFFB4C23EFF4948F4FE8706B5" bold="true" box="[151,274,1759,1786]" italics="true" pageId="3" pageNumber="116">Agoo keili</emphasis>
Bahder &amp; Bartlett
</taxonomicName>
<taxonomicNameLabel id="A098C931FFB4C23EFDD848F4FDF106B5" box="[518,612,1760,1786]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1C5FBD3FFB4C23EFF494910FEBA0751" box="[151,303,1796,1822]" pageId="3" pageNumber="116" type="description">
<paragraph id="8960A858FFB4C23EFF494910FEBA0751" blockId="3.[151,612,1759,1822]" box="[151,303,1796,1822]" pageId="3" pageNumber="116">
(
<figureCitation id="11E4B4DDFFB4C23EFF414910FEB20751" box="[159,295,1796,1822]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="4.[152,255,712,736]" captionStartId-1="5.[152,255,1228,1252]" captionStartId-2="6.[152,255,1284,1308]" captionStartId-3="7.[152,253,1104,1128]" captionStartId-4="7.[152,255,1951,1975]" captionTargetBox-0="[151,1435,234,686]" captionTargetBox-1="[177,1410,415,1195]" captionTargetBox-2="[151,1432,179,1251]" captionTargetBox-3="[152,1436,179,1079]" captionTargetBox-4="[151,1424,1152,1922]" captionTargetId-0="figure-51@4.[151,1435,234,686]" captionTargetId-1="figure-108@5.[170,1416,414,1202]" captionTargetId-2="figure-24@6.[151,1436,179,1258]" captionTargetId-3="figure-18@7.[151,1436,179,1079]" captionTargetId-4="figure-48@7.[151,1435,1151,1926]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="7" captionText-0="FIGURE 2. Adult male habitus of Agoo keili sp. nov.; A) pale form in lateral view, B) yellow form in lateral view, C) pale form in dorsal view and D) yellow form in dorsal view. Scale bar = 1 mm." captionText-1="FIGURE 3. Adult male Agoo keili sp. nov., pale form top row and yellow form bottom row;A) head, pronotum and mesonotum dorsal view, B) head, pronotum and mesonotum lateral view and C) head frontal view. Scale bar = 1 mm." captionText-2="FIGURE 4. Adult male Agoo keili sp. nov. forewing; A) pale form and B) yellow form, C) line art; black text = vein, italics = crossvein, and green text = cell. Scale bar = 1 mm." captionText-3="FIGURE 5. Adult male Agoo keili sp. nov. terminalia; A) lateral view, B) ventral view and C) dorsal view. Scale bar = 0.5 mm." captionText-4="FIGURE 6. Agoo keili sp. nov. aedeagus; A) right lateral view, B) left lateral view, C) dorsal view and D) ventral view. Scale bar = 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14812378" figureDoi-1="http://doi.org/10.5281/zenodo.14812380" figureDoi-2="http://doi.org/10.5281/zenodo.14812382" figureDoi-3="http://doi.org/10.5281/zenodo.14812384" figureDoi-4="http://doi.org/10.5281/zenodo.14812386" httpUri-0="https://zenodo.org/record/14812378/files/figure.png" httpUri-1="https://zenodo.org/record/14812380/files/figure.png" httpUri-2="https://zenodo.org/record/14812382/files/figure.png" httpUri-3="https://zenodo.org/record/14812384/files/figure.png" httpUri-4="https://zenodo.org/record/14812386/files/figure.png" pageId="3" pageNumber="116">Figures 2–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1C5FBD3FFB4C23EFF494958FCFF0729" box="[151,874,1868,1894]" pageId="3" pageNumber="116" type="materials_examined">
<paragraph id="8960A858FFB4C23EFF494958FCFF0729" blockId="3.[151,1436,1868,2038]" box="[151,874,1868,1894]" pageId="3" pageNumber="116">
<emphasis id="BBAB744AFFB4C23EFF494958FED50729" bold="true" box="[151,320,1868,1894]" pageId="3" pageNumber="116">Type Locality.</emphasis>
<collectingCountry id="F1C8E8C8FFB4C23EFE994958FE510729" box="[327,452,1868,1894]" name="Costa Rica" pageId="3" pageNumber="116">Costa Rica</collectingCountry>
,
<collectingRegion id="4B1B66BAFFB4C23EFE114958FD220729" box="[463,695,1868,1894]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="116">Puntarenas Province</collectingRegion>
, Osa Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="C1C5FBD3FFB4C239FF194964FC0E009A" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116" type="diagnosis">
<paragraph id="8960A858FFB4C239FF194964FC0E009A" blockId="3.[151,1436,1868,2038]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116">
<emphasis id="BBAB744AFFB4C23EFF194964FED707C5" bold="true" box="[199,322,1904,1930]" pageId="3" pageNumber="116">Diagnosis.</emphasis>
Moderate sized
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, body pallid to yellow, wings pale translucent with darker yellow band along trailing margin. Medioventral process of pygofer broadly triangular, wider than tall. Processes on inner margin of gonostyli in ventral view roughly quadrate, arising in distal half. Aedeagus bearing large, subtriangular processes on dorsolateral margins of shaft. Endosoma with four pairs of processes, bearing triangular projection on dorsal margin near apex and large pair of processes curved cephalad from apex.Anal tube broadly triangular, apex caudally projected (not downcurved), apex strongly bilobed from dorsal view.
</paragraph>
</subSubSection>
<caption id="DDA0F8D0FFB3C239FF464CDCFCC4034B" ID-DOI="http://doi.org/10.5281/zenodo.14812378" ID-Zenodo-Dep="14812378" httpUri="https://zenodo.org/record/14812378/files/figure.png" pageId="4" pageNumber="117" startId="4.[152,255,712,736]" targetBox="[151,1435,234,686]" targetPageId="4" targetType="figure">
<paragraph id="8960A858FFB3C239FF464CDCFCC4034B" blockId="4.[151,1436,712,772]" pageId="4" pageNumber="117">
<emphasis id="BBAB744AFFB3C239FF464CDCFE8202AF" bold="true" box="[152,279,712,736]" pageId="4" pageNumber="117">FIGURE 2.</emphasis>
Adult male habitus of
<taxonomicName id="4EDFD3DBFFB3C239FE224CDDFDF602AF" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2025" box="[508,611,712,736]" class="Insecta" family="Derbidae" genus="Agoo" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="keili" status="sp. nov.">
<emphasis id="BBAB744AFFB3C239FE224CDDFDF602AF" box="[508,611,712,736]" italics="true" pageId="4" pageNumber="117">Agoo keili</emphasis>
</taxonomicName>
<emphasis id="BBAB744AFFB3C239FDB64CDCFD2D02AF" bold="true" box="[616,696,712,736]" pageId="4" pageNumber="117">
<taxonomicNameLabel id="A098C931FFB3C239FDB64CDCFD2D02AF" box="[616,696,712,736]" pageId="4" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; A) pale form in lateral view, B) yellow form in lateral view, C) pale form in dorsal view and D) yellow form in dorsal view. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C1C5FBD3FFB3C23BFF194D42FC9105E3" lastPageId="6" lastPageNumber="119" pageId="4" pageNumber="117" type="description">
<paragraph id="8960A858FFB3C239FF194D42FA8A03F7" blockId="4.[151,1436,854,1277]" pageId="4" pageNumber="117">
<emphasis id="BBAB744AFFB3C239FF194D42FECC033F" bold="true" box="[199,345,854,880]" pageId="4" pageNumber="117">Description.</emphasis>
<emphasis id="BBAB744AFFB3C239FEBE4D42FE30033F" box="[352,421,854,880]" italics="true" pageId="4" pageNumber="117">Color.</emphasis>
Body pallid (nearly white) to yellow (
<figureCitation id="11E4B4DDFFB3C239FC8D4D42FC0D033E" box="[851,920,854,881]" captionStart="FIGURE 2" captionStartId="4.[152,255,712,736]" captionTargetBox="[151,1435,234,686]" captionTargetId="figure-51@4.[151,1435,234,686]" captionTargetPageId="4" captionText="FIGURE 2. Adult male habitus of Agoo keili sp. nov.; A) pale form in lateral view, B) yellow form in lateral view, C) pale form in dorsal view and D) yellow form in dorsal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812378" httpUri="https://zenodo.org/record/14812378/files/figure.png" pageId="4" pageNumber="117">Fig. 2</figureCitation>
), darker dorsally; mesonotum white to yellow (when yellow, pale on lateral margins, near tegula); legs pale or yellow and pale proximally (coxae to midlength of femur white). Forewings translucent, mostly clear to strongly washed with yellow (especially dorsally).
</paragraph>
<paragraph id="8960A858FFB3C239FF194DD7FAA504B3" blockId="4.[151,1436,854,1277]" pageId="4" pageNumber="117">
<emphasis id="BBAB744AFFB3C239FF194DD7FEBB0392" box="[199,302,963,989]" italics="true" pageId="4" pageNumber="117">Structure</emphasis>
. Body length: males with wings
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, body length (without wings)
<quantity id="4E2705BDFFB3C239FBA94DD7FA940392" box="[1143,1281,963,989]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="4" pageNumber="117" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
; females with wings
<quantity id="4E2705BDFFB3C239FF3F4DF2FEFF044E" box="[225,362,998,1025]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.6" metricValueMin="7.2" pageId="4" pageNumber="117" unit="mm" value="7.4" valueMax="7.6" valueMin="7.2">7.2–7.6 mm</quantity>
, body length
<quantity id="4E2705BDFFB3C239FDDC4DF2FD1C044E" box="[514,649,998,1025]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="4" pageNumber="117" unit="mm" value="3.4" valueMax="3.6" valueMin="3.2">3.2–3.6 mm</quantity>
(
<tableCitation id="C45D9DE3FFB3C239FD494DF2FD7B044E" box="[663,750,998,1025]" captionStart="TABLE 3" captionStartId="4.[152,244,1324,1348]" captionText="TABLE 3. Biometric data for Agoo keili sp. nov." pageId="4" pageNumber="117">Table 3</tableCitation>
). Head. Head narrower than pronotum. Vertex in dorsal view triangular, widest posteriorly; lateral margins foliate, bearing single row of sensorial pits, disc depressed, median carina absent; anterior margin narrow, concave (apical transverse carina absent); posterior margin quadrately concave (
<figureCitation id="11E4B4DDFFB3C239FEDF4A46FECC0422" box="[257,345,1106,1133]" captionStart="FIGURE 3" captionStartId="5.[152,255,1228,1252]" captionTargetBox="[177,1410,415,1195]" captionTargetId="figure-108@5.[170,1416,414,1202]" captionTargetPageId="5" captionText="FIGURE 3. Adult male Agoo keili sp. nov., pale form top row and yellow form bottom row;A) head, pronotum and mesonotum dorsal view, B) head, pronotum and mesonotum lateral view and C) head frontal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812380" httpUri="https://zenodo.org/record/14812380/files/figure.png" pageId="4" pageNumber="117">Fig. 3A</figureCitation>
); in lateral view, profile smoothly rounded from posterior margin of vertex to frontoclypeal sutures (
<figureCitation id="11E4B4DDFFB3C239FF414A62FF6204DE" box="[159,247,1142,1169]" captionStart="FIGURE 3" captionStartId="5.[152,255,1228,1252]" captionTargetBox="[177,1410,415,1195]" captionTargetId="figure-108@5.[170,1416,414,1202]" captionTargetPageId="5" captionText="FIGURE 3. Adult male Agoo keili sp. nov., pale form top row and yellow form bottom row;A) head, pronotum and mesonotum dorsal view, B) head, pronotum and mesonotum lateral view and C) head frontal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812380" httpUri="https://zenodo.org/record/14812380/files/figure.png" pageId="4" pageNumber="117">Fig. 3B</figureCitation>
). Frons in frontal view with lateral margins foliate and slightly sinuate, bearing single row of sensorial pits (
<figureCitation id="11E4B4DDFFB3C239FF414A8EFF6204FA" box="[159,247,1178,1205]" captionStart="FIGURE 3" captionStartId="5.[152,255,1228,1252]" captionTargetBox="[177,1410,415,1195]" captionTargetId="figure-108@5.[170,1416,414,1202]" captionTargetPageId="5" captionText="FIGURE 3. Adult male Agoo keili sp. nov., pale form top row and yellow form bottom row;A) head, pronotum and mesonotum dorsal view, B) head, pronotum and mesonotum lateral view and C) head frontal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812380" httpUri="https://zenodo.org/record/14812380/files/figure.png" pageId="4" pageNumber="117">Fig. 3C</figureCitation>
), disc depressed median carina absent. Clypeus elongate-triangular. Lateral margins carinate (not foliate), bearing distinct median carina. Eye hemispherical, emarginated at antenna. Lateral ocelli distinct below eye near midline. Antennal scape short, hidden, pedicel ovate mearing many irregularly arranged sensory plaques.
</paragraph>
<caption id="DDA0F8D0FFB3C239FF464B38FD3C050B" box="[152,681,1324,1348]" pageId="4" pageNumber="117" startId="4.[152,244,1324,1348]" targetType="table">
<paragraph id="8960A858FFB3C239FF464B38FD3C050B" blockId="4.[152,681,1324,1348]" box="[152,681,1324,1348]" pageId="4" pageNumber="117">
<emphasis id="BBAB744AFFB3C239FF464B38FE9A050B" bold="true" box="[152,271,1324,1348]" pageId="4" pageNumber="117">TABLE 3.</emphasis>
Biometric data for
<taxonomicName id="4EDFD3DBFFB3C239FE3E4B39FDD8050B" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2025" box="[480,589,1324,1348]" class="Insecta" family="Derbidae" genus="Agoo" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="keili" status="sp. nov.">
<emphasis id="BBAB744AFFB3C239FE3E4B39FDD8050B" box="[480,589,1324,1348]" italics="true" pageId="4" pageNumber="117">Agoo keili</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A098C931FFB3C239FD8D4B38FD3C050B" box="[595,681,1324,1348]" pageId="4" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8960A858FFB3C239FD4D4B41FAB907AF" pageId="4" pageNumber="117">
<table id="FBDF5AF8FFB33DC2FF414B41FACE07AF" box="[159,1371,1365,2016]" gridcols="5" gridrows="17" pageId="4" pageNumber="117">
<tr id="37EFAA1AFFB33DC2FF414B41FACE0524" box="[159,1371,1365,1387]" gridrow="0" pageId="4" pageNumber="117" rowspan-0="1" rowspan-2="1" rowspan-4="1">
<th id="743EC366FFB33DC2FD4D4B41FC970524" box="[659,770,1365,1387]" gridcol="1" gridrow="0" pageId="4" pageNumber="117">
Male,
<emphasis id="BBAB744AFFB3C239FD0E4B42FD490524" box="[720,732,1366,1387]" italics="true" pageId="4" pageNumber="117">n</emphasis>
=12
</th>
<th id="743EC366FFB33DC2FBC14B41FB0F0524" box="[1055,1178,1365,1387]" gridcol="3" gridrow="0" pageId="4" pageNumber="117">
Female,
<emphasis id="BBAB744AFFB3C239FBAC4B42FB1B0524" box="[1138,1166,1366,1387]" italics="true" pageId="4" pageNumber="117">n=</emphasis>
4
</th>
</tr>
<tr id="37EFAA1AFFB33DC2FF414B69FACE05DC" box="[159,1371,1405,1427]" gridrow="1" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414B69FE3D05DC" box="[159,424,1405,1427]" gridcol="0" gridrow="1" pageId="4" pageNumber="117">Character</th>
<td id="743EC366FFB33DC2FD4D4B69FC9705DC" box="[659,770,1405,1427]" gridcol="1" gridrow="1" pageId="4" pageNumber="117">Range</td>
<td id="743EC366FFB33DC2FC814B69FC4D05DC" box="[863,984,1405,1427]" gridcol="2" gridrow="1" pageId="4" pageNumber="117">Average±SE</td>
<td id="743EC366FFB33DC2FBC14B69FB0F05DC" box="[1055,1178,1405,1427]" gridcol="3" gridrow="1" pageId="4" pageNumber="117">Range</td>
<td id="743EC366FFB33DC2FB3C4B69FACE05DC" box="[1250,1371,1405,1427]" gridcol="4" gridrow="1" pageId="4" pageNumber="117">Average±SE</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414BB0FACE05F5" box="[159,1371,1444,1466]" gridrow="2" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414BB0FE3D05F5" box="[159,424,1444,1466]" gridcol="0" gridrow="2" pageId="4" pageNumber="117">Body length, with wings</th>
<td id="743EC366FFB33DC2FD4D4BB0FC9705F5" box="[659,770,1444,1466]" gridcol="1" gridrow="2" pageId="4" pageNumber="117">6.7–7.2</td>
<td id="743EC366FFB33DC2FC814BB0FC4D05F5" box="[863,984,1444,1466]" gridcol="2" gridrow="2" pageId="4" pageNumber="117">6.9±0.2</td>
<td id="743EC366FFB33DC2FBC14BB0FB0F05F5" box="[1055,1178,1444,1466]" gridcol="3" gridrow="2" pageId="4" pageNumber="117">7.2–7.6</td>
<td id="743EC366FFB33DC2FB3C4BB0FACE05F5" box="[1250,1371,1444,1466]" gridcol="4" gridrow="2" pageId="4" pageNumber="117">7.4±0.2</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414BDFFACE05AE" box="[159,1371,1483,1505]" gridrow="3" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414BDFFE3D05AE" box="[159,424,1483,1505]" gridcol="0" gridrow="3" pageId="4" pageNumber="117">Body length, no wings</th>
<td id="743EC366FFB33DC2FD4D4BDFFC9705AE" box="[659,770,1483,1505]" gridcol="1" gridrow="3" pageId="4" pageNumber="117">2.8–3.2</td>
<td id="743EC366FFB33DC2FC814BDFFC4D05AE" box="[863,984,1483,1505]" gridcol="2" gridrow="3" pageId="4" pageNumber="117">3.0±0.2</td>
<td id="743EC366FFB33DC2FBC14BDFFB0F05AE" box="[1055,1178,1483,1505]" gridcol="3" gridrow="3" pageId="4" pageNumber="117">3.2–3.6</td>
<td id="743EC366FFB33DC2FB3C4BDFFACE05AE" box="[1250,1371,1483,1505]" gridcol="4" gridrow="3" pageId="4" pageNumber="117">3.4±0.2</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414BE7FACE0646" box="[159,1371,1523,1545]" gridrow="4" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414BE7FE3D0646" box="[159,424,1523,1545]" gridcol="0" gridrow="4" pageId="4" pageNumber="117">Forewing length</th>
<td id="743EC366FFB33DC2FD4D4BE7FC970646" box="[659,770,1523,1545]" gridcol="1" gridrow="4" pageId="4" pageNumber="117">4.8–5.2</td>
<td id="743EC366FFB33DC2FC814BE7FC4D0646" box="[863,984,1523,1545]" gridcol="2" gridrow="4" pageId="4" pageNumber="117">5.0±0.2</td>
<td id="743EC366FFB33DC2FBC14BE7FB0F0646" box="[1055,1178,1523,1545]" gridcol="3" gridrow="4" pageId="4" pageNumber="117">5.2–5.7</td>
<td id="743EC366FFB33DC2FB3C4BE7FACE0646" box="[1250,1371,1523,1545]" gridcol="4" gridrow="4" pageId="4" pageNumber="117">5.5±0.2</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF41480EFACE067F" box="[159,1371,1562,1584]" gridrow="5" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF41480EFE3D067F" box="[159,424,1562,1584]" gridcol="0" gridrow="5" pageId="4" pageNumber="117">Vertex length</th>
<td id="743EC366FFB33DC2FD4D480EFC97067F" box="[659,770,1562,1584]" gridcol="1" gridrow="5" pageId="4" pageNumber="117">0.2–0.3</td>
<td id="743EC366FFB33DC2FC81480EFC4D067F" box="[863,984,1562,1584]" gridcol="2" gridrow="5" pageId="4" pageNumber="117">0.3±0.1</td>
<td id="743EC366FFB33DC2FBC1480EFB0F067F" box="[1055,1178,1562,1584]" gridcol="3" gridrow="5" pageId="4" pageNumber="117">0.3–0.4</td>
<td id="743EC366FFB33DC2FB3C480EFACE067F" box="[1250,1371,1562,1584]" gridcol="4" gridrow="5" pageId="4" pageNumber="117">0.4±0.1</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414855FACE0618" box="[159,1371,1601,1623]" gridrow="6" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414855FE3D0618" box="[159,424,1601,1623]" gridcol="0" gridrow="6" pageId="4" pageNumber="117">Vertex width, basal margin</th>
<td id="743EC366FFB33DC2FD4D4855FC970618" box="[659,770,1601,1623]" gridcol="1" gridrow="6" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FC814855FC4D0618" box="[863,984,1601,1623]" gridcol="2" gridrow="6" pageId="4" pageNumber="117">0.2±0.0</td>
<td id="743EC366FFB33DC2FBC14855FB0F0618" box="[1055,1178,1601,1623]" gridcol="3" gridrow="6" pageId="4" pageNumber="117">0.3–0.3</td>
<td id="743EC366FFB33DC2FB3C4855FACE0618" box="[1250,1371,1601,1623]" gridcol="4" gridrow="6" pageId="4" pageNumber="117">0.3±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF41487DFACE0630" box="[159,1371,1641,1663]" gridrow="7" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF41487DFE3D0630" box="[159,424,1641,1663]" gridcol="0" gridrow="7" pageId="4" pageNumber="117">Vertex width, distal margin</th>
<td id="743EC366FFB33DC2FD4D487DFC970630" box="[659,770,1641,1663]" gridcol="1" gridrow="7" pageId="4" pageNumber="117">0.1–0.1</td>
<td id="743EC366FFB33DC2FC81487DFC4D0630" box="[863,984,1641,1663]" gridcol="2" gridrow="7" pageId="4" pageNumber="117">0.1±0.0</td>
<td id="743EC366FFB33DC2FBC1487DFB0F0630" box="[1055,1178,1641,1663]" gridcol="3" gridrow="7" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FB3C487DFACE0630" box="[1250,1371,1641,1663]" gridcol="4" gridrow="7" pageId="4" pageNumber="117">0.2±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414884FACE06E9" box="[159,1371,1680,1702]" gridrow="8" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414884FE3D06E9" box="[159,424,1680,1702]" gridcol="0" gridrow="8" pageId="4" pageNumber="117">Pronotum length, midline</th>
<td id="743EC366FFB33DC2FD4D4884FC9706E9" box="[659,770,1680,1702]" gridcol="1" gridrow="8" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FC814884FC4D06E9" box="[863,984,1680,1702]" gridcol="2" gridrow="8" pageId="4" pageNumber="117">0.2±0.0</td>
<td id="743EC366FFB33DC2FBC14884FB0F06E9" box="[1055,1178,1680,1702]" gridcol="3" gridrow="8" pageId="4" pageNumber="117">0.3–0.3</td>
<td id="743EC366FFB33DC2FB3C4884FACE06E9" box="[1250,1371,1680,1702]" gridcol="4" gridrow="8" pageId="4" pageNumber="117">0.3±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF4148A3FACE0682" box="[159,1371,1719,1741]" gridrow="9" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF4148A3FE3D0682" box="[159,424,1719,1741]" gridcol="0" gridrow="9" pageId="4" pageNumber="117">Mesonotum length, midline</th>
<td id="743EC366FFB33DC2FD4D48A3FC970682" box="[659,770,1719,1741]" gridcol="1" gridrow="9" pageId="4" pageNumber="117">0.6–0.6</td>
<td id="743EC366FFB33DC2FC8148A3FC4D0682" box="[863,984,1719,1741]" gridcol="2" gridrow="9" pageId="4" pageNumber="117">0.6±0.0</td>
<td id="743EC366FFB33DC2FBC148A3FB0F0682" box="[1055,1178,1719,1741]" gridcol="3" gridrow="9" pageId="4" pageNumber="117">0.7–0.7</td>
<td id="743EC366FFB33DC2FB3C48A3FACE0682" box="[1250,1371,1719,1741]" gridcol="4" gridrow="9" pageId="4" pageNumber="117">0.7±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF4148CBFACE06BA" box="[159,1371,1759,1781]" gridrow="10" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF4148CBFE3D06BA" box="[159,424,1759,1781]" gridcol="0" gridrow="10" pageId="4" pageNumber="117">Mesonotum width</th>
<td id="743EC366FFB33DC2FD4D48CBFC9706BA" box="[659,770,1759,1781]" gridcol="1" gridrow="10" pageId="4" pageNumber="117">0.9–0.9</td>
<td id="743EC366FFB33DC2FC8148CBFC4D06BA" box="[863,984,1759,1781]" gridcol="2" gridrow="10" pageId="4" pageNumber="117">0.9±0.0</td>
<td id="743EC366FFB33DC2FBC148CBFB0F06BA" box="[1055,1178,1759,1781]" gridcol="3" gridrow="10" pageId="4" pageNumber="117">1.0–1.0</td>
<td id="743EC366FFB33DC2FB3C48CBFACE06BA" box="[1250,1371,1759,1781]" gridcol="4" gridrow="10" pageId="4" pageNumber="117">1.0±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414912FACE0753" box="[159,1371,1798,1820]" gridrow="11" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414912FE3D0753" box="[159,424,1798,1820]" gridcol="0" gridrow="11" pageId="4" pageNumber="117">Frons width, dorsal margin</th>
<td id="743EC366FFB33DC2FD4D4912FC970753" box="[659,770,1798,1820]" gridcol="1" gridrow="11" pageId="4" pageNumber="117">0.1–0.1</td>
<td id="743EC366FFB33DC2FC814912FC4D0753" box="[863,984,1798,1820]" gridcol="2" gridrow="11" pageId="4" pageNumber="117">0.1±0.0</td>
<td id="743EC366FFB33DC2FBC14912FB0F0753" box="[1055,1178,1798,1820]" gridcol="3" gridrow="11" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FB3C4912FACE0753" box="[1250,1371,1798,1820]" gridcol="4" gridrow="11" pageId="4" pageNumber="117">0.2±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414939FACE070C" box="[159,1371,1837,1859]" gridrow="12" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414939FE3D070C" box="[159,424,1837,1859]" gridcol="0" gridrow="12" pageId="4" pageNumber="117">Frons width, clypeal suture</th>
<td id="743EC366FFB33DC2FD4D4939FC97070C" box="[659,770,1837,1859]" gridcol="1" gridrow="12" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FC814939FC4D070C" box="[863,984,1837,1859]" gridcol="2" gridrow="12" pageId="4" pageNumber="117">0.2±0.0</td>
<td id="743EC366FFB33DC2FBC14939FB0F070C" box="[1055,1178,1837,1859]" gridcol="3" gridrow="12" pageId="4" pageNumber="117">0.3–0.3</td>
<td id="743EC366FFB33DC2FB3C4939FACE070C" box="[1250,1371,1837,1859]" gridcol="4" gridrow="12" pageId="4" pageNumber="117">0.3±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414941FACE0724" box="[159,1371,1877,1899]" gridrow="13" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414941FE3D0724" box="[159,424,1877,1899]" gridcol="0" gridrow="13" pageId="4" pageNumber="117">Frons width, widest</th>
<td id="743EC366FFB33DC2FD4D4941FC970724" box="[659,770,1877,1899]" gridcol="1" gridrow="13" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FC814941FC4D0724" box="[863,984,1877,1899]" gridcol="2" gridrow="13" pageId="4" pageNumber="117">0.2±0.0</td>
<td id="743EC366FFB33DC2FBC14941FB0F0724" box="[1055,1178,1877,1899]" gridcol="3" gridrow="13" pageId="4" pageNumber="117">0.3–0.3</td>
<td id="743EC366FFB33DC2FB3C4941FACE0724" box="[1250,1371,1877,1899]" gridcol="4" gridrow="13" pageId="4" pageNumber="117">0.3±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF414968FACE07DD" box="[159,1371,1916,1938]" gridrow="14" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF414968FE3D07DD" box="[159,424,1916,1938]" gridcol="0" gridrow="14" pageId="4" pageNumber="117">Frons width, narrowest</th>
<td id="743EC366FFB33DC2FD4D4968FC9707DD" box="[659,770,1916,1938]" gridcol="1" gridrow="14" pageId="4" pageNumber="117">0.1–0.1</td>
<td id="743EC366FFB33DC2FC814968FC4D07DD" box="[863,984,1916,1938]" gridcol="2" gridrow="14" pageId="4" pageNumber="117">0.1±0.0</td>
<td id="743EC366FFB33DC2FBC14968FB0F07DD" box="[1055,1178,1916,1938]" gridcol="3" gridrow="14" pageId="4" pageNumber="117">0.2–0.2</td>
<td id="743EC366FFB33DC2FB3C4968FACE07DD" box="[1250,1371,1916,1938]" gridcol="4" gridrow="14" pageId="4" pageNumber="117">0.2±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF4149B7FACE07F6" box="[159,1371,1955,1977]" gridrow="15" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF4149B7FE3D07F6" box="[159,424,1955,1977]" gridcol="0" gridrow="15" pageId="4" pageNumber="117">Frons length, midline</th>
<td id="743EC366FFB33DC2FD4D49B7FC9707F6" box="[659,770,1955,1977]" gridcol="1" gridrow="15" pageId="4" pageNumber="117">0.5–0.5</td>
<td id="743EC366FFB33DC2FC8149B7FC4D07F6" box="[863,984,1955,1977]" gridcol="2" gridrow="15" pageId="4" pageNumber="117">0.5±0.0</td>
<td id="743EC366FFB33DC2FBC149B7FB0F07F6" box="[1055,1178,1955,1977]" gridcol="3" gridrow="15" pageId="4" pageNumber="117">0.6–0.6</td>
<td id="743EC366FFB33DC2FB3C49B7FACE07F6" box="[1250,1371,1955,1977]" gridcol="4" gridrow="15" pageId="4" pageNumber="117">0.6±0.0</td>
</tr>
<tr id="37EFAA1AFFB33DC2FF4149DEFACE07AF" box="[159,1371,1994,2016]" gridrow="16" pageId="4" pageNumber="117">
<th id="743EC366FFB33DC2FF4149DEFE3D07AF" box="[159,424,1994,2016]" gridcol="0" gridrow="16" pageId="4" pageNumber="117">Clypeus length</th>
<td id="743EC366FFB33DC2FD4D49DEFC9707AF" box="[659,770,1994,2016]" gridcol="1" gridrow="16" pageId="4" pageNumber="117">0.6±0.0</td>
<td id="743EC366FFB33DC2FC8149DEFC4D07AF" box="[863,984,1994,2016]" gridcol="2" gridrow="16" pageId="4" pageNumber="117">0.6±0.0</td>
<td id="743EC366FFB33DC2FBC149DEFB0F07AF" box="[1055,1178,1994,2016]" gridcol="3" gridrow="16" pageId="4" pageNumber="117">0.7±0.0</td>
<td id="743EC366FFB33DC2FB3C49DEFACE07AF" box="[1250,1371,1994,2016]" gridcol="4" gridrow="16" pageId="4" pageNumber="117">0.7±0.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8960A858FFB2C238FF194E83FE3701C6" blockId="5.[151,1436,151,393]" pageId="5" pageNumber="118">
Thorax. Pronotum in dorsal view roughly trapezoidal (about equal in length as vertex at midline), anterior median portion (between eyes), quadrate with anterior margin linear, posterior margin concave, median carina obscure, lateral carinae appear concurrent with anterior lateral pronotal margin; paradiscal broadly foliate behind antennae forming large fossae, partially surrounding antennae. Mesonotum at midline exceeding length of pronotum and vertex combined, tricarinate, median carina becoming obsolete posteriorly, lateral carinae medially curved from anterior margin, approximately reaching hind margin (
<figureCitation id="11E4B4DDFFB2C238FCD64F5FFCF1012A" box="[776,868,331,357]" captionStart="FIGURE 3" captionStartId="5.[152,255,1228,1252]" captionTargetBox="[177,1410,415,1195]" captionTargetId="figure-108@5.[170,1416,414,1202]" captionTargetPageId="5" captionText="FIGURE 3. Adult male Agoo keili sp. nov., pale form top row and yellow form bottom row;A) head, pronotum and mesonotum dorsal view, B) head, pronotum and mesonotum lateral view and C) head frontal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812380" httpUri="https://zenodo.org/record/14812380/files/figure.png" pageId="5" pageNumber="118">Fig. 3A</figureCitation>
). Hind tibiae unarmed laterally, with apical spine ornamentation of 7-6-5.
</paragraph>
<caption id="DDA0F8D0FFB2C238FF464AD8FB570547" ID-DOI="http://doi.org/10.5281/zenodo.14812380" ID-Zenodo-Dep="14812380" httpUri="https://zenodo.org/record/14812380/files/figure.png" pageId="5" pageNumber="118" startId="5.[152,255,1228,1252]" targetBox="[177,1410,415,1195]" targetPageId="5" targetType="figure">
<paragraph id="8960A858FFB2C238FF464AD8FB570547" blockId="5.[151,1436,1228,1288]" pageId="5" pageNumber="118">
<emphasis id="BBAB744AFFB2C238FF464AD8FE8204AB" bold="true" box="[152,279,1228,1252]" pageId="5" pageNumber="118">FIGURE 3.</emphasis>
Adult male
<taxonomicName id="4EDFD3DBFFB2C238FE4C4AD9FE6D04AB" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2025" box="[402,504,1228,1252]" class="Insecta" family="Derbidae" genus="Agoo" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="keili" status="sp. nov.">
<emphasis id="BBAB744AFFB2C238FE4C4AD9FE6D04AB" box="[402,504,1228,1252]" italics="true" pageId="5" pageNumber="118">Agoo keili</emphasis>
</taxonomicName>
<emphasis id="BBAB744AFFB2C238FE234AD8FDC604AB" bold="true" box="[509,595,1228,1252]" pageId="5" pageNumber="118">
<taxonomicNameLabel id="A098C931FFB2C238FE234AD8FDDA04AB" box="[509,591,1228,1252]" pageId="5" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
pale form top row and yellow form bottom row; A) head, pronotum and mesonotum dorsal view, B) head, pronotum and mesonotum lateral view and C) head frontal view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8960A858FFB2C238FF194B46FACF05FB" blockId="5.[151,1437,1362,2037]" pageId="5" pageNumber="118">Forewing elongate oval; apex of clavus near wing midlength; MP fused with ScP+R to form short composite vein from apex of basal cell: Fork of RP from ScP+RA nearly at same level of fusion of Pcu+1A in clavus, these just proximad of CuA fork; branching pattern: RA 1-branched, RP 2-branched, MP 4-branched, CuA 2-branched.</paragraph>
<paragraph id="8960A858FFB2C238FF194BAAFDC707BB" blockId="5.[151,1437,1362,2037]" pageId="5" pageNumber="118">
Male Terminalia. Pygofer in lateral view narrow and irregular in shape; narrowest dorsally, widest ventrally, anterior and posterior margin irregularly sinuate (
<figureCitation id="11E4B4DDFFB2C238FD654BF6FC8105B2" box="[699,788,1506,1533]" captionStart="FIGURE 5" captionStartId="7.[152,253,1104,1128]" captionTargetBox="[152,1436,179,1079]" captionTargetId="figure-18@7.[151,1436,179,1079]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Agoo keili sp. nov. terminalia; A) lateral view, B) ventral view and C) dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14812384" httpUri="https://zenodo.org/record/14812384/files/figure.png" pageId="5" pageNumber="118">Fig. 5A</figureCitation>
); in ventral view, medioventral process subtriangular (
<figureCitation id="11E4B4DDFFB2C238FAB14BF6FF2E066E" captionStart="FIGURE 5" captionStartId="7.[152,253,1104,1128]" captionTargetBox="[152,1436,179,1079]" captionTargetId="figure-18@7.[151,1436,179,1079]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Agoo keili sp. nov. terminalia; A) lateral view, B) ventral view and C) dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14812384" httpUri="https://zenodo.org/record/14812384/files/figure.png" pageId="5" pageNumber="118">Fig. 5B</figureCitation>
), broader than tall with acutely pointed apex. Gonostyli in lateral view spatulate, apex broadly rounded (
<figureCitation id="11E4B4DDFFB2C238FAB14812FF28060A" captionStart="FIGURE 5" captionStartId="7.[152,253,1104,1128]" captionTargetBox="[152,1436,179,1079]" captionTargetId="figure-18@7.[151,1436,179,1079]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Agoo keili sp. nov. terminalia; A) lateral view, B) ventral view and C) dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14812384" httpUri="https://zenodo.org/record/14812384/files/figure.png" pageId="5" pageNumber="118">Fig. 5A</figureCitation>
); dorsal process small, bilobed, distal lobe sclerotized; in ventral view (
<figureCitation id="11E4B4DDFFB2C238FC3B483EFBA8060A" box="[997,1085,1578,1605]" captionStart="FIGURE 5" captionStartId="7.[152,253,1104,1128]" captionTargetBox="[152,1436,179,1079]" captionTargetId="figure-18@7.[151,1436,179,1079]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Agoo keili sp. nov. terminalia; A) lateral view, B) ventral view and C) dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14812384" httpUri="https://zenodo.org/record/14812384/files/figure.png" pageId="5" pageNumber="118">Fig. 5B</figureCitation>
), median margin bearing large, quadrate process just distad midlength, distally narrowed with apices medially hooked to blunt apices. Aedeagus simple (
<figureCitation id="11E4B4DDFFB2C238FF2E4866FEA606C3" box="[240,307,1650,1676]" captionStart="FIGURE 6" captionStartId="7.[152,255,1951,1975]" captionTargetBox="[151,1424,1152,1922]" captionTargetId="figure-48@7.[151,1435,1151,1926]" captionTargetPageId="7" captionText="FIGURE 6. Agoo keili sp. nov. aedeagus; A) right lateral view, B) left lateral view, C) dorsal view and D) ventral view. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14812386" httpUri="https://zenodo.org/record/14812386/files/figure.png" pageId="5" pageNumber="118">Fig. 6</figureCitation>
), approximately bilaterally symmetrical, shaft upcurved bearing a pair of apical processes (A1 &amp; A2), elongated and slender, gently curving ventrad, angled cephalad, approximately reaching midpoint; second pair large, subtriangular processes (A3 &amp; A4) arising on dorsolateral margins in basal 1/3 of shaft (
<figureCitation id="11E4B4DDFFB2C238FBB248AEFB70069A" box="[1132,1253,1722,1749]" captionStart="FIGURE 6" captionStartId="7.[152,255,1951,1975]" captionTargetBox="[151,1424,1152,1922]" captionTargetId="figure-48@7.[151,1435,1151,1926]" captionTargetPageId="7" captionText="FIGURE 6. Agoo keili sp. nov. aedeagus; A) right lateral view, B) left lateral view, C) dorsal view and D) ventral view. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14812386" httpUri="https://zenodo.org/record/14812386/files/figure.png" pageId="5" pageNumber="118">Fig. 6A, B</figureCitation>
), ventral margin serrate along midlength, endosoma with multiple processes present, first pair(E1 &amp; E2) arising subapically on lateral margins, gently curved dorsad, angled cephalad, reaching apex of A3 and A4; second pair (E3 &amp; E4) arising on dorsal margin along inner margin, nearly straight, angled directly caudad, extending just beyond aedeagal apex; third pair (E5 &amp; E6) arising on endosomal ventral margin, sinuate along basal region, angled dorsad and cephalad, curving directly dorsad at apex; and the fourth pair (E7 &amp; E8) arising at endosoma apex, immediately curving dorsad, approximately reaching apex of E5 and E6. Anal tube in lateral view roughly triangular and relatively broad, posteriorly projecting (not downcurved), narrowed to blunt apices; in ventral view broad, strongly bilobed distad of epiproct and paraproct; paraproct short.
</paragraph>
<caption id="DDA0F8D0FFB1C23BFF464B10FD0C050F" ID-DOI="http://doi.org/10.5281/zenodo.14812382" ID-Zenodo-Dep="14812382" httpUri="https://zenodo.org/record/14812382/files/figure.png" pageId="6" pageNumber="119" startId="6.[152,255,1284,1308]" targetBox="[151,1432,179,1251]" targetPageId="6" targetType="figure">
<paragraph id="8960A858FFB1C23BFF464B10FD0C050F" blockId="6.[151,1436,1284,1345]" pageId="6" pageNumber="119">
<emphasis id="BBAB744AFFB1C23BFF464B10FE8D0553" bold="true" box="[152,280,1284,1308]" pageId="6" pageNumber="119">FIGURE 4.</emphasis>
Adult male
<taxonomicName id="4EDFD3DBFFB1C23BFE484B10FE6B0553" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2025" box="[406,510,1284,1308]" class="Insecta" family="Derbidae" genus="Agoo" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="keili" status="sp. nov.">
<emphasis id="BBAB744AFFB1C23BFE484B10FE6B0553" box="[406,510,1284,1308]" italics="true" pageId="6" pageNumber="119">Agoo keili</emphasis>
</taxonomicName>
<emphasis id="BBAB744AFFB1C23BFDDA4B10FDC00553" bold="true" box="[516,597,1284,1308]" pageId="6" pageNumber="119">
<taxonomicNameLabel id="A098C931FFB1C23BFDDA4B10FDC00553" box="[516,597,1284,1308]" pageId="6" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
forewing; A) pale form and B) yellow form, C) line art; black text = vein, italics = crossvein, and green text = cell. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8960A858FFB1C23BFF194B86FC9105E3" blockId="6.[151,1436,1426,1741]" box="[199,772,1426,1452]" pageId="6" pageNumber="119">
<emphasis id="BBAB744AFFB1C23BFF194B86FE3205E3" bold="true" box="[199,423,1426,1452]" pageId="6" pageNumber="119">Plant Associations.</emphasis>
Sweeping undetermined palm.
</paragraph>
</subSubSection>
<subSubSection id="C1C5FBD3FFB1C23BFF194BA2FD7D059F" box="[199,744,1462,1488]" pageId="6" pageNumber="119" type="distribution">
<paragraph id="8960A858FFB1C23BFF194BA2FD7D059F" blockId="6.[151,1436,1426,1741]" box="[199,744,1462,1488]" pageId="6" pageNumber="119">
<emphasis id="BBAB744AFFB1C23BFF194BA2FEF4059F" bold="true" box="[199,353,1462,1488]" pageId="6" pageNumber="119">Distribution.</emphasis>
<collectingCountry id="F1C8E8C8FFB1C23BFEB94BA2FE74059F" box="[359,481,1462,1488]" name="Costa Rica" pageId="6" pageNumber="119">Costa Rica</collectingCountry>
(
<collectingRegion id="4B1B66BAFFB1C23BFE2E4BA2FD48059F" box="[496,733,1462,1488]" country="Costa Rica" name="Puntarenas" pageId="6" pageNumber="119">Puntarenas Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C1C5FBD3FFB1C23BFF194BCEFBF30673" pageId="6" pageNumber="119" type="etymology">
<paragraph id="8960A858FFB1C23BFF194BCEFBF30673" blockId="6.[151,1436,1426,1741]" pageId="6" pageNumber="119">
<emphasis id="BBAB744AFFB1C23BFF194BCEFEDB05BB" bold="true" box="[199,334,1498,1524]" pageId="6" pageNumber="119">Etymology.</emphasis>
The specific name is given in honor of Dr. Clifford Keil (retired faculty from the University of Delaware Department of Entomology and retired faculty member and director of the Museum of Invertebrates in the School of Biological Sciences, Pontifical Catholic University of
<collectingCountry id="F1C8E8C8FFB1C23BFC4A4836FC640673" box="[916,1009,1570,1596]" name="Ecuador" pageId="6" pageNumber="119">Ecuador</collectingCountry>
in Quito).
</paragraph>
</subSubSection>
<subSubSection id="C1C5FBD3FFB1C23BFF194852FE3B0683" pageId="6" pageNumber="119" type="materials_examined">
<paragraph id="8960A858FFB1C23BFF194852FC8A06CB" blockId="6.[151,1436,1426,1741]" pageId="6" pageNumber="119">
<emphasis id="BBAB744AFFB1C23BFF194852FE22062F" bold="true" box="[199,439,1606,1632]" pageId="6" pageNumber="119">Material Examined.</emphasis>
<materialsCitation id="39B7A205FFB1C23BFE614852FC8A06CB" collectingDate="2021-06-09" collectionCode="FLREC" collectorName="Coll. &amp; B. W. Bahder" country="Costa Rica" location="Hotel Keili" pageId="6" pageNumber="119" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="566416FAFFB1C23BFE614852FDB2062F" box="[447,551,1606,1632]" pageId="6" pageNumber="119" type="holotype">Holotype</typeStatus>
male, “
<collectingCountry id="F1C8E8C8FFB1C23BFDA34852FD68062F" box="[637,765,1606,1632]" name="Costa Rica" pageId="6" pageNumber="119">Costa Rica</collectingCountry>
,
<collectingRegion id="4B1B66BAFFB1C23BFCD74853FC11062E" box="[777,900,1607,1633]" country="Costa Rica" name="Puntarenas" pageId="6" pageNumber="119">Puntarenas</collectingRegion>
Pr. /
<location id="8C00FE83FFB1C23BFC1E4852FBD4062F" LSID="urn:lsid:plazi:treatment:0176194EFFB4C23BFF4948F4FE3B0683:8C00FE83FFB1C23BFC1E4852FBD4062F" box="[960,1089,1606,1632]" country="Costa Rica" name="Hotel Keili" pageId="6" pageNumber="119" stateProvince="Puntarenas">Hotel Keili</location>
, roadside /
<date id="FD618E98FFB1C23BFB1C4852FAD2062F" box="[1218,1351,1606,1632]" pageId="6" pageNumber="119" value="2021-06-09">
<collectingDate id="ED257770FFB1C23BFB1C4852FAD2062F" box="[1218,1351,1606,1632]" pageId="6" pageNumber="119" value="2021-06-09">09-VI-2021</collectingDate>
</date>
/
<collectorName id="242ACD8EFFB1C23BFA834852FA0C062F" box="[1373,1433,1606,1632]" pageId="6" pageNumber="119">Coll.</collectorName>
:
<collectorName id="242ACD8EFFB1C23BFF49487FFEBD06CB" box="[151,296,1642,1669]" pageId="6" pageNumber="119">B.W. Bahder</collectorName>
/ Sweeping palms /
<taxonomicName id="4EDFD3DBFFB1C23BFDD5487FFDEB06CB" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2025" box="[523,638,1642,1669]" class="Insecta" family="Derbidae" genus="Agoo" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="keili">
<emphasis id="BBAB744AFFB1C23BFDD5487FFDEB06CB" box="[523,638,1642,1669]" italics="true" pageId="6" pageNumber="119">Agoo keili</emphasis>
</taxonomicName>
<specimenCount id="9FD963D1FFB1C23BFD5B487EFD0E06CB" box="[645,667,1642,1668]" count="1" pageId="6" pageNumber="119" type="male">♂</specimenCount>
” (
<collectionCode id="EFCE309DFFB1C23BFD6B487EFC8606CB" box="[693,787,1642,1668]" pageId="6" pageNumber="119">FLREC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8960A858FFB1C23BFF19489BFE3B0683" blockId="6.[151,1436,1426,1741]" pageId="6" pageNumber="119">
<typeStatus id="566416FAFFB1C23BFF19489BFEA106E6" box="[199,308,1679,1705]" pageId="6" pageNumber="119" type="paratype">Paratypes</typeStatus>
same as
<typeStatus id="566416FAFFB1C23BFE44489AFE6906E7" box="[410,508,1678,1704]" pageId="6" pageNumber="119" type="holotype">holotype</typeStatus>
(
<specimenCount id="9FD963D1FFB1C23BFDD3489AFDE606E7" box="[525,627,1678,1704]" count="11" pageId="6" pageNumber="119" type="male">11 males</specimenCount>
,
<specimenCount id="9FD963D1FFB1C23BFDA1489AFD7A06E7" box="[639,751,1678,1704]" count="4" pageId="6" pageNumber="119" type="female">4 females</specimenCount>
;
<specimenCount id="9FD963D1FFB1C23BFD24489AFCC506E7" box="[762,848,1678,1704]" count="2" pageId="6" pageNumber="119" type="male">2 males</specimenCount>
and
<specimenCount id="9FD963D1FFB1C23BFC59489AFC7D06E7" box="[903,1000,1678,1704]" count="1" pageId="6" pageNumber="119" type="female">1 female</specimenCount>
were yellow morph and
<specimenCount id="9FD963D1FFB1C23BFADF489AFAC206E7" box="[1281,1367,1678,1704]" count="9" pageId="6" pageNumber="119" type="male">9 males</specimenCount>
and
<specimenCount id="9FD963D1FFB1C23BFA50489AFF760683" count="1" pageId="6" pageNumber="119" type="female">1 female</specimenCount>
were pale morph.
</paragraph>
</subSubSection>
</treatment>
</document>

102
data/01/76/19/0176194EFFB4C23EFF494ECBFA9E0213.xml Normal file
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@ -0,0 +1,102 @@
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<mods:mods id="21F5A733E2FDB072AC4555E3215CA4CD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="58DD62909F0FBC81AA718D4E6900CB33">
<mods:title id="D6E3B675469417298E00BEA7BF27275C">Multi-local analysis supports the transfer of Omolicna fulva to the genus Agoo and establishment of a new species in the genus from Costa Rica</mods:title>
</mods:titleInfo>
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<mods:role id="3ADE1B2B1E956D4273DE55010A8B9856">
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<mods:namePart id="359D3C4DACFD370760F03DA55629BE92">Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:affiliation id="2FB3B3D67C10831FE35D366239DDF142">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica</mods:affiliation>
</mods:name>
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<mods:namePart id="797FF43AEC6FB321A0593CE928FBD7E4">Zumbado Echavarria, Marco A.</mods:namePart>
</mods:name>
<mods:name id="22132BD5FA61A557CD1A0197CB03006A" type="personal">
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<mods:namePart id="0C577A5429FC3A0D31D9657D1A1706B0">Bartlett, Charles R.</mods:namePart>
<mods:affiliation id="0D2E00621B0E36B76F5AA42037996BF4">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA</mods:affiliation>
</mods:name>
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<mods:namePart id="BC2497BB0B51CD31EC5938A5B6332193">Helmick, Ericka E.</mods:namePart>
<mods:affiliation id="C4AE99C12AB275C70C358D5B151A23B6">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
</mods:name>
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Genus
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species:
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<emphasis id="BBAB744AFFB4C23EFEF84F30FE3D0174" box="[294,424,292,315]" italics="true" pageId="3" pageNumber="116">Agoo xavieri</emphasis>
<bibRefCitation id="ED4ED5A9FFB4C23EFE714F30FD310173" author="Bahder, B. W. &amp; Bartlett, C. R. &amp; Barrantes, E. A. &amp; Echavarria, M. A. Z. &amp; Humphries, A. R. &amp; Helmick, E. E. &amp; Ascunce, M. S. &amp; Goss, E. M." box="[431,676,292,316]" pageId="3" pageNumber="116" pagination="501 - 514" refId="ref6761" refString="Bahder, B. W., Bartlett, C. R., Barrantes, E. A. B, Echavarria, M. A. Z., Humphries, A. R., Helmick, E. E., Ascunce, M. S. &amp; Goss, E. M. (2019) A new species of Omolicna (Hemiptera: Auchenorrhyncha: Fulgoroidea: Derbidae) from coconut palm in Costa Rica and new country records for Omolicna brunnea and Omolicna triata. Zootaxa, 4577 (3), 501-514. https://doi.org/10.11646/zootaxa.4577.3.5" type="journal article" year="2019">Bahder &amp; Bartlett, 2019</bibRefCitation>
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<emphasis id="BBAB744AFFB4C23EFF494F7DFE8701CC" bold="true" box="[151,274,361,387]" pageId="3" pageNumber="116">Diagnosis.</emphasis>
Elongate cenchreines with narrow forewings. Head in lateral view narrowly projected anteriorly, with vertex + frons profile smoothly rounded. Frons narrow (but lateral margins not in contact), vertex subtriangular, with sensorial pits along lateral margins of frons and vertex; transverse carina at fastigium lacking; median carina of vertex and frons absent. Antennae short. Paranotal expansions behind antennae quadrate to semiquadrate in frontal view. In caudoventral view, medioventral lobe of pygofer simple, broad, subtriangular, distally attenuating to rounded apex. Aedeagus and endosoma nearly bilaterally symmetrical, aedeagus with at least one pair of elongate processes at apex, extremely complex endosoma with at least two pairs of large sclerotized processes.
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Larval key to the genus
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1. Posterolateral expansions of abdominal segments well developed on III–VIII and directed outwards (
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); pronotum greatly enlarged laterally and posteriorly (
<figureCitation id="13DE2A7EFFE9FF81DA5DB28188DBF90B" box="[639,713,1750,1772]" captionStart="FIGURES 96–103" captionStartId="17.[151,264,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-19@17.[151,1435,181,1859]" captionTargetPageId="17" captionText="FIGURES 96–103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.14798089" httpUri="https://zenodo.org/record/14798089/files/figure.png" pageId="22" pageNumber="269">Fig. 96</figureCitation>
); anterior margin of the labrum with a smooth, shallow depression in the middle area (
<figureCitation id="13DE2A7EFFE9FF81D942B2A48BB7F8EF" box="[352,421,1779,1801]" captionStart="FIGURES 96–103" captionStartId="17.[151,264,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-19@17.[151,1435,181,1859]" captionTargetPageId="17" captionText="FIGURES 96–103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.14798089" httpUri="https://zenodo.org/record/14798089/files/figure.png" pageId="22" pageNumber="269">Fig. 99</figureCitation>
)...........................................................................
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- Posterolateral expansions of abdominal segment III–VIII moderately developed (
<figureCitation id="13DE2A7EFFE9FF81DBDEB3468E46F8C1" box="[1020,1108,1808,1831]" captionStart="FIGURES 4–5" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 4–5. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." figureDoi="http://doi.org/10.5281/zenodo.14798057" httpUri="https://zenodo.org/record/14798057/files/figure.png" pageId="22" pageNumber="269">Figs 4–5</figureCitation>
,
<figureCitation id="13DE2A7EFFE9FF81DC42B3478EB3F8C1" box="[1120,1185,1808,1830]" captionStart="FIGURES 53–54" captionStartId="11.[151,264,1385,1409]" captionTargetBox="[164,1423,273,1362]" captionTargetId="figure-62@11.[164,1423,273,1362]" captionTargetPageId="11" captionText="FIGURES 53–54. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." figureDoi="http://doi.org/10.5281/zenodo.14798075" httpUri="https://zenodo.org/record/14798075/files/figure.png" pageId="22" pageNumber="269">53–54</figureCitation>
); pronotum is moderately enlarged laterally and posteriorly (
<figureCitation id="13DE2A7EFFE9FF81DA0EB3798890F8A3" box="[556,642,1838,1860]" captionStart="FIGURES 4–5" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 4–5. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." figureDoi="http://doi.org/10.5281/zenodo.14798057" httpUri="https://zenodo.org/record/14798057/files/figure.png" pageId="22" pageNumber="269">Figs 4–5</figureCitation>
,
<figureCitation id="13DE2A7EFFE9FF81DAAFB37988BAF8A3" box="[653,680,1838,1860]" captionStart="FIGURES 51–52" captionStartId="10.[151,264,1632,1656]" captionTargetBox="[164,1423,273,1609]" captionTargetId="figure-68@10.[164,1423,273,1609]" captionTargetPageId="10" captionText="FIGURES 51–52. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." figureDoi="http://doi.org/10.5281/zenodo.14798073" httpUri="https://zenodo.org/record/14798073/files/figure.png" pageId="22" pageNumber="269">51</figureCitation>
); labrum a different shape......................................... 2
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<paragraph id="8B5A36FBFFE9FF81D8B5B31C8F8EF899" blockId="22.[151,1437,1721,2007]" pageId="22" pageNumber="269">
<keyLead id="301495CEFFE9FF81D8B5B31C8F8EF899" pageId="22" pageNumber="269">
2. Posterior margins of terga with irregular pointed teeth and a row of microdenticles (
<figureCitation id="13DE2A7EFFE9FF81DC32B31C8E90F886" box="[1040,1154,1867,1889]" captionStart="FIGURES 78–87" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 78–87. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 83–84. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." figureDoi="http://doi.org/10.5281/zenodo.14798085" httpUri="https://zenodo.org/record/14798085/files/figure.png" pageId="22" pageNumber="269">Figs 84–86</figureCitation>
); hind tibia with two rows of thin setae on the dorsal face of hind femora arrow-shaped......................................................3
</keyLead>
</paragraph>
<paragraph id="8B5A36FBFFE9FF81D8B5B3D18F8EF831" blockId="22.[151,1437,1721,2007]" pageId="22" pageNumber="269">
<keyLead id="301495CEFFE9FF81D8B5B3D18F8EF831" pageId="22" pageNumber="269">
- Posterior margin of tergites with regular pointed teeth, and numerous microdenticles (
<bibRefCitation id="EF744B0AFFE9FF81DC07B3D18F1FF87C" author="Selvakumar, C. &amp; Chandra, K. &amp; Sivaramakrishnan, K. G. &amp; Jehamalar, E. E." box="[1061,1293,1926,1948]" pageId="22" pageNumber="269" pagination="84 - 90" refId="ref7896" refString="Selvakumar, C., Chandra, K., Sivaramakrishnan, K. G. &amp; Jehamalar, E. E. (2017) A new species of Thalerosphyrus Eaton 1881 (Ephemeroptera: Heptageniidae: Ecdyonurinae) from India. Zootaxa, 4350 (1), 84-90. https://doi.org/10.11646/zootaxa.4350.1.4" type="journal article" year="2017">
Selvakumar
<emphasis id="B991EAE9FFE9FF81DC82B3D08EC0F87C" box="[1184,1234,1926,1948]" italics="true" pageId="22" pageNumber="269">et al.</emphasis>
2017
</bibRefCitation>
:
<figureCitation id="13DE2A7EFFE9FF81DD35B3D18F4DF87C" box="[1303,1375,1926,1948]" captionStart="FIGURES 20–24" captionStartId="5.[151,264,1421,1445]" captionTargetBox="[189,1398,181,1397]" captionTargetId="figure-19@5.[189,1398,181,1397]" captionTargetPageId="5" captionText="FIGURES 20–24. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." figureDoi="http://doi.org/10.5281/zenodo.14798063" httpUri="https://zenodo.org/record/14798063/files/figure.png" pageId="22" pageNumber="269">Fig. 21</figureCitation>
); hind tibia with one row of thin setae; setae on the dorsal surface of hind femora truncate or rounded at apex (
<bibRefCitation id="EF744B0AFFE9FF81DC95B3F48F88F85F" author="Selvakumar, C. &amp; Chandra, K. &amp; Sivaramakrishnan, K. G. &amp; Jehamalar, E. E." box="[1207,1434,1955,1977]" pageId="22" pageNumber="269" pagination="84 - 90" refId="ref7896" refString="Selvakumar, C., Chandra, K., Sivaramakrishnan, K. G. &amp; Jehamalar, E. E. (2017) A new species of Thalerosphyrus Eaton 1881 (Ephemeroptera: Heptageniidae: Ecdyonurinae) from India. Zootaxa, 4350 (1), 84-90. https://doi.org/10.11646/zootaxa.4350.1.4" type="journal article" year="2017">
Selvakumar
<emphasis id="B991EAE9FFE9FF81DD13B3F38F49F85F" box="[1329,1371,1955,1977]" italics="true" pageId="22" pageNumber="269">et al</emphasis>
. 2017
</bibRefCitation>
:
<figureCitation id="13DE2A7EFFE9FF81D8C0B3968B34F831" box="[226,294,1984,2007]" captionStart="FIGURES 12–19" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 12–19. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." figureDoi="http://doi.org/10.5281/zenodo.14798061" httpUri="https://zenodo.org/record/14798061/files/figure.png" pageId="22" pageNumber="269">Fig. 15</figureCitation>
)...............................................................................
<taxonomicName id="4CE54D78FFE9FF81DCD5B3968F8EF831" authorityName="Selvakumar &amp; Chandra" authorityYear="2017" box="[1271,1436,1985,2006]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="269" phylum="Arthropoda" rank="species" species="meghalayensis">
<emphasis id="B991EAE9FFE9FF81DCD5B3968F8EF831" box="[1271,1436,1985,2006]" italics="true" pageId="22" pageNumber="269">T. meghalayensis</emphasis>
</taxonomicName>
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<keyStep id="30112E5EFFE8FF80D8B5B4CF8F8EFEC5" pageId="23" pageNumber="270">
<paragraph id="8B5A36FBFFE8FF80D8B5B4CF8F8EFF00" blockId="23.[151,1437,151,291]" pageId="23" pageNumber="270">
<keyLead id="301495CEFFE8FF80D8B5B4CF8F8EFF00" pageId="23" pageNumber="270">
3. Hind femur dorsal surface with arrow-shaped, blunt and strong setae (
<figureCitation id="13DE2A7EFFE8FF80DB55B4C089ACFF4A" box="[887,958,151,173]" captionStart="FIGURES 38–43" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 38–43. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." figureDoi="http://doi.org/10.5281/zenodo.14798069" httpUri="https://zenodo.org/record/14798069/files/figure.png" pageId="23" pageNumber="270">Fig. 40</figureCitation>
); labrum expanded laterally, 4.5× wider than long, anterior margin straight, medially bulged with a concave shape and small blunt crenations on either side, (
<figureCitation id="13DE2A7EFFE8FF80DCF9B4E28F33FF2D" box="[1243,1313,181,203]" captionStart="FIGURES 12–19" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 12–19. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." figureDoi="http://doi.org/10.5281/zenodo.14798061" httpUri="https://zenodo.org/record/14798061/files/figure.png" pageId="23" pageNumber="270">Fig. 12</figureCitation>
)...........................................................................................
<taxonomicName id="4CE54D78FFE8FF80DCE3B4848F5BFF00" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[1217,1353,210,232]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="270" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFE8FF80DCE3B4848F5BFF00" box="[1217,1353,210,232]" italics="true" pageId="23" pageNumber="270">T. bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFE8FF80DD72B4858F8EFF00" bold="true" box="[1360,1436,210,231]" pageId="23" pageNumber="270">
<taxonomicNameLabel id="A2A25792FFE8FF80DD72B4858F8EFF00" box="[1360,1436,210,231]" pageId="23" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<keyLead id="301495CEFFE8FF80D8B5B4A78F8EFEC5" pageId="23" pageNumber="270">
- Hind femur dorsal surface with arrow-shaped setae, clearly point (
<figureCitation id="13DE2A7EFFE8FF80DB45B4B889BDFEE2" box="[871,943,239,261]" captionStart="FIGURES 68–77" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 68–77. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." figureDoi="http://doi.org/10.5281/zenodo.14798083" httpUri="https://zenodo.org/record/14798083/files/figure.png" pageId="23" pageNumber="270">Fig. 75</figureCitation>
); Labrum moderately broad, 3.5× wider than long, anterior margin straight (
<figureCitation id="13DE2A7EFFE8FF80D9F3B55A882DFEC4" box="[465,575,269,291]" captionStart="FIGURES 55–62" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 55–62. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" figureDoi="http://doi.org/10.5281/zenodo.14798077" httpUri="https://zenodo.org/record/14798077/files/figure.png" pageId="23" pageNumber="270">Figs 57–58</figureCitation>
).......................................................
<taxonomicName id="4CE54D78FFE8FF80DCC8B55A8F5BFEC5" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[1258,1353,269,290]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="270" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFE8FF80DCC8B55A8F5BFEC5" box="[1258,1353,269,290]" italics="true" pageId="23" pageNumber="270">T. sartorii</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFE8FF80DD72B55A8F8EFEC5" bold="true" box="[1360,1436,269,290]" pageId="23" pageNumber="270">
<taxonomicNameLabel id="A2A25792FFE8FF80DD72B55A8F8EFEC5" box="[1360,1436,269,290]" pageId="23" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
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<taxonomicName id="4C3DAFF7FFC0FF9AFF15651C6186FEBD" ID-CoL="92P6W" authority="Walsingham, 1897" authorityName="Walsingham" authorityYear="1897" box="[151,673,330,357]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFF15651C6186FEBD" bold="true" box="[151,673,330,357]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF15651C6293FEBC" bold="true" box="[151,436,331,357]" italics="true" pageId="2" pageNumber="273">THAUMATOGRAPHA</emphasis>
<bibRefCitation id="EFACA985FFC0FF9AFE39651D6186FEBD" author="Walsingham, T. de" box="[443,673,330,356]" pageId="2" pageNumber="273" pagination="33 - 67" refId="ref12725" refString="Walsingham, T. de G. (1897) Western Equatorial African Micro-Lepidoptera. Transactions of the Entomological Society of London, 1897, 33-67, pls. 2 - 3. https://doi.org/10.1111/j.1365-2311.1897.tb01674.x" type="journal article" year="1897">Walsingham, 1897</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC0FF9AFF1565D8611FFE7E" box="[151,568,399,423]" pageId="2" pageNumber="273" type="reference_group">
<paragraph id="8B82D474FFC0FF9AFF1565D8611FFE7E" blockId="2.[151,1245,399,460]" box="[151,568,399,423]" pageId="2" pageNumber="273">
<treatmentCitationGroup id="AB2DF35AFFC0FF9AFF1565D8611FFE7E" box="[151,568,399,423]" pageId="2" pageNumber="273">
<taxonomicName id="4C3DAFF7FFC0FF9AFF1565D86113FE7E" ID-CoL="92P6W" authority="Walsignham, 1897: 52" authorityName="Walsignham" authorityPageNumber="52" authorityYear="1897" box="[151,564,399,423]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFF1565D86260FE7E" box="[151,327,399,423]" italics="true" pageId="2" pageNumber="273">Thaumatographa</emphasis>
Walsignham, 1897: 52
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<paragraph id="8B82D474FFC0FF9AFDBF65C76189FE12" blockId="2.[151,1245,399,460]" pageId="2" pageNumber="273">
<subSubSection id="C32787FFFFC0FF9AFDBF65C767FAFE7E" box="[573,1245,399,424]" pageId="2" pageNumber="273" type="type_taxon">
<treatmentCitationGroup id="AB2DF35AFFC0FF9AFDBF65C767FAFE7E" box="[573,1245,399,424]" pageId="2" pageNumber="273">
<typeStatus id="54866AD6FFC0FF9AFDBF65C76148FE71" box="[573,623,400,424]" pageId="2" pageNumber="273">Type</typeStatus>
species:
<taxonomicName id="4C3DAFF7FFC0FF9AFD4E65D86718FE7E" ID-CoL="9762K" authority="Meyrick, 1886" authorityName="Meyrick" authorityYear="1886" box="[716,1087,399,423]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="zapyra">
<emphasis id="B9490866FFC0FF9AFD4E65D86083FE7E" box="[716,932,399,423]" italics="true" pageId="2" pageNumber="273">Hilarographa zapyra</emphasis>
<bibRefCitation id="EFACA985FFC0FF9AFC2865D86718FE7E" author="Meyrick, E." box="[938,1087,399,423]" pageId="2" pageNumber="273" pagination="189 - 296" refId="ref12134" refString="Meyrick, E. (1886) Descriptions of Lepidoptera from the South Pacific. Transactions of the Entomological Society of London, 1886, 189-296. https://doi.org/10.1111/j.1365-2311.1886.tb01626.x" type="journal article" year="1886">Meyrick, 1886</bibRefCitation>
</taxonomicName>
(New
<collectingCountry id="F32A94E4FFC0FF9AFB0665D867F5FE7E" box="[1156,1234,399,423]" name="Guinea-Bissau" pageId="2" pageNumber="273">Guinea</collectingCountry>
).
</treatmentCitationGroup>
</subSubSection>
<subSubSection id="C32787FFFFC0FF9AFF1565E66189FE12" box="[151,686,433,460]" pageId="2" pageNumber="273" type="reference_group">
<treatmentCitationGroup id="AB2DF35AFFC0FF9AFF1565E66189FE12" box="[151,686,433,460]" pageId="2" pageNumber="273">
<taxonomicName id="4C3DAFF7FFC0FF9AFF1565E66271FE12" ID-CoL="B63FX" authorityName="Diakonoff" authorityYear="1977" box="[151,342,433,459]" class="Insecta" family="Tortricidae" genus="Tharmatographa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFF1565E66271FE12" box="[151,342,433,459]" italics="true" pageId="2" pageNumber="273">Tharmatographa</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9CF265FFC0FF9AFEE765E6617EFE15" author="Diakonoff, A." box="[357,601,433,460]" page="51" pageId="2" pageNumber="273" year="1977">
<bibRefCitation id="EFACA985FFC0FF9AFEE765E6617EFE15" author="Diakonoff, A." box="[357,601,433,460]" pageId="2" pageNumber="273" pagination="3 - 55" refId="ref10939" refString="Diakonoff, A. (1977 b) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 3-55." type="journal article" year="1977">Diakonoff, 1977b: 51</bibRefCitation>
</treatmentCitation>
, missp.
</treatmentCitationGroup>
</subSubSection>
</paragraph>
<subSubSection id="C32787FFFFC0FF9BFF1565AE61FAFD70" lastPageId="3" lastPageNumber="274" pageId="2" pageNumber="273" type="diagnosis">
<paragraph id="8B82D474FFC0FF9AFF1565AE6265FCD6" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF1565AE622DFDCA" bold="true" box="[151,266,505,531]" pageId="2" pageNumber="273">Diagnosis</emphasis>
. Wing expanse range, 7.5–27.0mm (average size:
<quantity id="4CC57991FFC0FF9AFCD565AE60F0FDCA" box="[855,983,505,531]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="2" pageNumber="273" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12–18 mm</quantity>
). The genus is defined in the tribe and distinguished from
<taxonomicName id="4C3DAFF7FFC0FF9AFEEC664A612EFDEE" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[366,521,541,567]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFEEC664A612EFDEE" box="[366,521,541,567]" italics="true" pageId="2" pageNumber="273">Hilarographa</emphasis>
</taxonomicName>
by the following particulars: the labial palpi are distinctly dorso-ventrally flattened (tubular in
<taxonomicName id="4C3DAFF7FFC0FF9AFE9A66166294FD82" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[280,435,577,603]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFE9A66166294FD82" box="[280,435,577,603]" italics="true" pageId="2" pageNumber="273">Hilarographa</emphasis>
</taxonomicName>
); in the male genitalia, the uncus is basally fused to the tegumen (membranous basally in
<taxonomicName id="4C3DAFF7FFC0FF9AFF3766326277FDA6" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[181,336,613,639]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFF3766326277FDA6" box="[181,336,613,639]" italics="true" pageId="2" pageNumber="273">Hilarographa</emphasis>
</taxonomicName>
), the hami are prominent and usually tubular, and the socii are sclerotic and usually slender, little setose, usually basally fused to the hami (socii wide, flattened and highly setose, and not fused to the hami in
<taxonomicName id="4C3DAFF7FFC0FF9AFF1566FA6215FD1E" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[151,306,685,711]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFF1566FA6215FD1E" box="[151,306,685,711]" italics="true" pageId="2" pageNumber="273">Hilarographa</emphasis>
</taxonomicName>
). The female genitalia usually have a bursal diverticular accessory sac, and the signum a bilobed fan-like form of numerous long spines curved inwardly, rarely absent or reduced (usually a scobinate patch in
<taxonomicName id="4C3DAFF7FFC0FF9AFF1566A26215FCD6" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[151,306,757,783]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC0FF9AFF1566A26215FCD6" box="[151,306,757,783]" italics="true" pageId="2" pageNumber="273">Hilarographa</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B82D474FFC0FF9AFF45674E60D8FC3E" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF45674E6285FCEA" bold="true" box="[199,418,793,819]" pageId="2" pageNumber="273">Descriptive Notes.</emphasis>
<emphasis id="B9490866FFC0FF9AFE29674E62CFFCEA" box="[427,488,793,819]" italics="true" pageId="2" pageNumber="273">Head</emphasis>
: small compared to most tortricids; scales relatively appressed except caudally on vertex; eyes large; ocellus posterior and moderate (sometimes enlarged); antenna very short (less than 1/2 length of forewing), filiform but thickened, with ventral bristles usually long in males (short in females); haustellum short; maxillary palpus very small, 1-segmented; labial palpus moderate and recurved, with the apical segment dorso-ventrally flattened, the median segment smooth-scaled but with few very fine and short bristles (but not always exposed among the scales), and the basal segment mostly also smooth-scaled.
</paragraph>
<paragraph id="8B82D474FFC0FF9AFF4567A66270FBF6" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF4567A6623FFBD2" box="[199,280,1009,1035]" italics="true" pageId="2" pageNumber="273">Thorax</emphasis>
: average tortricoid, with mostly appressed scales; tegula short. Legs average for body size, mostly with scales appressed.
</paragraph>
<paragraph id="8B82D474FFC0FF9AFF45606D6016FB3A" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF45606D6214FB8A" box="[199,307,1082,1107]" italics="true" pageId="2" pageNumber="273">Forewing</emphasis>
(
<figureCitation id="1306C8F1FFC0FF9AFEC4606E62A9FB8A" box="[326,398,1081,1107]" captionStart="FIGURE 7" captionStartId="5.[151,250,1110,1134]" captionTargetBox="[410,1178,305,1083]" captionTargetId="figure-97@5.[410,1178,305,1086]" captionTargetPageId="5" captionText="FIGURE 7. Wing venation of Thaumatographa zapyra (Meyrick) (New Guinea), type-species of the genus: redrawn from original sketch of Walsingham (1897). Note: the wing venation could not be compared to an actual specimen of T. zapyra for accuracy, so the venation may not be totally accurate in all details since the original sketch by Walsingham is small and somewhat simplified." figureDoi="http://doi.org/10.5281/zenodo.14798837" httpUri="https://zenodo.org/record/14798837/files/figure.png" pageId="2" pageNumber="273">Fig. 7</figureCitation>
): oblong, with costal margin slightly curved, termen truncated and apex and tornus more or less rounded; dorsal margin straight until anal curve; all veins free and mostly equally spaced, except R1 from middle of radius or from 2/3 of cell, and R2 and R3 sometimes close at end of chorda; chorda present, short (ca. 1/5 cell length), or vestigial; R5 to termen; median of cell vestigial; CuA1 from 2/3 end of cell; CuP at tornus, then vestigial toward base; A1+2 with relatively long basal fork.
</paragraph>
<paragraph id="8B82D474FFC0FF9AFF4560BA6234FA62" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF4560BA6212FADE" box="[199,309,1261,1287]" italics="true" pageId="2" pageNumber="273">Hindwing</emphasis>
(
<figureCitation id="1306C8F1FFC0FF9AFEC760BA62ADFADE" box="[325,394,1261,1287]" captionStart="FIGURE 7" captionStartId="5.[151,250,1110,1134]" captionTargetBox="[410,1178,305,1083]" captionTargetId="figure-97@5.[410,1178,305,1086]" captionTargetPageId="5" captionText="FIGURE 7. Wing venation of Thaumatographa zapyra (Meyrick) (New Guinea), type-species of the genus: redrawn from original sketch of Walsingham (1897). Note: the wing venation could not be compared to an actual specimen of T. zapyra for accuracy, so the venation may not be totally accurate in all details since the original sketch by Walsingham is small and somewhat simplified." figureDoi="http://doi.org/10.5281/zenodo.14798837" httpUri="https://zenodo.org/record/14798837/files/figure.png" pageId="2" pageNumber="273">Fig. 7</figureCitation>
): oblong-triangular to more oblong shape, with widely rounded tornus and anal margin; costa relatively straight; Sc to near apex; R to costa just before apex and M1 to termen just below apex and short-stalked with R near end of cell; cell from 1/2 to 3/5 length of hindwing; cell median vein vestigial; median veins equidistant at end of cell and at termen; M3 merged with CuA2 at end of cell; CuA1 from middle of cell or from 2/3 of cell; CuP vestigial or slightly at termen; 1A+2A with short basal fork; 3A long; male frenulum long; 2-bristle frenulum in females.
</paragraph>
<paragraph id="8B82D474FFC0FF9AFF45619266A8F9DD" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF4561926216FA06" box="[199,305,1477,1503]" italics="true" pageId="2" pageNumber="273">Abdomen</emphasis>
: average for body size, but sometimes slightly elongated; thoracic articulation tortricoid but apodemes somewhat thin; male coremata usually present and short, but can be prominent and as long as pregenital segment.
</paragraph>
<paragraph id="8B82D474FFC0FF9AFF45625A61B8F801" blockId="2.[151,1437,505,2008]" pageId="2" pageNumber="273">
<emphasis id="B9490866FFC0FF9AFF45625A624CF9FE" box="[199,363,1549,1575]" italics="true" pageId="2" pageNumber="273">Male genitalia</emphasis>
(
<figureCitation id="1306C8F1FFC0FF9AFEFA625A629BF9FE" box="[376,444,1549,1575]" captionStart="FIGURE 8" captionStartId="5.[151,250,1802,1826]" captionTargetBox="[156,1435,1263,1779]" captionTargetId="figure-160@5.[151,1436,1263,1779]" captionTargetPageId="5" captionText="FIGURE 8. Male genital structures in Thaumatographa, showing main characters for the complex male genitalia of this genus (modified after Diakonoff 1977, for T. zapyra, New Guinea) (the tegumen in this sketch is detached from the vinculum); coremata on pregenital segment (8a) (T. aurosa, Japan; after Diakonoff 1986). Note: the coremata typically are attached to the pregenital membrane structure of the abdomen in Thaumatographa (Fig. 8a), and rest within dorsal slits of each valva when not extruded, but in T. zapyra are attached to the costal margin of the valva itself. The aedeagus in Thaumatographa typically has a long ductus ejaculatorius (2 to 4 times length of aedeagus), with a long proximal hood (not shown in the drawing)." figureDoi="http://doi.org/10.5281/zenodo.14798839" httpUri="https://zenodo.org/record/14798839/files/figure.png" pageId="2" pageNumber="273">Fig. 8</figureCitation>
): hilarographine, with hami and socii prominent and usually long; tegumen and vinculum both generally triangular in shape, thin; uncus usually slender and very strongly sclerotized (rarely wide or short, or very long), and usually recurved caudally, basally fused to tegumen, and usually apically hooked; hami sclerotic and usually long and slender (rarely flattened or short); socii porrect or pendulate and long (rarely short and stubby), setose, usually slender and fused to base of hami; gnathos usually ovate to quadrate, and pendulate; transtilla usually present, band-like (or as valval stubs), sometimes caudally widened and even with caudal projections); juxta usually a small, caudally concave ovate plate; anellus a membranous and undeveloped ring; aedeagus tubular, short to long, sometimes with a distal lateral thorn; vesica finely scobinate, cornuti reduced or a single elongated tube (sometimes with an apical hook); phallobase often undeveloped, or small and bulbous, often ventrally decumbent; ductus ejaculatorius typically long, with prominent proximal elongated hood (rarely bulbous); valva oblong, usually quadratic with truncated termen and generally unadorned (sometimes with various ridges), but split open on dorsal margin (pouch-like split retains coremata in repose); saccus reduced and usually very short (rarely long), with acute apex, or sometimes a recurved truncated apex.
</paragraph>
<paragraph id="8B82D474FFC1FF9BFF4564EC61FAFD70" blockId="3.[151,1437,187,2013]" pageId="3" pageNumber="274">
<emphasis id="B9490866FFC1FF9BFF4564EC62A2FF0C" box="[199,389,187,213]" italics="true" pageId="3" pageNumber="274">Female genitalia</emphasis>
(
<figureCitation id="1306C8F1FFC1FF9BFE1064EC62F1FF0C" box="[402,470,187,213]" captionStart="FIGURE 9" captionStartId="6.[151,250,1897,1921]" captionTargetBox="[497,1080,181,1861]" captionTargetId="figure-26@6.[497,1090,180,1873]" captionTargetPageId="6" captionText="FIGURE 9. Female genital structures in Thaumatographa, showing main characters for the complex female genitalia of this genus (after Diakonoff 1986, for T. aurosa, Japan). Note: female genitalia in this genus typically also have a dorsal flap of unknown purpose on the tergal membrane of segment 8 (not shown in this sketch)." figureDoi="http://doi.org/10.5281/zenodo.14798841" httpUri="https://zenodo.org/record/14798841/files/figure.png" pageId="3" pageNumber="274">Fig. 9</figureCitation>
): complicated bursal and accessory sac form, usually with very long narrow but straight ductus bursae; ovipositor usually somewhat short, with setose and elongated papilla anales; apophyses usually subequal and average length; dorsum of tergite 8 on ovipositor sometimes with an upright flattened membranous flap (either laterally flattened or horizontally flattened); ostium a varied funnel-shape, on the intersegmental membrane between sternites 7 and 8; sterigma usually little developed, as a sclerotized or membranous margin; ductus bursae usually a long membranous tube uniformly narrow (rarely short and wide), but sometimes sclerotized on the pre-ostial section (rarely with some modification or colliculum-like formation), sometimes with a small ventral sac-like diverticulum (see
<figureCitation id="1306C8F1FFC1FF9BFD9665E06156FE08" box="[532,625,438,465]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798853" httpUri="https://zenodo.org/record/14798853/files/figure.png" pageId="3" pageNumber="274">Fig. 15a</figureCitation>
) internally spiculate and emergent from the ductus bursae by the antrum (rarely distally bifid, see
<figureCitation id="1306C8F1FFC1FF9BFE32658C612FFE2C" box="[432,520,475,501]" captionStart="FIGURE 14" captionStartId="13.[151,250,1341,1365]" captionTargetBox="[152,1436,181,1317]" captionTargetId="figure-20@13.[151,1436,181,1317]" captionTargetPageId="13" captionText="FIGURE 14. Thaumatographa fennchihwu Heppner &amp; Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798851" httpUri="https://zenodo.org/record/14798851/files/figure.png" pageId="3" pageNumber="274">Fig 14a</figureCitation>
); ductus seminalis varied, usually from corpus bursae junction with ductus bursae (or from near ostium); corpus busae usually semi-ovate and asymmetrical with one end somewhat elongated (rarely more uniformly elongate-ovate), with an elongate-ovate diverticular accessory sac on a short and slender tube from caudal end of bursa and near emergence of ductus seminalis and ductus bursae (ductus seminalis rarely from caudal end of ductus bursae); signum usually a fan-like cluster of dense and long spines curved inwardly (rarely as a single cluster of short spines, or signum diffuse or absent).
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC1FF9BFF4566E5613DFAA0" pageId="3" pageNumber="274" type="biology_ecology">
<paragraph id="8B82D474FFC1FF9BFF4566E56259FBA4" blockId="3.[151,1437,187,2013]" pageId="3" pageNumber="274">
<emphasis id="B9490866FFC1FF9BFF4566E56206FD15" bold="true" box="[199,289,690,716]" pageId="3" pageNumber="274">Biology</emphasis>
. Mostly unknown, but adults appear to all be diurnally active, yet may come to lights. Larvae are borers of shoots and roots as far as is known:
<taxonomicName id="4C3DAFF7FFC1FF9BFD0D6680676FFD28" authority="(Meyrick)" baseAuthorityName="Meyrick" baseAuthorityYear="1905" box="[655,1096,727,753]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="caminodes">
<emphasis id="B9490866FFC1FF9BFD0D668060F7FD28" box="[655,976,727,753]" italics="true" pageId="3" pageNumber="274">Thaumatographa caminodes</emphasis>
(Meyrick)
</taxonomicName>
feeding on roots of cardamom (
<taxonomicName id="4C3DAFF7FFC1FF9BFF2366AC6281FCCC" box="[161,422,763,789]" class="Liliopsida" family="Zingiberaceae" genus="Elettaria" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="cardamomum">
<emphasis id="B9490866FFC1FF9BFF2366AC6281FCCC" box="[161,422,763,789]" italics="true" pageId="3" pageNumber="274">Elettaria cardamomum</emphasis>
</taxonomicName>
) and also a wild
<taxonomicName id="4C3DAFF7FFC1FF9BFDF166AC6035FCCC" box="[627,786,763,789]" class="Liliopsida" family="Zingiberaceae" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Zingiberaceae</taxonomicName>
in
<collectingCountry id="F32A94E4FFC1FF9BFCB866AC608DFCCC" box="[826,938,763,789]" name="Sri Lanka" pageId="3" pageNumber="274">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFACA985FFC1FF9BFC3866AC6752FCCC" author="Diakonoff, A." box="[954,1141,763,789]" pageId="3" pageNumber="274" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff 1986</bibRefCitation>
),
<taxonomicName id="4C3DAFF7FFC1FF9BFB0866AC6276FCE0" authority="(Meyrick)" baseAuthorityName="Meyrick" baseAuthorityYear="1912" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="leucopyrga">
<emphasis id="B9490866FFC1FF9BFB0866AC63F0FCE1" italics="true" pageId="3" pageNumber="274">Thaumatographa leucopyrga</emphasis>
(Meyrick)
</taxonomicName>
in shoots of
<taxonomicName id="4C3DAFF7FFC1FF9BFE66674861B1FCE0" box="[484,662,799,825]" class="Magnoliopsida" family="Primulaceae" genus="Ardisia" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="sieboldi">
<emphasis id="B9490866FFC1FF9BFE66674861B1FCE0" box="[484,662,799,825]" italics="true" pageId="3" pageNumber="274">Ardisia sieboldi</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3DAFF7FFC1FF9BFD276748601FFCE0" box="[677,824,799,825]" class="Magnoliopsida" family="Myrsinaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Myrsinaceae</taxonomicName>
) in
<collectingRegion id="49F91A96FFC1FF9BFCE6674860B1FCE0" box="[868,918,799,825]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="274">Java</collectingRegion>
(
<collectingCountry id="F32A94E4FFC1FF9BFC276748673FFCE0" box="[933,1048,799,825]" name="Indonesia" pageId="3" pageNumber="274">Indonesia</collectingCountry>
), and
<taxonomicName id="4C3DAFF7FFC1FF9BFBDE6748622DFC84" authority="Diakonoff" authorityName="Diakonoff" authorityYear="1977" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="oenobapta">
<emphasis id="B9490866FFC1FF9BFBDE674866BBFCE0" box="[1116,1436,799,825]" italics="true" pageId="3" pageNumber="274">Thaumatographa oenobapta</emphasis>
Diakonoff
</taxonomicName>
in calyxes of
<taxonomicName id="4C3DAFF7FFC1FF9BFE26671462C6FC85" box="[420,481,835,860]" class="Magnoliopsida" family="Rubiaceae" genus="Ixora" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="genus">
<emphasis id="B9490866FFC1FF9BFE26671462C6FC85" box="[420,481,835,860]" italics="true" pageId="3" pageNumber="274">Ixora</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3DAFF7FFC1FF9BFE6D6714614DFC84" box="[495,618,835,861]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
), are the only examples known, but
<taxonomicName id="4C3DAFF7FFC1FF9BFC78671467B6FC84" baseAuthorityName="Meyrick" baseAuthorityYear="1912" box="[1018,1169,835,861]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="leucopyrga">
<emphasis id="B9490866FFC1FF9BFC78671467B6FC84" box="[1018,1169,835,861]" italics="true" pageId="3" pageNumber="274">T. leucopyrga</emphasis>
</taxonomicName>
is not a typical member of the genus. The other species with biological data that have been included among
<taxonomicName id="4C3DAFF7FFC1FF9BFBB9673067DAFC58" authorityName="Walsignham" authorityYear="1897" box="[1083,1277,871,897]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFBB9673067DAFC58" box="[1083,1277,871,897]" italics="true" pageId="3" pageNumber="274">Thaumatographa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFACA985FFC1FF9BFA8967306234FC7C" author="Diakonoff, A. &amp; Arita, Y." pageId="3" pageNumber="274" pagination="56 - 60" refId="ref11081" refString="Diakonoff, A. &amp; Arita, Y. (1981) The early stages of Thaumatographa eremnotorna Diakonoff &amp; Arita with remarks on the status of the Hilarographini (Lepidoptera: Tortricoidea). Entomologische Berichten, 41, 56-60." type="journal article" year="1981">Diakonoff &amp; Arita 1981</bibRefCitation>
;
<bibRefCitation id="EFACA985FFC1FF9BFE9C67DC62E6FC7C" author="Heppner, J. B." box="[286,449,907,933]" pageId="3" pageNumber="274" pagination="165 - 176" refId="ref11281" refString="Heppner, J. B. (1983) Revision of American Thaumatographa with a new species from Cuba (Lepidoptera: Tortricidae). Pan-Pacific Entomologist, 58, 165-176." type="journal article" year="1983">Heppner 1983</bibRefCitation>
,
<bibRefCitation id="EFACA985FFC1FF9BFE4E67DD6132FC7C" author="Heppner, J. B." box="[460,533,906,933]" pageId="3" pageNumber="274" refId="ref11535" refString="Heppner, J. B. (2025 b) Nearctic Hilarographa and a new genus for species in North America and Asia (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini). Lepidoptera Novae, 18, in press." type="journal volume" year="2025">2025b</bibRefCitation>
), appear to all be pine trunk feeders (cambium layer) but are atypical and will be transferred to their own genus:
<taxonomicName id="4C3DAFF7FFC1FF9BFE7767F86704FC10" authority="Diakonoff &amp; Arita" authorityName="Diakonoff &amp; Arita" box="[501,1059,943,969]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="eremnotorna">
<emphasis id="B9490866FFC1FF9BFE7767F86069FC11" box="[501,846,943,969]" italics="true" pageId="3" pageNumber="274">Thaumatographa eremnotorna</emphasis>
Diakonoff &amp; Arita
</taxonomicName>
from
<taxonomicName id="4C3DAFF7FFC1FF9BFBE567F86607FC10" box="[1127,1312,943,969]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="densiflora">
<emphasis id="B9490866FFC1FF9BFBE567F86607FC10" box="[1127,1312,943,969]" italics="true" pageId="3" pageNumber="274">Pinus densiflora</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3DAFF7FFC1FF9BFAAC67F866B2FC10" box="[1326,1429,943,969]" class="Pinopsida" family="Pinaceae" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Pinaceae</taxonomicName>
) in
<collectingCountry id="F32A94E4FFC1FF9BFF35678463DDFC34" box="[183,250,979,1005]" name="Japan" pageId="3" pageNumber="274">Japan</collectingCountry>
;
<taxonomicName id="4C3DAFF7FFC1FF9BFE8B678461EDFC34" authority="(Walsingham)" baseAuthorityName="Walsingham" baseAuthorityYear="1881" box="[265,714,979,1005]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="B9490866FFC1FF9BFE8B67846104FC34" box="[265,547,979,1005]" italics="true" pageId="3" pageNumber="274">Thaumatographa regalis</emphasis>
(Walsingham)
</taxonomicName>
from
<taxonomicName id="4C3DAFF7FFC1FF9BFC976784608EFC34" box="[789,937,979,1005]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="ponderosa">
<emphasis id="B9490866FFC1FF9BFC976784608EFC34" box="[789,937,979,1005]" italics="true" pageId="3" pageNumber="274">P. ponderosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3DAFF7FFC1FF9BFC6467846755FC34" box="[998,1138,979,1005]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="species" species="sabiniana">
<emphasis id="B9490866FFC1FF9BFC6467846755FC34" box="[998,1138,979,1005]" italics="true" pageId="3" pageNumber="274">P. sabiniana</emphasis>
</taxonomicName>
in
<collectingCountry id="F32A94E4FFC1FF9BFB1F678467F4FC34" box="[1181,1235,979,1005]" name="United States of America" pageId="3" pageNumber="274">USA</collectingCountry>
(
<collectingRegion id="49F91A96FFC1FF9BFB676784667AFC34" box="[1253,1373,979,1005]" country="United States of America" name="California" pageId="3" pageNumber="274">California</collectingRegion>
); and
<taxonomicName id="4C3DAFF7FFC1FF9BFF1567A0610FFBC8" authority="Heppner" authorityName="Heppner" authorityYear="1983" box="[151,552,1015,1041]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="cubensis">
<emphasis id="B9490866FFC1FF9BFF1567A062E7FBC8" box="[151,448,1015,1041]" italics="true" pageId="3" pageNumber="274">Thaumatographa cubensis</emphasis>
Heppner
</taxonomicName>
from
<taxonomicName id="4C3DAFF7FFC1FF9BFDEE67A061CFFBC8" authorityName="Heppner" authorityYear="1983" box="[620,744,1015,1041]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="cubensis">
<emphasis id="B9490866FFC1FF9BFDEE67A061CFFBC8" box="[620,744,1015,1041]" italics="true" pageId="3" pageNumber="274">P. cubensis</emphasis>
</taxonomicName>
in
<collectingCountry id="F32A94E4FFC1FF9BFC8967A0606DFBC8" box="[779,842,1015,1041]" name="Cuba" pageId="3" pageNumber="274">Cuba</collectingCountry>
. None of the other more tropical species of the genus have any host data available (likewise for species of
<taxonomicName id="4C3DAFF7FFC1FF9BFD69604C60A1FBEC" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[747,902,1051,1077]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFD69604C60A1FBEC" box="[747,902,1051,1077]" italics="true" pageId="3" pageNumber="274">Hilarographa</emphasis>
</taxonomicName>
from the Neotropics), so we do not know what tropical trees they may feed on, but perhaps the other Asian species are also on
<taxonomicName id="4C3DAFF7FFC1FF9BFBCE606867F9FB80" box="[1100,1246,1087,1113]" class="Magnoliopsida" family="Myrsinaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Myrsinaceae</taxonomicName>
,
<taxonomicName id="4C3DAFF7FFC1FF9BFB6F60686642FB80" box="[1261,1381,1087,1113]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
, and
<taxonomicName id="4C3DAFF7FFC1FF9BFF1560346211FBA4" box="[151,310,1123,1149]" class="Liliopsida" family="Zingiberaceae" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="274" phylum="Tracheophyta" rank="family">Zingiberaceae</taxonomicName>
hosts.
</paragraph>
<paragraph id="8B82D474FFC1FF9BFF4560D0613DFAA0" blockId="3.[151,1437,187,2013]" pageId="3" pageNumber="274">
The only larva and pupa of
<taxonomicName id="4C3DAFF7FFC1FF9BFD8C60D06194FB78" box="[526,691,1159,1185]" class="Insecta" family="Tortricidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">Hilarographini</taxonomicName>
with detailed descriptions are for
<taxonomicName id="4C3DAFF7FFC1FF9BFBC060D067D6FB79" box="[1090,1265,1159,1184]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="species" species="eremnotorna">
<emphasis id="B9490866FFC1FF9BFBC060D067D6FB79" box="[1090,1265,1159,1184]" italics="true" pageId="3" pageNumber="274">T. eremnotorna</emphasis>
</taxonomicName>
in
<collectingCountry id="F32A94E4FFC1FF9BFA9860D0667AFB78" box="[1306,1373,1159,1185]" name="Japan" pageId="3" pageNumber="274">Japan</collectingCountry>
, with details described by
<bibRefCitation id="EFACA985FFC1FF9BFEF860FC61BEFB1C" author="Diakonoff, A. &amp; Arita, Y." box="[378,665,1195,1221]" pageId="3" pageNumber="274" pagination="56 - 60" refId="ref11081" refString="Diakonoff, A. &amp; Arita, Y. (1981) The early stages of Thaumatographa eremnotorna Diakonoff &amp; Arita with remarks on the status of the Hilarographini (Lepidoptera: Tortricoidea). Entomologische Berichten, 41, 56-60." type="journal article" year="1981">Diakonoff &amp; Arita (1981)</bibRefCitation>
, but this species will be transferred to a related new genus of northern pine-feeding species (
<bibRefCitation id="EFACA985FFC1FF9BFE0E60986165FB30" author="Heppner, J. B." box="[396,578,1230,1257]" pageId="3" pageNumber="274" refId="ref11535" refString="Heppner, J. B. (2025 b) Nearctic Hilarographa and a new genus for species in North America and Asia (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini). Lepidoptera Novae, 18, in press." type="journal volume" year="2025">Heppner 2025b</bibRefCitation>
). The larval characters described for this species, nonetheless, may have the main characters applicable also to the tribe, since larvae of all the species are thought to be borers. The primary distinction of this borer larva from
<collectingCountry id="F32A94E4FFC1FF9BFD9C61406179FAE8" box="[542,606,1303,1329]" name="Japan" pageId="3" pageNumber="274">Japan</collectingCountry>
is the bisetose state of the prothoracic L-group setae, normally trisetose in all other
<taxonomicName id="4C3DAFF7FFC1FF9BFF7A616C6255FA8C" authorityName="Latreille" authorityYear="1803" box="[248,370,1339,1365]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
and including the other group of borer larvae in the subfamily
<taxonomicName id="4C3DAFF7FFC1FF9BFBA7616C679AFA8C" authorityName="Walsingham" authorityYear="1895" box="[1061,1213,1339,1365]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="subFamily" subFamily="Olethreutinae">Olethreutinae</taxonomicName>
(
<bibRefCitation id="EFACA985FFC1FF9BFB49616C6674FA8C" author="Horak, M." box="[1227,1363,1339,1365]" pageId="3" pageNumber="274" pagination="3 - 64" refId="ref11678" refString="Horak, M. (1984) Assessment of taxonomically significant structures in Tortricinae (Lep., Tortricidae). Mitteilungen der Schweizerische Entomologische Gessellschaft, 57, 3-64." type="journal article" year="1984">Horak 1984</bibRefCitation>
,
<bibRefCitation id="EFACA985FFC1FF9BFADF616C66B0FA8C" author="Horak, M." box="[1373,1431,1339,1365]" pageId="3" pageNumber="274" pagination="1 - 22" refId="ref11711" refString="Horak, M. (1991) Morphology, phylogeny and systematics. In: van der Geest, L. P. S. &amp; Evenhuis, H. H. (Eds.), Tortricid Pests, their Biology, Natural Enemies and Control. Elsevier Scientific Publishers, Amsterdam, pp. 1-22." type="book chapter" year="1991">1991</bibRefCitation>
,
<bibRefCitation id="EFACA985FFC1FF9BFF15610863F5FAA0" author="Horak, M." box="[151,210,1375,1401]" pageId="3" pageNumber="274" pagination="199 - 215" refId="ref11771" refString="Horak, M. (1998) The Tortricoidea. In: Kristensen, N. P. (Ed.), Lepidoptera, Moths and Butterflies. Vol. 1. Evolution, Systematics, and Biogeography, In: Handbuch der Zoologie. Band IV. Arthropoda: Insecta. Teilband 35. Walter De Gruyter, Berlin, pp. 199-215. [1999] https://doi.org/10.1515/9783110804744.199" type="book chapter" year="1998">1998</bibRefCitation>
;
<bibRefCitation id="EFACA985FFC1FF9BFF5C610862A7FAA0" author="MacKay, M. R." box="[222,384,1374,1401]" pageId="3" pageNumber="274" pagination="1 - 338" refId="ref12006" refString="MacKay, M. R. (1959) Larvae of the North American Olethreutidae (Lepidoptera). Canadian Entomologist, 91 (Supplement 10), 1-338. https://doi.org/10.4039/entm9110fv" type="journal article" year="1959">MacKay 1959</bibRefCitation>
,
<bibRefCitation id="EFACA985FFC1FF9BFE09610862E1FAA0" author="MacKay, M. R." box="[395,454,1375,1401]" pageId="3" pageNumber="274" pagination="1 - 182" refId="ref12050" refString="MacKay, M. R. (1962) Larvae of the North American Tortricinae (Lepidoptera: Tortricidae). Canadian Entomologist, Supplement 28, 1-182. https://doi.org/10.4039/entm9428fv" type="journal article" year="1962">1962</bibRefCitation>
,
<bibRefCitation id="EFACA985FFC1FF9BFE536108612AFAA0" author="MacKay, M. R." box="[465,525,1375,1401]" pageId="3" pageNumber="274" pagination="1321 - 1344" refId="ref12093" refString="MacKay, M. R. (1963) Evolution and adaptation of larval characters in the Tortricidae. Canadian Entomologist, 95, 1321-1344. https://doi.org/10.4039/Ent951321-12" type="journal article" year="1963">1963</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC1FF9BFF4561D562BCF964" pageId="3" pageNumber="274" type="distribution">
<paragraph id="8B82D474FFC1FF9BFF4561D562BCF964" blockId="3.[151,1437,187,2013]" pageId="3" pageNumber="274">
<emphasis id="B9490866FFC1FF9BFF4561D5627EFA45" bold="true" box="[199,345,1410,1436]" pageId="3" pageNumber="274">Distribution</emphasis>
. South Asia (
<collectingCountry id="F32A94E4FFC1FF9BFE7A61D46114FA44" box="[504,563,1411,1437]" name="India" pageId="3" pageNumber="274">India</collectingCountry>
and
<collectingCountry id="F32A94E4FFC1FF9BFDED61D461C6FA44" box="[623,737,1411,1437]" name="Sri Lanka" pageId="3" pageNumber="274">Sri Lanka</collectingCountry>
) to Southeast Asia, from
<collectingCountry id="F32A94E4FFC1FF9BFB8B61D4674AFA44" box="[1033,1133,1411,1437]" name="Thailand" pageId="3" pageNumber="274">Thailand</collectingCountry>
to
<collectingCountry id="F32A94E4FFC1FF9BFB1761D467D3FA44" box="[1173,1268,1411,1437]" name="Vietnam" pageId="3" pageNumber="274">Vietnam</collectingCountry>
and
<collectingCountry id="F32A94E4FFC1FF9BFAAD61D466BBFA44" box="[1327,1436,1411,1437]" name="Indonesia" pageId="3" pageNumber="274">Indonesia</collectingCountry>
(including Borneo,
<collectingRegion id="49F91A96FFC1FF9BFEF461F0628DFA18" box="[374,426,1447,1473]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="274">Java</collectingRegion>
,
<collectingCountry id="F32A94E4FFC1FF9BFE3B61F0610EFA18" box="[441,553,1447,1473]" name="Indonesia" pageId="3" pageNumber="274">Moluccas</collectingCountry>
,
<collectingRegion id="49F91A96FFC1FF9BFDB461F061B9FA18" box="[566,670,1447,1473]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="274">Sulawesi</collectingRegion>
, and
<collectingRegion id="49F91A96FFC1FF9BFD5F61F0601AFA18" box="[733,829,1447,1473]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="274">Sumatra</collectingRegion>
, and undoubtedly the other islands as well), north to southern
<collectingCountry id="F32A94E4FFC1FF9BFF7C619C6266FA3C" box="[254,321,1483,1509]" name="China" pageId="3" pageNumber="274">China</collectingCountry>
(
<collectingRegion id="49F91A96FFC1FF9BFECD619C6283FA3C" box="[335,420,1483,1509]" country="China" name="Hainan" pageId="3" pageNumber="274">Hainan</collectingRegion>
,
<collectingCountry id="F32A94E4FFC1FF9BFE2D619C6112FA3C" box="[431,565,1483,1509]" name="Hong Kong" pageId="3" pageNumber="274">Hong Kong</collectingCountry>
, Taiwan) and
<collectingCountry id="F32A94E4FFC1FF9BFD52619C6034FA3C" box="[720,787,1483,1509]" name="Japan" pageId="3" pageNumber="274">Japan</collectingCountry>
, and in northeastern
<collectingCountry id="F32A94E4FFC1FF9BFC7A619C6778FA3C" box="[1016,1119,1483,1509]" name="Australia" pageId="3" pageNumber="274">Australia</collectingCountry>
(
<collectingRegion id="49F91A96FFC1FF9BFBEF619C67D2FA3C" box="[1133,1269,1483,1509]" country="Australia" name="Queensland" pageId="3" pageNumber="274">Queensland</collectingRegion>
) and the South Pacific (New
<collectingCountry id="F32A94E4FFC1FF9BFEB061B862A4F9D0" box="[306,387,1519,1545]" name="Guinea-Bissau" pageId="3" pageNumber="274">Guinea</collectingCountry>
and Solomons, and an undescribed species from Tahiti) (
<bibRefCitation id="EFACA985FFC1FF9BFB8A61B86798F9D0" author="Diakonoff, A." box="[1032,1215,1519,1545]" pageId="3" pageNumber="274" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff 1986</bibRefCitation>
;
<bibRefCitation id="EFACA985FFC1FF9BFB4E61B8665CF9D0" author="Heppner, J. B." box="[1228,1403,1518,1545]" pageId="3" pageNumber="274" refId="ref11576" refString="Heppner, J. B. (2025 c) A new Thaumatographa specices from Tahiti (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini). Lepidoptera Novae, 18, in press." type="journal volume" year="2025">Heppner 2025c</bibRefCitation>
, d;
<bibRefCitation id="EFACA985FFC1FF9BFF1562446271F9F4" author="Razowski, J." box="[151,342,1555,1581]" pageId="3" pageNumber="274" pagination="261 - 287" refId="ref12491" refString="Razowski, J. (2009 a) The Old World Hilarographini (Lepidoptera: Tortricidae). SHILAP Revista Lepidopterologica, 37, 261-287." type="journal article" year="2009">Razowski 2009a</bibRefCitation>
). The genus is undoubtedly also in the
<collectingCountry id="F32A94E4FFC1FF9BFC89624460AEF9F4" box="[779,905,1555,1581]" name="Philippines" pageId="3" pageNumber="274">Philippines</collectingCountry>
but has not been recorded there yet (
<bibRefCitation id="EFACA985FFC1FF9BFAA5624463F4F988" author="Diakonoff, A." pageId="3" pageNumber="274" pagination="1 - 484" refId="ref10826" refString="Diakonoff, A. ([1968]) Microlepidoptera of the Philippine Islands. Bulletin of the United States National Museum, 257, 1-484. [1967] https://doi.org/10.5479/si.03629236.257.1" type="journal article" year="1968">Diakonoff 1968</bibRefCitation>
). Three species are recorded in North America (
<bibRefCitation id="EFACA985FFC1FF9BFD74626060BCF988" author="Heppner, J. B." box="[758,923,1591,1617]" pageId="3" pageNumber="274" pagination="165 - 176" refId="ref11281" refString="Heppner, J. B. (1983) Revision of American Thaumatographa with a new species from Cuba (Lepidoptera: Tortricidae). Pan-Pacific Entomologist, 58, 165-176." type="journal article" year="1983">Heppner 1983</bibRefCitation>
), and one related pine-feeder from
<collectingCountry id="F32A94E4FFC1FF9BFAAC6260664BF988" box="[1326,1388,1591,1617]" name="Cuba" pageId="3" pageNumber="274">Cuba</collectingCountry>
, but these are partially in a related new genus while two are in the genus
<taxonomicName id="4C3DAFF7FFC1FF9BFC1F620C671FF9AC" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[925,1080,1627,1653]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFC1F620C671FF9AC" box="[925,1080,1627,1653]" italics="true" pageId="3" pageNumber="274">Hilarographa</emphasis>
</taxonomicName>
. The pine feeders in the related genus occur in
<collectingCountry id="F32A94E4FFC1FF9BFEC3622862A3F940" box="[321,388,1663,1689]" name="Japan" pageId="3" pageNumber="274">Japan</collectingCountry>
, the Kuril Islands (Russian Far East) and North America (plus one in
<collectingCountry id="F32A94E4FFC1FF9BFB0C622867ECF940" box="[1166,1227,1663,1689]" name="Cuba" pageId="3" pageNumber="274">Cuba</collectingCountry>
) (
<bibRefCitation id="EFACA985FFC1FF9BFB63622866B0F940" author="Diakonoff, A." box="[1249,1431,1663,1689]" pageId="3" pageNumber="274" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff 1986</bibRefCitation>
;
<bibRefCitation id="EFACA985FFC1FF9BFF1562F4621DF964" author="Heppner, J. B." box="[151,314,1699,1725]" pageId="3" pageNumber="274" pagination="165 - 176" refId="ref11281" refString="Heppner, J. B. (1983) Revision of American Thaumatographa with a new species from Cuba (Lepidoptera: Tortricidae). Pan-Pacific Entomologist, 58, 165-176." type="journal article" year="1983">Heppner 1983</bibRefCitation>
,
<bibRefCitation id="EFACA985FFC1FF9BFEC762F562B7F964" author="Heppner, J. B." box="[325,400,1698,1725]" pageId="3" pageNumber="274" refId="ref11535" refString="Heppner, J. B. (2025 b) Nearctic Hilarographa and a new genus for species in North America and Asia (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini). Lepidoptera Novae, 18, in press." type="journal volume" year="2025">2025b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC1FF9CFF456291677EFEC4" lastPageId="4" lastPageNumber="275" pageId="3" pageNumber="274" type="discussion">
<paragraph id="8B82D474FFC1FF9BFF4562916102F804" blockId="3.[151,1437,187,2013]" pageId="3" pageNumber="274">
<emphasis id="B9490866FFC1FF9BFF4562916262F939" bold="true" box="[199,325,1734,1760]" pageId="3" pageNumber="274">Discussion</emphasis>
.
<bibRefCitation id="EFACA985FFC1FF9BFED662906132F938" author="Razowski, J." box="[340,533,1735,1761]" pageId="3" pageNumber="274" pagination="1 - 16" refId="ref12550" refString="Razowski, J. (2017) Diagnoses and remarks on the genera of Tortricidae (Lepidoptera). Part 5. Chlidanotinae. Acta Zoologica Cracovoensia, 60, 1-16. https://doi.org/10.3409/azc.60_1.01" type="journal article" year="2017">Razowski (2017)</bibRefCitation>
synonymized
<taxonomicName id="4C3DAFF7FFC1FF9BFD3862906732F938" authority="Walsingham, 1897" authorityName="Walsingham" authorityYear="1897" box="[698,1045,1735,1761]" class="Insecta" family="Tortricidae" genus="Idiothauma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFD386290601CF938" box="[698,827,1735,1761]" italics="true" pageId="3" pageNumber="274">Idiothauma</emphasis>
<bibRefCitation id="EFACA985FFC1FF9BFCC162906732F938" author="Walsingham, T. de" box="[835,1045,1735,1761]" pageId="3" pageNumber="274" pagination="33 - 67" refId="ref12725" refString="Walsingham, T. de G. (1897) Western Equatorial African Micro-Lepidoptera. Transactions of the Entomological Society of London, 1897, 33-67, pls. 2 - 3. https://doi.org/10.1111/j.1365-2311.1897.tb01674.x" type="journal article" year="1897">Walsingham, 1897</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="4C3DAFF7FFC1FF9BFBD5629067D5F938" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[1111,1266,1735,1761]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFBD5629067D5F938" box="[1111,1266,1735,1761]" italics="true" pageId="3" pageNumber="274">Hilarographa</emphasis>
</taxonomicName>
, but it remains a distinct genus of three African species (
<bibRefCitation id="EFACA985FFC1FF9BFDE962BC6006F8DC" author="Heppner, J. B." box="[619,801,1770,1797]" pageId="3" pageNumber="274" refId="ref11610" refString="Heppner, J. B. (2025 d) Notes on Hilarographine genera (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini). Lepidoptera Novae, 18, in press." type="journal volume" year="2025">Heppner 2025d</bibRefCitation>
). These two genera and
<taxonomicName id="4C3DAFF7FFC1FF9BFBB662BC67D1F8DC" authorityName="Walsignham" authorityYear="1897" box="[1076,1270,1771,1797]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFBB662BC67D1F8DC" box="[1076,1270,1771,1797]" italics="true" pageId="3" pageNumber="274">Thaumatographa</emphasis>
</taxonomicName>
are all closely related genera.
<bibRefCitation id="EFACA985FFC1FF9BFEC86358613CF8F0" author="Razowski, J." box="[330,539,1807,1833]" pageId="3" pageNumber="274" pagination="261 - 287" refId="ref12491" refString="Razowski, J. (2009 a) The Old World Hilarographini (Lepidoptera: Tortricidae). SHILAP Revista Lepidopterologica, 37, 261-287." type="journal article" year="2009">Razowski (2009a)</bibRefCitation>
described numerous new species from Southeast Asia (all as ‘
<taxonomicName id="4C3DAFF7FFC1FF9BFB78635866B2F8F0" baseAuthorityName="Razowski" baseAuthorityYear="2009" box="[1274,1429,1807,1833]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFB78635866B2F8F0" box="[1274,1429,1807,1833]" italics="true" pageId="3" pageNumber="274">Hilarographa</emphasis>
</taxonomicName>
’ but transferred to
<taxonomicName id="4C3DAFF7FFC1FF9BFEE663646101F894" authorityName="Walsignham" authorityYear="1897" box="[356,550,1843,1869]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFEE663646101F894" box="[356,550,1843,1869]" italics="true" pageId="3" pageNumber="274">Thaumatographa</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFACA985FFC1FF9BFDE163646032F894" author="Heppner, J. B." box="[611,789,1842,1869]" pageId="3" pageNumber="274" refId="ref11610" refString="Heppner, J. B. (2025 d) Notes on Hilarographine genera (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini). Lepidoptera Novae, 18, in press." type="journal volume" year="2025">Heppner 2025d</bibRefCitation>
), with varying wing shapes and somewhat varied genital morphology, so further study will be needed to verify if all are really in the same genus. Most species of the genus are known from only single or very few specimens. The 2-bristle frenulum of female
<taxonomicName id="4C3DAFF7FFC1FF9BFBCF632C6628F84C" authorityName="Walsignham" authorityYear="1897" box="[1101,1295,1915,1941]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC1FF9BFBCF632C6628F84C" box="[1101,1295,1915,1941]" italics="true" pageId="3" pageNumber="274">Thaumatographa</emphasis>
</taxonomicName>
conforms to this morphology in
<taxonomicName id="4C3DAFF7FFC1FF9BFEF063C86130F860" box="[370,535,1951,1977]" class="Insecta" family="Tortricidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">Hilarographini</taxonomicName>
and for the subfamily
<taxonomicName id="4C3DAFF7FFC1FF9BFC9163C86094F860" box="[787,947,1951,1977]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="subFamily" subFamily="Chlidanotinae">Chlidanotinae</taxonomicName>
, versus the 3-bristle norm for the remainder of tortricids (
<bibRefCitation id="EFACA985FFC1FF9BFEAB6394613EF804" author="Yang, A. &amp; Brown, J. W." box="[297,537,1987,2013]" pageId="3" pageNumber="274" pagination="743 - 751" refId="ref12779" refString="Yang, A. &amp; Brown, J. W. (2009) Variation in the female frenulum in Tortricidae (Lepidoptera) Part 1. Chlidanotinae. Proceedings of the Entomological Society of Washington, 111, 743-751. https://doi.org/10.4289/0013-8797-111.3.743" type="journal article" year="2009">Yang &amp; Brown 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8B82D474FFC6FF9CFF4564C0677EFEC4" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="275">
The thin ovate dorsal flap at the middle of tergite
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females is present in some
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<emphasis id="B9490866FFC6FF9CFBFD64C06666FF68" box="[1151,1345,151,177]" italics="true" pageId="4" pageNumber="275">Thaumatographa</emphasis>
species
</taxonomicName>
but the function of this structure is unknown. Likewise, some species have a bulbous flap (or pouch) ventrally at the base of the ostial funnel at the caudel end of the ductus bursae, seemingly emergent approximate to the antrum (sometimes even distally bifid), but the function of this sac or pouch is also unknown.
</paragraph>
</subSubSection>
</treatment>
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598
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<emphasis id="B9490866FFC6FF9CFF156539610DFE50" bold="true" box="[151,554,366,393]" italics="true" pageId="4" pageNumber="275">Thaumatographa mesostigmatias</emphasis>
Diakonoff, 1977
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<paragraph id="8B82D474FFC6FF9CFF1565C46241FE74" blockId="4.[151,761,366,429]" box="[151,358,403,429]" pageId="4" pageNumber="275">
(
<figureCitation id="1306C8F1FFC6FF9CFF1D65C4622CFE74" box="[159,267,403,429]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="4" pageNumber="275">Figs. 1–2</figureCitation>
,
<figureCitation id="1306C8F1FFC6FF9CFE9565C46212FE74" box="[279,309,403,429]" captionStart="FIGURE 10" captionStartId="8.[151,250,1905,1929]" captionTargetBox="[170,1418,151,1889]" captionTargetId="figure-26@8.[170,1418,151,1889]" captionTargetPageId="8" captionText="FIGURE 10. Thaumatographa mesostigmatias Diakonoff, male genitalia (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus in situ, and with ductus ejaculatorius (a, reversed), enlargement of uncus to juxta (b, with distal end of aedeagus and cornutus hook), and structures of the pregenital abdomen and coremata (c) (gen. slide YA-1159) (MGCL); and d) drawing of male holotype genitalia (Taiwan: [Kuantzuling, Tainan Co.] (gen. slide AD-9811, USNM) (after Diakonoff, 1977b)." figureDoi="http://doi.org/10.5281/zenodo.14798843" httpUri="https://zenodo.org/record/14798843/files/figure.png" pageId="4" pageNumber="275">10</figureCitation>
,
<figureCitation id="1306C8F1FFC6FF9CFEC365C4627AFE74" box="[321,349,403,429]" captionStart="FIGURE 13" captionStartId="11.[151,250,1455,1479]" captionTargetBox="[160,1436,237,1432]" captionTargetId="figure-57@11.[151,1436,237,1432]" captionTargetPageId="11" captionText="FIGURE 13. Thaumatographa mesostigmatias Diakonoff, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798849" httpUri="https://zenodo.org/record/14798849/files/figure.png" pageId="4" pageNumber="275">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC6FF9CFF1565806675FDCA" pageId="4" pageNumber="275" type="reference_group">
<paragraph id="8B82D474FFC6FF9CFF1565806675FDCA" blockId="4.[151,1379,471,531]" pageId="4" pageNumber="275">
<treatmentCitationGroup id="AB2DF35AFFC6FF9CFF1565806644FE36" box="[151,1379,471,495]" pageId="4" pageNumber="275">
<treatmentCitation id="0A9CF265FFC6FF9CFF15658061EAFE36" author="Diakonoff, A." box="[151,717,471,495]" page="42" pageId="4" pageNumber="275" year="1977">
<taxonomicName id="4C3DAFF7FFC6FF9CFF15658061EAFE36" authority="Diakonoff, 1977 b: 42" authorityName="Diakonoff &amp; Arita" authorityPageNumber="42" authorityYear="1977" box="[151,717,471,495]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFC6FF9CFF15658062CEFE36" box="[151,489,471,495]" italics="true" pageId="4" pageNumber="275">Thaumatographa mesostigmatias</emphasis>
<bibRefCitation id="EFACA985FFC6FF9CFE72658061EAFE36" author="Diakonoff, A." box="[496,717,471,495]" pageId="4" pageNumber="275" pagination="3 - 55" refId="ref10939" refString="Diakonoff, A. (1977 b) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 3-55." type="journal article" year="1977">Diakonoff, 1977b: 42</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type-locality:
<collectingCountry id="F32A94E4FFC6FF9CFCE865806093FE36" box="[874,948,471,495]" name="Taiwan" pageId="4" pageNumber="275">Taiwan</collectingCountry>
[
<collectingRegion id="49F91A96FFC6FF9CFC42658F672AFE36" box="[960,1037,472,495]" country="Taiwan" name="Nantou" pageId="4" pageNumber="275">Nantou</collectingRegion>
Co.]: “Rantaisan” [= Lantashan].
</treatmentCitationGroup>
<treatmentCitationGroup id="AB2DF35AFFC6FF9CFF1565AC6675FDCA" box="[151,1362,507,531]" pageId="4" pageNumber="275">
<treatmentCitation id="0A9CF265FFC6FF9CFF1565AC616EFDCA" author="Razowski, J." box="[151,585,507,531]" page="276" pageId="4" pageNumber="275" year="2009">
<taxonomicName id="4C3DAFF7FFC6FF9CFF1565AC616EFDCA" ID-CoL="B3TKQ" authority="Razowski, 2009a: 276" authorityName="Razowski" authorityPageNumber="276" authorityYear="2009" box="[151,585,507,531]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="species" species="calyx" status="syn. nov.">
<emphasis id="B9490866FFC6FF9CFF1565AC6279FDCA" box="[151,350,507,531]" italics="true" pageId="4" pageNumber="275">Hilarographa calyx</emphasis>
<bibRefCitation id="EFACA985FFC6FF9CFEE765AC616EFDCA" author="Razowski, J." box="[357,585,507,531]" pageId="4" pageNumber="275" pagination="261 - 287" refId="ref12491" refString="Razowski, J. (2009 a) The Old World Hilarographini (Lepidoptera: Tortricidae). SHILAP Revista Lepidopterologica, 37, 261-287." type="journal article" year="2009">Razowski, 2009a: 276</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B9490866FFC6FF9CFDD665AC6196FDCA" bold="true" box="[596,689,507,531]" pageId="4" pageNumber="275">
<taxonomicNameLabel id="A27AB51DFFC6FF9CFDD665AC6196FDCA" box="[596,689,507,531]" pageId="4" pageNumber="275">syn. nov.</taxonomicNameLabel>
</emphasis>
Type-locality:
<collectingCountry id="F32A94E4FFC6FF9CFCC965AC60B2FDCA" box="[843,917,507,531]" name="Taiwan" pageId="4" pageNumber="275">Taiwan</collectingCountry>
[
<collectingRegion id="49F91A96FFC6FF9CFC2365AC60CFFDCA" box="[929,1000,507,531]" country="Taiwan" name="Tainan" pageId="4" pageNumber="275">Tainan</collectingRegion>
Co.]: “Kanshirei” [= Kuantzuling].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF4284FCFFC6FF9CFF15624F619AF8F5" ID-DOI="http://doi.org/10.5281/zenodo.14798835" ID-Zenodo-Dep="14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="4" pageNumber="275" startId="4.[151,264,1560,1584]" targetBox="[151,1436,555,1536]" targetPageId="4" targetType="figure">
<paragraph id="8B82D474FFC6FF9CFF15624F6097F93D" blockId="4.[151,1436,1560,1836]" pageId="4" pageNumber="275">
<emphasis id="B9490866FFC6FF9CFF15624F621DF9E9" bold="true" box="[151,314,1560,1584]" pageId="4" pageNumber="275">FIGURES 1–6.</emphasis>
Adults of Taiwan
<taxonomicName id="4C3DAFF7FFC6FF9CFE71624F61D2F9E9" box="[499,757,1560,1584]" pageId="4" pageNumber="275">
<emphasis id="B9490866FFC6FF9CFE71624F6184F9E9" box="[499,675,1560,1584]" italics="true" pageId="4" pageNumber="275">Thaumatographa</emphasis>
species
</taxonomicName>
, with lateral views of heads (a) for each moth:
<emphasis id="B9490866FFC6FF9CFB4F624F67C5F9E9" bold="true" box="[1229,1250,1560,1584]" pageId="4" pageNumber="275">1)</emphasis>
<taxonomicName id="4C3DAFF7FFC6FF9CFB6A624E627FF98D" authority="Diakonoff &amp; Arita" authorityName="Diakonoff &amp; Arita" authorityYear="1977" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFC6FF9CFB6A624E66BBF9E9" box="[1256,1436,1561,1584]" italics="true" pageId="4" pageNumber="275">T. mesostigmatias</emphasis>
Diakonoff &amp; Arita
</taxonomicName>
, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm);
<emphasis id="B9490866FFC6FF9CFC6B626B60D9F98D" bold="true" box="[1001,1022,1596,1620]" pageId="4" pageNumber="275">2)</emphasis>
same data, female (photo14173) (WS = 14.8 mm) (MGCL).
<emphasis id="B9490866FFC6FF9CFEEA62376259F9A1" bold="true" box="[360,382,1632,1656]" pageId="4" pageNumber="275">3)</emphasis>
<taxonomicName id="4C3DAFF7FFC6FF9CFE06623661E0F9A1" authority="Heppner &amp; Arita" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[388,711,1632,1656]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="species" species="fennchihwu" status="sp. nov.">
<emphasis id="B9490866FFC6FF9CFE0662366135F9A1" box="[388,530,1632,1656]" italics="true" pageId="4" pageNumber="275">T. fennchihwu</emphasis>
Heppner &amp; Arita
</taxonomicName>
,
<emphasis id="B9490866FFC6FF9CFD5362376005F9A1" bold="true" box="[721,802,1632,1656]" pageId="4" pageNumber="275">
<taxonomicNameLabel id="A27AB51DFFC6FF9CFD5362376005F9A1" box="[721,802,1632,1656]" pageId="4" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm);
<emphasis id="B9490866FFC6FF9CFE8762D3623DF945" bold="true" box="[261,282,1668,1692]" pageId="4" pageNumber="275">4)</emphasis>
same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL).
<emphasis id="B9490866FFC6FF9CFC5262D360C1F945" bold="true" box="[976,998,1668,1692]" pageId="4" pageNumber="275">5)</emphasis>
<taxonomicName id="4C3DAFF7FFC6FF9CFC6962D26668F945" authority="Heppner &amp; Arita" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[1003,1359,1668,1692]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="subSpecies" species="aurosa" status="subsp. nov." subSpecies="taiwana">
<emphasis id="B9490866FFC6FF9CFC6962D267B8F945" box="[1003,1183,1669,1692]" italics="true" pageId="4" pageNumber="275">T. aurosa taiwana</emphasis>
Heppner &amp; Arita
</taxonomicName>
,
<emphasis id="B9490866FFC6FF9CFADA62D363E4F919" bold="true" pageId="4" pageNumber="275">
<taxonomicNameLabel id="A27AB51DFFC6FF9CFADA62D363E4F919" pageId="4" pageNumber="275" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL).
<emphasis id="B9490866FFC6FF9CFC6462FF60DCF919" bold="true" box="[998,1019,1704,1728]" pageId="4" pageNumber="275">6)</emphasis>
<taxonomicName id="4C3DAFF7FFC6FF9CFB8062FE661EF919" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2025" box="[1026,1337,1704,1728]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="species" species="kentingana" status="sp. nov.">
<emphasis id="B9490866FFC6FF9CFB8062FE67AAF919" box="[1026,1165,1704,1728]" italics="true" pageId="4" pageNumber="275">T. kentingana</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<emphasis id="B9490866FFC6FF9CFAC662FF66B2F919" bold="true" box="[1348,1429,1704,1728]" pageId="4" pageNumber="275">
<taxonomicNameLabel id="A27AB51DFFC6FF9CFAC662FF66B2F919" box="[1348,1429,1704,1728]" pageId="4" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM).
</paragraph>
<paragraph id="8B82D474FFC6FF9CFF1562A6619AF8F5" blockId="4.[151,1436,1560,1836]" pageId="4" pageNumber="275">Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a.</paragraph>
</caption>
<subSubSection id="C32787FFFFC6FF9FFF15630D673FF939" lastPageId="7" lastPageNumber="278" pageId="4" pageNumber="275" type="diagnosis">
<paragraph id="8B82D474FFC6FF9DFF15630D6739FEC3" blockId="4.[151,1436,1882,2017]" lastBlockId="5.[151,1436,151,283]" lastPageId="5" lastPageNumber="276" pageId="4" pageNumber="275">
<emphasis id="B9490866FFC6FF9CFF15630D622DF8AD" bold="true" box="[151,266,1882,1908]" pageId="4" pageNumber="275">Diagnosis</emphasis>
. This first species of
<taxonomicName id="4C3DAFF7FFC6FF9CFD8A630D61EDF8AD" authorityName="Walsignham" authorityYear="1897" box="[520,714,1882,1908]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC6FF9CFD8A630D61EDF8AD" box="[520,714,1882,1908]" italics="true" pageId="4" pageNumber="275">Thaumatographa</emphasis>
</taxonomicName>
described from
<collectingCountry id="F32A94E4FFC6FF9CFC0A630D60FDF8AD" box="[904,986,1882,1908]" name="Taiwan" pageId="4" pageNumber="275">Taiwan</collectingCountry>
already in 1977, is large for the genus (
<figureCitation id="1306C8F1FFC6FF9CFF1D6329622CF841" box="[159,267,1918,1944]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="4" pageNumber="275">Figs. 1–2</figureCitation>
), with a prominent preapical orange bar on the forewing, and five short subterminal black bar spots from the tornus to middle of forewing, and three prominent pale yellow-white curved lines from the dorsal margin; both sexes with equal maculation. The hindwing has a pale orange (nearly white in some specimens) and elongated median patch from middle of forewing to the wing base. The male genitalia (
<figureCitation id="1306C8F1FFC7FF9DFC7864C0671AFF68" box="[1018,1085,151,177]" captionStart="FIGURE 8" captionStartId="5.[151,250,1802,1826]" captionTargetBox="[156,1435,1263,1779]" captionTargetId="figure-160@5.[151,1436,1263,1779]" captionTargetPageId="5" captionText="FIGURE 8. Male genital structures in Thaumatographa, showing main characters for the complex male genitalia of this genus (modified after Diakonoff 1977, for T. zapyra, New Guinea) (the tegumen in this sketch is detached from the vinculum); coremata on pregenital segment (8a) (T. aurosa, Japan; after Diakonoff 1986). Note: the coremata typically are attached to the pregenital membrane structure of the abdomen in Thaumatographa (Fig. 8a), and rest within dorsal slits of each valva when not extruded, but in T. zapyra are attached to the costal margin of the valva itself. The aedeagus in Thaumatographa typically has a long ductus ejaculatorius (2 to 4 times length of aedeagus), with a long proximal hood (not shown in the drawing)." figureDoi="http://doi.org/10.5281/zenodo.14798839" httpUri="https://zenodo.org/record/14798839/files/figure.png" pageId="5" pageNumber="276">Fig. 8</figureCitation>
) are distinctive in the flattened hami and very short socii. The female genitalia (
<figureCitation id="1306C8F1FFC7FF9DFD3D64ED6035FF0D" box="[703,786,186,212]" captionStart="FIGURE 10" captionStartId="8.[151,250,1905,1929]" captionTargetBox="[170,1418,151,1889]" captionTargetId="figure-26@8.[170,1418,151,1889]" captionTargetPageId="8" captionText="FIGURE 10. Thaumatographa mesostigmatias Diakonoff, male genitalia (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus in situ, and with ductus ejaculatorius (a, reversed), enlargement of uncus to juxta (b, with distal end of aedeagus and cornutus hook), and structures of the pregenital abdomen and coremata (c) (gen. slide YA-1159) (MGCL); and d) drawing of male holotype genitalia (Taiwan: [Kuantzuling, Tainan Co.] (gen. slide AD-9811, USNM) (after Diakonoff, 1977b)." figureDoi="http://doi.org/10.5281/zenodo.14798843" httpUri="https://zenodo.org/record/14798843/files/figure.png" pageId="5" pageNumber="276">Fig. 10</figureCitation>
) are notable for the bulbous colliculum-like bulge on the caudal end of the very short and wide ductus bursae; and likewise the signum is very reduced to a single cluster of spines (not a dual cluster of curved spines as in most other species of the genus).
</paragraph>
<caption id="DF4284FCFFC7FF9DFF1560016255FB0E" ID-DOI="http://doi.org/10.5281/zenodo.14798837" ID-Zenodo-Dep="14798837" httpUri="https://zenodo.org/record/14798837/files/figure.png" pageId="5" pageNumber="276" startId="5.[151,250,1110,1134]" targetBox="[410,1178,305,1083]" targetPageId="5" targetType="figure">
<paragraph id="8B82D474FFC7FF9DFF1560016255FB0E" blockId="5.[151,1437,1110,1239]" pageId="5" pageNumber="276">
<emphasis id="B9490866FFC7FF9DFF1560016231FBB7" bold="true" box="[151,278,1110,1134]" pageId="5" pageNumber="276">FIGURE 7.</emphasis>
Wing venation of
<taxonomicName id="4C3DAFF7FFC7FF9DFE5F6001606BFBB7" authority="(Meyrick)" baseAuthorityName="Meyrick" baseAuthorityYear="1886" box="[477,844,1110,1134]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="zapyra">
<emphasis id="B9490866FFC7FF9DFE5F600161FBFBB7" box="[477,732,1110,1134]" italics="true" pageId="5" pageNumber="276">Thaumatographa zapyra</emphasis>
(Meyrick)
</taxonomicName>
(New Guinea), type-species of the genus: redrawn from original sketch of
<bibRefCitation id="EFACA985FFC7FF9DFED5602E6102FB48" author="Walsingham, T. de" box="[343,549,1145,1169]" pageId="5" pageNumber="276" pagination="33 - 67" refId="ref12725" refString="Walsingham, T. de G. (1897) Western Equatorial African Micro-Lepidoptera. Transactions of the Entomological Society of London, 1897, 33-67, pls. 2 - 3. https://doi.org/10.1111/j.1365-2311.1897.tb01674.x" type="journal article" year="1897">Walsingham (1897)</bibRefCitation>
. Note: the wing venation could not be compared to an actual specimen of
<taxonomicName id="4C3DAFF7FFC7FF9DFABE602D66BBFB48" baseAuthorityName="Meyrick" baseAuthorityYear="1886" box="[1340,1436,1146,1169]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="zapyra">
<emphasis id="B9490866FFC7FF9DFABE602D66BBFB48" box="[1340,1436,1146,1169]" italics="true" pageId="5" pageNumber="276">T. zapyra</emphasis>
</taxonomicName>
for accuracy, so the venation may not be totally accurate in all details since the original sketch by Walsingham is small and somewhat simplified.
</paragraph>
</caption>
<caption id="DF4284FCFFC7FF9DFF15635D616BF82C" ID-DOI="http://doi.org/10.5281/zenodo.14798839" ID-Zenodo-Dep="14798839" httpUri="https://zenodo.org/record/14798839/files/figure.png" pageId="5" pageNumber="276" startId="5.[151,250,1802,1826]" targetBox="[156,1435,1263,1779]" targetPageId="5" targetType="figure">
<paragraph id="8B82D474FFC7FF9DFF15635D6088F8B0" blockId="5.[151,1437,1802,2038]" pageId="5" pageNumber="276">
<emphasis id="B9490866FFC7FF9DFF15635D6234F8FB" bold="true" box="[151,275,1802,1826]" pageId="5" pageNumber="276">FIGURE 8.</emphasis>
Male genital structures in
<taxonomicName id="4C3DAFF7FFC7FF9DFDA3635D61F6F8FB" authorityName="Walsignham" authorityYear="1897" box="[545,721,1802,1826]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC7FF9DFDA3635D61F6F8FB" box="[545,721,1802,1826]" italics="true" pageId="5" pageNumber="276">Thaumatographa</emphasis>
</taxonomicName>
, showing main characters for the complex male genitalia of this genus (modified after Diakonoff 1977, for
<taxonomicName id="4C3DAFF7FFC7FF9DFD986379615CF89C" baseAuthorityName="Meyrick" baseAuthorityYear="1886" box="[538,635,1838,1861]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="zapyra">
<emphasis id="B9490866FFC7FF9DFD986379615CF89C" box="[538,635,1838,1861]" italics="true" pageId="5" pageNumber="276">T. zapyra</emphasis>
</taxonomicName>
, New Guinea) (the tegumen in this sketch is detached from the vinculum); coremata on pregenital segment (8a) (
<taxonomicName id="4C3DAFF7FFC7FF9DFD9A63066151F8B1" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[536,630,1873,1896]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFC7FF9DFD9A63066151F8B1" box="[536,630,1873,1896]" italics="true" pageId="5" pageNumber="276">T. aurosa</emphasis>
</taxonomicName>
, Japan; after
<bibRefCitation id="EFACA985FFC7FF9DFD7D63066083F8B0" author="Diakonoff, A." box="[767,932,1873,1897]" pageId="5" pageNumber="276" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff 1986</bibRefCitation>
).
</paragraph>
<paragraph id="8B82D474FFC7FF9DFF156323616BF82C" blockId="5.[151,1437,1802,2038]" pageId="5" pageNumber="276">
Note: the coremata typically are attached to the pregenital membrane structure of the abdomen in
<taxonomicName id="4C3DAFF7FFC7FF9DFB0463236611F855" authorityName="Walsignham" authorityYear="1897" box="[1158,1334,1908,1932]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC7FF9DFB0463236611F855" box="[1158,1334,1908,1932]" italics="true" pageId="5" pageNumber="276">Thaumatographa</emphasis>
</taxonomicName>
(Fig. 8a), and rest within dorsal slits of each valva when not extruded, but in
<taxonomicName id="4C3DAFF7FFC7FF9DFCDE63CF609BF876" baseAuthorityName="Meyrick" baseAuthorityYear="1886" box="[860,956,1944,1967]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="species" species="zapyra">
<emphasis id="B9490866FFC7FF9DFCDE63CF609BF876" box="[860,956,1944,1967]" italics="true" pageId="5" pageNumber="276">T. zapyra</emphasis>
</taxonomicName>
are attached to the costal margin of the valva itself. The aedeagus in
<taxonomicName id="4C3DAFF7FFC7FF9DFE0363ED6116F80B" authorityName="Walsignham" authorityYear="1897" box="[385,561,1978,2002]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC7FF9DFE0363ED6116F80B" box="[385,561,1978,2002]" italics="true" pageId="5" pageNumber="276">Thaumatographa</emphasis>
</taxonomicName>
typically has a long ductus ejaculatorius (2 to 4 times length of aedeagus), with a long proximal hood (not shown in the drawing).
</paragraph>
</caption>
<caption id="DF4284FCFFC4FF9EFF15633E60F9F813" ID-DOI="http://doi.org/10.5281/zenodo.14798841" ID-Zenodo-Dep="14798841" httpUri="https://zenodo.org/record/14798841/files/figure.png" pageId="6" pageNumber="277" startId="6.[151,250,1897,1921]" targetBox="[497,1080,181,1861]" targetPageId="6" targetType="figure">
<paragraph id="8B82D474FFC4FF9EFF15633E60F9F813" blockId="6.[151,1437,1897,1994]" pageId="6" pageNumber="277">
<emphasis id="B9490866FFC4FF9EFF15633E6233F858" bold="true" box="[151,276,1897,1921]" pageId="6" pageNumber="277">FIGURE 9.</emphasis>
Female genital structures in
<taxonomicName id="4C3DAFF7FFC4FF9EFDC2633D61D7F85B" authorityName="Walsignham" authorityYear="1897" box="[576,752,1898,1922]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFC4FF9EFDC2633D61D7F85B" box="[576,752,1898,1922]" italics="true" pageId="6" pageNumber="277">Thaumatographa</emphasis>
</taxonomicName>
, showing main characters for the complex female genitalia of this genus (after
<bibRefCitation id="EFACA985FFC4FF9EFE9963D962E7F87F" author="Diakonoff, A." box="[283,448,1934,1958]" pageId="6" pageNumber="277" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff 1986</bibRefCitation>
, for
<taxonomicName id="4C3DAFF7FFC4FF9EFE7163D96174F87C" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[499,595,1934,1957]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="277" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFC4FF9EFE7163D96174F87C" box="[499,595,1934,1957]" italics="true" pageId="6" pageNumber="277">T. aurosa</emphasis>
</taxonomicName>
, Japan). Note: female genitalia in this genus typically also have a dorsal flap of unknown purpose on the tergal membrane of segment 8 (not shown in this sketch).
</paragraph>
</caption>
<paragraph id="8B82D474FFC5FF9FFF4564C167B4FF68" blockId="7.[151,1437,150,1976]" box="[199,1171,150,177]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4564C162B4FF69" bold="true" box="[199,403,150,176]" pageId="7" pageNumber="278">Descriptive notes</emphasis>
. Wing expanse: male 13.0mm (n = 2), female 14.0–14.8mm (n = 5).
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF4564ED6036FEBD" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4564ED6223FF0D" box="[199,260,186,212]" italics="true" pageId="7" pageNumber="278">Head</emphasis>
(
<figureCitation id="1306C8F1FFC5FF9FFE9064ED62B2FF0D" box="[274,405,186,212]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="7" pageNumber="278">Figs. 1a, 2a</figureCitation>
): dark gray-brown, with median scales yellow on vertex and on frons; lateral tuft of yellow to orange scales laterally on neck; eyes large, dark brown to black; ocellus large and clear to white or amber; antennal scape pale orange to brown; antenna brown-purplish, mixed with scattered orange-yellow scales dorsally; labial palpus brown basally to pale yellow on dorsum, with remainder white with bluish iridesence (especially iridescent on apical segment), and venter white; head venter white.
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF45653967F1FE2D" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF456539623FFE51" box="[199,280,366,392]" italics="true" pageId="7" pageNumber="278">Thorax</emphasis>
(
<figureCitation id="1306C8F1FFC5FF9FFEAA653962B1FE51" box="[296,406,366,392]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="7" pageNumber="278">Figs. 1–2</figureCitation>
): dark brown, with orange-yellow pair of lines from patagia to caudal end, and metathorax also with median patch of orange-yellow; tegula short, dark brown, with orange-yellow medially; venter of thorax white; legs cream white, mesally white, with black-brown dorsal spots basad and apically on midtibia, with hindleg mostly black-brown except white ventrally; spurs white; tarsi alternating white and black-brown.
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF4565A86289FC59" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4565A86214FDC1" box="[199,307,511,536]" italics="true" pageId="7" pageNumber="278">Forewing</emphasis>
(
<figureCitation id="1306C8F1FFC5FF9FFEC065A96288FDC1" box="[322,431,510,536]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="7" pageNumber="278">Figs. 1–2</figureCitation>
): dark brown, with striae of pale yellow from dorsal margin curved to middle of forewing and directed toward apex, all reduced in length from longest on middle of wing to shortest before tornus; wingbase with a recurved pale yellow line from costal margin, a second more straight orange line from costa to near middle; costa with several angled striae alternating orange and silver-white (becoming more silver to middle of wing) and with dark brown between the striae, distally aimed to forewing black-brown spot at end of cell, and nearly convergent with curved striae from dorsal margin; apex with a white margin bordered with a thin black line before apical 1/5 of orange, divided by a silver angled subapical line from costa to mid-termen; tornal quarter with 5 black bar-spots surrounded by merged orange scales, and a silver-brown subtornal line; fringe brown, with orange along termen, divided by a white bar on termen just below apical white border; venter dark brown with dorsal striae diffusely repeated but dull, with cell area suffused with pale tan markings, and apical fringe white mark repeated, and dorsal margin white.
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF4567DD6190FC35" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4567DD6212FC7D" box="[199,309,906,932]" italics="true" pageId="7" pageNumber="278">Hindwing</emphasis>
(
<figureCitation id="1306C8F1FFC5FF9FFEC467DD6294FC7D" box="[326,435,906,932]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." figureDoi="http://doi.org/10.5281/zenodo.14798835" httpUri="https://zenodo.org/record/14798835/files/figure.png" pageId="7" pageNumber="278">Figs. 1–2</figureCitation>
): dark brown on distal half, with diffusely edged pale orange median patch to wingbase, and similar patch of tawny orange on anal sector; costa white from near apex to wingbase; fringe white with brown baseline; venter like dorsum nut marks suffused.
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF4567A16128FB81" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4567A16216FBC9" box="[199,305,1014,1040]" italics="true" pageId="7" pageNumber="278">Abdomen</emphasis>
: dark brown, with dark tan posterior margins on tergites; venter tan-white; genital tufts dark brown with white distally and on venter; pregenital segment with medially invaginated sternite and tergite, with long lateral coremata from pouch formations.
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF4560356231FA44" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4560356249FBA5" box="[199,366,1122,1148]" italics="true" pageId="7" pageNumber="278">Male genitalia</emphasis>
(
<figureCitation id="1306C8F1FFC5FF9FFEFD603562F1FBA5" box="[383,470,1122,1148]" captionStart="FIGURE 10" captionStartId="8.[151,250,1905,1929]" captionTargetBox="[170,1418,151,1889]" captionTargetId="figure-26@8.[170,1418,151,1889]" captionTargetPageId="8" captionText="FIGURE 10. Thaumatographa mesostigmatias Diakonoff, male genitalia (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus in situ, and with ductus ejaculatorius (a, reversed), enlargement of uncus to juxta (b, with distal end of aedeagus and cornutus hook), and structures of the pregenital abdomen and coremata (c) (gen. slide YA-1159) (MGCL); and d) drawing of male holotype genitalia (Taiwan: [Kuantzuling, Tainan Co.] (gen. slide AD-9811, USNM) (after Diakonoff, 1977b)." figureDoi="http://doi.org/10.5281/zenodo.14798843" httpUri="https://zenodo.org/record/14798843/files/figure.png" pageId="7" pageNumber="278">Fig. 10</figureCitation>
): tegumen-vinculum both subtriangular with thin margins; uncus very stout, slightly bent from base on tegumen, as long as hami; hamus long and flattened (subequal to uncus length); socius short and stubby elongate-triangular, with few setae midway; gnathos quadratic and membranous, with a thin distal margin; transtilla incomplete, as short triangular valval stubs; juxta a small subtriangular plate; anellus undeveloped (membranous); aedeagus large and thick, tube-like, narrowing to distal end; vesica with a strongly bent apical hook from large tubular and long cornutus; phallobase undeveloped; ductus ejaculatorius relatively short (subequal to aedeagus length), with short bulbous proximal hood; valva short and oblong, with rounded termen, setose over most of surface and with dense setae mid-valval tuft near base; saccus undeveloped, merged into vinculum form with acute apex.
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF4561F1673FF939" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<emphasis id="B9490866FFC5FF9FFF4561F162A1FA19" box="[199,390,1446,1472]" italics="true" pageId="7" pageNumber="278">Female genitalia</emphasis>
(
<figureCitation id="1306C8F1FFC5FF9FFE1761F162CFFA19" box="[405,488,1446,1472]" captionStart="FIGURE 13" captionStartId="11.[151,250,1455,1479]" captionTargetBox="[160,1436,237,1432]" captionTargetId="figure-57@11.[151,1436,237,1432]" captionTargetPageId="11" captionText="FIGURE 13. Thaumatographa mesostigmatias Diakonoff, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798849" httpUri="https://zenodo.org/record/14798849/files/figure.png" pageId="7" pageNumber="278">Fig. 13</figureCitation>
): ovipositor very short (less than length of 7
<superScript id="7C48793CFFC5FF9FFC5561F160C3FA6D" attach="left" box="[983,996,1446,1460]" fontSize="6" pageId="7" pageNumber="278">th</superScript>
abdominal segment), with an elongate spiculate patch midway on venter; papilla anales large; ostium (
<figureCitation id="1306C8F1FFC5FF9FFCFE619D60F9FA3D" box="[892,990,1482,1508]" captionStart="FIGURE 13" captionStartId="11.[151,250,1455,1479]" captionTargetBox="[160,1436,237,1432]" captionTargetId="figure-57@11.[151,1436,237,1432]" captionTargetPageId="11" captionText="FIGURE 13. Thaumatographa mesostigmatias Diakonoff, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798849" httpUri="https://zenodo.org/record/14798849/files/figure.png" pageId="7" pageNumber="278">Fig. 13a</figureCitation>
), a large shallow cup; sterigma with a central convex and setose bulge, ventrally with a curved margin; ductus bursae very short (less than length of ovipositor), with a large colliculum-like bulge proximal to antrum, the ductus bulge (
<figureCitation id="1306C8F1FFC5FF9FFBDE6245679DF9F5" box="[1116,1210,1554,1580]" captionStart="FIGURE 13" captionStartId="11.[151,250,1455,1479]" captionTargetBox="[160,1436,237,1432]" captionTargetId="figure-57@11.[151,1436,237,1432]" captionTargetPageId="11" captionText="FIGURE 13. Thaumatographa mesostigmatias Diakonoff, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798849" httpUri="https://zenodo.org/record/14798849/files/figure.png" pageId="7" pageNumber="278">Fig. 13c</figureCitation>
) having a dual duct interior of curved ducts (as if there were two antrums) and merging as one into the ductus bursae; the corpus bursae merges immediately proximal to a short ductus bursae section; corpus bursae very elongated, with ovate proximal end; ductus seminalis from near ductus bursae bulge; accessory diverticula large and oblong-ovate, emergent from corpus bursae near middle with short, thin ductus; signum (
<figureCitation id="1306C8F1FFC5FF9FFCBD62F56086F965" box="[831,929,1698,1724]" captionStart="FIGURE 13" captionStartId="11.[151,250,1455,1479]" captionTargetBox="[160,1436,237,1432]" captionTargetId="figure-57@11.[151,1436,237,1432]" captionTargetPageId="11" captionText="FIGURE 13. Thaumatographa mesostigmatias Diakonoff, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.14798849" httpUri="https://zenodo.org/record/14798849/files/figure.png" pageId="7" pageNumber="278">Fig. 13b</figureCitation>
), small with a single bunch of asymmetrical straight spines arrayed from short to long and directed to proximal end of bursa.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC5FF91FF4562BD67F0F98E" lastPageId="9" lastPageNumber="280" pageId="7" pageNumber="278" type="materials_examined">
<paragraph id="8B82D474FFC5FF9FFF4562BD629DF8F1" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<materialsCitation id="3B55DE29FFC5FF9FFF4562BD629DF8F1" collectingDate="1933-05-15" collectionCode="USNM" collectorName="S. Issiki" country="Taiwan" location="Lantashan" pageId="7" pageNumber="278" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="B9490866FFC5FF9FFF4562BD625FF8DD" bold="true" box="[199,376,1770,1796]" pageId="7" pageNumber="278">
<typeStatus id="54866AD6FFC5FF9FFF4562BD6213F8DD" box="[199,308,1770,1796]" pageId="7" pageNumber="278" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<taxonomicName id="4C3DAFF7FFC5FF9FFE0862BC6110F8DD" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[394,567,1771,1796]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="278" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFC5FF9FFE0862BC6110F8DD" box="[394,567,1771,1796]" italics="true" pageId="7" pageNumber="278">mesostigmatias</emphasis>
</taxonomicName>
).
<collectingCountry id="F32A94E4FFC5FF9FFDD262BD6181F8DD" box="[592,678,1770,1796]" name="Taiwan" pageId="7" pageNumber="278">Taiwan</collectingCountry>
: “Rantaisan” [
<location id="8EE282AFFFC5FF9FFCC962BD60E0F8DD" LSID="urn:lsid:plazi:treatment:03946562FFC6FF92FF1565396160FE50:8EE282AFFFC5FF9FFCC962BD60E0F8DD" box="[843,967,1770,1796]" country="Taiwan" name="Lantashan" pageId="7" pageNumber="278" stateProvince="Nantou">Lantashan</location>
], [
<collectingRegion id="49F91A96FFC5FF9FFC6362BC6710F8DC" box="[993,1079,1771,1797]" country="Taiwan" name="Nantou" pageId="7" pageNumber="278">Nantou</collectingRegion>
Co.],
<date id="FF83F2B4FFC5FF9FFB0062BD6638F8DD" box="[1154,1311,1770,1797]" pageId="7" pageNumber="278" value="1933-05-15">
<collectingDate id="EFC70B5CFFC5FF9FFB0062BD6638F8DD" box="[1154,1311,1770,1797]" pageId="7" pageNumber="278" value="1933-05-15">15 May 1933</collectingDate>
</date>
, [
<collectorName id="26C8B1A2FFC5FF9FFAB762BD66B1F8DD" box="[1333,1430,1770,1796]" pageId="7" pageNumber="278">S. Issiki</collectorName>
] (gen. AD-9811) (
<collectionCode id="ED2C4CB1FFC5FF9FFED46359628AF8F1" box="[342,429,1806,1832]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="278" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF456365611AF8A9" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<materialsCitation id="3B55DE29FFC5FF9FFF456365611AF8A9" collectingDate="1905-08-20" collectionCode="BMNH" collectorName="A. E. Wileman" country="Taiwan" elevation="250" location="Kuantzuling" pageId="7" pageNumber="278" specimenCount="1" stateProvince="Tainan" typeStatus="holotype">
<typeStatus id="54866AD6FFC5FF9FFF4563656208F895" box="[199,303,1842,1868]" pageId="7" pageNumber="278" type="holotype">Holotype</typeStatus>
female (
<taxonomicName id="4C3DAFF7FFC5FF9FFE1F636562FFF895" authorityName="Razowski" authorityYear="2009" box="[413,472,1842,1868]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="278" phylum="Arthropoda" rank="species" species="calyx">
<emphasis id="B9490866FFC5FF9FFE1F636562FFF895" box="[413,472,1842,1868]" italics="true" pageId="7" pageNumber="278">calyx</emphasis>
</taxonomicName>
).
<collectingCountry id="F32A94E4FFC5FF9FFE766365616DF895" box="[500,586,1842,1868]" name="Taiwan" pageId="7" pageNumber="278">Taiwan</collectingCountry>
: “Kanshirei” [
<location id="8EE282AFFFC5FF9FFD73636560A6F895" LSID="urn:lsid:plazi:treatment:03946562FFC6FF92FF1565396160FE50:8EE282AFFFC5FF9FFD73636560A6F895" box="[753,897,1842,1868]" country="Taiwan" name="Kuantzuling" pageId="7" pageNumber="278" stateProvince="Tainan">Kuantzuling</location>
] [
<quantity id="4CC57991FFC5FF9FFC18636560CAF894" box="[922,1005,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="278" unit="m" value="250.0">
<elevation id="00103347FFC5FF9FFC18636560CAF894" box="[922,1005,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="278" unit="m" value="250.0">250 m</elevation>
</quantity>
], [
<collectingRegion id="49F91A96FFC5FF9FFB896365677DF895" box="[1035,1114,1842,1868]" country="Taiwan" name="Tainan" pageId="7" pageNumber="278">Tainan</collectingRegion>
Co.],
<date id="FF83F2B4FFC5FF9FFB2863656668F894" box="[1194,1359,1842,1869]" pageId="7" pageNumber="278" value="1905-08-20">
<collectingDate id="EFC70B5CFFC5FF9FFB2863656668F894" box="[1194,1359,1842,1869]" pageId="7" pageNumber="278" value="1905-08-20">20 Aug. 1905</collectingDate>
</date>
,
<collectorName id="26C8B1A2FFC5FF9FFADF636563DAF8A9" pageId="7" pageNumber="278">A. E. Wileman</collectorName>
(gen. BM-31849) (
<collectionCode id="ED2C4CB1FFC5FF9FFE5563016116F8A9" box="[471,561,1878,1904]" pageId="7" pageNumber="278">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B82D474FFC5FF9FFF45632D625EF861" blockId="7.[151,1437,150,1976]" pageId="7" pageNumber="278">
<materialsCitation id="3B55DE29FFC5FF9FFF45632D6252F861" collectingDate="1934-11-25" collectionCode="RMNH" collectorName="Co. &amp; S. Issiki" country="Taiwan" elevation="350" location="Raisya" pageId="7" pageNumber="278" specimenCount="1" stateProvince="Hualien" typeStatus="paratype">
<emphasis id="B9490866FFC5FF9FFF45632D62AEF84D" bold="true" box="[199,393,1914,1940]" pageId="7" pageNumber="278">
<typeStatus id="54866AD6FFC5FF9FFF45632D6213F84D" box="[199,308,1914,1940]" pageId="7" pageNumber="278" type="paratype">Paratype</typeStatus>
female
</emphasis>
(
<taxonomicName id="4C3DAFF7FFC5FF9FFE1B632C6161F84D" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[409,582,1915,1940]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="278" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFC5FF9FFE1B632C6161F84D" box="[409,582,1915,1940]" italics="true" pageId="7" pageNumber="278">mesostigmatias</emphasis>
</taxonomicName>
).
<collectingCountry id="F32A94E4FFC5FF9FFDD9632D6196F84D" box="[603,689,1914,1940]" name="Taiwan" pageId="7" pageNumber="278">Taiwan</collectingCountry>
: “
<location id="8EE282AFFFC5FF9FFD47632D6031F84D" LSID="urn:lsid:plazi:treatment:03946562FFC6FF92FF1565396160FE50:8EE282AFFFC5FF9FFD47632D6031F84D" box="[709,790,1914,1940]" country="Taiwan" name="Raisya" pageId="7" pageNumber="278" stateProvince="Hualien">Raisya</location>
” [Juisui Hot Spgs.] [
<quantity id="4CC57991FFC5FF9FFC79632D6760F84C" box="[1019,1095,1914,1941]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="278" unit="m" value="350.0">
<elevation id="00103347FFC5FF9FFC79632D6760F84C" box="[1019,1095,1914,1941]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="278" unit="m" value="350.0">350 m</elevation>
</quantity>
], [
<collectingRegion id="49F91A96FFC5FF9FFBE2632D679BF84D" box="[1120,1212,1914,1940]" country="Taiwan" name="Hualien" pageId="7" pageNumber="278">Hualien</collectingRegion>
<collectorName id="26C8B1A2FFC5FF9FFB40632D67CAF84D" box="[1218,1261,1914,1940]" pageId="7" pageNumber="278">Co.</collectorName>
],
<date id="FF83F2B4FFC5FF9FFA82632D66BFF84D" box="[1280,1432,1914,1941]" pageId="7" pageNumber="278" value="1934-11-25">
<collectingDate id="EFC70B5CFFC5FF9FFA82632D66BFF84D" box="[1280,1432,1914,1941]" pageId="7" pageNumber="278" value="1934-11-25">25 Nov. 1934</collectingDate>
</date>
, [
<collectorName id="26C8B1A2FFC5FF9FFF2263C963D9F861" box="[160,254,1950,1976]" pageId="7" pageNumber="278">S. Issiki</collectorName>
] (
<collectionCode id="ED2C4CB1FFC5FF9FFE9163C96249F861" box="[275,366,1950,1976]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="278" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF4284FCFFCAFF90FF15632667D2F82F" ID-DOI="http://doi.org/10.5281/zenodo.14798843" ID-Zenodo-Dep="14798843" httpUri="https://zenodo.org/record/14798843/files/figure.png" pageId="8" pageNumber="279" startId="8.[151,250,1905,1929]" targetBox="[170,1418,151,1889]" targetPageId="8" targetType="figure">
<paragraph id="8B82D474FFCAFF90FF15632667D2F82F" blockId="8.[151,1436,1905,2038]" pageId="8" pageNumber="279">
<emphasis id="B9490866FFCAFF90FF1563266205F853" bold="true" box="[151,290,1905,1930]" pageId="8" pageNumber="279">FIGURE 10.</emphasis>
<taxonomicName id="4C3DAFF7FFCAFF90FEA8632561D6F853" authority="Diakonoff" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[298,753,1906,1930]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="279" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFCAFF90FEA863256159F850" box="[298,638,1906,1930]" italics="true" pageId="8" pageNumber="279">Thaumatographa mesostigmatias</emphasis>
Diakonoff
</taxonomicName>
, male genitalia (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus
<emphasis id="B9490866FFCAFF90FF1563C163F0F874" box="[151,215,1942,1965]" italics="true" pageId="8" pageNumber="279">in situ</emphasis>
, and with ductus ejaculatorius (a, reversed), enlargement of uncus to juxta (b, with distal end of aedeagus and cornutus hook), and structures of the pregenital abdomen and coremata (c) (gen. slide YA-1159) (MGCL); and d) drawing of male holotype genitalia (Taiwan: [Kuantzuling, Tainan Co.] (gen. slide AD-9811, USNM) (after
<bibRefCitation id="EFACA985FFCAFF90FBB0638967CDF82F" author="Diakonoff, A." box="[1074,1258,2014,2038]" pageId="8" pageNumber="279" pagination="3 - 55" refId="ref10939" refString="Diakonoff, A. (1977 b) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 3-55." type="journal article" year="1977">Diakonoff, 1977b</bibRefCitation>
).
</paragraph>
</caption>
<caption id="DF4284FCFFCBFF91FF156149621DFA7A" ID-DOI="http://doi.org/10.5281/zenodo.14798845" ID-Zenodo-Dep="14798845" httpUri="https://zenodo.org/record/14798845/files/figure.png" pageId="9" pageNumber="280" startId="9.[151,250,1310,1334]" targetBox="[151,1437,181,1287]" targetPageId="9" targetType="figure">
<paragraph id="8B82D474FFCBFF91FF156149621DFA7A" blockId="9.[151,1436,1310,1443]" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FF1561496239FAEF" bold="true" box="[151,286,1310,1334]" pageId="9" pageNumber="280">FIGURE 11.</emphasis>
<taxonomicName id="4C3DAFF7FFCBFF91FEA761486023FAEE" authority="Heppner &amp; Arita" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[293,772,1311,1335]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="fennchihwu" status="sp. nov.">
<emphasis id="B9490866FFCBFF91FEA761486177FAEE" box="[293,592,1311,1335]" italics="true" pageId="9" pageNumber="280">Thaumatographa fennchihwu</emphasis>
Heppner &amp; Arita
</taxonomicName>
,
<emphasis id="B9490866FFCBFF91FC8C61486079FAEE" bold="true" box="[782,862,1311,1335]" pageId="9" pageNumber="280">
<taxonomicNameLabel id="A27AB51DFFCBFF91FC8C61486079FAEE" box="[782,862,1311,1335]" pageId="9" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus enlarged (a) and with ductus ejaculatorius (with typical proximal elongated hood found in many
<taxonomicName id="4C3DAFF7FFCBFF91FB6E611463C3FAA6" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FB6E611466BBFA82" box="[1260,1436,1347,1371]" italics="true" pageId="9" pageNumber="280">Thaumatographa</emphasis>
species
</taxonomicName>
), with enlargement of uncus to juxta (b), and structures of the pregenital abdomen and coremata (c) (gen. slide JBH-4646) (MGCL).
</paragraph>
</caption>
<paragraph id="8B82D474FFCBFF91FF45618667F0F98E" blockId="9.[151,1437,1489,2019]" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FF45618662B0FA32" bold="true" box="[199,407,1489,1515]" pageId="9" pageNumber="280">Added specimens</emphasis>
. (n = 4):
<materialsCitation id="3B55DE29FFCBFF91FD8061866147F9D6" collectingDate="1984-07-18" collectingDateMax="1984-07-19" collectingDateMin="1984-07-18" collectionCode="MGCL" country="Taiwan" elevation="1400" location="Fennchihwu" pageId="9" pageNumber="280" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiayi">
<collectingCountry id="F32A94E4FFCBFF91FD8061866170FA32" box="[514,599,1489,1515]" name="Taiwan" pageId="9" pageNumber="280">Taiwan</collectingCountry>
,
<collectingRegion id="49F91A96FFCBFF91FDE161866189FA32" box="[611,686,1489,1515]" country="Taiwan" name="Chiayi" pageId="9" pageNumber="280">Chiayi</collectingRegion>
Co.:
<location id="8EE282AFFFCBFF91FD6E61866051FA32" LSID="urn:lsid:plazi:treatment:03946562FFC6FF92FF1565396160FE50:8EE282AFFFCBFF91FD6E61866051FA32" box="[748,886,1489,1515]" country="Taiwan" name="Fennchihwu" pageId="9" pageNumber="280" stateProvince="Chiayi">Fennchihwu</location>
(
<quantity id="4CC57991FFCBFF91FC04618660C7FA32" box="[902,992,1489,1515]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="280" unit="m" value="1400.0">
<elevation id="00103347FFCBFF91FC04618660C7FA32" box="[902,992,1489,1515]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="280" unit="m" value="1400.0">1400 m</elevation>
</quantity>
),
<date id="FF83F2B4FFCBFF91FC7661866758FA32" box="[1012,1151,1489,1515]" pageId="9" pageNumber="280" value="1984-07-18">
<collectingDate id="EFC70B5CFFCBFF91FC7661866758FA32" box="[1012,1151,1489,1515]" pageId="9" pageNumber="280" value="1984-07-18">18 Jul. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFCBFF91FB1061866796FA32" box="[1170,1201,1489,1515]" count="1" pageId="9" pageNumber="280" type="female">1♀</specimenCount>
) (gen. YA-1193),
<date id="FF83F2B4FFCBFF91FA02618663D9F9D6" pageId="9" pageNumber="280" value="1984-07-19">
<collectingDate id="EFC70B5CFFCBFF91FA02618663D9F9D6" pageId="9" pageNumber="280" value="1984-07-19">19 Jul. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFCBFF91FE8D61A26215F9D6" box="[271,306,1525,1551]" count="1" pageId="9" pageNumber="280" type="male">1♂</specimenCount>
) (gen. YA-1159) (
<collectionCode id="ED2C4CB1FFCBFF91FD8261A26170F9D6" box="[512,599,1525,1551]" pageId="9" pageNumber="280">MGCL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B55DE29FFCBFF91FDE861A26065F9EA" collectingDate="1984-10-15" collectingDateMax="1984-10-17" collectingDateMin="1984-10-15" collectionCode="MGCL" collectorName="J. B. Heppner &amp; H. - Y. Wang" country="Taiwan" county="Ilan Co." elevation="480" location="For. Rec. Area" pageId="9" pageNumber="280" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F32A94E4FFCBFF91FDE861A26198F9D6" box="[618,703,1525,1551]" name="Taiwan" pageId="9" pageNumber="280">Taiwan</collectingCountry>
,
<collectingCounty id="62E3ACF8FFCBFF91FD4861A2600EF9D6" box="[714,809,1525,1551]" pageId="9" pageNumber="280">Ilan Co.</collectingCounty>
: Chilan Nsy. [
<location id="8EE282AFFFCBFF91FC4461A26749F9D6" LSID="urn:lsid:plazi:treatment:03946562FFC6FF92FF1565396160FE50:8EE282AFFFCBFF91FC4461A26749F9D6" box="[966,1134,1525,1551]" country="Taiwan" county="Ilan Co." name="For. Rec. Area" pageId="9" pageNumber="280">For. Rec. Area</location>
] (
<quantity id="4CC57991FFCBFF91FB0661A267F7F9D6" box="[1156,1232,1525,1551]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="280" unit="m" value="480.0">
<elevation id="00103347FFCBFF91FB0661A267F7F9D6" box="[1156,1232,1525,1551]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="280" unit="m" value="480.0">480 m</elevation>
</quantity>
),
<date id="FF83F2B4FFCBFF91FB6761A266BBF9D6" box="[1253,1436,1525,1552]" pageId="9" pageNumber="280" value="1984-10-15" valueMax="1984-10-17" valueMin="1984-10-15">
<collectingDate id="EFC70B5CFFCBFF91FB6761A266BBF9D6" box="[1253,1436,1525,1552]" pageId="9" pageNumber="280" value="1984-10-15" valueMax="1984-10-17" valueMin="1984-10-15">15-17 Oct. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFCBFF91FF21624E63E5F9EA" box="[163,194,1561,1587]" count="1" pageId="9" pageNumber="280" type="female">1♀</specimenCount>
),
<collectorName id="26C8B1A2FFCBFF91FF59624E6253F9EA" box="[219,372,1561,1587]" pageId="9" pageNumber="280">J. B. Heppner</collectorName>
&amp;
<collectorName id="26C8B1A2FFCBFF91FE1A624E613CF9EA" box="[408,539,1561,1587]" pageId="9" pageNumber="280">H.-Y. Wang</collectorName>
(gen. JBH4645) (
<collectionCode id="ED2C4CB1FFCBFF91FD61624E601DF9EA" box="[739,826,1561,1587]" pageId="9" pageNumber="280">MGCL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B55DE29FFCBFF91FCCE624E67F4F98E" collectingDate="1989-04-29" collectingDateMax="1989-05-03" collectingDateMin="1989-04-29" collectionCode="MGCL" collectorName="J. B. Heppner &amp; H. - Y. Wang" country="Taiwan" elevation="750" location="Liukuei" pageId="9" pageNumber="280" specimenCount="1" specimenCount-female="1" stateProvince="Kaohsiung">
<collectingCountry id="F32A94E4FFCBFF91FCCE624E6086F9EA" box="[844,929,1561,1587]" name="Taiwan" pageId="9" pageNumber="280">Taiwan</collectingCountry>
,
<collectingRegion id="49F91A96FFCBFF91FC2E624E6702F9EA" box="[940,1061,1561,1587]" country="Taiwan" name="Kaohsiung" pageId="9" pageNumber="280">Kaohsiung</collectingRegion>
Co.: [Shanpin] For. Sta. (
<quantity id="4CC57991FFCBFF91FAC6624E66A8F9EA" box="[1348,1423,1561,1588]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="280" unit="m" value="750.0">
<elevation id="00103347FFCBFF91FAC6624E66A8F9EA" box="[1348,1423,1561,1588]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="280" unit="m" value="750.0">750 m</elevation>
</quantity>
), [nr.]
<location id="8EE282AFFFCBFF91FF4C626A620FF98E" LSID="urn:lsid:plazi:treatment:03946562FFC6FF92FF1565396160FE50:8EE282AFFFCBFF91FF4C626A620FF98E" box="[206,296,1597,1623]" country="Taiwan" name="Liukuei" pageId="9" pageNumber="280" stateProvince="Kaohsiung">Liukuei</location>
,
<date id="FF83F2B4FFCBFF91FEB1626A6131F98E" box="[307,534,1597,1623]" pageId="9" pageNumber="280" value="1989-04-29" valueMax="1989-05-03" valueMin="1989-04-29">
<collectingDate id="EFC70B5CFFCBFF91FEB1626A6131F98E" box="[307,534,1597,1623]" pageId="9" pageNumber="280" value="1989-04-29" valueMax="1989-05-03" valueMin="1989-04-29">29 Apr.-3 May 1989</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFCBFF91FDAA626A616FF98E" box="[552,584,1597,1623]" count="1" pageId="9" pageNumber="280" type="female">1♀</specimenCount>
),
<collectorName id="26C8B1A2FFCBFF91FDE2626A61DDF98E" box="[608,762,1597,1623]" pageId="9" pageNumber="280">J. B. Heppner</collectorName>
&amp;
<collectorName id="26C8B1A2FFCBFF91FC9C626A6085F98E" box="[798,930,1597,1623]" pageId="9" pageNumber="280">H.-Y. Wang</collectorName>
(gen. JBH-4641) (
<collectionCode id="ED2C4CB1FFCBFF91FBF1626A67EDF98E" box="[1139,1226,1597,1623]" pageId="9" pageNumber="280">MGCL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCBFF91FF4562366754F9A2" box="[199,1139,1633,1659]" pageId="9" pageNumber="280" type="etymology">
<paragraph id="8B82D474FFCBFF91FF4562366754F9A2" blockId="9.[151,1437,1489,2019]" box="[199,1139,1633,1659]" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FF456236626FF9A2" bold="true" box="[199,328,1633,1659]" pageId="9" pageNumber="280">Etymology</emphasis>
. The species was named for “middle” and “spotted” (
<bibRefCitation id="EFACA985FFCBFF91FC2362366740F9A2" author="Diakonoff, A." box="[929,1127,1633,1659]" pageId="9" pageNumber="280" pagination="3 - 55" refId="ref10939" refString="Diakonoff, A. (1977 b) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 3-55." type="journal article" year="1977">Diakonoff 1977b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCBFF91FF4562D261D3F91A" pageId="9" pageNumber="280" type="biology_ecology">
<paragraph id="8B82D474FFCBFF91FF4562D261D3F91A" blockId="9.[151,1437,1489,2019]" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FF4562D26206F946" bold="true" box="[199,289,1669,1695]" pageId="9" pageNumber="280">Biology</emphasis>
. Unknown. Habitat is montane forest (
<quantity id="4CC57991FFCBFF91FD5962D26048F946" box="[731,879,1669,1696]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="14.0" metricValueMin="2.5" pageId="9" pageNumber="280" unit="m" value="825.0" valueMax="1400.0" valueMin="250.0">250–1400 m</quantity>
). Flight period (n = 7 records): late April to mid-July, mid-August, and mid-October to late November.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCBFF91FF45629A67B1F8D2" pageId="9" pageNumber="280" type="distribution">
<paragraph id="8B82D474FFCBFF91FF45629A67B1F8D2" blockId="9.[151,1437,1489,2019]" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FF45629A627EF93E" bold="true" box="[199,345,1741,1767]" pageId="9" pageNumber="280">Distribution</emphasis>
. Known only from
<collectingCountry id="F32A94E4FFCBFF91FDB7629A61A0F93E" box="[565,647,1741,1767]" name="Taiwan" pageId="9" pageNumber="280">Taiwan</collectingCountry>
mountains. The old Japanese locality names in
<collectingCountry id="F32A94E4FFCBFF91FB1E629A67C9F93E" box="[1180,1262,1741,1767]" name="Taiwan" pageId="9" pageNumber="280">Taiwan</collectingCountry>
are verified for their current Chinese names in the
<collectingCountry id="F32A94E4FFCBFF91FD9F62A66148F8D2" box="[541,623,1777,1803]" name="Taiwan" pageId="9" pageNumber="280">Taiwan</collectingCountry>
gazeteer for
<taxonomicName id="4C3DAFF7FFCBFF91FC8062A660AEF8D2" authorityName="Linnaeus" authorityYear="1758" box="[770,905,1777,1803]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
sites (
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).
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCBFF92FF4563426160FE50" lastPageId="10" lastPageNumber="281" pageId="9" pageNumber="280" type="discussion">
<paragraph id="8B82D474FFCBFF92FF4563426160FE50" blockId="9.[151,1437,1489,2019]" lastBlockId="10.[151,1437,151,393]" lastPageId="10" lastPageNumber="281" pageId="9" pageNumber="280">
<emphasis id="B9490866FFCBFF91FF4563426262F8F6" bold="true" box="[199,325,1813,1839]" pageId="9" pageNumber="280">Discussion</emphasis>
.
<bibRefCitation id="EFACA985FFCBFF91FED76342610EF8F6" author="Diakonoff, A." box="[341,553,1813,1839]" pageId="9" pageNumber="280" pagination="3 - 55" refId="ref10939" refString="Diakonoff, A. (1977 b) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 3-55." type="journal article" year="1977">Diakonoff (1977b)</bibRefCitation>
described this species as the first hilarographine with a
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type-locality, collected by Japanese in the 1930s but not then named. This species is not a typical
<taxonomicName id="4C3DAFF7FFCBFF91FBE7636E6600F88A" authorityName="Walsignham" authorityYear="1897" box="[1125,1319,1849,1875]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFCBFF91FBE7636E6600F88A" box="[1125,1319,1849,1875]" italics="true" pageId="9" pageNumber="280">Thaumatographa</emphasis>
</taxonomicName>
, since the female genitalia are highly anomalous for the genus, with the short ductus bursae and its caudal bulge. The male genitalia are also anomalous in the genus, with the flattened hami and very short socii.
<bibRefCitation id="EFACA985FFCBFF91FB0763D66674F842" author="Razowski, J." box="[1157,1363,1921,1947]" pageId="9" pageNumber="280" pagination="261 - 287" refId="ref12491" refString="Razowski, J. (2009 a) The Old World Hilarographini (Lepidoptera: Tortricidae). SHILAP Revista Lepidopterologica, 37, 261-287." type="journal article" year="2009">Razowski (2009a)</bibRefCitation>
wrote of some differences for his new species,
<taxonomicName id="4C3DAFF7FFCBFF91FDE663F2618CF866" authorityName="Razowski" authorityYear="2009" box="[612,683,1957,1983]" class="Insecta" family="Tortricidae" genus="Hilarographa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="calyx">
“
<emphasis id="B9490866FFCBFF91FDF263F2618CF866" box="[624,683,1957,1983]" italics="true" pageId="9" pageNumber="280">calyx</emphasis>
</taxonomicName>
”, but there are no significant differences to discern between it and
<taxonomicName id="4C3DAFF7FFCBFF91FF15639D6244F83A" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[151,355,1994,2019]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFCBFF91FF15639D6244F83A" box="[151,355,1994,2019]" italics="true" pageId="9" pageNumber="280">T. mesostigmatias</emphasis>
</taxonomicName>
, thus the new synonymy. In
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,
<taxonomicName id="4C3DAFF7FFCBFF91FC9D639D60CCF83A" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[799,1003,1994,2019]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="280" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFCBFF91FC9D639D60CCF83A" box="[799,1003,1994,2019]" italics="true" pageId="9" pageNumber="280">T. mesostigmatias</emphasis>
</taxonomicName>
has been the only previously known
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species of
<taxonomicName id="4C3DAFF7FFC8FF92FEEB64C0606CFF68" authority="(Kawabe and Komai 1992)" baseAuthorityName="Kawabe and Komai" baseAuthorityYear="1992" box="[361,843,151,177]" class="Insecta" family="Tortricidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="281" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">
Hilarographini (
<bibRefCitation id="EFACA985FFC8FF92FD9F64C06064FF68" author="Kawabe, A. &amp; Komai, F." box="[541,835,151,177]" pageId="10" pageNumber="281" pagination="103 - 109" refId="ref11922" refString="Kawabe, A. &amp; Komai, F. (1992) Tortricidae. In: Heppner, J. B. &amp; Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103-109." type="book chapter" year="1992">Kawabe and Komai 1992</bibRefCitation>
)
</taxonomicName>
. The female genitalia are very different from typical
<taxonomicName id="4C3DAFF7FFC8FF92FF1564EC629DFF0C" box="[151,442,187,213]" pageId="10" pageNumber="281">
<emphasis id="B9490866FFC8FF92FF1564EC627EFF0C" box="[151,345,187,213]" italics="true" pageId="10" pageNumber="281">Thaumatographa</emphasis>
species
</taxonomicName>
, with a short ovipositor, a very short ductus bursae and its colliculum-like bulbous expansion, and in the bursa a reduced signum of short spines. The female genitalia vary a bit on the ostial-sterigmal bulge, being somewhat straight to more convex. Among all the
<taxonomicName id="4C3DAFF7FFC8FF92FCD96554675FFEC4" box="[859,1144,259,285]" pageId="10" pageNumber="281">
<emphasis id="B9490866FFC8FF92FCD96554673AFEC4" box="[859,1053,259,285]" italics="true" pageId="10" pageNumber="281">Thaumatographa</emphasis>
species
</taxonomicName>
,
<taxonomicName id="4C3DAFF7FFC8FF92FB016554666CFEC5" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[1155,1355,259,284]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="281" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFC8FF92FB016554666CFEC5" box="[1155,1355,259,284]" italics="true" pageId="10" pageNumber="281">T. mesostigmatias</emphasis>
</taxonomicName>
should have its own monobasic species-group due to anomalies of the male and female genitalia; and no related species are known of yet that would be included in this species-group. Some darker forms have wing maculation where the typical dark brown is more prominent.
</paragraph>
</subSubSection>
</treatment>
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134
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<subSubSection id="C31840F0FF9DA9636505FC78FE5CFC56" box="[114,462,963,983]" pageId="4" pageNumber="15" type="nomenclature">
<paragraph id="8BBD137BFF9DA9636505FC78FE5CFC56" blockId="4.[114,462,963,983]" box="[114,462,963,983]" pageId="4" pageNumber="15">
<heading id="D0F5A417FF9DA9636505FC78FE5CFC56" box="[114,462,963,983]" fontSize="36" level="2" pageId="4" pageNumber="15" reason="3">
<emphasis id="B976CF69FF9DA9636505FC78FE5CFC56" box="[114,462,963,983]" italics="true" pageId="4" pageNumber="15">
2.3. Characterization of
<taxonomicName id="4C0268F8FF9DA963642EFC78FE3EFC56" authority="EH" authorityName="EH" box="[345,428,963,983]" class="Insecta" family="Cicadidae" genus="Karenia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="brevis">K. brevis</taxonomicName>
<taxonomicName id="4C0268F8FF9DA963642EFC78FE3EFC56" box="[345,428,963,983]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">K. brevis</taxonomicName>
EH
</emphasis>
</heading>
@ -100,13 +100,13 @@ and
/
<emphasis id="B976CF69FF9DA9636760FC47FDA3FB8E" bold="true" box="[535,561,1020,1039]" pageId="4" pageNumber="15">5a</emphasis>
were used for initial characterization of the expressed
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<taxonomicName id="4C0268F8FF9DA96364AFFBACFDBCFBAA" box="[472,558,1047,1067]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9DA96364AFFBACFDBCFBAA" box="[472,558,1047,1067]" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
</taxonomicName>
EH (
<figureCitation id="13390FFEFF9DA963671EFBA3FD3CFBAA" box="[617,686,1048,1067]" captionStart="Fig" captionStartId="4.[113,139,1953,1967]" captionTargetBox="[204,1382,1176,1924]" captionTargetPageId="4" captionText="Fig. 3. (A) Alignment of putative K.brevis EH1 with Aspergillus niger EH.Catalytic residues are indicated by.. (B) SDS PAGE of expressed protein after purification and dialysis. Lanes M: Precision Plus Protein Unstained Standards (Biorad 161-0363) lane 1: lysate; lanes 2–3: column flow-through; lanes 4–5: column washes; lane 6: elution.C.EH (5b) and esterase (3b) assay of purified K. brevis EH." figureDoi="http://doi.org/10.5281/zenodo.10485363" httpUri="https://zenodo.org/record/10485363/files/figure.png" pageId="4" pageNumber="15">Fig. 3C</figureCitation>
). The EH was optimally active at pH 6.8–7.1, with half-maximal activity at pH 6.5 and 8.3. Consistent with the selectivity observed in the
<taxonomicName id="4C0268F8FF9DA963678AFBF4FCEAFF48" authority="CFE" authorityName="CFE" box="[765,888,181,1123]" class="Insecta" family="Cicadidae" genus="Karenia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="brevis">
<taxonomicName id="4C0268F8FF9DA963678AFBF4FCEAFF48" box="[765,888,181,1123]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9DA963678AFBF4FCEAFF48" box="[765,888,181,1123]" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
</taxonomicName>
CFE, the purified EH showed a preference for epoxide
@ -130,7 +130,7 @@ and
</paragraph>
<paragraph id="8BBD137BFF9DA9636629FEC2FC1EFCE6" blockId="4.[831,1502,181,1122]" pageId="4" pageNumber="15">
Substrate and enantioselectivity of the
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<emphasis id="B976CF69FF9DA9636195FEC3FAA1FE0D" box="[1250,1331,376,396]" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
</taxonomicName>
EH was assessed by kinetic studies. Kinetic parameters of the enzyme using substrates
@ -140,7 +140,7 @@ and
were performed using the sensitive fluorescent assay. Other substrates having no fluorescent reporter group were evaluated by GC–MS analysis of the reaction mixtures using 1- naphthol as an internal standard. The data shown in
<tableCitation id="C68026C0FF9DA963602CFDBEFA37FD99" box="[1371,1445,517,536]" captionStart="Table 1" captionStartId="5.[87,131,183,197]" captionTargetPageId="5" captionText="Table 1 Kinetic parameters of the K. brevis EH with various substrates and inhibitors." httpUri="http://table.plazi.org/id/DF7D43F3FF9CA9626520FF0CFD2FFF5D" pageId="4" pageNumber="15" tableUuid="DF7D43F3FF9CA9626520FF0CFD2FFF5D">Table 1</tableCitation>
indicates that the
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<taxonomicName id="4C0268F8FF9DA96366AEFD9BFBBDFDB5" box="[985,1071,544,564]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9DA96366AEFD9BFBBDFDB5" box="[985,1071,544,564]" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
</taxonomicName>
EH has a strong preference for terminal epoxides over internal epoxides (Entries 3–9 vs. Entries 1–2). Aliphatic substituents are preferred over aromatic substituents by approximately 5:1 (Entries 7–9 vs. Entries 3–6) and the enzyme shows only a slight preference for
@ -148,7 +148,7 @@ EH has a strong preference for terminal epoxides over internal epoxides (Entries
over
<emphasis id="B976CF69FF9DA9636183FD34FB6DFD22" box="[1268,1279,655,675]" italics="true" pageId="4" pageNumber="15">S</emphasis>
enantiomers (Entries 3 and 4). The reaction rates using 1,2-disubstituted epoxides as substrates are too slow to be determined using the GC–MS method. However, disubstituted epoxides proved to be inhibitors of the
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<taxonomicName id="4C0268F8FF9DA96360BDFD58FCEAFC92" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9DA96360BDFD58FCEAFC92" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
</taxonomicName>
EH only when an aromatic substituent is present (Entries 10–12 vs. Entries 13–14) and
@ -186,7 +186,7 @@ cyclization of the terminal ring (
<paragraph id="8BBD137BFF9DA9636506F81AFE66F85C" blockId="4.[113,1502,1953,2013]" pageId="4" pageNumber="15">
<emphasis id="B976CF69FF9DA9636506F81AFF30F82E" bold="true" box="[113,162,1953,1967]" pageId="4" pageNumber="15">Fig. 3.</emphasis>
(A) Alignment of putative
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<taxonomicName id="4C0268F8FF9DA963640EF81AFE4BF82E" box="[377,473,1953,1968]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9DA963640EF81AFE28F831" box="[377,442,1953,1968]" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
EH
</taxonomicName>
@ -202,7 +202,7 @@ residues are indicated by .. (B) SDS PAGE of expressed protein after purificatio
) and esterase (
<emphasis id="B976CF69FF9DA9636594F874FF6AF85C" bold="true" box="[227,248,1999,2013]" pageId="4" pageNumber="15">3b</emphasis>
) assay of purified
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<taxonomicName id="4C0268F8FF9DA96364E7F875FE66F85C" box="[400,500,1998,2013]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="4" pageNumber="15" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9DA96364E7F875FE40F85C" box="[400,466,1998,2013]" italics="true" pageId="4" pageNumber="15">K. brevis</emphasis>
EH.
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@ -214,7 +214,7 @@ EH.
</paragraph>
<paragraph id="8BBD137BFF9CA9626520FF75FD2FFF5D" blockId="5.[87,701,183,220]" box="[87,701,205,220]" pageId="5" pageNumber="16">
Kinetic parameters of the
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<taxonomicName id="4C0268F8FF9CA9626450FF76FE11FF5D" box="[295,387,205,220]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="5" pageNumber="16" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9CA9626450FF76FEFAFF5D" box="[295,360,205,220]" italics="true" pageId="5" pageNumber="16">K. brevis</emphasis>
EH
</taxonomicName>
@ -407,7 +407,7 @@ Regioisomeric cyclizations of
</caption>
<paragraph id="8BBD137BFF9CA9626501FBA7FED8FAAB" blockId="5.[87,757,857,1322]" pageId="5" pageNumber="16">
The ability of the
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<taxonomicName id="4C0268F8FF9CA9626447FBA0FE16FBAE" box="[304,388,1051,1071]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Protista" order="Hemiptera" pageId="5" pageNumber="16" phylum="Myzozoa" rank="species" species="brevis">
<emphasis id="B976CF69FF9CA9626447FBA0FE16FBAE" box="[304,388,1051,1071]" italics="true" pageId="5" pageNumber="16">K. brevis</emphasis>
</taxonomicName>
EH to perform cyclization was also evaluated. The substrate

202
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<treatment id="03B387837752B83AFF7DFC07E20AF846" ID-DOI="http://doi.org/10.5281/zenodo.14812335" ID-Zenodo-Dep="14812335" LSID="urn:lsid:plazi:treatment:03B387837752B83AFF7DFC07E20AF846" httpUri="http://treatment.plazi.org/id/03B387837752B83AFF7DFC07E20AF846" lastPageNumber="89" pageId="2" pageNumber="89">
<subSubSection id="C300651E7752B83AFF7DFC07E3AAFC12" pageId="2" pageNumber="89" type="nomenclature">
<paragraph id="8BA536957752B83AFF7DFC07E366FC36" blockId="2.[151,970,942,1005]" box="[151,774,942,969]" pageId="2" pageNumber="89">
<heading id="D0ED81F97752B83AFF7DFC07E366FC36" bold="true" box="[151,774,942,969]" fontSize="11" level="1" pageId="2" pageNumber="89" reason="1">
<emphasis id="B96EEA877752B83AFF7DFC07E366FC36" bold="true" box="[151,774,942,969]" pageId="2" pageNumber="89">
Genus
<taxonomicName id="4C1A4D167752B83AFF04FC07E2F1FC36" authority="Markhaseva &amp; Renz" authorityName="Markhaseva &amp; Renz" authorityYear="2025" box="[238,657,942,969]" class="Copepoda" family="Diaixidae" genus="Mixtodiaixis" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96EEA877752B83AFF04FC07E1E6FC36" bold="true" box="[238,390,942,969]" italics="true" pageId="2" pageNumber="89">Mixtodiaixis</emphasis>
Markhaseva &amp; Renz
</taxonomicName>
<taxonomicNameLabel id="A25D57FC7752B83AFD72FC06E366FC36" box="[664,774,943,969]" pageId="2" pageNumber="89" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA536957752B83AFF7DFC7BE3AAFC12" blockId="2.[151,970,942,1005]" box="[151,970,978,1005]" pageId="2" pageNumber="89">
<uri id="FF8B3A977752B83AFF7DFC7BE3AAFC12" box="[151,970,978,1005]" pageId="2" pageNumber="89">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C300651E7752B83AFF7DFBB3E47CFA3E" pageId="2" pageNumber="89" type="diagnosis">
<paragraph id="8BA536957752B83AFF7DFBB3E47CFA3E" blockId="2.[151,1437,1050,1977]" pageId="2" pageNumber="89">
<emphasis id="B96EEA877752B83AFF7DFBB3E16AFBCB" bold="true" box="[151,266,1050,1076]" pageId="2" pageNumber="89">Diagnosis</emphasis>
(females and males). The combination of 4 setae with 5 teeth at the mandible gnathobase cutting edge is an apomorphie for the genus. The following combination of characters define
<taxonomicName id="4C1A4D167752B83AFBA8FB96E4AEFBA6" authorityName="Markhaseva &amp; Renz" authorityYear="2025" box="[1090,1230,1087,1113]" class="Copepoda" family="Diaixidae" genus="Mixtodiaixis" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96EEA877752B83AFBA8FB96E4AEFBA6" box="[1090,1230,1087,1113]" italics="true" pageId="2" pageNumber="89">Mixtodiaixis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57FC7752B83AFB32FB96E526FBA6" box="[1240,1350,1087,1113]" pageId="2" pageNumber="89" rank="genus">
<emphasis id="B96EEA877752B83AFB32FB96E55FFBA6" bold="true" box="[1240,1343,1087,1113]" pageId="2" pageNumber="89">gen. nov</emphasis>
.
</taxonomicNameLabel>
among “Bradfordians”: 1) rostrum as a bilobed plate; 2) oral parts nearly identical in females and males; 3) antenna exopod setal formula 0,0-0-0,1,1,1,1,1,3; 4) mandible, basis with 3 setae; exopod 5-segmented with 1, 1, 1, 1, and 2 setae; endopod segment 2 with 9 setae; 5) maxillule praecoxal arthrite with 7 terminal spines, anterior and posterior setae absent, coxal endite with 2 setae; coxal epipodite with 9 setae; proximal basal endite with 4 setae, distal basal endite with 4 setae; endopod with 8 setae; exopod with 7–8 setae; 6) maxilla endites setal formula 4, 3, 3, 3 and 4 setae, proximal basal endite with 2sc+1w setae and distal basal endite with 2sc+1w setae, endopod with 3 worm-like and 6 brush-like setae, and 7) maxilliped syncoxa with praecoxal setal formula: 1w, 2 (1sc+1w), and 3 (2sc+1br) setae and 2 setae on coxal endite, endopod segment 1 with 3 setae. P1–P4 of typical clausocalanoidean setation and segmentation. P
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females uniramous, symmetrical, 3-segmented, lacking terminal and lateral spines. P
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males nearly reaching urosome somite 3, of simple structure, uniramous, 5-segmented.
</paragraph>
</subSubSection>
<subSubSection id="C300651E7752B83AFF2DFA62E54DF9F6" pageId="2" pageNumber="89" type="etymology">
<paragraph id="8BA536957752B83AFF2DFA62E54DF9F6" blockId="2.[151,1437,1050,1977]" pageId="2" pageNumber="89">
<emphasis id="B96EEA877752B83AFF2DFA62E12EFA1A" bold="true" box="[199,334,1483,1509]" pageId="2" pageNumber="89">Etymology.</emphasis>
The generic name is combined from the Latin “mixto”, meaning mixture, that indicates that the described genus possesses characters typical for
<taxonomicName id="4C1A4D167752B83AFD5DFA46E344F9F6" authorityName="Sars" authorityYear="1902" box="[695,804,1519,1545]" class="Copepoda" family="Diaixidae" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="family">Diaixidae</taxonomicName>
and selected features of other “Bradfordians”.
</paragraph>
</subSubSection>
<subSubSection id="C300651E7752B83AFF2DF9BAE3DCF9D2" box="[199,956,1554,1581]" pageId="2" pageNumber="89" type="type_taxon">
<paragraph id="8BA536957752B83AFF2DF9BAE3DCF9D2" blockId="2.[151,1437,1050,1977]" box="[199,956,1554,1581]" pageId="2" pageNumber="89">
<emphasis id="B96EEA877752B83AFF2DF9BAE13BF9D3" bold="true" box="[199,347,1554,1581]" pageId="2" pageNumber="89">
<typeStatus id="54A188377752B83AFF2DF9BAE162F9D2" box="[199,258,1555,1581]" pageId="2" pageNumber="89">Type</typeStatus>
species
</emphasis>
<taxonomicName id="4C1A4D167752B83AFE88F9BAE2D3F9D2" authorityName="Markhaseva &amp; Renz" authorityYear="2025" box="[354,691,1555,1581]" class="Copepoda" family="Diaixidae" genus="Mixtodiaixis" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="kurambiohadalis" status="sp. nov.">
<emphasis id="B96EEA877752B83AFE88F9BAE2D3F9D2" box="[354,691,1555,1581]" italics="true" pageId="2" pageNumber="89">Mixtodiaixis kurambiohadalis</emphasis>
</taxonomicName>
<emphasis id="B96EEA877752B83AFD50F9BAE374F9D2" bold="true" box="[698,788,1555,1581]" pageId="2" pageNumber="89">
<taxonomicNameLabel id="A25D57FC7752B83AFD50F9BAE374F9D2" box="[698,788,1555,1581]" pageId="2" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C300651E7752B83AFF2DF99EE20AF846" pageId="2" pageNumber="89" type="discussion">
<paragraph id="8BA536957752B83AFF2DF99EE20AF846" blockId="2.[151,1437,1050,1977]" pageId="2" pageNumber="89">
<emphasis id="B96EEA877752B83AFF2DF99EE15BF9AE" bold="true" box="[199,315,1591,1617]" pageId="2" pageNumber="89">Remarks.</emphasis>
The new genus
<taxonomicName id="4C1A4D167752B83AFE17F99EE2E9F9AE" authorityName="Markhaseva &amp; Renz" authorityYear="2025" box="[509,649,1591,1617]" class="Copepoda" family="Diaixidae" genus="Mixtodiaixis" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96EEA877752B83AFE17F99EE2E9F9AE" box="[509,649,1591,1617]" italics="true" pageId="2" pageNumber="89">Mixtodiaixis</emphasis>
</taxonomicName>
<emphasis id="B96EEA877752B83AFD79F99EE29EF9AE" bold="true" box="[659,766,1591,1617]" pageId="2" pageNumber="89">
<taxonomicNameLabel id="A25D57FC7752B83AFD79F99EE29EF9AE" box="[659,766,1591,1617]" pageId="2" pageNumber="89" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
fits the diagnosis of
<taxonomicName id="4C1A4D167752B83AFC1CF99EE5F8F9AE" authority="(Markhaseva et al., 2014)" baseAuthorityName="Markhaseva" baseAuthorityYear="2014" box="[1014,1432,1591,1617]" class="Copepoda" family="Diaixidae" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="family">
Diaixidae (
<bibRefCitation id="EF8B4B647752B83AFB9EF99EE5F0F9AE" author="Markhaseva, E. L. &amp; Laakmann, S. &amp; Renz, J." box="[1140,1424,1591,1617]" pageId="2" pageNumber="89" pagination="63 - 88" refId="ref6720" refString="Markhaseva, E. L., Laakmann, S. &amp; Renz, J. (2014) An interim synopsis of the Bradfordian families with a description of Thoxancalanus spinatus (Copepoda: Calanoida), a new diaixid genus and species from the deep Atlantic Ocean. Marine Biodiversity, 44, 63-88. https://doi.org/10.1007/s12526-013-0185-0" type="journal article" year="2014">
Markhaseva
<emphasis id="B96EEA877752B83AFAE2F99EE55CF9AE" box="[1288,1340,1591,1617]" italics="true" pageId="2" pageNumber="89">et al</emphasis>
., 2014
</bibRefCitation>
)
</taxonomicName>
, however, shares some characters with other Bradfordian genera, e.g., 1) maxillule praecoxal arthrite with 7 terminal spines a new genus shares with
<taxonomicName id="4C1A4D167752B83AFE04F9D6E444F966" authority="Markhaseva, Schulz &amp; Renz, 2010" authorityName="Markhaseva, Schulz &amp; Renz" authorityYear="2010" box="[494,1060,1663,1689]" class="Copepoda" family="Scolecitrichidae" genus="Diaiscolecithrix" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFE04F9D6E2C2F966" box="[494,674,1663,1689]" italics="true" pageId="2" pageNumber="89">Diaiscolecithrix</emphasis>
Markhaseva, Schulz &amp; Renz, 2010
</taxonomicName>
(
<taxonomicName id="4C1A4D167752B83AFBDAF9D6E488F966" authorityName="Giesbrecht" authorityYear="1893" box="[1072,1256,1663,1689]" class="Copepoda" family="Scolecitrichidae" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="family">Scolecitrichidae</taxonomicName>
) and unresolved genus
<taxonomicName id="4C1A4D167752B83AFF09F90AE193F942" authority="Sars, 1907" authorityName="Sars" authorityYear="1907" box="[227,499,1699,1725]" class="Magnoliopsida" family="Hydrangeaceae" genus="Heteromalla" kingdom="Plantae" order="Cornales" pageId="2" pageNumber="89" phylum="Tracheophyta" rank="genus">
<emphasis id="B96EEA877752B83AFF09F90AE110F942" box="[227,368,1699,1725]" italics="true" pageId="2" pageNumber="89">Heteromalla</emphasis>
Sars, 1907
</taxonomicName>
<emphasis id="B96EEA877752B83AFE14F90AE275F943" box="[510,533,1699,1724]" italics="true" pageId="2" pageNumber="89">vs</emphasis>
9 terminal spines present in nearly all other “Bradfordians”; 2) maxilla basal endites with 2sc+1w setae
<taxonomicName id="4C1A4D167752B83AFE25F96EE23BF91E" authorityName="Markhaseva &amp; Renz" authorityYear="2025" box="[463,603,1735,1761]" class="Copepoda" family="Diaixidae" genus="Mixtodiaixis" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96EEA877752B83AFE25F96EE23BF91E" box="[463,603,1735,1761]" italics="true" pageId="2" pageNumber="89">Mixtodiaixis</emphasis>
</taxonomicName>
<emphasis id="B96EEA877752B83AFD8FF96EE2B1F91E" bold="true" box="[613,721,1735,1761]" pageId="2" pageNumber="89">
<taxonomicNameLabel id="A25D57FC7752B83AFD8FF96EE2B1F91E" box="[613,721,1735,1761]" pageId="2" pageNumber="89" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
shares with some species of
<taxonomicName id="4C1A4D167752B83AFBC2F96EE5FCF91E" authority="Giesbrecht, 1892" authorityName="Giesbrecht" authorityYear="1892" box="[1064,1436,1735,1761]" class="Copepoda" family="Phaennidae" genus="Xanthocalanus" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFBC2F96EE4B0F91E" box="[1064,1232,1735,1761]" italics="true" pageId="2" pageNumber="89">Xanthocalanus</emphasis>
Giesbrecht, 1892
</taxonomicName>
and
<taxonomicName id="4C1A4D167752B83AFF23F942E26CF8FA" authority="Farran, 1905" authorityName="Farran" authorityYear="1905" box="[201,524,1770,1797]" class="Copepoda" family="Phaennidae" genus="Brachycalanus" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFF23F942E111F8FA" box="[201,369,1771,1797]" italics="true" pageId="2" pageNumber="89">Brachycalanus</emphasis>
Farran, 1905
</taxonomicName>
(
<taxonomicName id="4C1A4D167752B83AFDF7F942E2C4F8FA" baseAuthorityName="Ferrari &amp; Steinberg" baseAuthorityYear="1993" box="[541,676,1771,1797]" class="Copepoda" family="Phaennidae" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="family">Phaennidae</taxonomicName>
), unresolved genus
<taxonomicName id="4C1A4D167752B83AFC62F942E487F8FA" authority="Vyshkvartzeva, 1989" authorityName="Vyshkvartzeva" authorityYear="1989" box="[904,1255,1771,1797]" class="Copepoda" family="Scolecitrichidae" genus="Puchinia" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFC62F942E38DF8FA" box="[904,1005,1771,1797]" italics="true" pageId="2" pageNumber="89">Puchinia</emphasis>
<bibRefCitation id="EF8B4B647752B83AFC1CF942E487F8FA" author="Vyshkvartzeva, N. V." box="[1014,1255,1771,1797]" pageId="2" pageNumber="89" pagination="29 - 38" refId="ref6959" refString="Vyshkvartzeva, N. V. (1989) Puchinia obtusa gen. et sp. n. (Copepoda, Calanoida) from the ultra-abyssal of the Kuril-Kamchatka Trench and the position of the genus in the family Scolecithricidae. Zoologicheskii Zhurnal, 68, 29-38." type="journal article" year="1989">Vyshkvartzeva, 1989</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C1A4D167752B83AFACCF942E1B9F8D6" authority="Markhaseva &amp; Schulz, 2010" authorityName="Markhaseva &amp; Schulz" authorityYear="2010" class="Copepoda" family="Diaixidae" genus="Ranthaxus" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFACCF942E5FCF8FA" box="[1318,1436,1771,1797]" italics="true" pageId="2" pageNumber="89">Ranthaxus</emphasis>
Markhaseva &amp; Schulz, 2010
</taxonomicName>
(
<taxonomicName id="4C1A4D167752B83AFE02F8A6E238F8D6" box="[488,600,1807,1833]" class="Copepoda" family="Diaixidae" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="family">Diaixidae</taxonomicName>
)
<emphasis id="B96EEA877752B83AFD82F8A6E21FF8D7" box="[616,639,1807,1832]" italics="true" pageId="2" pageNumber="89">vs</emphasis>
other “Bradfordians” lacking worm-like sensory setae at both endites; 3) maxilliped coxal endite with 2 setae a new genus shares with
<taxonomicName id="4C1A4D167752B83AFC6AF89AE14BF88E" authority="Markhaseva, Schulz &amp; Martinez Arbizu, 2008" authorityName="Markhaseva, Schulz &amp; Martinez Arbizu" authorityYear="2008" class="Copepoda" family="Rostrocalanidae" genus="Rostrocalanus" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFC6AF89AE441F8B2" box="[896,1057,1843,1869]" italics="true" pageId="2" pageNumber="89">Rostrocalanus</emphasis>
<bibRefCitation id="EF8B4B647752B83AFBC0F89AE14BF88E" author="Markhaseva, E. L. &amp; Schulz, K. &amp; Martinez Arbizu, P." pageId="2" pageNumber="89" pagination="2417 - 2441" refId="ref6796" refString="Markhaseva, E. L., Schulz, K. &amp; Martinez Arbizu, P. (2008) New family and genus Rostrocalanus gen. nov. (Crustacea: Calanoida: Rostrocalanidae fam. nov.) from deep Atlantic waters. Journal of Natural History, 42, 2417-2441. https://doi.org/10.1080/00222930802254771" type="journal article" year="2008">Markhaseva, Schulz &amp; Martinez Arbizu, 2008</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C1A4D167752B83AFED1F8FEE191F88E" authorityName="Markhaseva, Schulz &amp; Martinez Arbizu" authorityYear="2009" box="[315,497,1879,1905]" class="Copepoda" family="Rostrocalanidae" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="family">Rostrocalanidae</taxonomicName>
)
<emphasis id="B96EEA877752B83AFDEAF8FEE277F88F" box="[512,535,1879,1904]" italics="true" pageId="2" pageNumber="89">vs</emphasis>
3 setae typical of nearly all “Bradfordians”, and 4) maxilliped endopod segment 1 with 3 setae a new genus shares with
<taxonomicName id="4C1A4D167752B83AFD8EF8D2E365F86A" authorityName="Markhaseva, Schulz &amp; Martinez Arbizu" authorityYear="2008" box="[612,773,1915,1941]" class="Copepoda" family="Rostrocalanidae" genus="Rostrocalanus" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFD8EF8D2E365F86A" box="[612,773,1915,1941]" italics="true" pageId="2" pageNumber="89">Rostrocalanus</emphasis>
</taxonomicName>
and unresolved genus
<taxonomicName id="4C1A4D167752B83AFBE4F8D2E505F86A" authority="Andronov, 2007" authorityName="Andronov" authorityYear="2007" box="[1038,1381,1915,1941]" class="Copepoda" family="Scolecitrichidae" genus="Bradfordiella" kingdom="Animalia" order="Calanoida" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEA877752B83AFBE4F8D2E4C5F86A" box="[1038,1189,1915,1941]" italics="true" pageId="2" pageNumber="89">Bradfordiella</emphasis>
Andronov, 2007
</taxonomicName>
<emphasis id="B96EEA877752B83AFA84F8D2E5E5F86B" box="[1390,1413,1915,1940]" italics="true" pageId="2" pageNumber="89">vs</emphasis>
4 setae typical of nearly all “Bradfordians”.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="D5BA319D3F494D2095D147C54BD5FBD7" ID-CLB-Dataset="307990" ID-DOI="10.11646/zootaxa.5584.1.10" ID-GBIF-Dataset="95bf28f2-a096-48bc-ba89-f13d26d93ed1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14812198" ID-ZooBank="14F06A74-266F-4145-AE76-C7C2C3EA0FFC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738776662074" checkinUser="plazi" docAuthor="Li, Shenlei, Ye, Fei, Lin, Xiaojun, Cheng, Fengxian, Wang, Zhaohong &amp; Tan, Jinfang" docDate="2025" docId="03DB87BCFF938C3DFF5FF9A3FAB8B814" docLanguage="en" docName="zootaxa.5584.1.10.pdf" docOrigin="Zootaxa 5584 (1)" docSource="https://doi.org/10.11646/zootaxa.5584.1.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Solothrips brevisetis Li &amp; Ye &amp; Lin &amp; Cheng &amp; Wang &amp; Tan 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="149" masterDocId="FFE2FFC4FF928C3EFFC8FF95FF86BB49" masterDocTitle="Solothrips brevisetis gen. et sp. nov. with a single pair of ocellar setae (Thysanoptera, Thripidae)" masterLastPageNumber="150" masterPageNumber="146" pageNumber="147" updateTime="1738777637348" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7D9613F187D5837C42177ED412215448">Solothrips brevisetis gen. et sp. nov. with a single pair of ocellar setae (Thysanoptera, Thripidae)</mods:title>
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<mods:namePart id="C7FA7DC6918E61EE995CB1AD31C08C5F">Li, Shenlei</mods:namePart>
<mods:affiliation id="1E3C39A01128872EDFBBDC8C7835EAC9">State Key Laboratory of Biocontrol, School of Agriculture and Biotechnology, Sun Yat-sen University, Shenzhen 518107, China &amp; Guangzhou Ruifeng Biotechnology limited company, Guangzhou, 510663.</mods:affiliation>
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<mods:namePart id="C5D9347F1241B82D4E45E9581AC0D73D">Ye, Fei</mods:namePart>
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<mods:namePart id="29BAF479A3FCB723C59EEDC4F2093BC9">Lin, Xiaojun</mods:namePart>
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<mods:affiliation id="BD43DAE32862D8D942D20303CB0A9E98">Guangzhou Ruifeng Biotechnology limited company, Guangzhou, 510663.</mods:affiliation>
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<mods:affiliation id="4C09F5F0F83141AA7554C5EA77A2E4A3">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, Guangzhou 510260, China.</mods:affiliation>
<mods:nameIdentifier id="D5429BFCDE15F55742CAA75AA2EA36A4" type="email">wangzh@giz.gd.cn</mods:nameIdentifier>
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<mods:affiliation id="3E3C00198DDAA4A6ADE7F4291B68A1AA">State Key Laboratory of Biocontrol, School of Agriculture and Biotechnology, Sun Yat-sen University, Shenzhen 518107, China</mods:affiliation>
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<treatment id="03DB87BCFF938C3DFF5FF9A3FAB8B814" ID-DOI="http://doi.org/10.5281/zenodo.14812196" ID-Zenodo-Dep="14812196" LSID="urn:lsid:plazi:treatment:03DB87BCFF938C3DFF5FF9A3FAB8B814" httpUri="http://treatment.plazi.org/id/03DB87BCFF938C3DFF5FF9A3FAB8B814" lastPageId="3" lastPageNumber="149" pageId="1" pageNumber="147" scope_family="Thripidae" scope_order="Thysanoptera">
<subSubSection id="C3686521FF938C3FFF5FF9A3FD0FBD18" box="[151,649,1590,1617]" pageId="1" pageNumber="147" type="nomenclature">
<paragraph id="8BCD36AAFF938C3FFF5FF9A3FD0FBD18" blockId="1.[151,649,1590,1653]" box="[151,649,1590,1617]" pageId="1" pageNumber="147">
<heading id="D08581C6FF938C3FFF5FF9A3FD0FBD18" bold="true" box="[151,649,1590,1617]" fontSize="11" level="1" pageId="1" pageNumber="147" reason="1">
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<taxonomicName id="4C724D29FF938C3FFF5FF9A3FDA3BD19" authority="Wang, 2025" authorityName="Li &amp; Ye &amp; Lin &amp; Cheng &amp; Wang &amp; Tan" authorityYear="2025" box="[151,549,1590,1617]" class="Insecta" family="Thripidae" genus="Solothrips" kingdom="Animalia" order="Thysanoptera" pageId="1" pageNumber="147" phylum="Arthropoda" rank="species" species="brevisetis" status="sp. nov.">
<emphasis id="B906EAB8FF938C3FFF5FF9A3FE0CBD18" bold="true" box="[151,394,1590,1617]" italics="true" pageId="1" pageNumber="147">Solothrips brevisetis</emphasis>
Wang, 2025
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<taxonomicNameLabel id="A23557C3FF938C3FFDE4F9A2FD0FBD18" box="[556,649,1591,1617]" pageId="1" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCD36AAFF938C3FFF5FF9CEFE8DBD3C" blockId="1.[151,649,1590,1653]" box="[151,267,1627,1653]" pageId="1" pageNumber="147">
(
<figureCitation id="13492A2FFF938C3FFF57F9CEFE84BD3C" box="[159,258,1627,1653]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="1" pageNumber="147">Figs 1–7</figureCitation>
)
</paragraph>
<paragraph id="8BCD36AAFF938C3FFF5FF936FAC6BC4C" blockId="1.[151,1437,1699,2013]" pageId="1" pageNumber="147">
Female macroptera. Body bicolored (
<figureCitation id="13492A2FFF938C3FFDF4F936FDF8BDF4" box="[572,638,1699,1725]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="1" pageNumber="147">Fig. 1</figureCitation>
), head and thorax brown, abdominal tergite I brown, II–VIII pale with median brown patch, IX pale brown, X brown.Antennal segment I pale with base brown, II–III pale, IV–V pale with apex brown, VI–VIII brown (
<figureCitation id="13492A2FFF938C3FFE2AF97EFDA3BC4C" box="[482,549,1771,1797]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="1" pageNumber="147">Fig. 3</figureCitation>
); fore wing brown with one pale area near base, clavus brown (
<figureCitation id="13492A2FFF938C3FFB26F97EFAB5BC4C" box="[1262,1331,1771,1797]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="1" pageNumber="147">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BCD36AAFF938C3FFF0FF89AFD57BC94" blockId="1.[151,1437,1699,2013]" pageId="1" pageNumber="147">
Head sculptured with reticles dorsally, posterior fourth with transverse reticles; ocellar setae III situated near the outside of anterior margins of ocellar triangle; 5 pairs of postocular setae arranged close to compound eyes (
<figureCitation id="13492A2FFF938C3FFAA7F8A6FF2EBC38" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="1" pageNumber="147">Fig. 2</figureCitation>
). Antennae 8-segmented, segments III–IV with short forked sense cone; segment III longest and asymmetrical, dorsal side with 8–9 microtrichial rows, ventral with 6–8 rows; segment IV with 6–8 microtrichial rows; segment V as long as or longer than VI, with 6–7 microtrichial rows; segment VI with one long inner sense cone and two short outer sense cones, 4–6 microtrichial rows. (
<figureCitation id="13492A2FFF938C3FFDB7F856FD45BC94" box="[639,707,1987,2013]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="1" pageNumber="147">Fig. 3</figureCitation>
).
</paragraph>
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<paragraph id="8BCD36AAFF908C3CFF5FF94EFD61BC5E" blockId="2.[151,1436,1755,1816]" pageId="2" pageNumber="148">
<emphasis id="B906EAB8FF908C3CFF5FF94EFEBDBDBD" bold="true" box="[151,315,1755,1780]" pageId="2" pageNumber="148">FIGURES 1–7.</emphasis>
<taxonomicName id="4C724D29FF908C3CFE8BF949FD94BDBD" authorityName="Li &amp; Ye &amp; Lin &amp; Cheng &amp; Wang &amp; Tan" authorityYear="2025" box="[323,530,1756,1780]" class="Insecta" family="Thripidae" genus="Solothrips" kingdom="Animalia" order="Thysanoptera" pageId="2" pageNumber="148" phylum="Arthropoda" rank="species" species="brevisetis" status="sp. nov.">
<emphasis id="B906EAB8FF908C3CFE8BF949FD94BDBD" box="[323,530,1756,1780]" italics="true" pageId="2" pageNumber="148">Solothrips brevisetis</emphasis>
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<taxonomicNameLabel id="A23557C3FF908C3CFDD1F949FDEDBDBD" box="[537,619,1756,1780]" pageId="2" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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(1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII.
</paragraph>
</caption>
<paragraph id="8BCD36AAFF908C3DFF0FF8FFFAB1BB9C" blockId="2.[151,1437,1898,2033]" lastBlockId="3.[151,1437,151,861]" lastPageId="3" lastPageNumber="149" pageId="2" pageNumber="148">
Pronotum surface with sculptured reticulations and about 30 discal setae, all setae short and flattened, no long setae (
<figureCitation id="13492A2FFF908C3CFF15F81BFEA7BCE1" box="[221,289,1934,1960]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="2" pageNumber="148">Fig. 2</figureCitation>
). Mesonotum with regular reticles, medial and submedial setae short and flattened, situated on posterior part, CPS present anteromedially. Metanotum reticulate, median pair of setae flattened and behind anterior margin, submedian setae near anterior margin, CPS present. Mesosternum and metasternum setae fine (
<figureCitation id="13492A2FFF908C3CFB2CF843FAAABCB8" box="[1252,1324,2006,2033]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="2" pageNumber="148">Fig. 5</figureCitation>
). Tarsi 2- segmented. Fore wing vein setae short and flattened, costal vein setae clearly shorter than costal cilia, apex with 1 terminal seta; first vein with 6–7 basal setae and 3 distal setae, second vein with 7 setae scattered (
<figureCitation id="13492A2FFF918C3DFB2CFF2EFAAFBB9C" box="[1252,1321,187,213]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="3" pageNumber="149">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BCD36AAFF918C3DFF0FFF4AFA8FBA98" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="149">
Abdominal tergite I sculptured with transverse reticles, tergites II–VIII smooth medially, laterally with transverse reticulations. Tergites discal setae S1 and S2 small, setae S1 arising from anterior half, setae S3 on tergites II–IV stout but small on tergites V–VIII, posteroangular setae arising mesad of posterior angle. Tergites I–VII medially with short posteromarginal craspeda. Tergite VIII with complete posteromarginal comb, CPS very close to posterior margin (
<figureCitation id="13492A2FFF918C3DFF30FEFAFEC6BAC0" box="[248,320,367,393]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="3" pageNumber="149">Fig. 6</figureCitation>
). A patch of microtrichia present on the anterior angle of tergite VII–VIII. Tergite IX MD setae longest; tergite X with median split complete. Pleurites without discal setae. Sternites with weak transverse reticles, no discal setae; sternite VII with 3 pairs of posteromarginal setae in front of posterior margin (
<figureCitation id="13492A2FFF918C3DFB7EFE22FB7CBA98" box="[1206,1274,439,465]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1755,1779]" captionTargetBox="[167,1419,181,1732]" captionTargetId="figure-20@2.[167,1419,181,1732]" captionTargetPageId="2" captionText="FIGURES 1–7. Solothrips brevisetis sp. nov. (1) female; (2) Head and thorax; (3) antennae; (4) fore wing; (5) thorax ventral surface; (6) Abdominal tergites VI–X; (7) sternites V–VII." figureDoi="http://doi.org/10.5281/zenodo.14812200" httpUri="https://zenodo.org/record/14812200/files/figure.png" pageId="3" pageNumber="149">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BCD36AAFF918C3DFF0FFE4EFEFEBABC" blockId="3.[151,1437,151,861]" box="[199,376,475,501]" pageId="3" pageNumber="149">Male unknown.</paragraph>
<paragraph id="8BCD36AAFF918C3DFF0FFE6AFD9FB9CC" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="149">
<emphasis id="B906EAB8FF918C3DFF0FFE6AFEF0B950" bold="true" box="[199,374,511,537]" pageId="3" pageNumber="149">Measurements</emphasis>
(
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female in microns). Total distended body length 960. Head length (width) 100 (130); eye length (width) 55 (40). Pronotum length (width) 110 (180). Length of antenna 190; length (width) of antennal segments I 22 (24), II 32 (21), III 34 (18), IV 32 (20), V 30 (16), VI 28 (15), VII 6 (6) and VIII 8 (5). Fore wing length 550. Ovipositor length 220.
</paragraph>
</subSubSection>
<subSubSection id="C3686521FF918C3DFF0FFD1BFAB8B814" pageId="3" pageNumber="149" type="materials_examined">
<paragraph id="8BCD36AAFF918C3DFF0FFD1BFE89B9B8" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="149">
<emphasis id="B906EAB8FF918C3DFF0FFD1BFE3FB9E1" bold="true" box="[199,441,654,680]" pageId="3" pageNumber="149">Material examined.</emphasis>
<materialsCitation id="3B1A3CF7FF918C3DFE0FFD1AFE89B9B8" collectingDate="2021-08-17" collectionCode="GIZGAS" collectorName="Fei Ye." country="China" county="Guangzhou" elevation="630" latitude="21.025833" location="National Forest Park" longLatPrecision="21" longitude="110.29222" municipality="Mt" pageId="3" pageNumber="149" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<typeStatus id="54C98808FF918C3DFE0FFD1AFDA9B9E0" box="[455,559,655,681]" pageId="3" pageNumber="149" type="holotype">Holotype</typeStatus>
female:
<collectingCountry id="F365763AFF918C3DFD6AFD1AFD78B9E0" box="[674,766,655,681]" name="China" pageId="3" pageNumber="149">CHINA</collectingCountry>
,
<collectingRegion id="49B6F848FF918C3DFCD8FD1AFB81B9E0" box="[784,1031,655,681]" country="China" name="Guangdong" pageId="3" pageNumber="149">Guangdong province</collectingRegion>
,
<collectingCounty id="62AC4E26FF918C3DFBD2FD1AFB18B9E0" box="[1050,1182,655,681]" pageId="3" pageNumber="149">Guangzhou</collectingCounty>
,
<collectingMunicipality id="6BA9ACD0FF918C3DFB79FD1AFB54B9E0" box="[1201,1234,655,681]" pageId="3" pageNumber="149">Mt</collectingMunicipality>
baiyun
<location id="8EAD6071FF918C3DFAF4FD1AFE9FB984" LSID="urn:lsid:plazi:treatment:03DB87BCFF938C3DFF5FF9A3FAB8B814:8EAD6071FF918C3DFAF4FD1AFE9FB984" country="China" county="Guangzhou" latitude="21.025833" longLatPrecision="21" longitude="110.29222" municipality="Mt" name="National Forest Park" pageId="3" pageNumber="149" stateProvince="Guangdong">
<location id="8EAD6071FF918C3DFAF4FD1AFF5BB984" LSID="urn:lsid:plazi:treatment:03DB87BCFF938C3DFF5FF9A3FAB8B814:8EAD6071FF918C3DFAF4FD1AFF5BB984" country="China" county="Guangzhou" latitude="21.025833" longLatPrecision="21" longitude="110.29222" municipality="Mt" name="National Forest" pageId="3" pageNumber="149" stateProvince="Guangdong">National Forest</location>
Park
</location>
(
<geoCoordinate id="EE46506DFF918C3DFEE3FD26FE34B984" box="[299,434,691,717]" degrees="21" direction="north" minutes="01" orientation="latitude" pageId="3" pageNumber="149" precision="15" seconds="33" value="21.025833">21°01'33&quot;N</geoCoordinate>
,
<geoCoordinate id="EE46506DFF918C3DFE08FD26FDD6B984" box="[448,592,691,717]" degrees="110" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="149" precision="15" seconds="32" value="110.29222">110°17'32&quot;E</geoCoordinate>
, alt.
<quantity id="4C8A9B4FFF918C3DFD42FD26FD4BB984" box="[650,717,691,717]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="149" unit="m" value="630.0">
<elevation id="005FD199FF918C3DFD42FD26FD4BB984" box="[650,717,691,717]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="149" unit="m" value="630.0">630m</elevation>
</quantity>
),
<collectingMethod id="52334EBDFF918C3DFD2BFD26FB87B984" box="[739,1025,691,717]" pageId="3" pageNumber="149">collected by malaise trap</collectingMethod>
under tree,
<date id="FFCC106AFF918C3DFB44FD26FA90B984" box="[1164,1302,691,717]" pageId="3" pageNumber="149" value="2021-08-17">
<collectingDate id="EF88E982FF918C3DFB44FD26FA90B984" box="[1164,1302,691,717]" pageId="3" pageNumber="149" value="2021-08-17">17.viii.2021</collectingDate>
</date>
,
<collectorName id="2687537CFF918C3DFAEBFD26FAF3B984" box="[1315,1397,691,717]" pageId="3" pageNumber="149">Fei Ye.</collectorName>
(in
<collectionCode id="ED63AE6FFF918C3DFF5FFD42FE85B9B8" box="[151,259,727,753]" pageId="3" pageNumber="149">GIZGAS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCD36AAFF918C3DFF0FFD6EFC0CB85C" blockId="3.[151,1437,151,861]" box="[199,906,762,789]" pageId="3" pageNumber="149">
<materialsCitation id="3B1A3CF7FF918C3DFF0FFD6EFC00B85C" box="[199,902,762,789]" collectingDate="2021-08-17" collectionCode="GIZGAS" collectorName="Fei Ye." country="China" county="Guangzhou" elevation="630" latitude="21.025833" location="National Forest Park" longLatPrecision="21" longitude="110.29222" municipality="Mt" pageId="3" pageNumber="149" specimenCount="5" specimenCount-female="5" stateProvince="Guangdong" typeStatus="paratype">
<typeStatus id="54C98808FF918C3DFF0FFD6EFEBEB85C" box="[199,312,763,789]" pageId="3" pageNumber="149" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D74FD23FF918C3DFE8BFD6FFE29B85C" box="[323,431,762,789]" count="5" pageId="3" pageNumber="149" type="female">5 females</specimenCount>
with same data as holotype.(in
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.
</paragraph>
<paragraph id="8BCD36AAFF918C3DFF0FFC8AFAB8B814" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="149">
<emphasis id="B906EAB8FF918C3DFF0FFC8AFD92B871" bold="true" box="[199,532,798,825]" pageId="3" pageNumber="149">Molecular characterization.</emphasis>
A partial mtCOI was sequenced and the annotated gene sequence was deposited in the National Centre for Biotechnology Information (
<collectionCode id="ED63AE6FFF918C3DFD36FCD6FCC2B814" box="[766,836,835,861]" pageId="3" pageNumber="149">NCBI</collectionCode>
) database, accession number
<collectionCode id="ED63AE6FFF918C3DFB58FCD6FB40B814" box="[1168,1222,835,861]" country="Germany" httpUri="http://grbio.org/cool/iz3p-2r7s" name="Universitat Bonn" pageId="3" pageNumber="149">SUB</collectionCode>
14680672.
</paragraph>
</subSubSection>
</treatment>
</document>

129
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species.
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<emphasis id="B906EAB8FF938C3FFEF1FB8EFD9BBF7C" box="[313,541,1051,1077]" italics="true" pageId="1" pageNumber="147">Solothrips brevisetis</emphasis>
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<taxonomicNameLabel id="A23557C3FF938C3FFDEDFB8EFDF8BF7C" box="[549,638,1051,1077]" pageId="1" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="B906EAB8FF938C3FFF0FFBABFEC4BF11" bold="true" box="[199,322,1086,1112]" pageId="1" pageNumber="147">Diagnosis.</emphasis>
Small macroptera. Head wider than long, mouth-cone short and pointed with 3-segmented maxillary palpi. Compound eyes with 4 pigmented facets ventrally. Ocellar setae I and II absent, setae III short; 5 pairs of postocular setae present. Antennae 8-segmented, segment I without dorso-apical setae, III and IV with forked sense cone. Pronotum wider than long without long posteromarginal or posteroangular setae. Prosternal ferna undivided, probasisternum without setae. Meso- and metanota sculptured with reticles, campaniform sensillum (CPS) present. Mesosternal furca with spinula, metasternal furca absent. Fore wing first vein setae discontinuous, second vein setae scattered, apex with 1 terminal seta; posteromarginal cilia wavy; clavus with 5 veinal setae and 1 discal seta. Tarsi 2-segmented. Abdominal tergites median pair of setae (S1) small, arising in anterior part of tergites; paired campaniform sensilla situated behind setae S1 and S2, close to posterior margin; tergites without ctenidia; tergite VIII with a patch of microtrichia anterior to the spiracle, complete posteromarginal comb present; tergite IX with two pairs of CPS. Sternite II with 2 pairs of posteromarginal setae, III–VII with 3 pairs of posteromarginal setae.
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<paragraph id="8BCD36AAFF938C3FFF0FFA5EFCDCBEAC" blockId="1.[151,1437,1050,1509]" box="[199,858,1483,1509]" pageId="1" pageNumber="147">
<emphasis id="B906EAB8FF938C3FFF0FFA5EFED4BEAC" bold="true" box="[199,338,1483,1509]" pageId="1" pageNumber="147">Etymology:</emphasis>
In reference to the single pair of ocellar setae.
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112
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(16)
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<taxonomicName id="4C484D4FFF8EFFEEA730F465FB36F8F4" authority="(Miyake and Kominami ex Oda)" baseAuthorityName="Oda" box="[587,1247,1841,1870]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Chromista" order="Hemiptera" pageId="7" pageNumber="53" phylum="Myzozoa" rank="species" species="mikimotoi">
<emphasis id="B93CEADEFF8EFFEEA730F465FCD5F8F4" bold="true" box="[587,828,1842,1870]" italics="true" pageId="7" pageNumber="53">Karenia mikimotoi</emphasis>
(Miyake and Kominami ex Oda)
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During the surveys considered here,
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<taxonomicName id="4C484D4FFF8EFFEEA693F4EAFB22F863" box="[1000,1227,1980,2009]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Chromista" order="Hemiptera" pageId="7" pageNumber="53" phylum="Myzozoa" rank="species" species="mikimotoi">
<emphasis id="B93CEADEFF8EFFEEA693F4EAFB22F863" box="[1000,1227,1980,2009]" italics="true" pageId="7" pageNumber="53">Karenia mikimotoi</emphasis>
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has been recorded from Scapa Flow on a single occasion, from a phytoplankton sample in 2015. However, it has frequently been identified in Scottish waters and was responsible for significant blooms recorded in 1999, 2003 and
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<taxonomicName id="4C484D4FFF81FFE1A4DDF061FDB2FCF0" box="[422,603,821,843]" class="Insecta" family="Cicadidae" genus="Karenia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="mikimotoi">
<taxonomicName id="4C484D4FFF81FFE1A4DDF061FDB2FCF0" box="[422,603,821,843]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Chromista" order="Hemiptera" pageId="8" pageNumber="54" phylum="Myzozoa" rank="species" species="mikimotoi">
<emphasis id="B93CEADEFF81FFE1A4DDF061FDB2FCF0" box="[422,603,821,843]" italics="true" pageId="8" pageNumber="54">Karenia mikimotoi</emphasis>
</taxonomicName>
</td>
@ -790,7 +790,7 @@ that transplantation of Japanese oysters to
from 1966 onwards may have aided its invasion. Transport in ballast water represents another potential vector. However, contrary to these opinions,
<bibRefCitation id="EFD94B3DFF80FFE0A148F02CFB08FC22" author="Gomez F" box="[1075,1249,891,920]" pageId="9" pageNumber="55" pagination="620 - 628" refId="ref16579" refString="Gomez F (2008) Phytoplankton invasions: Comments on the validity of categorizing the non-indigenous dinoflagellates and diatoms in European Seas. Marine Pollution Bulletin 56: 620 - 628, https: // doi. org / 10.1016 / j. marpolbul. 2007.12.014" type="journal article" year="2008">Gómez (2008)</bibRefCitation>
simply regarded
<taxonomicName id="4C484D4FFF80FFE0A76BF0F1FD5FFC78" box="[528,694,933,962]" class="Insecta" family="Cicadidae" genus="Karenia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="mikimotoi">
<taxonomicName id="4C484D4FFF80FFE0A76BF0F1FD5FFC78" box="[528,694,933,962]" class="Insecta" family="Cicadidae" genus="Karenia" higherTaxonomySource="CoL" kingdom="Chromista" order="Hemiptera" pageId="9" pageNumber="55" phylum="Myzozoa" rank="species" species="mikimotoi">
<emphasis id="B93CEADEFF80FFE0A76BF0F1FD5FFC78" box="[528,694,933,962]" italics="true" pageId="9" pageNumber="55">K. mikimotoi</emphasis>
</taxonomicName>
as cosmopolitan and likely transported by water currents. Given this conflict we herein regard it as cryptogenic pending further investigation.

230
data/03/FA/87/03FA87F7FFC0FF0D78EF9A7AFA03FE4B.xml Normal file
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@ -0,0 +1,230 @@
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<paragraph id="8BEC36E1FFC0FF0C78EF9A7AFDEAF912" blockId="14.[151,616,1646,1709]" box="[151,616,1646,1673]" pageId="14" pageNumber="15">
<heading id="D0A4818DFFC0FF0C78EF9A7AFDEAF912" bold="true" box="[151,616,1646,1673]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C534D62FFC0FF0C78EF9A7AFDEAF912" ID-CoL="4Z3VY" authority="(Ringdahl, 1932)" baseAuthorityName="Ringdahl" baseAuthorityYear="1932" box="[151,616,1646,1673]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lapponica">
<emphasis id="B927EAF3FFC0FF0C78EF9A7AFDEAF912" bold="true" box="[151,616,1646,1673]" pageId="14" pageNumber="15">
<emphasis id="B927EAF3FFC0FF0C78EF9A7AFE12F913" bold="true" box="[151,400,1646,1673]" italics="true" pageId="14" pageNumber="15">Spilogona lapponica</emphasis>
(
<bibRefCitation id="EFC24B10FFC0FF0C79E79A7AFDDDF912" author="Ringdahl, O." box="[415,607,1646,1672]" pageId="14" pageNumber="15" pagination="156 - 160" refId="ref19017" refString="Ringdahl, O. (1932) Einige neue Musciden aus Schweden. Entomologisk Tidskrift, 53, 156-160." type="journal article" year="1932">Ringdahl, 1932</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
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<paragraph id="8BEC36E1FFC0FF0C78EF9A87FE9AF937" blockId="14.[151,616,1646,1709]" box="[151,280,1682,1709]" pageId="14" pageNumber="15">
<figureCitation id="13682A64FFC0FF0C78EF9A87FE9AF937" box="[151,280,1682,1709]" captionStart="FIGURE 5" captionStartId="16.[151,250,1785,1809]" captionTargetBox="[151,1429,181,1761]" captionTargetId="figure-41@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURE 5. Spilogona ssp. A. Spilogona fulvibasis, lateral view, holotype (photo: Garry Ouellette). B–G. Spilogona lapponica (photos of paratypes of Spilogona compacta). B. Sternite 5. C. Terminalia, lateral view. D. Male, lateral view. E. Female, lateral view. F. Male abdomen apex, ventral view. G. Cercal plate, dorsal view. H–J. Spilogona padlei. H. Terminalia, lateral view. I. Sternite 5. J. Female abdomen apex, lateral view. K–M. Spilogona scutulata. K. Male, lateral view. L. Female, lateral view. M. Sternite 5." figureDoi="http://doi.org/10.5281/zenodo.14812247" httpUri="https://zenodo.org/record/14812247/files/figure.png" pageId="14" pageNumber="15">Figs 5 B–G</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BEC36E1FFC0FF0C78EF9AC3FA96F894" blockId="14.[151,1436,1751,1898]" pageId="14" pageNumber="15">
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<taxonomicName id="4C534D62FFC0FF0C78EF9AC3FC87F975" ID-CoL="8CKS5" authority="Ringdahl, 1932: 158" authorityName="Ringdahl" authorityPageNumber="158" authorityYear="1932" box="[151,773,1751,1775]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lapponica" subGenus="Spilogonoides">
<emphasis id="B927EAF3FFC0FF0C78EF9AC3FE96F975" box="[151,276,1751,1775]" italics="true" pageId="14" pageNumber="15">Limnophora</emphasis>
(
<emphasis id="B927EAF3FFC0FF0C795C9AC3FE36F975" box="[292,436,1751,1775]" italics="true" pageId="14" pageNumber="15">Spilogonoides</emphasis>
)
<emphasis id="B927EAF3FFC0FF0C79BB9AC3FDAAF975" box="[451,552,1751,1775]" italics="true" pageId="14" pageNumber="15">lapponica</emphasis>
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. Type-locality: “Ich fing die Art in zwei männlichen Stücken am Strande vom Torneträsk (Regio subarkt.) in dem nördlichsten Lappland, Juli” [
<collectingCountry id="F3447671FFC0FF0C7B9B9AE2FBD0F894" box="[995,1106,1782,1806]" name="Sweden" pageId="14" pageNumber="15">SWEDEN</collectingCountry>
, Torne Lappmark].
</treatmentCitationGroup>
</paragraph>
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<treatmentCitation id="0AF210F0FFC0FF0C78EF9B00FDBCF8B7" author="Huckett, H. C." box="[151,574,1812,1837]" page="198" pageId="14" pageNumber="15" year="1965">
<taxonomicName id="4C534D62FFC0FF0C78EF9B00FDBCF8B7" ID-CoL="4Z3TK" authority="Huckett, 1965: 198" authorityName="Huckett" authorityPageNumber="198" authorityYear="1965" box="[151,574,1812,1837]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="B927EAF3FFC0FF0C78EF9B00FEEAF8B6" box="[151,360,1812,1836]" italics="true" pageId="14" pageNumber="15">Spilogona compacta</emphasis>
<materialsCitation id="3B3B3CBCFFC0FF0C790A9B00FDBCF8B7" box="[370,574,1812,1837]" collectingDate="1954-07-30" collectionCode="R" country="Canada" location="Indian House Lake" municipality="Richards" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Quebec">
<bibRefCitation id="EFC24B10FFC0FF0C790A9B00FDBCF8B7" author="Huckett, H. C." box="[370,574,1812,1837]" pageId="14" pageNumber="15" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 198</bibRefCitation>
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, 208 (key), and 214.
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locality: “
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,
<collectingRegion id="4997F803FFC0FF0C7CD99B00FB73F8B6" box="[1185,1265,1812,1836]" country="Canada" name="Quebec" pageId="14" pageNumber="15">Quebec</collectingRegion>
, 7.30 ... 54 [i.e.
<date id="FFED1021FFC0FF0C78BF9B27FEBDF8D1" box="[199,319,1843,1867]" pageId="14" pageNumber="15" value="1954-07-30">
<collectingDate id="EFA9E9C9FFC0FF0C78BF9B27FEBDF8D1" box="[199,319,1843,1867]" pageId="14" pageNumber="15" value="1954-07-30">30.vii.1954</collectingDate>
</date>
], W.R.
<collectingMunicipality id="6B88AC9BFFC0FF0C79F19B27FE67F8D1" box="[393,485,1843,1867]" pageId="14" pageNumber="15">Richards</collectingMunicipality>
” [
<collectingCountry id="F3447671FFC0FF0C79829B27FDEBF8D1" box="[506,617,1843,1867]" name="Canada" pageId="14" pageNumber="15">CANADA</collectingCountry>
,
<collectingRegion id="4997F803FFC0FF0C7A0E9B27FD44F8D1" box="[630,710,1843,1867]" country="Canada" name="Quebec" pageId="14" pageNumber="15">Quebec</collectingRegion>
] [seen].
<emphasis id="B927EAF3FFC0FF0C7B659B27FCFEF8D1" bold="true" box="[797,892,1843,1867]" pageId="14" pageNumber="15">Syn. nov.</emphasis>
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<paragraph id="8BEC36E1FFC0FF0C78EF9B46FBE0F8F0" blockId="14.[151,1436,1751,1898]" box="[151,1122,1874,1898]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB4311CFFFC0FF0C78EF9B46FBE0F8F0" box="[151,1122,1874,1898]" pageId="14" pageNumber="15">
<taxonomicName id="4C534D62FFC0FF0C78EF9B46FDA2F8F0" ID-CoL="4Z3VY" authority="(Ringdahl, 1932)" baseAuthorityName="Ringdahl" baseAuthorityYear="1932" box="[151,544,1874,1898]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lapponica">
<emphasis id="B927EAF3FFC0FF0C78EF9B46FEEBF8F0" box="[151,361,1874,1898]" italics="true" pageId="14" pageNumber="15">Spilogona lapponica</emphasis>
(
<bibRefCitation id="EFC24B10FFC0FF0C79009B46FD9BF8F0" author="Ringdahl, O." box="[376,537,1874,1898]" pageId="14" pageNumber="15" pagination="156 - 160" refId="ref19017" refString="Ringdahl, O. (1932) Einige neue Musciden aus Schweden. Entomologisk Tidskrift, 53, 156-160." type="journal article" year="1932">Ringdahl, 1932</bibRefCitation>
)
</taxonomicName>
; Tridrikh &amp;
<treatmentCitation id="0AF210F0FFC0FF0C7AE69B46FCECF8F0" author="Sorokina, V. S. &amp; Tridrikh, N. N." box="[670,878,1874,1898]" page="827" pageId="14" pageNumber="15" year="2021">
<bibRefCitation id="EFC24B10FFC0FF0C7AE69B46FCECF8F0" author="Sorokina, V. S. &amp; Tridrikh, N. N." box="[670,878,1874,1898]" pageId="14" pageNumber="15" pagination="205 - 234" refId="ref19868" refString="Sorokina, V. S. &amp; Tridrikh, N. N. (2021) An annotated checklist of the Muscidae (Diptera) of Chukotka (Russia), with new records. Annales de la Societe entomologique de France, New Series, 57 (3), 205-234. https://doi.org/10.1080/00379271.2021.1939161" type="journal article" year="2021">Sorokina, 2021: 827</bibRefCitation>
</treatmentCitation>
, 830 (
<collectingRegion id="4997F803FFC0FF0C7BD49B46FBD5F8F0" box="[940,1111,1874,1898]" country="Russia" name="Magadan" pageId="14" pageNumber="15">Magadan region</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC0FF0D78EF9B8CFDC0FE37" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BEC36E1FFC0FF0D78EF9B8CFCA1FF63" blockId="14.[151,1436,1944,2043]" lastBlockId="15.[151,1437,150,465]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B927EAF3FFC0FF0C78EF9B8CFF58F828" bold="true" box="[151,218,1944,1970]" pageId="14" pageNumber="15">Notes</emphasis>
. The
<typeStatus id="54E88843FFC0FF0C795D9B8DFED7F829" box="[293,341,1945,1971]" pageId="14" pageNumber="15">type</typeStatus>
material of
<taxonomicName id="4C534D62FFC0FF0C79959B8CFD54F828" authorityName="Huckett" authorityYear="1965" box="[493,726,1944,1970]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="B927EAF3FFC0FF0C79959B8CFD54F828" box="[493,726,1944,1970]" italics="true" pageId="14" pageNumber="15">Spilogona compacta</emphasis>
</taxonomicName>
(in Canadian National Collection) and European and Asian specimens of
<taxonomicName id="4C534D62FFC0FF0C79489BA8FD9BF84C" baseAuthorityName="Ringdahl" baseAuthorityYear="1932" box="[304,537,1980,2006]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lapponica">
<emphasis id="B927EAF3FFC0FF0C79489BA8FD9BF84C" box="[304,537,1980,2006]" italics="true" pageId="14" pageNumber="15">Spilogona lapponica</emphasis>
</taxonomicName>
were examined. The descriptions of both species and the male terminalia are the same. Males of this species are very distinctive with a very broad frons; a stout thick abdomen with an unusually large sternite 5 (
<figureCitation id="13682A64FFC1FF0D79379C83FE45FF2B" box="[335,455,150,177]" captionStart="FIGURE 5" captionStartId="16.[151,250,1785,1809]" captionTargetBox="[151,1429,181,1761]" captionTargetId="figure-41@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURE 5. Spilogona ssp. A. Spilogona fulvibasis, lateral view, holotype (photo: Garry Ouellette). B–G. Spilogona lapponica (photos of paratypes of Spilogona compacta). B. Sternite 5. C. Terminalia, lateral view. D. Male, lateral view. E. Female, lateral view. F. Male abdomen apex, ventral view. G. Cercal plate, dorsal view. H–J. Spilogona padlei. H. Terminalia, lateral view. I. Sternite 5. J. Female abdomen apex, lateral view. K–M. Spilogona scutulata. K. Male, lateral view. L. Female, lateral view. M. Sternite 5." figureDoi="http://doi.org/10.5281/zenodo.14812247" httpUri="https://zenodo.org/record/14812247/files/figure.png" pageId="15" pageNumber="16">Fig. 5B, F</figureCitation>
); costa with a strong costal spine in both sexes (
<figureCitation id="13682A64FFC1FF0D7B939C83FBE5FF2B" box="[1003,1127,150,177]" captionStart="FIGURE 5" captionStartId="16.[151,250,1785,1809]" captionTargetBox="[151,1429,181,1761]" captionTargetId="figure-41@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURE 5. Spilogona ssp. A. Spilogona fulvibasis, lateral view, holotype (photo: Garry Ouellette). B–G. Spilogona lapponica (photos of paratypes of Spilogona compacta). B. Sternite 5. C. Terminalia, lateral view. D. Male, lateral view. E. Female, lateral view. F. Male abdomen apex, ventral view. G. Cercal plate, dorsal view. H–J. Spilogona padlei. H. Terminalia, lateral view. I. Sternite 5. J. Female abdomen apex, lateral view. K–M. Spilogona scutulata. K. Male, lateral view. L. Female, lateral view. M. Sternite 5." figureDoi="http://doi.org/10.5281/zenodo.14812247" httpUri="https://zenodo.org/record/14812247/files/figure.png" pageId="15" pageNumber="16">Figs 5D, E</figureCitation>
); mid tibia usually with
<emphasis id="B927EAF3FFC1FF0D7DF89C83FA1EFF2B" box="[1408,1436,151,177]" italics="true" pageId="15" pageNumber="16">ad</emphasis>
setae and hind femur usually with several short
<emphasis id="B927EAF3FFC1FF0D7ADE9CAFFD42FF4E" box="[678,704,187,212]" italics="true" pageId="15" pageNumber="16">pv</emphasis>
setae. I conclude that
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<emphasis id="B927EAF3FFC1FF0D7BCE9CAFFB1BFF4E" box="[950,1177,187,213]" italics="true" pageId="15" pageNumber="16">Spilogona compacta</emphasis>
<bibRefCitation id="EFC24B10FFC1FF0D7CE79CAFFABEFF4F" author="Huckett, H. C." box="[1183,1340,186,213]" pageId="15" pageNumber="16" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965</bibRefCitation>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="4C534D62FFC1FF0D79159CCBFC9DFF63" authority="(Ringdahl, 1932)" baseAuthorityName="Ringdahl" baseAuthorityYear="1932" box="[365,799,223,249]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="lapponica">
<emphasis id="B927EAF3FFC1FF0D79159CCBFDD4FF63" box="[365,598,223,249]" italics="true" pageId="15" pageNumber="16">Spilogona lapponica</emphasis>
(
<bibRefCitation id="EFC24B10FFC1FF0D7A1E9CCBFC95FF63" author="Ringdahl, O." box="[614,791,223,249]" pageId="15" pageNumber="16" pagination="156 - 160" refId="ref19017" refString="Ringdahl, O. (1932) Einige neue Musciden aus Schweden. Entomologisk Tidskrift, 53, 156-160." type="journal article" year="1932">Ringdahl, 1932</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
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There is no description of the female of
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<emphasis id="B927EAF3FFC1FF0D7A239D17FCC6FE87" box="[603,836,259,285]" italics="true" pageId="15" pageNumber="16">Spilogona lapponica</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFC24B10FFC1FF0D7B119D17FB9FFE87" author="Hennig, W." box="[873,1053,258,285]" pageId="15" pageNumber="16" pagination="289 - 336" refId="ref18560" refString="Hennig, W. (1959 b) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 205. E. Schweizerbart, Stuttgart, pp. 289-336." type="book chapter" year="1959">Hennig (1959b)</bibRefCitation>
, but there is the description of the female of
<taxonomicName id="4C534D62FFC1FF0D79719D33FE6EFEDA" authorityName="Huckett" authorityYear="1965" box="[265,492,295,321]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="compacta">
<emphasis id="B927EAF3FFC1FF0D79719D33FE6EFEDA" box="[265,492,295,321]" italics="true" pageId="15" pageNumber="16">Spilogona compacta</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFC24B10FFC1FF0D7A689D33FD74FEDB" author="Huckett, H. C." box="[528,758,294,321]" pageId="15" pageNumber="16" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett (1965: 215)</bibRefCitation>
.
</paragraph>
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<emphasis id="B927EAF3FFC1FF0D78BF9D5EFDB5FEFE" bold="true" box="[199,567,330,356]" pageId="15" pageNumber="16">Additional material examined</emphasis>
.
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<collectingCountry id="F3447671FFC1FF0D7A339D5FFD31FEFF" box="[587,691,331,357]" name="Russia" pageId="15" pageNumber="16">RUSSIA</collectingCountry>
,
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<collectingRegion id="4997F803FFC1FF0D7ABF9D5FFCB0FEFF" box="[711,818,331,357]" country="Russia" name="Magadan" pageId="15" pageNumber="16">Magadan</collectingRegion>
area
</emphasis>
:
<specimenCount id="9D55FD68FFC1FF0D7BFC9D5FFC3EFEFF" box="[900,956,331,357]" count="3" pageId="15" pageNumber="16" type="male">3♂♂</specimenCount>
<specimenCount id="9D55FD68FFC1FF0D7BB19D5FFB80FEFF" box="[969,1026,331,357]" count="2" pageId="15" pageNumber="16" type="female">2♀♀</specimenCount>
, ~
<locationDeviation id="9A5E2FC3FFC1FF0D7C5A9D5FFABDFEFF" box="[1058,1343,331,357]" location="Seimchan" pageId="15" pageNumber="16">
<quantity id="4CAB9B04FFC1FF0D7C5A9D5FFBFFFEFF" box="[1058,1149,331,357]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="km" value="100.0">100 km</quantity>
NNE Seimchan
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,
<collectingMunicipality id="6B88AC9BFFC1FF0D7D289D5FFF73FE13" pageId="15" pageNumber="16">Nature Reserve</collectingMunicipality>
“Magadansky”, valley of
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,
<geoCoordinate id="EE675026FFC1FF0D7AA99D7BFCB0FE13" box="[721,818,367,393]" degrees="63" direction="north" minutes="39" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="63.65">63°39’N</geoCoordinate>
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, thickets of young willow, sweeping in grass,
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<collectingDate id="EFA9E9C9FFC1FF0D78EF9D87FE9BFE37" box="[151,281,403,429]" pageId="15" pageNumber="16" value="2019-07-17">17.vii.2019</collectingDate>
</date>
, leg.
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(
<collectionCode id="ED42AE24FFC1FF0D799A9D87FDB4FE37" box="[482,566,403,429]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="15" pageNumber="16">SZMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC1FF0D78BF9DA2FA03FE4B" box="[199,1409,438,465]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BEC36E1FFC1FF0D78BF9DA2FA03FE4B" blockId="15.[151,1437,150,465]" box="[199,1409,438,465]" pageId="15" pageNumber="16">
<emphasis id="B927EAF3FFC1FF0D78BF9DA2FEE3FE4A" bold="true" box="[199,353,438,464]" pageId="15" pageNumber="16">Distribution.</emphasis>
PALAEARCTIC:
<collectingCountry id="F3447671FFC1FF0D7A4D9DA3FD12FE4B" box="[565,656,439,465]" name="Norway" pageId="15" pageNumber="16">Norway</collectingCountry>
,
<collectingCountry id="F3447671FFC1FF0D7AE39DA3FD64FE4B" box="[667,742,439,465]" name="Russia" pageId="15" pageNumber="16">Russia</collectingCountry>
(Europe, Far East),
<collectingCountry id="F3447671FFC1FF0D7BB19DA3FBA7FE4B" box="[969,1061,439,465]" name="Sweden" pageId="15" pageNumber="16">Sweden</collectingCountry>
. NEARCTIC: Alaska, Quebec.
</paragraph>
</subSubSection>
</treatment>
</document>

326
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<taxonomicName id="4C534D62FFC4FF0878EF994AFDE9FAE2" ID-CoL="4Z3TF" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,619,1374,1401]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
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<emphasis id="B927EAF3FFC4FF0878EF994AFE20FAE3" bold="true" box="[151,418,1374,1401]" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
(Schnabl, 1915)
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<taxonomicName id="4C534D62FFC4FF0878EF99B7FDECFA21" authority="Schnabl" authorityName="Schnabl" box="[151,622,1443,1467]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta" subGenus="Spilogona">
<emphasis id="B927EAF3FFC4FF0878EF99B7FE96FA21" box="[151,276,1443,1467]" italics="true" pageId="10" pageNumber="11">Limnophora</emphasis>
(
<emphasis id="B927EAF3FFC4FF08795D99B7FE0EFA21" box="[293,396,1443,1467]" italics="true" pageId="10" pageNumber="11">Spilogona</emphasis>
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<emphasis id="B927EAF3FFC4FF0879E599B7FD91FA21" box="[413,531,1443,1467]" italics="true" pageId="10" pageNumber="11">ciliatocosta</emphasis>
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</taxonomicName>
in
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Becker
<emphasis id="B927EAF3FFC4FF087A9A99B0FC95FA21" box="[738,791,1443,1467]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
, 1915: 31
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. Type-locality: “in der Karskaja Tundra” [
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,
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,
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peninsula,
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;
<date id="FFED1021FFC4FF087AC399D6FC93FA40" box="[699,785,1474,1498]" pageId="10" pageNumber="11" value="1909-07">
<collectingDate id="EFA9E9C9FFC4FF087AC399D6FC93FA40" box="[699,785,1474,1498]" pageId="10" pageNumber="11" value="1909-07">vii.1909</collectingDate>
</date>
; collector
<collectorName id="26A65337FFC4FF087B0399D6FC62FA40" box="[891,992,1474,1498]" pageId="10" pageNumber="11">F. Zaitsev</collectorName>
] [seen].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878EF99F4FCDDF98D" blockId="10.[151,1436,1443,1621]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4311CFFFC4FF0878EF99F4FCDDF98D" pageId="10" pageNumber="11">
<treatmentCitation id="0AF210F0FFC4FF0878EF99F4FD9BFA63" author="Huckett, H. C." box="[151,537,1504,1529]" page="18" pageId="10" pageNumber="11" year="1965">
<taxonomicName id="4C534D62FFC4FF0878EF99F4FD9BFA63" ID-CoL="4Z3TQ" authority="Huckett, 1965: 18" authorityName="Huckett" authorityPageNumber="18" authorityYear="1965" box="[151,537,1504,1529]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF0878EF99F4FEDFFA62" box="[151,349,1504,1528]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
<materialsCitation id="3B3B3CBCFFC4FF08791A99F4FD9BFA63" box="[354,537,1504,1529]" collectingDate="1951-07-14" collectorName="J. G. Chillcott" country="Canada" location="Spence Bay" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Northwest Territories">
<bibRefCitation id="EFC24B10FFC4FF08791A99F4FD9BFA63" author="Huckett, H. C." box="[354,537,1504,1529]" pageId="10" pageNumber="11" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 18</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, 190, 201 (key), and 216. Type-locality: “
<location id="8E8C603AFFC4FF087BC399F5FBB4FA63" LSID="urn:lsid:plazi:treatment:03FA87F7FFC4FF0978EF994AFF76FB5B:8E8C603AFFC4FF087BC399F5FBB4FA63" box="[955,1078,1505,1529]" country="Canada" name="Spence Bay" pageId="10" pageNumber="11" stateProvince="Northwest Territories">Spence Bay</location>
,
<collectingRegion id="4997F803FFC4FF087C4799F4FA98FA62" box="[1087,1306,1504,1528]" country="Canada" name="Northwest Territories" pageId="10" pageNumber="11">Northwest Territories</collectingRegion>
, 7.14.51 [i.e.
<date id="FFED1021FFC4FF0878BF99EBFEBCF98D" box="[199,318,1535,1559]" pageId="10" pageNumber="11" value="1951-07-14">
<collectingDate id="EFA9E9C9FFC4FF0878BF99EBFEBCF98D" box="[199,318,1535,1559]" pageId="10" pageNumber="11" value="1951-07-14">14.vii.1951</collectingDate>
</date>
],
<collectorName id="26A65337FFC4FF08792899EBFE58F98D" box="[336,474,1535,1559]" pageId="10" pageNumber="11">J.G. Chillcott</collectorName>
” [
<collectingCountry id="F3447671FFC4FF08799799EBFDDCF98D" box="[495,606,1535,1559]" name="Canada" pageId="10" pageNumber="11">CANADA</collectingCountry>
, NWT] [seen].
<emphasis id="B927EAF3FFC4FF087A8699EBFCDDF98D" bold="true" box="[766,863,1535,1559]" pageId="10" pageNumber="11">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878EF9A0AFE57F9CE" blockId="10.[151,1436,1443,1621]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4311CFFFC4FF0878EF9A0AFE57F9CE" pageId="10" pageNumber="11">
<taxonomicName id="4C534D62FFC4FF0878EF9A0AFDA4F9AC" ID-CoL="4Z3TF" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,550,1566,1590]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0878EF9A0AFEF8F9AC" box="[151,378,1566,1590]" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
(Schnabl, 1915)
</taxonomicName>
;
<materialsCitation id="3B3B3CBCFFC4FF087A489A0AFE52F9CE" collectorName="Sorokina &amp; Thidrikh" location="Wrangel Island" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Chukotka">
<treatmentCitation id="0AF210F0FFC4FF087A489A0AFC8BF9AC" author="Sorokina, V. S." box="[560,777,1566,1590]" page="331" pageId="10" pageNumber="11" year="2012">
<bibRefCitation id="EFC24B10FFC4FF087A489A0AFC8BF9AC" author="Sorokina, V. S." box="[560,777,1566,1590]" pageId="10" pageNumber="11" pagination="328 - 332" refId="ref19142" refString="Sorokina, V. S. (2012 a) The Muscidae (Diptera) of the Russian tundra zones. Report 1. Caucasian Entomological Bulletin, 8 (2), 328-332. [in Russian, with English abstract]" type="journal article" year="2012">Sorokina, 2012a: 331</bibRefCitation>
</treatmentCitation>
(Taymyr Peninsula);
<collectorName id="26A65337FFC4FF087B9D9A0AFBC2F9AC" box="[997,1088,1566,1590]" pageId="10" pageNumber="11">Sorokina</collectorName>
&amp;
<collectorName id="26A65337FFC4FF087C189A0AFB39F9AC" box="[1120,1211,1566,1590]" pageId="10" pageNumber="11">Thidrikh</collectorName>
, 2021: 219 (
<collectingRegion id="4997F803FFC4FF087D409A0AFA1EF9AC" box="[1336,1436,1566,1590]" country="Russia" name="Chukotka" pageId="10" pageNumber="11">Chukotka</collectingRegion>
AO, incl.
<location id="8E8C603AFFC4FF0879539A28FE4BF9CE" LSID="urn:lsid:plazi:treatment:03FA87F7FFC4FF0978EF994AFF76FB5B:8E8C603AFFC4FF0879539A28FE4BF9CE" box="[299,457,1596,1620]" name="Wrangel Island" pageId="10" pageNumber="11" stateProvince="Chukotka">Wrangel Island</location>
)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC4FF0978EF9A97FEDCFCA3" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BEC36E1FFC4FF0978EF9A97FE80FE87" blockId="10.[151,1437,1667,2017]" lastBlockId="11.[151,1437,151,825]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878EF9A97FF58F907" bold="true" box="[151,218,1667,1693]" pageId="10" pageNumber="11">Notes</emphasis>
. The type material of
<taxonomicName id="4C534D62FFC4FF0879B59A97FD59F907" box="[461,731,1667,1693]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0879B59A97FD59F907" box="[461,731,1667,1693]" italics="true" pageId="10" pageNumber="11">Limnophora ciliatocosta</emphasis>
</taxonomicName>
(in
<collectionCode id="ED42AE24FFC4FF087B7E9A97FCC3F907" box="[774,833,1667,1693]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="11">ZISP</collectionCode>
) and
<taxonomicName id="4C534D62FFC4FF087B059A97FBD0F906" authorityName="Huckett" authorityYear="1965" box="[893,1106,1667,1693]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF087B059A97FBD0F906" box="[893,1106,1667,1693]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
</taxonomicName>
(in
<collectionCode id="ED42AE24FFC4FF087C059A97FB3AF907" box="[1149,1208,1667,1693]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="10" pageNumber="11">CNC</collectionCode>
) was examined. The descriptions of both species and the male terminalia are the same. However, the illustrations of the male terminalia of
<taxonomicName id="4C534D62FFC4FF0878CF9ADFFE2DF97F" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[183,431,1739,1765]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF0878CF9ADFFE2DF97F" box="[183,431,1739,1765]" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC24B10FFC4FF0879A49ADFFD04F97F" author="Hennig, W." box="[476,646,1738,1765]" pageId="10" pageNumber="11" pagination="233 - 288" refId="ref18511" refString="Hennig, W. (1959 a) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 204. E. Schweizerbart, Stuttgart, pp. 233-288." type="book chapter" year="1959">Hennig (1959a</bibRefCitation>
: Taf. XV, Fig. 299) and of
<taxonomicName id="4C534D62FFC4FF087BCF9ADFFB0DF97E" authorityName="Huckett" authorityYear="1965" box="[951,1167,1739,1765]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF087BCF9ADFFB0DF97E" box="[951,1167,1739,1765]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC24B10FFC4FF087CC49ADFFA1AF97F" author="Huckett, H. C." box="[1212,1432,1738,1765]" pageId="10" pageNumber="11" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett (1965: 352</bibRefCitation>
, Fig. 165) are slightly different in the width of the surstylus. I examined many specimens of Palaearctic
<taxonomicName id="4C534D62FFC4FF087D549AFBFE95F8B7" baseAuthorityName="Schnabl" baseAuthorityYear="1915" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF087D549AFBFE95F8B7" italics="true" pageId="10" pageNumber="11">Spilogona ciliatocosta</emphasis>
</taxonomicName>
and Nearctic
<taxonomicName id="4C534D62FFC4FF0879CF9B07FD12F8B6" authorityName="Huckett" authorityYear="1965" box="[439,656,1811,1837]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF0879CF9B07FD12F8B6" box="[439,656,1811,1837]" italics="true" pageId="10" pageNumber="11">Spilogona contigua</emphasis>
</taxonomicName>
and I have concluded that the width of the surstylus depends on the angle of viewing the terminalia and is variable. The cercal plate of these species is very distinctive because of the different length of right and left cerci (
<bibRefCitation id="EFC24B10FFC4FF087A339B4FFCABF8EF" author="Huckett, H. C." box="[587,809,1882,1909]" pageId="10" pageNumber="11" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 346</bibRefCitation>
, Fig. 96). Hennig in his monograph illustrated the male terminalia (
<bibRefCitation id="EFC24B10FFC4FF0879619B6BFE44F803" author="Hennig, W." box="[281,454,1918,1945]" pageId="10" pageNumber="11" pagination="233 - 288" refId="ref18511" refString="Hennig, W. (1959 a) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 204. E. Schweizerbart, Stuttgart, pp. 233-288." type="book chapter" year="1959">Hennig, 1959a</bibRefCitation>
: Taf. XIII, Fig. 276; 1959b: Taf. XVIII, Fig. 361) of a specimen from Wrangel Island named as “
<taxonomicName id="4C534D62FFC4FF08796C9BB7FDD8F827" authority="Frey" authorityName="Frey" authorityYear="1915" box="[276,602,1955,1981]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macropyga">
<emphasis id="B927EAF3FFC4FF08796C9BB7FD9CF826" box="[276,542,1955,1981]" italics="true" pageId="10" pageNumber="11">Limnophora macropyga</emphasis>
Frey
</taxonomicName>
” (det. A. Stackelberg). These figures correspond well with both
<taxonomicName id="4C534D62FFC4FF087D649BB7FA1EF827" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1308,1436,1955,1981]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC4FF087D649BB7FA1EF827" box="[1308,1436,1955,1981]" italics="true" pageId="10" pageNumber="11">ciliatocosta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFC4FF0878BE9BD3FEA4F87A" authorityName="Huckett" authorityYear="1965" box="[198,294,1991,2016]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC4FF0878BE9BD3FEA4F87A" box="[198,294,1991,2016]" italics="true" pageId="10" pageNumber="11">contigua</emphasis>
</taxonomicName>
. I examined this specimen and found that the terminalia is mounted on a permanent slide preparation, which is why it is slightly compressed, and the surstylus thus appears wider than that of the Hennig’s figure of the type of
<taxonomicName id="4C534D62FFC5FF0979609CAFFDAAFF4F" box="[280,552,187,213]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0979609CAFFDAAFF4F" box="[280,552,187,213]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
. Unfortunately, the terminalia of the
<typeStatus id="54E88843FFC5FF097BBC9CAFFB98FF4F" box="[964,1050,187,213]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF097C479CAFFACDFF4F" box="[1087,1359,187,213]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097C479CAFFACDFF4F" box="[1087,1359,187,213]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
is also mounted on a poorly prepared permanent mount, and it was impossible to describe the form and exact width of the surstylus.
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978BF9D33FEBEFE4B" blockId="11.[151,1437,151,825]" pageId="11" pageNumber="12">
Specimens of both sexes of these species are distinctive, with a completely black body, facial edge projecting beyond level of profrons, mid tibia with
<emphasis id="B927EAF3FFC5FF097A0B9D5FFDFDFEFE" box="[627,639,331,356]" italics="true" pageId="11" pageNumber="12">v</emphasis>
setae and hind femur without
<emphasis id="B927EAF3FFC5FF097B9F9D5FFB83FEFE" box="[999,1025,331,356]" italics="true" pageId="11" pageNumber="12">pv</emphasis>
setae, scutellum with downwardly-directed preapical setulae on upper border of declivities, shining prementum, and costa with costal spinules. I therefore conclude that
<taxonomicName id="4C534D62FFC5FF0979D39D87FCB2FE37" authority="Huckett, 1965" authorityName="Huckett" authorityYear="1965" box="[427,816,402,429]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="contigua">
<emphasis id="B927EAF3FFC5FF0979D39D87FD04FE36" box="[427,646,403,429]" italics="true" pageId="11" pageNumber="12">Spilogona contigua</emphasis>
<bibRefCitation id="EFC24B10FFC5FF097AE89D87FCB2FE37" author="Huckett, H. C." box="[656,816,402,429]" pageId="11" pageNumber="12" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965</bibRefCitation>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="4C534D62FFC5FF097CF19D87FEBAFE4B" authority="Schnabl, 1915" authorityName="Schnabl" authorityYear="1915" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097CF19D87FA1EFE37" box="[1161,1436,403,429]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
Schnabl, 1915
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978BF9DCFFA93FD6B" blockId="11.[151,1437,151,825]" pageId="11" pageNumber="12">
In the
<collectionCode id="ED42AE24FFC5FF09796B9DCFFECEFE6F" box="[275,332,475,501]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="12">ZISP</collectionCode>
, apart from the male
<typeStatus id="54E88843FFC5FF097A3C9DCFFD19FE6F" box="[580,667,475,501]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF097ABD9DCFFC54FE6F" box="[709,982,475,501]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097ABD9DCFFC54FE6F" box="[709,982,475,501]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
there is a female named as
<taxonomicName id="4C534D62FFC5FF097D6C9DCFFE95FD83" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097D6C9DCFFE95FD83" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
. The locality is the same as the male (Karskaja Tundra). However, this specimen does not agree with the male
<typeStatus id="54E88843FFC5FF0978879E37FED4FDA7" box="[255,342,547,573]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF0979049E37FD0EFDA7" box="[380,652,547,573]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0979049E37FD0EFDA7" box="[380,652,547,573]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC24B10FFC5FF097AE29E37FCCAFDA7" author="Hennig, W." box="[666,840,546,573]" pageId="11" pageNumber="12" pagination="233 - 288" refId="ref18511" refString="Hennig, W. (1959 a) Muscidae. [Part.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 63 b, Lieferung 204. E. Schweizerbart, Stuttgart, pp. 233-288." type="book chapter" year="1959">Hennig (1959a)</bibRefCitation>
also had doubts about the identity of this female. The female
<typeStatus id="54E88843FFC5FF0978939E53FEC0FDFB" box="[235,322,583,609]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF0979109E53FDF5FDFB" box="[360,631,583,609]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0979109E53FDF5FDFB" box="[360,631,583,609]" italics="true" pageId="11" pageNumber="12">Limnophora ciliatocosta</emphasis>
</taxonomicName>
does not have the facial edge projecting beyond level of profrons, costa with very strong costal spines, mid tibia without
<emphasis id="B927EAF3FFC5FF097AB89E7FFD4EFD1E" box="[704,716,619,644]" italics="true" pageId="11" pageNumber="12">v</emphasis>
setae and hind femur with several
<emphasis id="B927EAF3FFC5FF097C209E7FFBF0FD1E" box="[1112,1138,619,644]" italics="true" pageId="11" pageNumber="12">pv</emphasis>
setae, proboscis short and thickened, prementum dusted, body with grey dust, not black, thorax grey with three distinct brown longitudinal stripes and with 2+3
<emphasis id="B927EAF3FFC5FF0979F89EA7FE18FD57" box="[384,410,691,717]" italics="true" pageId="11" pageNumber="12">dc</emphasis>
setae, abdomen wet, but with traces of paired brown spots, postgenital plate with hairs. This female is definitely a different species and the specimen runs to
<taxonomicName id="4C534D62FFC5FF097B1E9EC3FA8EFD6B" authority="(Ringdahl, 1926)" baseAuthorityName="Ringdahl" baseAuthorityYear="1926" box="[870,1292,727,753]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tornensis">
<emphasis id="B927EAF3FFC5FF097B1E9EC3FBC1FD6A" box="[870,1091,727,753]" italics="true" pageId="11" pageNumber="12">Spilogona tornensis</emphasis>
(Ringdahl, 1926)
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978BF9EEFFEDCFCA3" blockId="11.[151,1437,151,825]" pageId="11" pageNumber="12">
<taxonomicName id="4C534D62FFC5FF0978BF9EEFFE3DFC8F" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[199,447,763,789]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF0978BF9EEFFE3DFC8F" box="[199,447,763,789]" italics="true" pageId="11" pageNumber="12">Spilogona ciliatocosta</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C534D62FFC5FF097AF29EEFFB91FC8F" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[650,1043,762,789]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tundrae">
<emphasis id="B927EAF3FFC5FF097AF29EEFFCDAFC8F" box="[650,856,763,789]" italics="true" pageId="11" pageNumber="12">Spilogona tundrae</emphasis>
(Schnabl, 1915)
</taxonomicName>
. These species can be separated as following couple:
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0978EF9F70FABFFBE3" pageId="11" pageNumber="12" type="key">
<keyStep id="30A72E44FFC5FF0978EF9F70FA1EFB91" pageId="11" pageNumber="12">
<key id="FFC261D0FFC5FF0978EF9F70FA1EFB91" pageId="11" pageNumber="12">
<paragraph id="8BEC36E1FFC5FF0978EF9F70FA1FFC29" blockId="11.[151,1437,867,1036]" pageId="11" pageNumber="12">
<keyLead id="30A295D4FFC5FF0978EF9F70FA1FFC29" pageId="11" pageNumber="12">
1 Larger species (
<quantity id="4CAB9B04FFC5FF0979029F77FE4CFCE3" box="[378,462,867,889]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="11" pageNumber="12" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
), costal setulae weak. Male: fore tibia with
<emphasis id="B927EAF3FFC5FF097B0A9F70FCFCFCE3" box="[882,894,868,889]" italics="true" pageId="11" pageNumber="12">p</emphasis>
seta in middle; fronto-orbital plates separated by a wide frontal vitta, which is as wide as width of outer margins of ocelli or wider; abdomen shining; anal segment swollen, when viewed in profile bulging towards the dorsal region. Female: abdomen blackish.............................
<taxonomicName id="4C534D62FFC5FF097D439F8BFA1FFC29" ID-CoL="4Z427" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1339,1437,926,948]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tundrae">
<emphasis id="B927EAF3FFC5FF097D439F8BFA1FFC29" box="[1339,1437,926,948]" italics="true" pageId="11" pageNumber="12">S. tundrae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978EF9FA8FA1EFB91" blockId="11.[151,1437,867,1036]" pageId="11" pageNumber="12">
<keyLead id="30A295D4FFC5FF0978EF9FA8FA1EFB91" pageId="11" pageNumber="12">
- Smaller species (
<quantity id="4CAB9B04FFC5FF0979FA9FAFFE65FC4B" box="[386,487,955,977]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="11" pageNumber="12" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
), costal setulae coarsened proximad of costal spine. Male: fore tibia without
<emphasis id="B927EAF3FFC5FF097CC19FA8FB47FC4B" box="[1209,1221,956,977]" italics="true" pageId="11" pageNumber="12">p</emphasis>
seta in middle; fronto-orbital plates touching or separated by a narrow frontal vitta; abdomen subshining; anal segment flattish or appressed dorsad, not bulging toward the dorsal region. Female: abdomen bronzy.......................................
<taxonomicName id="4C534D62FFC5FF097D6F9FE3FA1EFB91" ID-CoL="4Z3TF" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1303,1436,1014,1036]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097D6F9FE3FA1EFB91" box="[1303,1436,1014,1036]" italics="true" pageId="11" pageNumber="12">S. ciliatocosta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
<paragraph id="8BEC36E1FFC5FF0978EF982FFABFFBE3" blockId="11.[151,1437,1083,1217]" pageId="11" pageNumber="12">
All of the characters given above for
<taxonomicName id="4C534D62FFC5FF097A20982FFCD0FBCF" ID-CoL="4Z3TF" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[600,850,1083,1109]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliatocosta">
<emphasis id="B927EAF3FFC5FF097A20982FFCD0FBCF" box="[600,850,1083,1109]" italics="true" pageId="11" pageNumber="12">Spilogona ciliatocosta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFC5FF097BE8982FFBE3FBCF" ID-CoL="4Z427" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[912,1121,1083,1109]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tundrae">
<emphasis id="B927EAF3FFC5FF097BE8982FFBE3FBCF" box="[912,1121,1083,1109]" italics="true" pageId="11" pageNumber="12">Spilogona tundrae</emphasis>
</taxonomicName>
are variable, but are more commonly as stated in the couplet. The male terminalia provides the most reliable identification character.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0978BF9896FF76FB5B" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BEC36E1FFC5FF0978BF9896FF76FB5B" blockId="11.[151,1437,1083,1217]" pageId="11" pageNumber="12">
<emphasis id="B927EAF3FFC5FF0978BF9896FEE3FB06" bold="true" box="[199,353,1154,1180]" pageId="11" pageNumber="12">Distribution.</emphasis>
PALAEARCTIC:
<collectingCountry id="F3447671FFC5FF097A469897FD0BFB07" box="[574,649,1155,1181]" name="Russia" pageId="11" pageNumber="12">Russia</collectingCountry>
(Arctic Siberia and Far East). NEARCTIC: Alaska, Yukon, NWT, Quebec.
</paragraph>
</subSubSection>
</treatment>
</document>

235
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<paragraph id="8BEC36E1FFC5FF0978EF9906FDF8FAB6" blockId="11.[151,634,1298,1361]" box="[151,634,1298,1325]" pageId="11" pageNumber="12">
<heading id="D0A4818DFFC5FF0978EF9906FDF8FAB6" bold="true" box="[151,634,1298,1325]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C534D62FFC5FF0978EF9906FDF8FAB6" ID-CoL="6YZTC" authority="Huchett, 1965" authorityName="Huchett" authorityYear="1965" box="[151,634,1298,1325]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC5FF0978EF9906FDF8FAB6" bold="true" box="[151,634,1298,1325]" pageId="11" pageNumber="12">
<emphasis id="B927EAF3FFC5FF0978EF9906FE46FAB7" bold="true" box="[151,452,1298,1325]" italics="true" pageId="11" pageNumber="12">Spilogona churchillensis</emphasis>
Huchett, 1965
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0978EF9923FE95FACB" box="[151,279,1335,1361]" pageId="11" pageNumber="12" type="description">
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<figureCitation id="13682A64FFC5FF0978EF9923FE95FACB" box="[151,279,1335,1361]" captionStart="FIGURE 4" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[161,1436,502,1903]" captionTargetId="figure-177@12.[151,1436,489,1913]" captionTargetPageId="12" captionText="FIGURE 4. Spilogona ssp. A–C. Spilogona churchillensis. A. Terminalia, lateral view. B. Cercal plate, dorsal view. C. Sternite 5. D. Spilogona quinquesetosa, sternite 5. E–K. Spilogona cordyluraeformis, scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14812242" httpUri="https://zenodo.org/record/14812242/files/figure.png" pageId="11" pageNumber="12">Figs 4 A–C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0978EF996FFD04F95D" pageId="11" pageNumber="12" type="reference_group">
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<treatmentCitationGroup id="AB4311CFFFC5FF0978EF996FFE7FFA2B" pageId="11" pageNumber="12">
<treatmentCitation id="0AF210F0FFC5FF0978EF996FFDE5FA09" author="Huckett, H. C." box="[151,615,1403,1427]" page="213" pageId="11" pageNumber="12" year="1965">
<taxonomicName id="4C534D62FFC5FF0978EF996FFDE5FA09" ID-CoL="6YZTC" authority="Huckett, 1965: 213" authorityName="Huckett" authorityPageNumber="213" authorityYear="1965" box="[151,615,1403,1427]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC5FF0978EF996FFE16FA09" box="[151,404,1403,1427]" italics="true" pageId="11" pageNumber="12">Spilogona churchillensis</emphasis>
<materialsCitation id="3B3B3CBCFFC5FF0979E5996FFDE5FA09" box="[413,615,1403,1427]" collectingDate="1947-07-22" collectionCode="T" collectorName="N. Freeman" country="Canada" location="Manitoba" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Manitoba">
<bibRefCitation id="EFC24B10FFC5FF0979E5996FFDE5FA09" author="Huckett, H. C." box="[413,615,1403,1427]" pageId="11" pageNumber="12" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 213</bibRefCitation>
</materialsCitation>
</taxonomicName>
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. Type-locality: “Churchill,
<collectingRegion id="4997F803FFC5FF097BFB996FFC65FA09" box="[899,999,1403,1427]" country="Canada" name="Manitoba" pageId="11" pageNumber="12">Manitoba</collectingRegion>
, 7.22.47 [i.e.
<date id="FFED1021FFC5FF097C01996FFB72FA09" box="[1145,1264,1403,1427]" pageId="11" pageNumber="12" value="1947-07-22">
<collectingDate id="EFA9E9C9FFC5FF097C01996FFB72FA09" box="[1145,1264,1403,1427]" pageId="11" pageNumber="12" value="1947-07-22">22.vii.1947</collectingDate>
</date>
], T.
<collectorName id="26A65337FFC5FF097D6F9968FA11FA09" box="[1303,1427,1403,1427]" pageId="11" pageNumber="12">N. Freeman</collectorName>
” [
<collectingCountry id="F3447671FFC5FF0978B5998EFEBEFA28" box="[205,316,1434,1458]" name="Canada" pageId="11" pageNumber="12">CANADA</collectingCountry>
,
<collectingRegion id="4997F803FFC5FF097931998EFE2FFA28" box="[329,429,1434,1458]" country="Canada" name="Manitoba" pageId="11" pageNumber="12">Manitoba</collectingRegion>
] [seen].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978EF99ACFEFFFA75" blockId="11.[151,1437,1403,1519]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB4311CFFFC5FF0978EF99ACFEFFFA75" pageId="11" pageNumber="12">
<taxonomicName id="4C534D62FFC5FF0978EF99ACFDE5FA4A" ID-CoL="4Z3Y9" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,615,1464,1488]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC5FF0978EF99ACFE20FA4A" box="[151,418,1464,1488]" italics="true" pageId="11" pageNumber="12">Spilogona quinquesetosa</emphasis>
(Schnabl, 1915)
</taxonomicName>
;
<treatmentCitation id="0AF210F0FFC5FF097A0699ACFCF0FA4B" author="Sorokina, V. S." box="[638,882,1464,1489]" page="212" pageId="11" pageNumber="12" year="2012">
<bibRefCitation id="EFC24B10FFC5FF097A0699ACFCF0FA4B" author="Sorokina, V. S." box="[638,882,1464,1489]" pageId="11" pageNumber="12" pagination="193 - 222" refId="ref19187" refString="Sorokina, V. S. (2012 b) Fauna of Muscidae (Diptera) of the Altai Mountains. Proceedings of the Russian Entomological Society, 83 (1), 193-222. [in Russian, with English abstract]" type="journal article" year="2012">Sorokina, 2012b: 212</bibRefCitation>
</treatmentCitation>
(
<collectingRegion id="4997F803FFC5FF097BF599ACFC42FA4A" box="[909,960,1464,1488]" country="Russia" name="Altay" pageId="11" pageNumber="12">Altai</collectingRegion>
Mts); Sorokina, 2018: 244 (
<collectingRegion id="4997F803FFC5FF097D5899ACFAD1FA4A" box="[1312,1363,1464,1488]" country="Russia" name="Altay" pageId="11" pageNumber="12">Altai</collectingRegion>
Mts). Misidentification.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978EF9A09FD04F95D" blockId="11.[151,1437,1565,1736]" pageId="11" pageNumber="12">
<emphasis id="B927EAF3FFC5FF0978EF9A09FF63F9AD" bold="true" box="[151,225,1565,1591]" pageId="11" pageNumber="12">Notes.</emphasis>
<taxonomicName id="4C534D62FFC5FF09789D9A09FE75F9AD" ID-CoL="6YZTC" authorityName="Huckett" authorityYear="1965" box="[229,503,1565,1591]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC5FF09789D9A09FE75F9AD" box="[229,503,1565,1591]" italics="true" pageId="11" pageNumber="12">Spilogona churchillensis</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName id="4C534D62FFC5FF097A829A09FB92F9AD" ID-CoL="4Z3Y9" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[762,1040,1565,1591]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC5FF097A829A09FB92F9AD" box="[762,1040,1565,1591]" italics="true" pageId="11" pageNumber="12">Spilogona quinquesetosa</emphasis>
</taxonomicName>
in 2018 (Sorokina, 2018). However, the re-examination of the types of both species and DNA analyses shows that
<taxonomicName id="4C534D62FFC5FF097B839A55FA92F9C1" ID-CoL="6YZTC" authorityName="Huckett" authorityYear="1965" box="[1019,1296,1601,1627]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC5FF097B839A55FA92F9C1" box="[1019,1296,1601,1627]" italics="true" pageId="11" pageNumber="12">Spilogona churchillensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C534D62FFC5FF0978EF9A71FE32F9E5" ID-CoL="4Z3Y9" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,432,1637,1663]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC5FF0978EF9A71FE32F9E5" box="[151,432,1637,1663]" italics="true" pageId="11" pageNumber="12">Spilogona quinquesetosa</emphasis>
</taxonomicName>
. The
<typeStatus id="54E88843FFC5FF0979889A71FDD0F9E5" box="[496,594,1637,1663]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C534D62FFC5FF097A009A71FC13F9E5" ID-CoL="4Z3Y9" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[632,913,1637,1663]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC5FF097A009A71FC13F9E5" box="[632,913,1637,1663]" italics="true" pageId="11" pageNumber="12">Spilogona quinquesetosa</emphasis>
</taxonomicName>
is in bad condition: without abdomen and mid and hind legs. According to the original descriptions and examination of the types some morphological differences between these two species have been found:
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC5FF0E78EF9AE6FAF6FE87" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="key">
<keyStep id="30A72E44FFC5FF0978EF9AE6FA1EF822" pageId="11" pageNumber="12">
<key id="FFC261D0FFC5FF0978EF9AE6FA1EF822" pageId="11" pageNumber="12">
<paragraph id="8BEC36E1FFC5FF0978EF9AE6FA1EF8C5" blockId="11.[151,1437,1778,1976]" pageId="11" pageNumber="12">
<keyLead id="30A295D4FFC5FF0978EF9AE6FA1EF8C5" pageId="11" pageNumber="12">
1 Scutellum without downwardly-directed preapical setulae on upper border of declivities; posthumeral and notopleural regions pale grey, in contract to darker adjoining surface; mid tibia usually with one
<emphasis id="B927EAF3FFC5FF097BC69B04FC4BF8BF" box="[958,969,1808,1829]" italics="true" pageId="11" pageNumber="12">v</emphasis>
seta; hind femur with short
<emphasis id="B927EAF3FFC5FF097CAE9B04FB6FF8BF" box="[1238,1261,1808,1829]" italics="true" pageId="11" pageNumber="12">pv</emphasis>
setae in basal 1/3, sometimes these setae elongated; sternite 5 with a wide rectangular median notch (
<figureCitation id="13682A64FFC5FF097B839B39FBCAF8D8" box="[1019,1096,1837,1859]" captionStart="FIGURE 4" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[161,1436,502,1903]" captionTargetId="figure-177@12.[151,1436,489,1913]" captionTargetPageId="12" captionText="FIGURE 4. Spilogona ssp. A–C. Spilogona churchillensis. A. Terminalia, lateral view. B. Cercal plate, dorsal view. C. Sternite 5. D. Spilogona quinquesetosa, sternite 5. E–K. Spilogona cordyluraeformis, scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14812242" httpUri="https://zenodo.org/record/14812242/files/figure.png" pageId="11" pageNumber="12">Fig. 4C</figureCitation>
); male cerci narrower (
<figureCitation id="13682A64FFC5FF097D5E9B39FA0FF8D8" box="[1318,1421,1837,1859]" captionStart="FIGURE 4" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[161,1436,502,1903]" captionTargetId="figure-177@12.[151,1436,489,1913]" captionTargetPageId="12" captionText="FIGURE 4. Spilogona ssp. A–C. Spilogona churchillensis. A. Terminalia, lateral view. B. Cercal plate, dorsal view. C. Sternite 5. D. Spilogona quinquesetosa, sternite 5. E–K. Spilogona cordyluraeformis, scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14812242" httpUri="https://zenodo.org/record/14812242/files/figure.png" pageId="11" pageNumber="12">Fig. 4A, B</figureCitation>
).......................................................................................
<taxonomicName id="4C534D62FFC5FF097C869B5FFA1EF8C5" ID-CoL="6YZTC" authorityName="Huckett" authorityYear="1965" box="[1278,1436,1866,1888]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC5FF097C869B5FFA1EF8C5" box="[1278,1436,1866,1888]" italics="true" pageId="11" pageNumber="12">S. churchillensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36E1FFC5FF0978EF9B7CFA1EF822" blockId="11.[151,1437,1778,1976]" pageId="11" pageNumber="12">
<keyLead id="30A295D4FFC5FF0978EF9B7CFA1EF822" pageId="11" pageNumber="12">
- Scutellum with downwardly-directed preapical setulae on upper border of declivities; posthumeral and notopleural regions dark brown; mid tibia usually with two or more
<emphasis id="B927EAF3FFC5FF097A079B92FD08F801" box="[639,650,1926,1947]" italics="true" pageId="11" pageNumber="12">v</emphasis>
setae; hind femur without
<emphasis id="B927EAF3FFC5FF097BF39B92FC20F801" box="[907,930,1926,1947]" italics="true" pageId="11" pageNumber="12">pv</emphasis>
setae; sternite 5 with a deep triangular median notch (
<figureCitation id="13682A64FFC5FF0978919BB6FEB4F82D" box="[233,310,1954,1976]" captionStart="FIGURE 4" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[161,1436,502,1903]" captionTargetId="figure-177@12.[151,1436,489,1913]" captionTargetPageId="12" captionText="FIGURE 4. Spilogona ssp. A–C. Spilogona churchillensis. A. Terminalia, lateral view. B. Cercal plate, dorsal view. C. Sternite 5. D. Spilogona quinquesetosa, sternite 5. E–K. Spilogona cordyluraeformis, scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14812242" httpUri="https://zenodo.org/record/14812242/files/figure.png" pageId="11" pageNumber="12">Fig. 4D</figureCitation>
); male cerci broader................................................................
<taxonomicName id="4C534D62FFC5FF097C839BB7FA1EF822" ID-CoL="4Z3Y9" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1275,1436,1955,1976]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC5FF097C839BB7FA1EF822" box="[1275,1436,1955,1976]" italics="true" pageId="11" pageNumber="12">S. quinquesetosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
<paragraph id="8BEC36E1FFC2FF0E78EF9C83FAF6FE87" blockId="12.[151,1437,151,465]" pageId="12" pageNumber="13">
The DNA barcoding confirmed the differences between
<taxonomicName id="4C534D62FFC2FF0E7B7D9C83FB9BFF2B" ID-CoL="6YZTC" authorityName="Huckett" authorityYear="1965" box="[773,1049,151,177]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC2FF0E7B7D9C83FB9BFF2B" box="[773,1049,151,177]" italics="true" pageId="12" pageNumber="13">Spilogona churchillensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFC2FF0E7C359C83FAE6FF2A" ID-CoL="4Z3Y9" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1101,1380,151,177]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC2FF0E7C359C83FAE6FF2A" box="[1101,1380,151,177]" italics="true" pageId="12" pageNumber="13">Spilogona quinquesetosa</emphasis>
</taxonomicName>
. The DNA barcodes for specimens under the name
<taxonomicName id="4C534D62FFC2FF0E7AF19CAFFCB4FF4E" ID-CoL="4Z3Y9" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[649,822,187,213]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quinquesetosa">
“
<emphasis id="B927EAF3FFC2FF0E7AEE9CAFFCB4FF4E" box="[662,822,187,212]" italics="true" pageId="12" pageNumber="13">quinquesetosa</emphasis>
</taxonomicName>
” from the Chukotka population and the Altai population were very different, with an intraspecific p-distance of from 4.26% to 4.59%. Altai specimens of
<taxonomicName id="4C534D62FFC2FF0E7C9B9CCBFA12FF62" ID-CoL="4Z3Y9" authority="(Sorokina, 2018)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1251,1424,223,249]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quinquesetosa">
“
<emphasis id="B927EAF3FFC2FF0E7C889CCBFA12FF62" box="[1264,1424,223,248]" italics="true" pageId="12" pageNumber="13">quinquesetosa</emphasis>
</taxonomicName>
” (Sorokina, 2018) proved to be genetically closest to
<taxonomicName id="4C534D62FFC2FF0E7A999D17FC77FE87" ID-CoL="6YZTC" authorityName="Huckett" authorityYear="1965" box="[737,1013,259,285]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC2FF0E7A999D17FC77FE87" box="[737,1013,259,285]" italics="true" pageId="12" pageNumber="13">Spilogona churchillensis</emphasis>
</taxonomicName>
from
<collectingRegion id="4997F803FFC2FF0E7C429D17FB2AFE87" box="[1082,1192,259,285]" country="Canada" name="Manitoba" pageId="12" pageNumber="13">Manitoba</collectingRegion>
,
<collectingCountry id="F3447671FFC2FF0E7CCB9D17FA85FE87" box="[1203,1287,259,285]" name="Canada" pageId="12" pageNumber="13">Canada</collectingCountry>
(BOLD).
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC2FF0E78EF9D33FEE7FE13" pageId="12" pageNumber="13" type="description">
<paragraph id="8BEC36E1FFC2FF0E78EF9D33FEE7FE13" blockId="12.[151,1437,151,465]" pageId="12" pageNumber="13">
Thus,
<taxonomicName id="4C534D62FFC2FF0E78A39D33FD0FFEDB" authority="Huckett, 1965" authorityName="Huckett" authorityYear="1965" box="[219,653,294,321]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC2FF0E78A39D33FE6FFEDB" box="[219,493,295,321]" italics="true" pageId="12" pageNumber="13">Spilogona churchillensis</emphasis>
<bibRefCitation id="EFC24B10FFC2FF0E79899D33FD0FFEDB" author="Huckett, H. C." box="[497,653,294,321]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965</bibRefCitation>
</taxonomicName>
is valid species and is not a junior synonym of
<taxonomicName id="4C534D62FFC2FF0E7CFE9D33FECEFEFF" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC2FF0E7CFE9D33FA1EFEDA" box="[1158,1436,295,321]" italics="true" pageId="12" pageNumber="13">Spilogona quinquesetosa</emphasis>
(Schnabl, 1915)
</taxonomicName>
.
<taxonomicName id="4C534D62FFC2FF0E792E9D5FFDE8FEFF" authorityName="Huckett" authorityYear="1965" box="[342,618,331,357]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="churchillensis">
<emphasis id="B927EAF3FFC2FF0E792E9D5FFDE8FEFF" box="[342,618,331,357]" italics="true" pageId="12" pageNumber="13">Spilogona churchillensis</emphasis>
</taxonomicName>
has an arcto-alpine distribution, whereas
<taxonomicName id="4C534D62FFC2FF0E7C409D5FFAD2FEFE" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[1080,1360,331,357]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFC2FF0E7C409D5FFAD2FEFE" box="[1080,1360,331,357]" italics="true" pageId="12" pageNumber="13">Spilogona quinquesetosa</emphasis>
</taxonomicName>
has an arctic distribution.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC2FF0E78BF9D86FB58FE4B" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BEC36E1FFC2FF0E78BF9D86FBA6FE37" blockId="12.[151,1437,151,465]" box="[199,1060,402,429]" pageId="12" pageNumber="13">
<emphasis id="B927EAF3FFC2FF0E78BF9D86FEE3FE36" bold="true" box="[199,353,402,428]" pageId="12" pageNumber="13">Distribution.</emphasis>
PALAEARCTIC: Siberia (
<collectingRegion id="4997F803FFC2FF0E7AEC9D87FD4EFE37" box="[660,716,403,429]" country="Russia" name="Altay" pageId="12" pageNumber="13">Altai</collectingRegion>
Mts). NEARCTIC:
<collectingRegion id="4997F803FFC2FF0E7BC99D87FBA2FE37" box="[945,1056,403,429]" country="Canada" name="Manitoba" pageId="12" pageNumber="13">Manitoba</collectingRegion>
.
</paragraph>
<paragraph id="8BEC36E1FFC2FF0E78BF9DA2FB58FE4B" blockId="12.[151,1437,151,465]" box="[199,1242,438,465]" pageId="12" pageNumber="13">
<emphasis id="B927EAF3FFC2FF0E78BF9DA2FEEBFE4B" bold="true" box="[199,361,438,465]" pageId="12" pageNumber="13">DNA barcode</emphasis>
. BOLD BIN: BOLD:AAB7605. See
<tableCitation id="C6D1035AFFC2FF0E7B759DA3FCE2FE4B" box="[781,864,439,465]" captionStart="TABLE 1" captionStartId="2.[152,245,657,681]" captionText="TABLE 1. List of Russian Spilogona species with barcode index numbers (BIN) and GenBank accession numbers." pageId="12" pageNumber="13">Table 1</tableCitation>
for GenBank accession numbers.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="4C534D62FFC6FF0A78EF9E9AFE19FD33" authorityName="Sorokina" authorityYear="2025" box="[151,411,654,681]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="latizygoma" status="sp. nov.">
<emphasis id="B927EAF3FFC6FF0A78EF9E9AFE19FD33" bold="true" box="[151,411,654,681]" italics="true" pageId="8" pageNumber="9">Spilogona latizygoma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2145788FFC6FF0A79DA9E9BFD82FD33" box="[418,512,655,681]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0A78EF9EA7FE8CFD57" box="[151,270,691,717]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BEC36E1FFC6FF0A78EF9EA7FE8CFD57" blockId="8.[151,512,654,717]" box="[151,270,691,717]" pageId="8" pageNumber="9">
<figureCitation id="13682A64FFC6FF0A78EF9EA7FE8CFD57" box="[151,270,691,717]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="8" pageNumber="9">Figs 3 A–I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0A78EF9EE3FB3AFC95" box="[151,1208,759,783]" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BEC36E1FFC6FF0A78EF9EE3FB3AFC95" blockId="8.[151,1208,759,783]" box="[151,1208,759,783]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB4311CFFFC6FF0A78EF9EE3FB3AFC95" box="[151,1208,759,783]" pageId="8" pageNumber="9">
<taxonomicName id="4C534D62FFC6FF0A78EF9EE3FE7DFC95" ID-CoL="4Z3ZP" authority="(Malloch, 1920)" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[151,511,759,783]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFC6FF0A78EF9EE3FED7FC95" box="[151,341,759,783]" italics="true" pageId="8" pageNumber="9">Spilogona tendipes</emphasis>
(Malloch, 1920)
</taxonomicName>
;
<treatmentCitation id="0AF210F0FFC6FF0A7A729EE3FCDAFC95" ID-DOI="http://doi.org/10.5281/zenodo.6125953" ID-Zenodo-Dep="6125953" author="Sorokina, V. S. &amp; Michelsen, V." box="[522,856,759,783]" httpUri="http://treatment.plazi.org/id/DD40B94FE0414200FF0CF92294DAFE5E" page="516" pageId="8" pageNumber="9" year="2014">
<bibRefCitation id="EFC24B10FFC6FF0A7A729EE3FCDAFC95" author="Sorokina, V. S. &amp; Michelsen, V." box="[522,856,759,783]" pageId="8" pageNumber="9" pagination="512 - 520" refId="ref19692" refString="Sorokina, V. S. &amp; Michelsen, V. (2014) Contributions to the taxonomy and faunistics of some arctic species of Spilogona Schnabl (Diptera: Muscidae). Zootaxa, 3814 (4), 512-520. https://doi.org/10.11646/zootaxa.3814.4.4" type="journal article" year="2014">Sorokina &amp; Michelsen 2014: 516</bibRefCitation>
</treatmentCitation>
. Wrangel Island. Misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0A78EF9F29FD2CFA9D" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BEC36E1FFC6FF0A78EF9F29FB54FCE1" blockId="8.[151,1437,829,2008]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78EF9F29FEC9FCCD" bold="true" box="[151,331,829,855]" pageId="8" pageNumber="9">Type material:</emphasis>
<materialsCitation id="3B3B3CBCFFC6FF0A792E9F29FB54FCE1" collectingDate="1966-07-19" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" county="Mineeva Mts" location="Wrangel Island" municipality="River Somnitil'naya" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54E88843FFC6FF0A792E9F29FE3CFCCD" box="[342,446,829,855]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
<specimenCount id="9D55FD68FFC6FF0A79B19F2AFE62FCC2" box="[457,480,830,856]" count="1" pageId="8" pageNumber="9" type="male">♂</specimenCount>
,
<collectingCountry id="F3447671FFC6FF0A79889F29FDD4FCCD" box="[496,598,829,855]" name="Russia" pageId="8" pageNumber="9">RUSSIA</collectingCountry>
,
<emphasis id="B927EAF3FFC6FF0A7A1E9F29FC97FCCD" box="[614,789,829,855]" italics="true" pageId="8" pageNumber="9">
<location id="8E8C603AFFC6FF0A7A1E9F29FC97FCCD" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7A1E9F29FC97FCCD" box="[614,789,829,855]" country="Russia" county="Mineeva Mts" municipality="River Somnitil'naya" name="Wrangel Island" pageId="8" pageNumber="9">Wrangel Island</location>
</emphasis>
:
<location id="8E8C603AFFC6FF0A7B509F29FB82FCC2" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7B509F29FB82FCC2" box="[808,1024,829,856]" country="Russia" county="Mineeva Mts" municipality="River Somnitil'naya" name="Somnitil'naya Bay" pageId="8" pageNumber="9">Somnitil’naya Bay</location>
, valley of the
<collectingMunicipality id="6B88AC9BFFC6FF0A7CD59F29FA1AFCCD" box="[1197,1432,829,855]" pageId="8" pageNumber="9">River Somnitil’naya</collectingMunicipality>
,
<collectingCounty id="628D4E6DFFC6FF0A78EF9F75FEACFCE6" box="[151,302,865,892]" pageId="8" pageNumber="9">Mineeva Mts</collectingCounty>
, 150 м, on flowers of
<taxonomicName id="4C534D62FFC6FF0A7A5F9F75FC9EFCE1" box="[551,796,865,891]" class="Magnoliopsida" family="Rosaceae" genus="Potentilla" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="emarginata">
<emphasis id="B927EAF3FFC6FF0A7A5F9F75FC9EFCE1" box="[551,796,865,891]" italics="true" pageId="8" pageNumber="9">Potentilla emarginata</emphasis>
</taxonomicName>
,
<date id="FFED1021FFC6FF0A7B529F75FC2FFCE1" box="[810,941,865,891]" pageId="8" pageNumber="9" value="1966-07-19">
<collectingDate id="EFA9E9C9FFC6FF0A7B529F75FC2FFCE1" box="[810,941,865,891]" pageId="8" pageNumber="9" value="1966-07-19">19.vii.1966</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A7B909F76FBFEFCE1" box="[1000,1148,865,892]" pageId="8" pageNumber="9">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A7CF39F75FB48FCE1" box="[1163,1226,865,891]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="9">ZISP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9F92FD2CFA9D" blockId="8.[151,1437,829,2008]" pageId="8" pageNumber="9">
<materialsCitation id="3B3B3CBCFFC6FF0A78BF9F92FDDEFC05" box="[199,604,901,928]" country="Russia" location="Wrangel Island" pageId="8" pageNumber="9" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E88843FFC6FF0A78BF9F92FEBAFC3A" box="[199,312,902,928]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3447671FFC6FF0A79399F91FE27FC05" box="[321,421,901,927]" name="Russia" pageId="8" pageNumber="9">RUSSIA</collectingCountry>
,
<emphasis id="B927EAF3FFC6FF0A79CA9F91FDDEFC05" box="[434,604,901,927]" italics="true" pageId="8" pageNumber="9">
<location id="8E8C603AFFC6FF0A79CA9F91FDDEFC05" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A79CA9F91FDDEFC05" box="[434,604,901,927]" country="Russia" name="Wrangel Island" pageId="8" pageNumber="9">Wrangel Island</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B3B3CBCFFC6FF0A7A119F91FC9FFC59" collectingDate="1966-07-22" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" county="Mineeva Mts" location="Somnitil'naya Bay" municipality="River Somnitil'naya" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A7A119F91FD0EFC05" box="[617,652,901,927]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
,
<location id="8E8C603AFFC6FF0A7AE19F91FCE9FC3A" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7AE19F91FCE9FC3A" box="[665,875,901,928]" country="Russia" county="Mineeva Mts" municipality="River Somnitil'naya" name="Somnitil'naya Bay" pageId="8" pageNumber="9">Somnitil’naya Bay</location>
, valley of the
<collectingMunicipality id="6B88AC9BFFC6FF0A7C7C9F91FB6BFC05" box="[1028,1257,901,927]" pageId="8" pageNumber="9">River Somnitil’naya</collectingMunicipality>
, south of island,
<collectingCounty id="628D4E6DFFC6FF0A78EF9FBDFEA9FC5E" box="[151,299,937,964]" pageId="8" pageNumber="9">Mineeva Mts</collectingCounty>
, 250 м,
<date id="FFED1021FFC6FF0A79F89FBDFD80FC59" box="[384,514,937,963]" pageId="8" pageNumber="9" value="1966-07-22">
<collectingDate id="EFA9E9C9FFC6FF0A79F89FBDFD80FC59" box="[384,514,937,963]" pageId="8" pageNumber="9" value="1966-07-22">22.vii.1966</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A7A409FBEFD48FC59" box="[568,714,937,964]" pageId="8" pageNumber="9">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A7AAE9FBDFC97FC59" box="[726,789,937,963]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="9">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A7B5E9FBDFE48FC7D" collectingDate="1966-07-26" collectionCode="SZMN" collectorName="K. Gorodkov" country="Russia" location="Somnitil'naya Bay" pageId="8" pageNumber="9" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A7B5E9FBDFCE2FC59" box="[806,864,937,963]" count="2" pageId="8" pageNumber="9" type="female">2♀♀</specimenCount>
,
<location id="8E8C603AFFC6FF0A7B129FBDFBB9FC5E" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7B129FBDFBB9FC5E" box="[874,1083,937,964]" country="Russia" name="Somnitil'naya Bay" pageId="8" pageNumber="9">Somnitil’naya Bay</location>
, south of island, 22,
<date id="FFED1021FFC6FF0A7D6E9FBDFA15FC59" box="[1302,1431,937,963]" pageId="8" pageNumber="9" value="1966-07-26">
<collectingDate id="EFA9E9C9FFC6FF0A7D6E9FBDFA15FC59" box="[1302,1431,937,963]" pageId="8" pageNumber="9" value="1966-07-26">26.vii.1966</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A78B19FDAFEDCFC7D" box="[201,350,973,1000]" pageId="8" pageNumber="9">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A79169FD9FE40FC7D" box="[366,450,973,999]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A79AE9FD9FB1CFB91" collectingDate="2016-07-12" collectingDateMax="2016-07-16" collectingDateMin="2016-07-12" collectionCode="SZMN" collectorName="L. Volkova" country="Russia" latitude="71.01667" location="River Neozhidannaya" longLatPrecision="973" longitude="-179.15" pageId="8" pageNumber="9" specimenCode="BUIC-DIP1761, BUIC-DIP1762" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A79AE9FD9FD8CFC7D" box="[470,526,973,999]" count="2" pageId="8" pageNumber="9" type="male">2♂♂</specimenCount>
<specimenCount id="9D55FD68FFC6FF0A7A6E9FD9FDCDFC7D" box="[534,591,973,999]" count="4" pageId="8" pageNumber="9" type="female">4♀♀</specimenCount>
, middle part of the
<location id="8E8C603AFFC6FF0A7B499FD9FBABFC7D" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7B499FD9FBABFC7D" box="[817,1065,973,999]" country="Russia" latitude="71.01667" longLatPrecision="973" longitude="-179.15" name="River Neozhidannaya" pageId="8" pageNumber="9">River Neozhidannaya</location>
,
<geoCoordinate id="EE675026FFC6FF0A7C4D9FD9FB17FC7D" box="[1077,1173,973,999]" degrees="71" direction="north" minutes="01" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="71.01667">71°01’N</geoCoordinate>
<geoCoordinate id="EE675026FFC6FF0A7CE69FD9FA91FC7D" box="[1182,1299,973,999]" degrees="179" direction="west" minutes="09" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-179.15">179°09’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7D679FD9FF52FB96" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
,
<date id="FFED1021FFC6FF0A78A29FE5FE04FB91" box="[218,390,1009,1035]" pageId="8" pageNumber="9" value="2016-07-12" valueMax="2016-07-16" valueMin="2016-07-12">
<collectingDate id="EFA9E9C9FFC6FF0A78A29FE5FE04FB91" box="[218,390,1009,1035]" pageId="8" pageNumber="9" value="2016-07-12" valueMax="2016-07-16" valueMin="2016-07-12">12–16.vii.2016</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A79B89FE6FDB8FB91" box="[448,570,1009,1036]" pageId="8" pageNumber="9">L. Volkova</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A7A309FE5FD18FB91" box="[584,666,1009,1035]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
,
<specimenCount id="9D55FD68FFC6FF0A7ADC9FE5FD3DFB91" box="[676,703,1009,1035]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
—
<specimenCode id="DBF59E9AFFC6FF0A7AA69FE5FC15FB91" box="[734,919,1009,1035]" collectionCode="BUIC-DIP" pageId="8" pageNumber="9">BUIC-DIP1761</specimenCode>
,
<specimenCount id="9D55FD68FFC6FF0A7BD99FE5FC35FB91" box="[929,951,1009,1035]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
—
<specimenCode id="DBF59E9AFFC6FF0A7BAE9FE5FB14FB91" box="[982,1174,1009,1035]" collectionCode="BUIC-DIP" pageId="8" pageNumber="9">BUIC-DIP1762</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A7CD09FE5FF79FBC9" collectingDate="2016-06-10" collectingDateMax="2016-07-27" collectingDateMin="2016-06-10" collectionCode="SZMN" collectorName="L. Khruleva" country="Russia" latitude="71.01667" location="River Neozhidannaya" longLatPrecision="973" longitude="-179.13333" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A7CD09FE5FB49FB91" box="[1192,1227,1009,1035]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
, middle part of the
<location id="8E8C603AFFC6FF0A78EF9801FE0DFBB5" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A78EF9801FE0DFBB5" box="[151,399,1045,1071]" country="Russia" latitude="71.01667" longLatPrecision="973" longitude="-179.13333" name="River Neozhidannaya" pageId="8" pageNumber="9">River Neozhidannaya</location>
, valley of river,
<geoCoordinate id="EE675026FFC6FF0A7A359801FD2FFBB5" box="[589,685,1045,1071]" degrees="71" direction="north" minutes="01" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="71.01667">71°01’N</geoCoordinate>
<geoCoordinate id="EE675026FFC6FF0A7ACE9801FCA9FBB5" box="[694,811,1045,1071]" degrees="179" direction="west" minutes="08" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-179.13333">179°08’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7B409801FC75FBAA" box="[824,1015,1045,1072]" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
,
<date id="FFED1021FFC6FF0A7C7C9801FB56FBB5" box="[1028,1236,1045,1071]" pageId="8" pageNumber="9" value="2016-06-10" valueMax="2016-07-27" valueMin="2016-06-10">
<collectingDate id="EFA9E9C9FFC6FF0A7C7C9801FB56FBB5" box="[1028,1236,1045,1071]" pageId="8" pageNumber="9" value="2016-06-10" valueMax="2016-07-27" valueMin="2016-06-10">10.vi.–27.vii.2016</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A7D6B9802FA1EFBB5" box="[1299,1436,1045,1072]" pageId="8" pageNumber="9">L. Khruleva</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A78E7982DFF71FBC9" box="[159,243,1081,1107]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A7971982DFE08FBED" collectingDate="2019-07-09" collectionCode="SZMN" collectorName="L. Khruleva" country="Russia" latitude="71.01667" location="River Neozhidannaya" longLatPrecision="973" longitude="-179.15" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A7971982DFEAEFBC9" box="[265,300,1081,1107]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
, middle part of the
<location id="8E8C603AFFC6FF0A7A6B982DFC8EFBC9" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7A6B982DFC8EFBC9" box="[531,780,1081,1107]" country="Russia" latitude="71.01667" longLatPrecision="973" longitude="-179.15" name="River Neozhidannaya" pageId="8" pageNumber="9">River Neozhidannaya</location>
,
<geoCoordinate id="EE675026FFC6FF0A7B61982DFCFBFBC9" box="[793,889,1081,1107]" degrees="71" direction="north" minutes="01" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="71.01667">71°01’N</geoCoordinate>
<geoCoordinate id="EE675026FFC6FF0A7BFB982DFC7AFBC9" box="[899,1016,1081,1107]" degrees="179" direction="west" minutes="09" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-179.15">179°09’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7C7E982DFB44FBCE" box="[1030,1222,1081,1108]" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
, 8,
<date id="FFED1021FFC6FF0A7C89982DFAE4FBC9" box="[1265,1382,1081,1107]" pageId="8" pageNumber="9" value="2019-07-09">
<collectingDate id="EFA9E9C9FFC6FF0A7C89982DFAE4FBC9" box="[1265,1382,1081,1107]" pageId="8" pageNumber="9" value="2019-07-09">9.vii.2019</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A78EF984AFE9DFBED" box="[151,287,1117,1144]" pageId="8" pageNumber="9">L. Khruleva</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A79569849FE00FBED" box="[302,386,1117,1143]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A79EE9849FF79FB01" collectingDate="2018-07-08" collectingDateMax="2018-07-10" collectingDateMin="2018-07-08" collectionCode="SZMN" collectorName="U. Babii" country="Russia" latitude="71.26667" location="Tundrovii Mt." longLatPrecision="972" longitude="-179.88333" pageId="8" pageNumber="9" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A79EE9849FE4CFBED" box="[406,462,1117,1143]" count="2" pageId="8" pageNumber="9" type="male">2♂♂</specimenCount>
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,
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,
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<geoCoordinate id="EE675026FFC6FF0A7B629849FC0DFBE2" box="[794,911,1117,1144]" degrees="179" direction="west" minutes="53" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-179.88333">179°53’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7BE39849FBD5FBE2" box="[923,1111,1117,1144]" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
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<date id="FFED1021FFC6FF0A7C1B9849FA83FBED" box="[1123,1281,1117,1143]" pageId="8" pageNumber="9" value="2018-07-08" valueMax="2018-07-10" valueMin="2018-07-08">
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</date>
, leg.
<collectorName id="26A65337FFC6FF0A7D45984AFA1FFBED" box="[1341,1437,1117,1144]" pageId="8" pageNumber="9">U. Babii</collectorName>
(
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)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A797F9895FF79FB25" collectingDate="2019-07-07" collectingDateMax="2019-07-11" collectingDateMin="2019-07-07" collectionCode="SZMN" collectorName="U. Babii &amp; O. Khruleva" country="Russia" latitude="71.3" location="Tundrovii Mt." longLatPrecision="972" longitude="-179.8" pageId="8" pageNumber="9" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A797F9895FEC3FB01" box="[263,321,1153,1179]" count="2" pageId="8" pageNumber="9" type="female">2♀♀</specimenCount>
,
<location id="8E8C603AFFC6FF0A79369895FE6DFB06" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A79369895FE6DFB06" box="[334,495,1153,1180]" country="Russia" latitude="71.3" longLatPrecision="972" longitude="-179.8" name="Tundrovii Mt." pageId="8" pageNumber="9">Tundrovii Mt.</location>
,
<geoCoordinate id="EE675026FFC6FF0A79849895FDDFFB01" box="[508,605,1153,1179]" degrees="71" direction="north" minutes="18" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="71.3">71°18’N</geoCoordinate>
<geoCoordinate id="EE675026FFC6FF0A7A1D9895FD58FB01" box="[613,730,1153,1179]" degrees="179" direction="west" minutes="48" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-179.8">179°48’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7A9E9895FC26FB06" box="[742,932,1153,1180]" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
,
<date id="FFED1021FFC6FF0A7BC89895FBCFFB01" box="[944,1101,1153,1179]" pageId="8" pageNumber="9" value="2019-07-07" valueMax="2019-07-11" valueMin="2019-07-07">
<collectingDate id="EFA9E9C9FFC6FF0A7BC89895FBCFFB01" box="[944,1101,1153,1179]" pageId="8" pageNumber="9" value="2019-07-07" valueMax="2019-07-11" valueMin="2019-07-07">7–11.vii.2019</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A7CF39896FB69FB01" box="[1163,1259,1153,1180]" pageId="8" pageNumber="9">U. Babii</collectorName>
&amp;
<collectorName id="26A65337FFC6FF0A7D699895FA1EFB01" box="[1297,1436,1153,1179]" pageId="8" pageNumber="9">O. Khruleva</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A78E798B1FF71FB25" box="[159,243,1189,1215]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A797F98B1FDA9FB79" collectingDate="2015-07-02" collectingDateMax="2015-07-07" collectingDateMin="2015-07-02" collectionCode="SZMN" collectorName="O. Khruleva" country="Russia" latitude="71.166664" location="River Mamontovaya" longLatPrecision="972" longitude="-179.75" pageId="8" pageNumber="9" specimenCode="BUIC-DIP1763" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A797F98B1FEA8FB25" box="[263,298,1189,1215]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
, middle part of the
<location id="8E8C603AFFC6FF0A7A7398B1FD74FB5A" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7A7398B1FD74FB5A" box="[523,758,1189,1216]" country="Russia" latitude="71.166664" longLatPrecision="972" longitude="-179.75" name="River Mamontovaya" pageId="8" pageNumber="9">River Mamontovaya</location>
,
<geoCoordinate id="EE675026FFC6FF0A7B7998B1FCE0FB25" box="[769,866,1189,1215]" degrees="71" direction="north" minutes="10" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="71.166664">71°10’N</geoCoordinate>
<geoCoordinate id="EE675026FFC6FF0A7B1198B1FC5CFB5A" box="[873,990,1189,1216]" degrees="179" direction="west" minutes="45" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-179.75">179°45’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7B9398B1FB2AFB5A" box="[1003,1192,1189,1216]" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
,
<date id="FFED1021FFC6FF0A7CCC98B1FAC6FB5A" box="[1204,1348,1189,1216]" pageId="8" pageNumber="9" value="2015-07-02" valueMax="2015-07-07" valueMin="2015-07-02">
<collectingDate id="EFA9E9C9FFC6FF0A7CCC98B1FAC6FB5A" box="[1204,1348,1189,1216]" pageId="8" pageNumber="9" value="2015-07-02" valueMax="2015-07-07" valueMin="2015-07-02">2–7.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A7DF998B1FF7DFB79" pageId="8" pageNumber="9">O. Khruleva</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A797698DDFEE2FB79" box="[270,352,1225,1251]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
,
<specimenCode id="DBF59E9AFFC6FF0A791498DDFDA1FB79" box="[364,547,1225,1251]" collectionCode="BUIC-DIP" pageId="8" pageNumber="9">BUIC-DIP1763</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC6FF0A7A4E98DDFD28FA9D" collectingDate="2015-07-04" collectingDateMax="2015-07-11" collectingDateMin="2015-07-04" collectionCode="SZMN" collectorName="O. Khruleva" country="Russia" latitude="71.21667" location="River Neizvestnaya" longLatPrecision="972" longitude="-170.31667" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC6FF0A7A4E98DDFDDBFB79" box="[566,601,1225,1251]" count="1" pageId="8" pageNumber="9" type="female">1♀</specimenCount>
, upper reaches of the
<location id="8E8C603AFFC6FF0A7B2A98DDFBB3FB79" LSID="urn:lsid:plazi:treatment:03FA87F7FFC6FF0878EF9E9AFCC3FB5F:8E8C603AFFC6FF0A7B2A98DDFBB3FB79" box="[850,1073,1225,1251]" country="Russia" latitude="71.21667" longLatPrecision="972" longitude="-170.31667" name="River Neizvestnaya" pageId="8" pageNumber="9">River Neizvestnaya</location>
,
<geoCoordinate id="EE675026FFC6FF0A7C4598DDFB19FB79" box="[1085,1179,1225,1251]" degrees="71" direction="north" minutes="13" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="71.21667">71º13’N</geoCoordinate>
<geoCoordinate id="EE675026FFC6FF0A7CDA98DDFA97FB79" box="[1186,1301,1225,1251]" degrees="170" direction="west" minutes="19" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-170.31667">170º19’W</geoCoordinate>
,
<collectingMethod id="52124EF6FFC6FF0A7D5898DDFF52FA92" pageId="8" pageNumber="9">yellow pan traps</collectingMethod>
,
<date id="FFED1021FFC6FF0A78A498F9FEFBFA92" box="[220,377,1261,1288]" pageId="8" pageNumber="9" value="2015-07-04" valueMax="2015-07-11" valueMin="2015-07-04">
<collectingDate id="EFA9E9C9FFC6FF0A78A498F9FEFBFA92" box="[220,377,1261,1288]" pageId="8" pageNumber="9" value="2015-07-04" valueMax="2015-07-11" valueMin="2015-07-04">4–11.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC6FF0A79CC98F9FDBDFA9D" box="[436,575,1261,1288]" pageId="8" pageNumber="9">O. Khruleva</collectorName>
(
<collectionCode id="ED42AE24FFC6FF0A7A3698F9FD20FA9D" box="[590,674,1261,1287]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="9">SZMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0A78BF9905FDADFAD5" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BEC36E1FFC6FF0A78BF9905FDADFAD5" blockId="8.[151,1437,829,2008]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9905FECCFAB1" bold="true" box="[199,334,1297,1323]" pageId="8" pageNumber="9">Etymology.</emphasis>
The species name is based on the Latin words “latus” and “zygoma” meaning “wide cheekbones”, because the parafacial is very broad.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0A78BF994DFD61F9F5" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BEC36E1FFC6FF0A78BF994DFD61F9F5" blockId="8.[151,1437,829,2008]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF994DFEC0FAE9" bold="true" box="[199,322,1369,1395]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
The new species is very similar to
<taxonomicName id="4C534D62FFC6FF0A7A9B994DFBF5FAE9" authority="(Malloch, 1920)" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[739,1143,1369,1395]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFC6FF0A7A9B994DFC3AFAE9" box="[739,952,1369,1395]" italics="true" pageId="8" pageNumber="9">Spilogona tendipes</emphasis>
(Malloch, 1920)
</taxonomicName>
because of its large size, covering of dense grey dusting with a bluish or greenish tinge and having paired dark marks visible on abdomen, a projecting lower face, small eyes, very broad parafacial, long and slender haustellum with a shiny prementum and slender and elongate labella, 4 pairs of postsutural dorsocentral setae, hind tarsus with a
<emphasis id="B927EAF3FFC6FF0A7CCB99D2FB3DFA45" box="[1203,1215,1478,1503]" italics="true" pageId="8" pageNumber="9">v</emphasis>
seta near base, and hypoproct with recurrent spines. But the new species differs substantially from
<taxonomicName id="4C534D62FFC6FF0A7C6399FDFB6CF999" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[1051,1262,1513,1539]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFC6FF0A7C6399FDFB6CF999" box="[1051,1262,1513,1539]" italics="true" pageId="8" pageNumber="9">Spilogona tendipes</emphasis>
</taxonomicName>
by the absence of ventral seta on mid tibia, distinct long
<emphasis id="B927EAF3FFC6FF0A7A179A1AFD0BF9BD" box="[623,649,1550,1575]" italics="true" pageId="8" pageNumber="9">pv</emphasis>
in basal half of hind femur, numerous hair-like
<emphasis id="B927EAF3FFC6FF0A7CD69A1AFB4AF9BD" box="[1198,1224,1550,1575]" italics="true" pageId="8" pageNumber="9">av</emphasis>
setae in basal half of all femora, long ground-setulae of scutum, long setae on sternites 3–5, more greenish tinge of body, almost inconspicuous hairs on eye, and the male terminalia.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0878BF9A6DFCE0FB8B" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="8BEC36E1FFC6FF0A78BF9A6DFCCBF90E" blockId="8.[151,1437,829,2008]" box="[199,841,1657,1684]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9A6DFED3F909" bold="true" box="[199,337,1657,1683]" pageId="8" pageNumber="9">Description</emphasis>
: Male: body
<quantity id="4CAB9B04FFC6FF0A79909A6DFDF3F90E" box="[488,625,1657,1684]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="8" pageNumber="9" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.3–5.7 mm</quantity>
, wing 4.8–5.0 mm.
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9A89FD0EF84D" blockId="8.[151,1437,829,2008]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9A89FE85F92D" bold="true" box="[199,263,1693,1719]" pageId="8" pageNumber="9">Head</emphasis>
: Ground-colour black. Frons, fronto-orbital plate, gena and parafacial shining silvery-grey or silverybrown, with dark reflections when viewed in profile. In lateral view, facial edge projecting beyond level of profrons (
<figureCitation id="13682A64FFC6FF0A78E79AF1FF75F89A" box="[159,247,1765,1792]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="8" pageNumber="9">Fig. 3A</figureCitation>
). Eye small, bare or with very sparse and inconspicuous hairs. Fronto-orbital plates separated by a wide frontal vitta. Distance between eye margins on upper part of frons about as wide as ocellar triangle (
<figureCitation id="13682A64FFC6FF0A7C999B1DFABAF8B9" box="[1249,1336,1801,1827]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
). Frontal triangle extending to middle of frons. Frons with 12–14 pairs of strong frontal setae. Parafacial at level of base of antenna 1.5–2 times width of postpedicel, not narrowing below. Height of gena about 1/3–1/4 of height of eye and 1.5–2 times width of postpedicel. Antenna black, postpedicel approximately 1.5 times as long as wide. Arista black, bare, thickened in basal half. Palpus black. Proboscis long, slender, prementum shining. Labella slender and elongate, crocheted relative to the proboscis.
</paragraph>
<caption id="DF2C6669FFC7FF0B78EF9A0CFE19F906" ID-DOI="http://doi.org/10.5281/zenodo.14812240" ID-Zenodo-Dep="14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1560,1584]" targetBox="[166,1422,181,1525]" targetPageId="9" targetType="figure">
<paragraph id="8BEC36E1FFC7FF0B78EF9A0CFE19F906" blockId="9.[151,1437,1560,1693]" pageId="9" pageNumber="10">
<emphasis id="B927EAF3FFC7FF0B78EF9A0CFE97F9AA" bold="true" box="[151,277,1560,1584]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
<taxonomicName id="4C534D62FFC7FF0B79659A0CFE79F9AA" authorityName="Sorokina" authorityYear="2025" box="[285,507,1560,1584]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="latizygoma" status="sp. nov.">
<emphasis id="B927EAF3FFC7FF0B79659A0CFE79F9AA" box="[285,507,1560,1584]" italics="true" pageId="9" pageNumber="10">Spilogona latizygoma</emphasis>
</taxonomicName>
<emphasis id="B927EAF3FFC7FF0B7A7C9A0DFDF5F9AA" bold="true" box="[516,631,1560,1585]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A2145788FFC7FF0B7A7C9A0DFDD4F9AB" box="[516,598,1561,1585]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Holotype, male, lateral view.
<emphasis id="B927EAF3FFC7FF0B7BCA9A0DFC4BF9AB" bold="true" box="[946,969,1561,1585]" pageId="9" pageNumber="10">B.</emphasis>
Holotype, frons and scutum, dorsal view.
<emphasis id="B927EAF3FFC7FF0B7DFC9A0CFA1EF9AA" bold="true" box="[1412,1436,1560,1584]" pageId="9" pageNumber="10">C.</emphasis>
Paratype, female, lateral view.
<emphasis id="B927EAF3FFC7FF0B79A39A29FE76F9CF" bold="true" box="[475,500,1597,1621]" pageId="9" pageNumber="10">D.</emphasis>
Terminalia, lateral view, digital photo, paratype.
<emphasis id="B927EAF3FFC7FF0B7B839A29FB93F9CF" bold="true" box="[1019,1041,1597,1621]" pageId="9" pageNumber="10">E.</emphasis>
Sternite 5, digital photo, paratype.
<emphasis id="B927EAF3FFC7FF0B7DF19A29FA1EF9CF" bold="true" box="[1417,1436,1597,1621]" pageId="9" pageNumber="10">F.</emphasis>
Ovipositor, lateral, dorsal and ventral views, digital photo.
<emphasis id="B927EAF3FFC7FF0B7A8B9A74FC8CF9E2" bold="true" box="[755,782,1632,1656]" pageId="9" pageNumber="10">G.</emphasis>
Terminalia, lateral view, paratype.
<emphasis id="B927EAF3FFC7FF0B7C009A75FB10F9E3" bold="true" box="[1144,1170,1633,1657]" pageId="9" pageNumber="10">H.</emphasis>
Cercal plate, dorsal view.
<emphasis id="B927EAF3FFC7FF0B78EF9A91FF25F907" bold="true" box="[151,167,1669,1693]" pageId="9" pageNumber="10">I.</emphasis>
Sternite 5. Bare: 1 mm.
</paragraph>
</caption>
<paragraph id="8BEC36E1FFC7FF0B78BF9AFBFDBDF803" blockId="9.[151,1437,1775,2017]" pageId="9" pageNumber="10">
<emphasis id="B927EAF3FFC7FF0B78BF9AFBFEA2F893" bold="true" box="[199,288,1775,1801]" pageId="9" pageNumber="10">Thorax</emphasis>
: Ground-colour black, densely grey dusted with a bluish or greenish tinge, with two brown indistinct presutural longitudinal stripes when viewed from in front. Ground-setulae of scutum relatively long, hair-like. Presutural
<emphasis id="B927EAF3FFC7FF0B79699B23FEB4F8CA" box="[273,310,1847,1872]" italics="true" pageId="9" pageNumber="10">acr</emphasis>
setae in 3–4 uneven rows, 2+4
<emphasis id="B927EAF3FFC7FF0B7ADC9B23FD3CF8CB" box="[676,702,1847,1873]" italics="true" pageId="9" pageNumber="10">dc</emphasis>
setae. Notopleuron bare, or with individual hairs. Anepisternum with 2–3 interspatial setae. Katepisternal setae 1+1. Scutellum grey dusted, with downwardly-directed preapical setulae on upper border of declivities.
</paragraph>
<paragraph id="8BEC36E1FFC7FF0B78BF9BB6FC93F87B" blockId="9.[151,1437,1775,2017]" pageId="9" pageNumber="10">
<emphasis id="B927EAF3FFC7FF0B78BF9BB6FE8AF826" bold="true" box="[199,264,1954,1980]" pageId="9" pageNumber="10">Wing</emphasis>
: Light brownish tinged, with dark base. Costa without spinules or costal spine. Calypters whitish, with light yellow margins. Haltere brown with blackish knob.
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9C83FF52FE13" blockId="10.[151,1438,151,1221]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9C83FF7DFF2B" bold="true" box="[199,255,151,177]" pageId="10" pageNumber="11">Legs</emphasis>
: Ground colour black, grey dusted. Fore tibia with 2–4
<emphasis id="B927EAF3FFC4FF087B0D9C83FC01FF2A" box="[885,899,151,176]" italics="true" pageId="10" pageNumber="11">p</emphasis>
setae. Fore femur with numerous long hair-like setae on posterior surface, with a row of short hair-like
<emphasis id="B927EAF3FFC4FF087B7F9CAFFCA3FF4E" box="[775,801,187,212]" italics="true" pageId="10" pageNumber="11">av</emphasis>
setae. Mid femur with a row of long
<emphasis id="B927EAF3FFC4FF087CBE9CAFFB62FF4E" box="[1222,1248,187,212]" italics="true" pageId="10" pageNumber="11">pv</emphasis>
setae and with a row of short hair-like
<emphasis id="B927EAF3FFC4FF0879F79CCBFE3DFF62" box="[399,447,223,248]" italics="true" pageId="10" pageNumber="11">av-v</emphasis>
setae, with 2
<emphasis id="B927EAF3FFC4FF087A269CCBFD17FF63" box="[606,661,223,249]" italics="true" pageId="10" pageNumber="11">pd–p</emphasis>
preapical setae. Mid tibia with 2–3
<emphasis id="B927EAF3FFC4FF087C569CCBFBC8FF63" box="[1070,1098,223,249]" italics="true" pageId="10" pageNumber="11">ad</emphasis>
and 3
<emphasis id="B927EAF3FFC4FF087CE09CCBFB36FF63" box="[1176,1204,223,249]" italics="true" pageId="10" pageNumber="11">pd</emphasis>
setae, without any
<emphasis id="B927EAF3FFC4FF087DE89CCBFA1EFF62" box="[1424,1436,223,248]" italics="true" pageId="10" pageNumber="11">v</emphasis>
setae. Hind femur with a row of long
<emphasis id="B927EAF3FFC4FF087A459D17FDD5FE86" box="[573,599,259,284]" italics="true" pageId="10" pageNumber="11">av</emphasis>
setae becoming shorter on basal third, with a row of long
<emphasis id="B927EAF3FFC4FF087C9D9D17FB7DFE86" box="[1253,1279,259,284]" italics="true" pageId="10" pageNumber="11">pv</emphasis>
setae in basal half, with numerous long hair like setae on basal part of posterior surface. Hind tibia with 3–5
<emphasis id="B927EAF3FFC4FF087CBB9D33FB5FFEDA" box="[1219,1245,295,320]" italics="true" pageId="10" pageNumber="11">av</emphasis>
, 4–6
<emphasis id="B927EAF3FFC4FF087D669D33FAB8FEDB" box="[1310,1338,295,321]" italics="true" pageId="10" pageNumber="11">ad</emphasis>
, with an uneven row of long
<emphasis id="B927EAF3FFC4FF08790E9D5FFE2BFEFF" box="[374,425,331,357]" italics="true" pageId="10" pageNumber="11">pd-p</emphasis>
setae in middle part, without
<emphasis id="B927EAF3FFC4FF087A8A9D5FFC8EFEFE" box="[754,780,331,356]" italics="true" pageId="10" pageNumber="11">pv</emphasis>
setae. First tarsomere of hind tarsus with strong
<emphasis id="B927EAF3FFC4FF087D509D5FFAB6FEFE" box="[1320,1332,331,356]" italics="true" pageId="10" pageNumber="11">v</emphasis>
seta near base.
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9D86FC2EFD83" blockId="10.[151,1438,151,1221]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9D86FEBEFE36" bold="true" box="[199,316,402,428]" pageId="10" pageNumber="11">Abdomen</emphasis>
: Black with dense bluish-grey dust, conical. Tergites 1+2–4 with large brown triangular marks separated by a brown median stripe, sometimes fused, tergite 5 with a brown median stripe. Sternite 1 bare or with individual hairs, sternites 2–5 with numerous long setae, especially sternite 5, sternite 5 large, longer then sternite 4, with a deep median notch, and a pointed caudal margin (
<figureCitation id="13682A64FFC4FF087B549DEBFC1FFD83" box="[812,925,511,537]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="10" pageNumber="11">Figs 3E, I</figureCitation>
).
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9E36FE2AFD33" blockId="10.[151,1438,151,1221]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9E36FEC8FDA6" bold="true" box="[199,330,546,572]" pageId="10" pageNumber="11">Terminalia</emphasis>
: In lateral view cerci wide in proximal half and narrow, rod-shaped in distal half, curved; surstylus rounded at apex, with a notch in ventral margin, shorter than cerci; epandrium large, hemispherical; hypandrium 2 times shorter then epandrium, with a wavy ventral margin (
<figureCitation id="13682A64FFC4FF087B0A9E7FFC7EFD1F" box="[882,1020,619,645]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="10" pageNumber="11">Figs 3D, G</figureCitation>
). Cercal plate oval with two apical prolongations (
<figureCitation id="13682A64FFC4FF0879389E9BFE19FD33" box="[320,411,655,681]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="10" pageNumber="11">Fig. 3H</figureCitation>
).
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9EA7FD4CFD57" blockId="10.[151,1438,151,1221]" box="[199,718,690,717]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9EA7FE9CFD57" bold="true" box="[199,286,691,717]" pageId="10" pageNumber="11">Female</emphasis>
: body
<quantity id="4CAB9B04FFC4FF0879149EA7FE77FD57" box="[364,501,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.3" metricValueMin="6.2" pageId="10" pageNumber="11" unit="mm" value="6.75" valueMax="7.3" valueMin="6.2">6.2–7.3 mm</quantity>
, wing
<quantity id="4CAB9B04FFC4FF087A389EA6FD48FD57" box="[576,714,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="10" pageNumber="11" unit="mm" value="6.2" valueMax="6.7" valueMin="5.7">5.7–6.7 mm</quantity>
.
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9EC3FD0BFD6B" blockId="10.[151,1438,151,1221]" box="[199,649,727,753]" pageId="10" pageNumber="11">Similar to male but differing as follows:</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9EEFFB6CFC3F" blockId="10.[151,1438,151,1221]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9EEFFE85FC8F" bold="true" box="[199,263,763,789]" pageId="10" pageNumber="11">Head</emphasis>
: Frons, fronto-orbital plate, gena and parafacial densely brown dusted, contrasting with the greenish-bluish-grey occiput (
<figureCitation id="13682A64FFC4FF0879059F0BFE56FCA3" box="[381,468,799,825]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="10" pageNumber="11">Fig. 3C</figureCitation>
). Frontal vitta matt grey, contrasting with brown fronto-orbital plate and frontal triangle when viewed from above. Width of frontal vitta 2–2.3 times width of a fronto-orbital plate. Frontal triangle brown dusted, extending to lunule. Fronto-orbital plate with an uneven row of frontal and orbital setae. Parafacial at level of base of antenna 2–2.3 times width of postpedicel. Height of gena about 1/2–1/3 of height of eye.
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9FBBFDBDFC77" blockId="10.[151,1438,151,1221]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9FBBFEA2FC53" bold="true" box="[199,288,943,969]" pageId="10" pageNumber="11">Thorax</emphasis>
: Colour lighter, with greenish-bluish-grey dust. Scutum with 3 brown longitudinal stripes, sometimes indistinct, not extending to scutellum.
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF9FE2FCE0FB8B" blockId="10.[151,1438,151,1221]" box="[199,866,1014,1041]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9FE2FEBEFB8A" bold="true" box="[199,316,1014,1040]" pageId="10" pageNumber="11">Abdomen</emphasis>
: Postgenital plate with weak individual spinules.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC4FF0878BF980EFB5BFBC3" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BEC36E1FFC4FF0878BF980EFC15FBAF" blockId="10.[151,1438,151,1221]" box="[199,919,1050,1077]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF980EFEE3FBAE" bold="true" box="[199,353,1050,1076]" pageId="10" pageNumber="11">Distribution.</emphasis>
PALAEARCTIC: Arctic
<collectingCountry id="F3447671FFC4FF087AF8980FFD49FBAF" box="[640,715,1051,1077]" name="Russia" pageId="10" pageNumber="11">Russia</collectingCountry>
(Wrangel Island).
</paragraph>
<paragraph id="8BEC36E1FFC4FF0878BF982AFB5BFBC3" blockId="10.[151,1438,151,1221]" box="[199,1241,1086,1113]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF982AFEEBFBC3" bold="true" box="[199,361,1086,1113]" pageId="10" pageNumber="11">DNA barcode</emphasis>
. BOLD BIN: BOLD:ADZ7218. See
<tableCitation id="C6D1035AFFC4FF087B74982BFCDDFBC3" box="[780,863,1087,1113]" captionStart="TABLE 1" captionStartId="2.[152,245,657,681]" captionText="TABLE 1. List of Russian Spilogona species with barcode index numbers (BIN) and GenBank accession numbers." pageId="10" pageNumber="11">Table 1</tableCitation>
for GenBank accession numbers.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC4FF0878BF9877FCC3FB5F" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BEC36E1FFC4FF0878BF9877FCC3FB5F" blockId="10.[151,1438,151,1221]" pageId="10" pageNumber="11">
<emphasis id="B927EAF3FFC4FF0878BF9877FEB9FBE7" bold="true" box="[199,315,1123,1149]" pageId="10" pageNumber="11">Remarks.</emphasis>
<taxonomicName id="4C534D62FFC4FF08793F9877FDC2FBE7" authorityName="Sorokina" authorityYear="2025" box="[327,576,1123,1149]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="latizygoma">
<emphasis id="B927EAF3FFC4FF08793F9877FDC2FBE7" box="[327,576,1123,1149]" italics="true" pageId="10" pageNumber="11">Spilogona latizygoma</emphasis>
</taxonomicName>
morphological is very similar to
<taxonomicName id="4C534D62FFC4FF087BAB9877FB28FBE7" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[979,1194,1123,1149]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFC4FF087BAB9877FB28FBE7" box="[979,1194,1123,1149]" italics="true" pageId="10" pageNumber="11">Spilogona tendipes</emphasis>
</taxonomicName>
. But based on DNA barcodes, these species are genetically different; the p-distance between them is 3.36%. The intraspecific distance of
<taxonomicName id="4C534D62FFC4FF0878EF98BFFE09FB5F" authorityName="Sorokina" authorityYear="2025" box="[151,395,1195,1221]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="latizygoma">
<emphasis id="B927EAF3FFC4FF0878EF98BFFE09FB5F" box="[151,395,1195,1221]" italics="true" pageId="10" pageNumber="11">Spilogona latizygoma</emphasis>
</taxonomicName>
specimens ranges from 0.0% to 0.16%.
</paragraph>
</subSubSection>
</treatment>
</document>

415
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<mods:title id="31E20C2B909DF403654AF93DFF38D079">New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species</mods:title>
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<paragraph id="8BEC36E1FFC8FF0478EF9E7EFE74FD1F" blockId="6.[151,502,618,681]" box="[151,502,618,645]" pageId="6" pageNumber="7">
<heading id="D0A4818DFFC8FF0478EF9E7EFE74FD1F" bold="true" box="[151,502,618,645]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<taxonomicName id="4C534D62FFC8FF0478EF9E7EFE10FD1F" authorityName="Sorokina" authorityYear="2025" box="[151,402,618,645]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gorodkovi" status="sp. nov.">
<emphasis id="B927EAF3FFC8FF0478EF9E7EFE10FD1F" bold="true" box="[151,402,618,645]" italics="true" pageId="6" pageNumber="7">Spilogona gorodkovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2145788FFC8FF0479E19E7FFE74FD1F" box="[409,502,619,645]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C349656AFFC8FF0478EF9E9BFE8CFD33" box="[151,270,655,681]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BEC36E1FFC8FF0478EF9E9BFE8CFD33" blockId="6.[151,502,618,681]" box="[151,270,655,681]" pageId="6" pageNumber="7">
<figureCitation id="13682A64FFC8FF0478EF9E9BFE8CFD33" box="[151,270,655,681]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="6" pageNumber="7">Figs 2 A–I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC8FF0478EF9EC3FBC2FC77" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BEC36E1FFC8FF0478EF9EC3FBC2FC77" blockId="6.[151,1437,726,1941]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478EF9EC3FEC7FD6A" bold="true" box="[151,325,726,753]" pageId="6" pageNumber="7">Type material:</emphasis>
<materialsCitation id="3B3B3CBCFFC8FF0479319EC3FC69FC8F" collectingDate="1972-07-28" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" location="Wrangel Island" municipality="River Somnitil'naya" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54E88843FFC8FF0479319EC3FE33FD6B" box="[329,433,727,753]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
<specimenCount id="9D55FD68FFC8FF0479CE9EC3FE4FFD6B" box="[438,461,727,753]" count="1" pageId="6" pageNumber="7" type="male">♂</specimenCount>
,
<collectingCountry id="F3447671FFC8FF0479AE9EC3FDBEFD6B" box="[470,572,727,753]" name="Russia" pageId="6" pageNumber="7">RUSSIA</collectingCountry>
,
<emphasis id="B927EAF3FFC8FF047A3D9EC3FD6DFD6B" box="[581,751,727,753]" italics="true" pageId="6" pageNumber="7">
<location id="8E8C603AFFC8FF047A3D9EC3FD6DFD6B" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF047A3D9EC3FD6DFD6B" box="[581,751,727,753]" country="Russia" municipality="River Somnitil'naya" name="Wrangel Island" pageId="6" pageNumber="7">Wrangel Island</location>
</emphasis>
:
<locationDeviation id="9A5E2FC3FFC8FF047A839EC3FBA2FD6B" box="[763,1056,727,753]" location="Somnitil'naya Bay" pageId="6" pageNumber="7">
<quantity id="4CAB9B04FFC8FF047A839EC3FCB0FD6B" box="[763,818,727,753]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="km" value="4.0">4 km</quantity>
N Somnitil’naya Bay
</locationDeviation>
, valley of the
<collectingMunicipality id="6B88AC9BFFC8FF047CCC9EC3FA1AFD6B" box="[1204,1432,727,753]" pageId="6" pageNumber="7">River Somnitil’naya</collectingMunicipality>
, on flowers of
<taxonomicName id="4C534D62FFC8FF0479409EEFFDACFC8E" box="[312,558,763,789]" class="Magnoliopsida" family="Rosaceae" genus="Potentilla" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="emarginata">
<emphasis id="B927EAF3FFC8FF0479409EEFFDACFC8E" box="[312,558,763,789]" italics="true" pageId="6" pageNumber="7">Potentilla emarginata</emphasis>
</taxonomicName>
,
<date id="FFED1021FFC8FF047A469EEFFD42FC8F" box="[574,704,763,789]" pageId="6" pageNumber="7" value="1972-07-28">
<collectingDate id="EFA9E9C9FFC8FF047A469EEFFD42FC8F" box="[574,704,763,789]" pageId="6" pageNumber="7" value="1972-07-28">28.vii.1972</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC8FF047A869EEFFC16FC8F" box="[766,916,763,789]" pageId="6" pageNumber="7">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC8FF047BDC9EEFFC61FC8F" box="[932,995,763,789]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="7">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3B3CBCFFC8FF047B8F9EEFFA16FC8F" box="[1015,1428,763,789]" country="Russia" location="Wrangel Island" pageId="6" pageNumber="7" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E88843FFC8FF047B8F9EEFFBEAFC8F" box="[1015,1128,763,789]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3447671FFC8FF047C0C9EEFFB5AFC8F" box="[1140,1240,763,789]" name="Russia" pageId="6" pageNumber="7">RUSSIA</collectingCountry>
,
<emphasis id="B927EAF3FFC8FF047C9F9EEFFA16FC8F" box="[1255,1428,763,789]" italics="true" pageId="6" pageNumber="7">
<location id="8E8C603AFFC8FF047C9F9EEFFA16FC8F" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF047C9F9EEFFA16FC8F" box="[1255,1428,763,789]" country="Russia" name="Wrangel Island" pageId="6" pageNumber="7">Wrangel Island</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B3B3CBCFFC8FF0478EF9F0BFDF6FCA3" box="[151,628,799,825]" collectingDate="1972-07-28" collectionCode="SZMN" collectorName="K. Gorodkov" country="Russia" location="Wrangel Island" municipality="River Somnitil'naya" pageId="6" pageNumber="7" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D55FD68FFC8FF0478EF9F0BFF4DFCA3" box="[151,207,799,825]" count="2" pageId="6" pageNumber="7" type="male">2♂♂</specimenCount>
<specimenCount id="9D55FD68FFC8FF0478AD9F0BFE8CFCA3" box="[213,270,799,825]" count="2" pageId="6" pageNumber="7" type="female">2♀♀</specimenCount>
, same data as holotype (
<collectionCode id="ED42AE24FFC8FF047A609F0BFDEEFCA3" box="[536,620,799,825]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="6" pageNumber="7">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC8FF047A069F0BFDF2FCC7" collectingDate="1972-07-25" collectionCode="SZMN" collectorName="K. Gorodkov" country="Russia" elevation="150" location="River Somnitil'naya" pageId="6" pageNumber="7" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="9D55FD68FFC8FF047A069F0BFD35FCA3" box="[638,695,799,825]" count="3" pageId="6" pageNumber="7" type="female">3♀♀</specimenCount>
,
<locationDeviation id="9A5E2FC3FFC8FF047ABB9F0AFC69FCA3" box="[707,1003,798,825]" location="Somnitil'naya Bay" pageId="6" pageNumber="7">
<quantity id="4CAB9B04FFC8FF047ABB9F0AFD78FCA3" box="[707,762,798,825]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="km" value="5.0">5 km</quantity>
N Somnitil’naya Bay
</locationDeviation>
, valley of the
<location id="8E8C603AFFC8FF047CFB9F0BFAEAFCA3" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF047CFB9F0BFAEAFCA3" box="[1155,1384,799,825]" country="Russia" name="River Somnitil'naya" pageId="6" pageNumber="7">River Somnitil’naya</location>
,
<quantity id="4CAB9B04FFC8FF047D0A9F0AFF2DFCC7" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="m" value="150.0">
<elevation id="007ED1D2FFC8FF047D0A9F0AFF2DFCC7" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="FFED1021FFC8FF0478C19F56FEBEFCC7" box="[185,316,834,861]" pageId="6" pageNumber="7" value="1972-07-25">
<collectingDate id="EFA9E9C9FFC8FF0478C19F56FEBEFCC7" box="[185,316,834,861]" pageId="6" pageNumber="7" value="1972-07-25">25.vii.1972</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC8FF04790C9F57FD85FCC7" box="[372,519,835,861]" pageId="6" pageNumber="7">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC8FF047A6C9F57FDEAFCC7" box="[532,616,835,861]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="6" pageNumber="7">SZMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC8FF047A019F57FC72FC1B" collectingDate="1966-07-26" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" location="Somnitil'naya Bay" municipality="River Somnitil'naya" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC8FF047A019F57FD1CFCC7" box="[633,670,835,861]" count="1" pageId="6" pageNumber="7" type="male">1♂</specimenCount>
,
<location id="8E8C603AFFC8FF047AD19F57FCF9FCC7" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF047AD19F57FCF9FCC7" box="[681,891,835,861]" country="Russia" municipality="River Somnitil'naya" name="Somnitil'naya Bay" pageId="6" pageNumber="7">Somnitil’naya Bay</location>
, south of island, bank of the
<collectingMunicipality id="6B88AC9BFFC8FF047CCB9F57FA1AFCC7" box="[1203,1432,835,861]" pageId="6" pageNumber="7">River Somnitil’naya</collectingMunicipality>
, on flowers of
<taxonomicName id="4C534D62FFC8FF0479419F73FDB3FC1A" box="[313,561,871,897]" class="Magnoliopsida" family="Rosaceae" genus="Potentilla" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="emarginata">
<emphasis id="B927EAF3FFC8FF0479419F73FDB3FC1A" box="[313,561,871,897]" italics="true" pageId="6" pageNumber="7">Potentilla emarginata</emphasis>
</taxonomicName>
,
<date id="FFED1021FFC8FF047A399F73FD40FC1B" box="[577,706,871,897]" pageId="6" pageNumber="7" value="1966-07-26">
<collectingDate id="EFA9E9C9FFC8FF047A399F73FD40FC1B" box="[577,706,871,897]" pageId="6" pageNumber="7" value="1966-07-26">26.vii.1966</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC8FF047B7A9F73FC1AFC1B" box="[770,920,871,897]" pageId="6" pageNumber="7">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC8FF047BD19F73FC6AFC1B" box="[937,1000,871,897]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="7">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC8FF047B869F73FBD6FC3F" collectingDate="1966-07-19" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" location="Somnitil'naya Bay" pageId="6" pageNumber="7" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC8FF047B869F73FBA3FC1B" box="[1022,1057,871,897]" count="1" pageId="6" pageNumber="7" type="male">1♂</specimenCount>
<specimenCount id="9D55FD68FFC8FF047C529F73FBCFFC1B" box="[1066,1101,871,897]" count="1" pageId="6" pageNumber="7" type="female">1♀</specimenCount>
,
<location id="8E8C603AFFC8FF047C259F73FAB0FC1B" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF047C259F73FAB0FC1B" box="[1117,1330,871,897]" country="Russia" name="Somnitil'naya Bay" pageId="6" pageNumber="7">Somnitil’naya Bay</location>
, south of island, tundra, on flowers of
<taxonomicName id="4C534D62FFC8FF0479AF9F9FFD22FC3F" box="[471,672,907,933]" class="Insecta" family="Nymphalidae" genus="Dryas" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="integrifolia">
<emphasis id="B927EAF3FFC8FF0479AF9F9FFD22FC3F" box="[471,672,907,933]" italics="true" pageId="6" pageNumber="7">Dryas integrifolia</emphasis>
</taxonomicName>
,
<date id="FFED1021FFC8FF047AD69F9FFCB2FC3F" box="[686,816,907,933]" pageId="6" pageNumber="7" value="1966-07-19">
<collectingDate id="EFA9E9C9FFC8FF047AD69F9FFCB2FC3F" box="[686,816,907,933]" pageId="6" pageNumber="7" value="1966-07-19">19.vii.1966</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC8FF047B129F9FFC7CFC3F" box="[874,1022,907,933]" pageId="6" pageNumber="7">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC8FF047C749F9FFBC9FC3F" box="[1036,1099,907,933]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="7">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC8FF047C279F9EFA1AFC53" collectingDate="1971-07-23" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" location="River Somnitil'naya" municipality="Mineeva Mts" pageId="6" pageNumber="7" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
<specimenCount id="9D55FD68FFC8FF047C279F9EFB1AFC3F" box="[1119,1176,906,933]" count="5" pageId="6" pageNumber="7" type="female">5♀♀</specimenCount>
,
<locationDeviation id="9A5E2FC3FFC8FF047CDC9F9EFF47FC53" location="Somnitil'naya Bay" pageId="6" pageNumber="7">
<quantity id="4CAB9B04FFC8FF047CDC9F9EFB5FFC3F" box="[1188,1245,906,933]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="km" value="5.0">5 km</quantity>
N Somnitil’naya Bay
</locationDeviation>
, south of island, middle part of the
<location id="8E8C603AFFC8FF047A2F9FBBFCBFFC53" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF047A2F9FBBFCBFFC53" box="[599,829,943,969]" country="Russia" municipality="Mineeva Mts" name="River Somnitil'naya" pageId="6" pageNumber="7">River Somnitil’naya</location>
,
<collectingMunicipality id="6B88AC9BFFC8FF047B329FBBFC63FC53" box="[842,993,943,969]" pageId="6" pageNumber="7">Mineeva Mts</collectingMunicipality>
,
<date id="FFED1021FFC8FF047B959FBBFBEDFC53" box="[1005,1135,943,969]" pageId="6" pageNumber="7" value="1971-07-23">
<collectingDate id="EFA9E9C9FFC8FF047B959FBBFBEDFC53" box="[1005,1135,943,969]" pageId="6" pageNumber="7" value="1971-07-23">23.vii.1971</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC8FF047CD49FBBFAC3FC53" box="[1196,1345,943,969]" pageId="6" pageNumber="7">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC8FF047D289FBBFA0DFC53" box="[1360,1423,943,969]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="7">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3B3CBCFFC8FF0478EF9FC7FBB9FC77" box="[151,1083,978,1005]" collectingDate="1971-07-18" collectionCode="ZISP" collectorName="K. Gorodkov" country="Russia" location="Rodzhers Bay" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D55FD68FFC8FF0478EF9FC7FF3EFC77" box="[151,188,979,1005]" count="1" pageId="6" pageNumber="7" type="male">1♂</specimenCount>
,
<quantity id="4CAB9B04FFC8FF0478B09FC6FE8DFC77" box="[200,271,978,1005]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="km" value="15.0">15 km</quantity>
W
<location id="8E8C603AFFC8FF04794E9FC7FE57FC77" LSID="urn:lsid:plazi:treatment:03FA87F7FFC8FF0A78EF9E7EFC15FDA7:8E8C603AFFC8FF04794E9FC7FE57FC77" box="[310,469,979,1005]" country="Russia" name="Rodzhers Bay" pageId="6" pageNumber="7">Rodzhers Bay</location>
, south of island,
<date id="FFED1021FFC8FF047AEB9FC7FC97FC77" box="[659,789,979,1005]" pageId="6" pageNumber="7" value="1971-07-18">
<collectingDate id="EFA9E9C9FFC8FF047AEB9FC7FC97FC77" box="[659,789,979,1005]" pageId="6" pageNumber="7" value="1971-07-18">18.vii.1971</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFC8FF047B299FC7FC67FC77" box="[849,997,979,1005]" pageId="6" pageNumber="7">K. Gorodkov</collectorName>
(
<collectionCode id="ED42AE24FFC8FF047B8C9FC7FBB1FC77" box="[1012,1075,979,1005]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="7">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC8FF0478BF9FE3FBFAFBAF" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BEC36E1FFC8FF0478BF9FE3FBFAFBAF" blockId="6.[151,1437,726,1941]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478BF9FE3FECCFB8B" bold="true" box="[199,334,1015,1041]" pageId="6" pageNumber="7">Etymology.</emphasis>
The species name is a patronym in honour of the celebrated Soviet dipterist Kirill B. Gorodkov (
<collectingCountry id="F3447671FFC8FF0478D9980FFF72FBAF" box="[161,240,1051,1077]" name="Russia" pageId="6" pageNumber="7">Russia</collectingCountry>
), who collected the
<typeStatus id="54E88843FFC8FF0479B7980FFE7DFBAF" box="[463,511,1051,1077]" pageId="6" pageNumber="7">type</typeStatus>
series of this species as well many other new
<taxonomicName id="4C534D62FFC8FF047C7C980FFBF7FBAF" baseAuthorityName="Hennig" baseAuthorityYear="1959" box="[1028,1141,1051,1077]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Muscidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC8FF0478BF982AFB07FAAB" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BEC36E1FFC8FF0478BF982AFB07FAAB" blockId="6.[151,1437,726,1941]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478BF982AFEC0FBC2" bold="true" box="[199,322,1086,1112]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
The new species is similar to Nearctic
<taxonomicName id="4C534D62FFC8FF047B63982BFB1BFBC3" authority="(Stein, 1920)" baseAuthorityName="Stein" baseAuthorityYear="1920" box="[795,1177,1087,1113]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B927EAF3FFC8FF047B63982BFC78FBC3" box="[795,1018,1087,1113]" italics="true" pageId="6" pageNumber="7">Spilogona rufitarsis</emphasis>
(Stein, 1920)
</taxonomicName>
because of 4 pairs of postsutural dorsocentral setae, facial edge not projecting beyond level of profrons, broad frons, anepisternum without interspatial setae, mid tibia without
<emphasis id="B927EAF3FFC8FF047AF39893FD15FB3A" box="[651,663,1159,1184]" italics="true" pageId="6" pageNumber="7">v</emphasis>
setae, hind femur with
<emphasis id="B927EAF3FFC8FF047BDC9893FC3CFB3A" box="[932,958,1159,1184]" italics="true" pageId="6" pageNumber="7">pv</emphasis>
setae on proximal half, scutellum without downwardly-directed preapical setulae on upper border of declivities, grey abdomen with black spots and a narrow brown median stripe between them. But the new species differs from
<taxonomicName id="4C534D62FFC8FF047BE698DBFBFAFB73" baseAuthorityName="Stein" baseAuthorityYear="1920" box="[926,1144,1231,1257]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B927EAF3FFC8FF047BE698DBFBFAFB73" box="[926,1144,1231,1257]" italics="true" pageId="6" pageNumber="7">Spilogona rufitarsis</emphasis>
</taxonomicName>
by the wider frons, dusted prementum, black fore tibia and tarsi, darker colour of thorax, blackish haltere, wide parafacial which is 1.2–2.0 times width of postpedicel at level of base of antenna, and postgenital plate with spinules.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC8FF0A78BF992EFCFDFD83" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BEC36E1FFC8FF0478BF992EFCCBFACF" blockId="6.[151,1437,726,1941]" box="[199,841,1338,1365]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478BF992EFED3FACE" bold="true" box="[199,337,1338,1364]" pageId="6" pageNumber="7">Description</emphasis>
: Male: body
<quantity id="4CAB9B04FFC8FF047990992EFDF3FACF" box="[488,625,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="5.8" metricValueMin="5.6" pageId="6" pageNumber="7" unit="mm" value="5.699999999999999" valueMax="5.8" valueMin="5.6">5.6–5.8 mm</quantity>
, wing 5.0–
<quantity id="4CAB9B04FFC8FF047A95992EFCC7FACF" box="[749,837,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="7" unit="mm" value="5.2">5.2 mm</quantity>
.
</paragraph>
<paragraph id="8BEC36E1FFC8FF0478BF994BFCDEF927" blockId="6.[151,1437,726,1941]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478BF994BFE85FAE3" bold="true" box="[199,263,1375,1401]" pageId="6" pageNumber="7">Head</emphasis>
: Ground-colour black. Frons, fronto-orbital plate, gena and parafacial shining silvery-grey, with dark reflections when viewed in profile. In lateral view, facial edge not projecting beyond level of profrons (
<figureCitation id="13682A64FFC8FF047D4C9997FA0EFA07" box="[1332,1420,1411,1437]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="6" pageNumber="7">Fig. 2A</figureCitation>
). Eye large, bare or with very sparse and inconspicuous hairs. Fronto-orbital plates separated by a wide black frontal vitta. Distance between eye margins on upper part of frons about as wide as distance between the first pair of
<emphasis id="B927EAF3FFC8FF047DFA99DFFA1EFA7F" box="[1410,1436,1483,1509]" italics="true" pageId="6" pageNumber="7">dc</emphasis>
setae, sometimes a little narrower. Frontal triangle small, grey dusted, not extending to middle of frons. Frons with 8–9 pairs of strong frontal setae and hair-like setae between them. Parafacial at level of base of antenna 1.2–1.3 times width of postpedicel, very weakly narrowing below. Height of gena about 1/4 of height of eye and 1.8–2 times width of postpedicel.Antenna black, postpedicel approximately 1.5 times as long as wide. Arista black, thickened at base, short pubescent, with the longest individual hairs shorter than basal diameter of arista. Palpus black. Proboscis short, thin, prementum broadly dusted in middle. Labella large.
</paragraph>
<paragraph id="8BEC36E1FFC8FF0478BF9AD3FAB5F8D7" blockId="6.[151,1437,726,1941]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478BF9AD3FEA2F97B" bold="true" box="[199,288,1735,1761]" pageId="6" pageNumber="7">Thorax</emphasis>
: Ground-colour black, densely grey dusted, with three distinct brown longitudinal stripes widening diffusely to scutellum (
<figureCitation id="13682A64FFC8FF0479DD9AFFFD83F89F" box="[421,513,1771,1797]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="6" pageNumber="7">Fig. 2B</figureCitation>
). Presutural
<emphasis id="B927EAF3FFC8FF047AED9AFFFD38F89E" box="[661,698,1771,1796]" italics="true" pageId="6" pageNumber="7">acr</emphasis>
setae in 2 even rows, 2+4
<emphasis id="B927EAF3FFC8FF047B879AFFFB9BF89F" box="[1023,1049,1771,1797]" italics="true" pageId="6" pageNumber="7">dc</emphasis>
setae. Notopleuron bare, or with individual hairs in upper part. Anepisternum without interspatial setae. Katepisternal setae 1+1. Scutellum brown dusted except grey at apex, without downwardly-directed preapical setulae on upper border of declivities.
</paragraph>
<paragraph id="8BEC36E1FFC8FF0478BF9B42FD55F80F" blockId="6.[151,1437,726,1941]" pageId="6" pageNumber="7">
<emphasis id="B927EAF3FFC8FF0478BF9B42FE8AF8EA" bold="true" box="[199,264,1878,1904]" pageId="6" pageNumber="7">Wing</emphasis>
: Light brownish tinged. Costa without spinules or costal spine. Calypters whitish, lower one with broadly yellow margins. Haltere brown with blackish knob.
</paragraph>
<caption id="DF2C6669FFC9FF0578EF997AFE19FA6B" ID-DOI="http://doi.org/10.5281/zenodo.14812236" ID-Zenodo-Dep="14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1390,1414]" targetBox="[189,1398,181,1356]" targetPageId="7" targetType="figure">
<paragraph id="8BEC36E1FFC9FF0578EF997AFE19FA6B" blockId="7.[151,1437,1390,1522]" pageId="7" pageNumber="8">
<emphasis id="B927EAF3FFC9FF0578EF997AFE97FA1C" bold="true" box="[151,277,1390,1414]" pageId="7" pageNumber="8">FIGURE 2.</emphasis>
<taxonomicName id="4C534D62FFC9FF057966997AFE76FA1C" authorityName="Sorokina" authorityYear="2025" box="[286,500,1390,1414]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gorodkovi" status="sp. nov.">
<emphasis id="B927EAF3FFC9FF057966997AFE76FA1C" box="[286,500,1390,1414]" italics="true" pageId="7" pageNumber="8">Spilogona gorodkovi</emphasis>
</taxonomicName>
<emphasis id="B927EAF3FFC9FF057984997AFDF3FA1C" bold="true" box="[508,625,1390,1414]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A2145788FFC9FF057984997AFDCDFA1C" box="[508,591,1390,1414]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Holotype, male, lateral view.
<emphasis id="B927EAF3FFC9FF057BD6997AFC47FA1C" bold="true" box="[942,965,1390,1414]" pageId="7" pageNumber="8">B.</emphasis>
Holotype, frons and scutum, dorsal view.
<emphasis id="B927EAF3FFC9FF057DFC997AFA1EFA1C" bold="true" box="[1412,1436,1390,1414]" pageId="7" pageNumber="8">C.</emphasis>
Paratype, female, lateral view.
<emphasis id="B927EAF3FFC9FF0579A39986FE76FA30" bold="true" box="[475,500,1426,1450]" pageId="7" pageNumber="8">D.</emphasis>
Terminalia, lateral view, digital photo, paratype.
<emphasis id="B927EAF3FFC9FF057B839986FB93FA30" bold="true" box="[1019,1041,1426,1450]" pageId="7" pageNumber="8">E.</emphasis>
Sternite 5, digital photo, paratype.
<emphasis id="B927EAF3FFC9FF057DF19986FA1EFA30" bold="true" box="[1417,1436,1426,1450]" pageId="7" pageNumber="8">F.</emphasis>
Ovipositor, lateral, dorsal and ventral views, digital photo.
<emphasis id="B927EAF3FFC9FF057A8B99A2FC8CFA54" bold="true" box="[755,782,1462,1486]" pageId="7" pageNumber="8">G.</emphasis>
Terminalia, lateral view, paratype.
<emphasis id="B927EAF3FFC9FF057C0099A2FB10FA54" bold="true" box="[1144,1170,1462,1486]" pageId="7" pageNumber="8">H.</emphasis>
Cercal plate, dorsal view.
<emphasis id="B927EAF3FFC9FF0578EF99CEFF25FA68" bold="true" box="[151,167,1498,1522]" pageId="7" pageNumber="8">I.</emphasis>
Sternite 5. Bare: 1 mm.
</paragraph>
</caption>
<paragraph id="8BEC36E1FFC9FF0578BF9A50FEFDF975" blockId="7.[151,1437,1604,2026]" pageId="7" pageNumber="8">
<emphasis id="B927EAF3FFC9FF0578BF9A50FF7DF9C4" bold="true" box="[199,255,1604,1630]" pageId="7" pageNumber="8">Legs</emphasis>
: Ground colour black, grey dusted. Fore tibia with 1
<emphasis id="B927EAF3FFC9FF057B2E9A51FCE6F9C4" box="[854,868,1605,1630]" italics="true" pageId="7" pageNumber="8">p</emphasis>
seta. Mid femur with a row of hair-like
<emphasis id="B927EAF3FFC9FF057D5F9A51FAC3F9C4" box="[1319,1345,1605,1630]" italics="true" pageId="7" pageNumber="8">pv</emphasis>
and
<emphasis id="B927EAF3FFC9FF057D009A51FF21F918" italics="true" pageId="7" pageNumber="8">avv</emphasis>
setae, which become more numerous and elongate towards base of
<emphasis id="B927EAF3FFC9FF057BDE9A7DFC42F918" box="[934,960,1641,1666]" italics="true" pageId="7" pageNumber="8">pv</emphasis>
surface, with 2
<emphasis id="B927EAF3FFC9FF057C009A7CFB32F918" box="[1144,1200,1640,1666]" italics="true" pageId="7" pageNumber="8">pd–p</emphasis>
preapical setae. Mid tibia with 0–1
<emphasis id="B927EAF3FFC9FF0579449A98FEDAF93C" box="[316,344,1676,1702]" italics="true" pageId="7" pageNumber="8">ad</emphasis>
and 3
<emphasis id="B927EAF3FFC9FF0579DD9A98FE43F93C" box="[421,449,1676,1702]" italics="true" pageId="7" pageNumber="8">pd</emphasis>
setae, without any
<emphasis id="B927EAF3FFC9FF057AE59A99FD2BF93C" box="[669,681,1677,1702]" italics="true" pageId="7" pageNumber="8">v</emphasis>
setae. Hind femur with 6–7 strong
<emphasis id="B927EAF3FFC9FF057C429A99FBD6F93C" box="[1082,1108,1677,1702]" italics="true" pageId="7" pageNumber="8">av</emphasis>
setae in distal third and with hair-like setae at base on
<emphasis id="B927EAF3FFC9FF0579C89AA5FD86F950" box="[432,516,1713,1738]" italics="true" pageId="7" pageNumber="8">av-v-pv</emphasis>
surfaces which are longer on
<emphasis id="B927EAF3FFC9FF057B2A9AA5FCEEF950" box="[850,876,1713,1738]" italics="true" pageId="7" pageNumber="8">pv</emphasis>
surface. Hind tibia with 1–3 short
<emphasis id="B927EAF3FFC9FF057C889AA5FA88F950" box="[1264,1290,1713,1738]" italics="true" pageId="7" pageNumber="8">av</emphasis>
, 4–5
<emphasis id="B927EAF3FFC9FF057D319AA4FAE7F950" box="[1353,1381,1712,1738]" italics="true" pageId="7" pageNumber="8">ad</emphasis>
, 3–4
<emphasis id="B927EAF3FFC9FF0578EF9AC0FF31F974" box="[151,179,1748,1774]" italics="true" pageId="7" pageNumber="8">pd</emphasis>
, without
<emphasis id="B927EAF3FFC9FF0579669AC1FEBAF974" box="[286,312,1749,1774]" italics="true" pageId="7" pageNumber="8">pv</emphasis>
setae.
</paragraph>
<paragraph id="8BEC36E1FFC9FF0578BF9AECFE9CF8E4" blockId="7.[151,1437,1604,2026]" pageId="7" pageNumber="8">
<emphasis id="B927EAF3FFC9FF0578BF9AECFEBEF888" bold="true" box="[199,316,1784,1810]" pageId="7" pageNumber="8">Abdomen</emphasis>
: Black with dense grey dust, conical. Tergites 1+2–4 with black triangular spots separated by a narrow brown median stripe, tergite 5 with a diffuse brown median stripe. Sternite 1 with hairs, sternite 5 somewhat protruding, a little longer then sternite 4, with a deep median notch and a pointed caudal margin, with shining apex (
<figureCitation id="13682A64FFC9FF0578E79B70FE92F8E4" box="[159,272,1892,1919]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="7" pageNumber="8">Figs 2E, I</figureCitation>
).
</paragraph>
<paragraph id="8BEC36E1FFC9FF0578BF9B9CFD1DF870" blockId="7.[151,1437,1604,2026]" pageId="7" pageNumber="8">
<emphasis id="B927EAF3FFC9FF0578BF9B9CFEC8F838" bold="true" box="[199,330,1928,1954]" pageId="7" pageNumber="8">Terminalia</emphasis>
: In lateral view cerci wide, curved, sharply tapered towards the apex; surstylus rounded at apex, with hairs inside, not longer then cerci; epandrium large, hemispherical; hypandrium 2 times shorter then epandrium (
<figureCitation id="13682A64FFC9FF0578E79BC4FEA2F870" box="[159,288,2000,2026]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="7" pageNumber="8">Figs 2D, G</figureCitation>
); cercal plate triangular (
<figureCitation id="13682A64FFC9FF057A409BC4FD10F870" box="[568,658,2000,2026]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="7" pageNumber="8">Fig. 2H</figureCitation>
).
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9C83FD4CFF2B" blockId="8.[151,1436,150,573]" box="[199,718,150,177]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9C83FE9CFF2B" bold="true" box="[199,286,151,177]" pageId="8" pageNumber="9">Female</emphasis>
: body
<quantity id="4CAB9B04FFC6FF0A79149C82FE77FF2B" box="[364,501,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="8" pageNumber="9" unit="mm" value="6.15" valueMax="6.5" valueMin="5.8">5.8–6.5 mm</quantity>
, wing
<quantity id="4CAB9B04FFC6FF0A7A389C82FD48FF2B" box="[576,714,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.8" metricValueMin="5.5" pageId="8" pageNumber="9" unit="mm" value="5.65" valueMax="5.8" valueMin="5.5">5.5–5.8 mm</quantity>
.
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9CAFFD0BFF4F" blockId="8.[151,1436,150,573]" box="[199,649,187,213]" pageId="8" pageNumber="9">Similar to male but differing as follows:</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9CCBFD2CFEFF" blockId="8.[151,1436,150,573]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9CCBFE85FF63" bold="true" box="[199,263,223,249]" pageId="8" pageNumber="9">Head</emphasis>
: Frons, fronto-orbital plate, gena and parafacial grey or brown dusted (
<figureCitation id="13682A64FFC6FF0A7C609CCBFBEDFF63" box="[1048,1135,223,249]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
). Frontal triangle brownish dusted, extending to middle of frons. Width of frontal vitta 3–4 times width of a fronto-orbital plate. Fronto-orbital plate with 2 pairs of orbital setae. Parafacial at level of base of antenna 2 times width of postpedicel. Height of gena about 1/3 of height of eye. Proboscis thickened.
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9D7BFB89FE13" blockId="8.[151,1436,150,573]" box="[199,1035,367,393]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9D7BFEA2FE13" bold="true" box="[199,288,367,393]" pageId="8" pageNumber="9">Thorax</emphasis>
: Scutum more matt. Scutellum with brown spots on lateral angles.
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9D87FB75FE6F" blockId="8.[151,1436,150,573]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9D87FF7DFE37" bold="true" box="[199,255,403,429]" pageId="8" pageNumber="9">Legs</emphasis>
: Mid femur with a row of hair-like
<emphasis id="B927EAF3FFC6FF0A7AF69D87FD2AFE36" box="[654,680,403,428]" italics="true" pageId="8" pageNumber="9">pv</emphasis>
which become more elongate towards base; only several short
<emphasis id="B927EAF3FFC6FF0A7D149D87FA1EFE36" box="[1388,1436,403,428]" italics="true" pageId="8" pageNumber="9">av-v</emphasis>
hairs at base. Mid tibia with 0
<emphasis id="B927EAF3FFC6FF0A79969DA3FD88FE4B" box="[494,522,439,465]" italics="true" pageId="8" pageNumber="9">ad</emphasis>
and 2–3
<emphasis id="B927EAF3FFC6FF0A7A0A9DA3FD0CFE4B" box="[626,654,439,465]" italics="true" pageId="8" pageNumber="9">pd</emphasis>
. Hind femur with 4–5 strong
<emphasis id="B927EAF3FFC6FF0A7BA59DA3FC75FE4A" box="[989,1015,439,464]" italics="true" pageId="8" pageNumber="9">av</emphasis>
setae in distal third; hair-like setae at base short and sparse on
<emphasis id="B927EAF3FFC6FF0A79D69DCFFE5CFE6E" box="[430,478,475,500]" italics="true" pageId="8" pageNumber="9">av-v</emphasis>
surface, longer on
<emphasis id="B927EAF3FFC6FF0A7ACE9DCFFD52FE6E" box="[694,720,475,500]" italics="true" pageId="8" pageNumber="9">pv</emphasis>
surface. Hind tibia with 3–4
<emphasis id="B927EAF3FFC6FF0A7C609DCFFBB6FE6F" box="[1048,1076,475,501]" italics="true" pageId="8" pageNumber="9">ad</emphasis>
, 2–3
<emphasis id="B927EAF3FFC6FF0A7C0B9DCFFB32FE6E" box="[1139,1200,475,501]" italics="true" pageId="8" pageNumber="9">pd pv</emphasis>
setae.
</paragraph>
<paragraph id="8BEC36E1FFC6FF0A78BF9DEAFCFDFD83" blockId="8.[151,1436,150,573]" box="[199,895,510,537]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9DEAFEBEFD82" bold="true" box="[199,316,510,536]" pageId="8" pageNumber="9">Abdomen</emphasis>
: Postgenital plate with spinules and hairs (
<figureCitation id="13682A64FFC6FF0A7B649DEBFCF0FD83" box="[796,882,511,537]" captionStart="FIGURE 2" captionStartId="7.[151,250,1390,1414]" captionTargetBox="[189,1398,181,1356]" captionTargetId="figure-17@7.[189,1398,181,1366]" captionTargetPageId="7" captionText="FIGURE 2. Spilogona gorodkovi sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812236" httpUri="https://zenodo.org/record/14812236/files/figure.png" pageId="8" pageNumber="9">Fig. 2F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFC6FF0A78BF9E36FC15FDA7" box="[199,919,546,573]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BEC36E1FFC6FF0A78BF9E36FC15FDA7" blockId="8.[151,1436,150,573]" box="[199,919,546,573]" pageId="8" pageNumber="9">
<emphasis id="B927EAF3FFC6FF0A78BF9E36FEE3FDA6" bold="true" box="[199,353,546,572]" pageId="8" pageNumber="9">Distribution.</emphasis>
PALAEARCTIC: Arctic
<collectingCountry id="F3447671FFC6FF0A7AF89E37FD49FDA7" box="[640,715,547,573]" name="Russia" pageId="8" pageNumber="9">Russia</collectingCountry>
(Wrangel Island).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="F54367297B96973A66FAC444095FEBB5">New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species</mods:title>
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<treatment id="03FA87F7FFD8FF1478EF9C83FABCFAB9" ID-DOI="http://doi.org/10.5281/zenodo.14812287" ID-Zenodo-Dep="14812287" LSID="urn:lsid:plazi:treatment:03FA87F7FFD8FF1478EF9C83FABCFAB9" httpUri="http://treatment.plazi.org/id/03FA87F7FFD8FF1478EF9C83FABCFAB9" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C349656AFFD8FF1478EF9C83FDADFF2B" box="[151,559,151,178]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BEC36E1FFD8FF1478EF9C83FDADFF2B" blockId="22.[151,559,151,214]" box="[151,559,151,178]" pageId="22" pageNumber="23">
<heading id="D0A4818DFFD8FF1478EF9C83FDADFF2B" bold="true" box="[151,559,151,178]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="4C534D62FFD8FF1478EF9C83FDADFF2B" ID-CoL="4Z42H" authority="Hennig, 1959" authorityName="Hennig" authorityYear="1959" box="[151,559,151,178]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF1478EF9C83FDADFF2B" bold="true" box="[151,559,151,178]" pageId="22" pageNumber="23">
<emphasis id="B927EAF3FFD8FF1478EF9C83FE01FF28" bold="true" box="[151,387,151,178]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
Hennig, 1959
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFD8FF1478EF9CAFFBBFFEAC" pageId="22" pageNumber="23" type="description">
<paragraph id="8BEC36E1FFD8FF1478EF9CAFFE91FF4C" blockId="22.[151,559,151,214]" box="[151,275,187,214]" pageId="22" pageNumber="23">
<figureCitation id="13682A64FFD8FF1478EF9CAFFE91FF4C" box="[151,275,187,214]" captionStart="FIGURE 6" captionStartId="19.[151,250,1228,1252]" captionTargetBox="[201,1386,345,1195]" captionTargetId="figure-103@19.[201,1386,345,1204]" captionTargetPageId="19" captionText="FIGURE 6. Spilogona ssp. A. Spilogona humeralis, sternite 5. B. Spilogona pusilla, sternite 5. C. Spilogona subfuscisquama, sternite 5. D. Spilogona malaisei, terminalia, lateral view. E, F. Spilogona wrangeli. E. Terminalia, lateral view. F. Cercal plate, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14812249" httpUri="https://zenodo.org/record/14812249/files/figure.png" pageId="22" pageNumber="23">Figs 6 E, F</figureCitation>
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478EF9D14FBBFFEAC" blockId="22.[151,1436,255,433]" pageId="22" pageNumber="23">
<taxonomicName id="4C534D62FFD8FF1478EF9D14FDA4FE8D" authority="Hennig, 1959: 360" authorityName="Hennig" authorityPageNumber="360" authorityYear="1959" box="[151,550,255,280]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF1478EF9D14FEDDFE82" box="[151,351,256,280]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
Hennig, 1959: 360
</taxonomicName>
. Type-locality: “O-w Wrangelja, werch. r. Klen k SSW ot pika Berri, 19.–21.VIII.
<quantity id="4CAB9B04FFD8FF147D109D14FA1AFE82" box="[1384,1432,256,280]" metricMagnitude="-2" metricUnit="kg" metricValue="3.8" pageId="22" pageNumber="23" unit="g" value="38.0">38 g</quantity>
. Gekker” [
<collectingCountry id="F3447671FFD8FF1479549D0AFE04FEAC" box="[300,390,286,310]" name="Russia" pageId="22" pageNumber="23">RUSSIA</collectingCountry>
, Far East, Wrangelya I., upper Klen river SSW of Mt. Berry] [seen].
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFD8FF1478EF9D29FC70FE2B" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="8BEC36E1FFD8FF1478EF9D29FD56FEEE" blockId="22.[151,1436,255,433]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB4311CFFFD8FF1478EF9D29FD56FEEE" pageId="22" pageNumber="23">
<treatmentCitation id="0AF210F0FFD8FF1478EF9D29FDBCFECF" author="Huckett, H. C." box="[151,574,317,341]" page="214" pageId="22" pageNumber="23" year="1965">
<taxonomicName id="4C534D62FFD8FF1478EF9D29FDBCFECF" ID-CoL="8962V" authority="Huckett, 1965: 214" authorityName="Huckett" authorityPageNumber="214" authorityYear="1965" box="[151,574,317,341]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF1478EF9D29FEE2FECF" box="[151,352,317,341]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
<materialsCitation id="3B3B3CBCFFD8FF1479149D29FDBCFECF" box="[364,574,317,341]" collectingDate="1953-07-11" collectorName="J. S. Waterhouse" country="Canada" location="Herschel Island" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Yukon">
<bibRefCitation id="EFC24B10FFD8FF1479149D29FDBCFECF" author="Huckett, H. C." box="[364,574,317,341]" pageId="22" pageNumber="23" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 214</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. Type-locality: “
<location id="8E8C603AFFD8FF147A979D29FC1AFECF" LSID="urn:lsid:plazi:treatment:03FA87F7FFD8FF1478EF9C83FABCFAB9:8E8C603AFFD8FF147A979D29FC1AFECF" box="[751,920,317,341]" country="Canada" name="Herschel Island" pageId="22" pageNumber="23" stateProvince="Yukon">Herschel Island</location>
,
<collectingRegion id="4997F803FFD8FF147BDF9D29FBD2FECF" box="[935,1104,317,341]" country="Canada" name="Yukon" pageId="22" pageNumber="23">Yukon Territory</collectingRegion>
, 7.11.53 [i.e.
<date id="FFED1021FFD8FF147C929D29FAE2FECF" box="[1258,1376,317,341]" pageId="22" pageNumber="23" value="1953-07-11">
<collectingDate id="EFA9E9C9FFD8FF147C929D29FAE2FECF" box="[1258,1376,317,341]" pageId="22" pageNumber="23" value="1953-07-11">11.vii.1953</collectingDate>
</date>
],
<collectorName id="26A65337FFD8FF147D009D29FEC0FEEE" pageId="22" pageNumber="23">J.S. Waterhouse</collectorName>
)” [
<collectingCountry id="F3447671FFD8FF1479269D48FE4FFEEE" box="[350,461,348,372]" name="Canada" pageId="22" pageNumber="23">CANADA</collectingCountry>
,
<collectingRegion id="4997F803FFD8FF1479A19D48FD9CFEEE" box="[473,542,348,372]" country="Canada" name="Yukon" pageId="22" pageNumber="23">Yukon</collectingRegion>
] [seen].
<emphasis id="B927EAF3FFD8FF147A0C9D4FFD56FEEE" bold="true" box="[628,724,347,372]" pageId="22" pageNumber="23">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478EF9D6EFB54FE08" blockId="22.[151,1436,255,433]" box="[151,1238,378,402]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB4311CFFFD8FF1478EF9D6EFB54FE08" box="[151,1238,378,402]" pageId="22" pageNumber="23">
<taxonomicName id="4C534D62FFD8FF1478EF9D6EFD92FE08" ID-CoL="4Z3WJ" authority="(Ringdahl, 1920)" authorityName="F" baseAuthorityName="Ringdahl" baseAuthorityYear="1920" box="[151,528,378,402]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B927EAF3FFD8FF1478EF9D6EFED8FE08" box="[151,346,378,402]" italics="true" pageId="22" pageNumber="23">Spilogona malaisei</emphasis>
(Ringdahl, 1920)
</taxonomicName>
;
<treatmentCitation id="0AF210F0FFD8FF147A639D6EFCE3FE08" author="Sorokina, V. S. &amp; Khruleva, O. A." box="[539,865,378,402]" page="556" pageId="22" pageNumber="23" year="2012">
<bibRefCitation id="EFC24B10FFD8FF147A639D6EFCE3FE08" author="Sorokina, V. S. &amp; Khruleva, O. A." box="[539,865,378,402]" pageId="22" pageNumber="23" pagination="553 - 564" refId="ref19632" refString="Sorokina, V. S. &amp; Khruleva, O. A. (2012) Details of species composition and distribution of house-flies (Diptera, Muscidae) of the Wrangel Island, Russia. Euroasian Entomological Journal, 11 (6), 553-564. [in Russian, with English summary and species list]" type="journal article" year="2012">Sorokina &amp; Khruleva 2012: 556</bibRefCitation>
</treatmentCitation>
(Wrangel Island). Misidentification.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478EF9D8DFC70FE2B" blockId="22.[151,1436,255,433]" box="[151,1010,409,433]" pageId="22" pageNumber="23">
<taxonomicName id="4C534D62FFD8FF1478EF9D8DFE70FE2B" ID-CoL="4Z42H" authority="Hennig, 1959" authorityName="Hennig" authorityYear="1959" box="[151,498,409,433]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF1478EF9D8DFEDCFE2B" box="[151,350,409,433]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
Hennig, 1959
</taxonomicName>
; Sorokina &amp; Thidrikh, 2021: 229 (Wrangel Island).
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFD8FF1478EF9DCEFBCFFB8A" pageId="22" pageNumber="23" type="description">
<paragraph id="8BEC36E1FFD8FF1478EF9DCEFC7CFC1A" blockId="22.[151,1437,474,1040]" pageId="22" pageNumber="23">
<emphasis id="B927EAF3FFD8FF1478EF9DCEFF58FE6E" bold="true" box="[151,218,474,500]" pageId="22" pageNumber="23">Notes</emphasis>
. The
<typeStatus id="54E88843FFD8FF1479659DCEFE08FE6E" box="[285,394,474,500]" pageId="22" pageNumber="23" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="4C534D62FFD8FF1479CA9DCEFD09FE6E" authorityName="Huckett" authorityYear="1965" box="[434,651,474,500]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF1479CA9DCEFD09FE6E" box="[434,651,474,500]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
</taxonomicName>
(in Canadian National Collection) and
<taxonomicName id="4C534D62FFD8FF147C349DCEFAA8FE6E" authorityName="Hennig" authorityYear="1959" box="[1100,1322,474,500]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF147C349DCEFAA8FE6E" box="[1100,1322,474,500]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
</taxonomicName>
(in ZISP) were examined. Both species had been described from arctic Islands, Wrangel Island and Herschel Island, and had not since been found in other localities.
<taxonomicName id="4C534D62FFD8FF147A1D9E36FCBDFDA6" authorityName="Huckett" authorityYear="1965" box="[613,831,546,572]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF147A1D9E36FCBDFDA6" box="[613,831,546,572]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54E88843FFD8FF147C269E36FB42FDA6" box="[1118,1216,546,572]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
and one additional male. A series of
<taxonomicName id="4C534D62FFD8FF1479239E52FDBAFDFA" authorityName="Hennig" authorityYear="1959" box="[347,568,582,608]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF1479239E52FDBAFDFA" box="[347,568,582,608]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
</taxonomicName>
has been found recently on Wrangel Island (Sorokina &amp; Thidrikh, 2021). The descriptions and the male terminalia are the same for both species. The exception was a
<emphasis id="B927EAF3FFD8FF147CEC9E7FFB22FD1E" box="[1172,1184,619,644]" italics="true" pageId="22" pageNumber="23">v</emphasis>
seta on the mid tibia, which Huckett described for
<taxonomicName id="4C534D62FFD8FF1479A09E9AFD2CFD32" authorityName="Huckett" authorityYear="1965" box="[472,686,654,680]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF1479A09E9AFD2CFD32" box="[472,686,654,680]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
</taxonomicName>
;
<taxonomicName id="4C534D62FFD8FF147AC49E9AFC15FD32" authorityName="Hennig" authorityYear="1959" box="[700,919,654,680]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF147AC49E9AFC15FD32" box="[700,919,654,680]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
</taxonomicName>
is without a
<emphasis id="B927EAF3FFD8FF147C5B9E9BFBADFD32" box="[1059,1071,655,680]" italics="true" pageId="22" pageNumber="23">v</emphasis>
seta on the mid tibia. I examined the
<typeStatus id="54E88843FFD8FF1478BB9EA6FEA7FD56" box="[195,293,690,716]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C534D62FFD8FF1479289EA6FDA8FD56" authorityName="Huckett" authorityYear="1965" box="[336,554,690,716]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF1479289EA6FDA8FD56" box="[336,554,690,716]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
</taxonomicName>
. The specimen is a little crumpled, legs pressed against the thorax, one mid tibia in glue; with difficulty I was able to see that the mid tibia is in fact without
<emphasis id="B927EAF3FFD8FF147C669EC3FBA8FD6A" box="[1054,1066,727,752]" italics="true" pageId="22" pageNumber="23">v</emphasis>
setae. Males of these species are small in size (
<quantity id="4CAB9B04FFD8FF1479499EEEFE25FC8E" box="[305,423,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="22" pageNumber="23" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 mm</quantity>
), facial edge often a little projecting beyond level of profrons, but sometimes not, subovate abdomen, blackish thorax without stripes, anepisternum without interspatial setae, scutellum with downwardly-directed preapical setulae on upper border of declivities, costa with spinules, mid tibia without
<emphasis id="B927EAF3FFD8FF147CBE9F57FB50FCC6" box="[1222,1234,835,860]" italics="true" pageId="22" pageNumber="23">v</emphasis>
setae, hind femur with weak
<emphasis id="B927EAF3FFD8FF14796A9F73FEAEFC1A" box="[274,300,871,896]" italics="true" pageId="22" pageNumber="23">pv</emphasis>
setae, and abdomen dark with indistinct black expansive marks.
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478BF9F9EFEB5FC53" blockId="22.[151,1437,474,1040]" pageId="22" pageNumber="23">
I therefore conclude that
<taxonomicName id="4C534D62FFD8FF1479929F9EFCECFC3F" authority="Huckett, 1965" authorityName="Huckett" authorityYear="1965" box="[490,878,906,933]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF1479929F9EFD46FC3E" box="[490,708,906,932]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
<bibRefCitation id="EFC24B10FFD8FF147AB59F9EFCECFC3F" author="Huckett, H. C." box="[717,878,906,933]" pageId="22" pageNumber="23" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965</bibRefCitation>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="4C534D62FFD8FF147CC69F9EFEB1FC53" authority="Hennig, 1959" authorityName="Hennig" authorityYear="1959" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF147CC69F9EFA1FFC3E" box="[1214,1437,906,932]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
Hennig, 1959
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478BF9FC6FBCFFB8A" blockId="22.[151,1437,474,1040]" pageId="22" pageNumber="23">
The female of both
<taxonomicName id="4C534D62FFD8FF1479D59FC6FD05FC76" authorityName="Huckett" authorityYear="1965" box="[429,647,978,1004]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coactilis">
<emphasis id="B927EAF3FFD8FF1479D59FC6FD05FC76" box="[429,647,978,1004]" italics="true" pageId="22" pageNumber="23">Spilogona coactilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFD8FF147AB99FC6FC22FC76" authorityName="Hennig" authorityYear="1959" box="[705,928,978,1004]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF147AB99FC6FC22FC76" box="[705,928,978,1004]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
</taxonomicName>
has been not described. It is very similar to
<taxonomicName id="4C534D62FFD8FF1478EF9FE2FEEDFB8A" authorityName="F" baseAuthorityName="Ringdahl" baseAuthorityYear="1920" box="[151,367,1014,1040]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B927EAF3FFD8FF1478EF9FE2FEEDFB8A" box="[151,367,1014,1040]" italics="true" pageId="22" pageNumber="23">Spilogona malaisei</emphasis>
</taxonomicName>
, like the male, and both can be separated as in following couplet:
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFD8FF1478EF982FFA1EFBF7" pageId="22" pageNumber="23" type="key">
<keyStep id="30A72E44FFD8FF1478EF982FFA1EFBF7" pageId="22" pageNumber="23">
<key id="FFC261D0FFD8FF1478EF982FFA1EFBF7" pageId="22" pageNumber="23">
<paragraph id="8BEC36E1FFD8FF1478EF982FFA1EFBCA" blockId="22.[151,1436,1083,1134]" box="[151,1436,1083,1105]" pageId="22" pageNumber="23">
<keyLead id="30A295D4FFD8FF1478EF982FFA1EFBCA" box="[151,1436,1083,1105]" pageId="22" pageNumber="23">
1 Costa without spinules, with short setae............................................................
<taxonomicName id="4C534D62FFD8FF147D4B982FFA1EFBCA" ID-CoL="4Z3WJ" authorityName="F" baseAuthorityName="Ringdahl" baseAuthorityYear="1920" box="[1331,1436,1083,1104]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B927EAF3FFD8FF147D4B982FFA1EFBCA" box="[1331,1436,1083,1104]" italics="true" pageId="22" pageNumber="23">S. malaisei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478EF984CFA1EFBF7" blockId="22.[151,1436,1083,1134]" box="[151,1436,1112,1134]" pageId="22" pageNumber="23">
<keyLead id="30A295D4FFD8FF1478EF984CFA1EFBF7" box="[151,1436,1112,1134]" pageId="22" pageNumber="23">
- Costa with spinules...........................................................................
<taxonomicName id="4C534D62FFD8FF147D57984CFA1EFBF7" ID-CoL="4Z42H" authorityName="Hennig" authorityYear="1959" box="[1327,1436,1112,1133]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF147D57984CFA1EFBF7" box="[1327,1436,1112,1133]" italics="true" pageId="22" pageNumber="23">S. wrangeli</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
<subSubSection id="C349656AFFD8FF1478BF9888FB5BFB40" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BEC36E1FFD8FF1478BF9888FA83FB2C" blockId="22.[151,1436,1180,1315]" box="[199,1281,1180,1207]" pageId="22" pageNumber="23">
<emphasis id="B927EAF3FFD8FF1478BF9888FEE3FB2C" bold="true" box="[199,353,1180,1206]" pageId="22" pageNumber="23">Distribution.</emphasis>
PALAEARCTIC:
<collectingCountry id="F3447671FFD8FF147A4D9888FD02FB2C" box="[565,640,1180,1206]" name="Russia" pageId="22" pageNumber="23">Russia</collectingCountry>
(Wrangel Island). NEARCTIC: Yukon (Herschel Island).
</paragraph>
<paragraph id="8BEC36E1FFD8FF1478BF98D4FB5BFB40" blockId="22.[151,1436,1180,1315]" box="[199,1241,1216,1243]" pageId="22" pageNumber="23">
<emphasis id="B927EAF3FFD8FF1478BF98D4FEEBFB41" bold="true" box="[199,361,1216,1243]" pageId="22" pageNumber="23">DNA barcode</emphasis>
. BOLD BIN: BOLD:AED9512. See
<tableCitation id="C6D1035AFFD8FF147B7498D4FCDDFB41" box="[780,863,1216,1243]" captionStart="TABLE 1" captionStartId="2.[152,245,657,681]" captionText="TABLE 1. List of Russian Spilogona species with barcode index numbers (BIN) and GenBank accession numbers." pageId="22" pageNumber="23">Table 1</tableCitation>
for GenBank accession numbers.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFD8FF1478BF98F1FABCFAB9" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BEC36E1FFD8FF1478BF98F1FABCFAB9" blockId="22.[151,1436,1180,1315]" pageId="22" pageNumber="23">
<emphasis id="B927EAF3FFD8FF1478BF98F1FEB9FB65" bold="true" box="[199,315,1253,1279]" pageId="22" pageNumber="23">Remarks.</emphasis>
<taxonomicName id="4C534D62FFD8FF14793C98F0FD9CFB64" authorityName="Hennig" authorityYear="1959" box="[324,542,1252,1278]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="wrangeli">
<emphasis id="B927EAF3FFD8FF14793C98F0FD9CFB64" box="[324,542,1252,1278]" italics="true" pageId="22" pageNumber="23">Spilogona wrangeli</emphasis>
</taxonomicName>
is morphologically very similar to
<taxonomicName id="4C534D62FFD8FF147BC998F0FB09FB64" authorityName="F" baseAuthorityName="Ringdahl" baseAuthorityYear="1920" box="[945,1163,1252,1278]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B927EAF3FFD8FF147BC998F0FB09FB64" box="[945,1163,1252,1278]" italics="true" pageId="22" pageNumber="23">Spilogona malaisei</emphasis>
</taxonomicName>
, especially females. But based on DNA barcodes, these species are genetically different; the p-distance between them is 2.04–2.15%.
</paragraph>
</subSubSection>
</treatment>
</document>

351
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<subSubSection id="C349656AFFDAFF1678EF9FFFFDC5FB9F" box="[151,583,1003,1030]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BEC36E1FFDAFF1678EF9FFFFDC5FB9F" blockId="20.[151,583,1003,1066]" box="[151,583,1003,1030]" pageId="20" pageNumber="21">
<heading id="D0A4818DFFDAFF1678EF9FFFFDC5FB9F" bold="true" box="[151,583,1003,1030]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="4C534D62FFDAFF1678EF9FFFFDC5FB9F" ID-CoL="4Z3ZP" authority="(Malloch, 1920)" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[151,583,1003,1030]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF1678EF9FFFFDC5FB9F" bold="true" box="[151,583,1003,1030]" pageId="20" pageNumber="21">
<emphasis id="B927EAF3FFDAFF1678EF9FFFFEFEFB9C" bold="true" box="[151,380,1003,1030]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
(Malloch, 1920)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C349656AFFDAFF1678EF9804FA1EF81E" pageId="20" pageNumber="21" type="description">
<paragraph id="8BEC36E1FFDAFF1678EF9804FE9BFBB0" blockId="20.[151,583,1003,1066]" box="[151,281,1040,1066]" pageId="20" pageNumber="21">
<figureCitation id="13682A64FFDAFF1678EF9804FE9BFBB0" box="[151,281,1040,1066]" captionStart="FIGURE 7" captionStartId="21.[151,250,1845,1869]" captionTargetBox="[153,1434,525,1814]" captionTargetId="figure-149@21.[153,1434,525,1822]" captionTargetPageId="21" captionText="FIGURE 7. Spilogona tendipes. A. Female, lateral view. B. Female, lateral view. C. Terminalia, lateral view, digital photo. D. Sternite 5, digital photo. E. Ovipositor, lateral, dorsal and ventral views, digital photo. F. Terminalia, lateral view. H. Cercal plate and surstylus, dorsal view. H. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812253" httpUri="https://zenodo.org/record/14812253/files/figure.png" pageId="20" pageNumber="21">Figs 7 A–H</figureCitation>
</paragraph>
<paragraph id="8BEC36E1FFDAFF1678EF984CFB47FA98" blockId="20.[151,1437,1112,1714]" pageId="20" pageNumber="21">
<emphasis id="B927EAF3FFDAFF1678EF984CFF58FBE8" bold="true" box="[151,218,1112,1138]" pageId="20" pageNumber="21">Notes</emphasis>
. In
<bibRefCitation id="EFC24B10FFDAFF167973984CFDDEFBE8" author="Sorokina, V. S. &amp; Michelsen, V." box="[267,604,1112,1138]" pageId="20" pageNumber="21" pagination="512 - 520" refId="ref19692" refString="Sorokina, V. S. &amp; Michelsen, V. (2014) Contributions to the taxonomy and faunistics of some arctic species of Spilogona Schnabl (Diptera: Muscidae). Zootaxa, 3814 (4), 512-520. https://doi.org/10.11646/zootaxa.3814.4.4" type="journal article" year="2014">Sorokina &amp; Michelsen (2014)</bibRefCitation>
<taxonomicName id="4C534D62FFDAFF167A1D984CFCBFFBE8" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[613,829,1112,1138]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF167A1D984CFCBFFBE8" box="[613,829,1112,1138]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
was recorded for the first time from Wrangel Island and from NE
<collectingCountry id="F3447671FFDAFF16794A9868FE25FB0C" box="[306,423,1148,1174]" name="Greenland" pageId="20" pageNumber="21">Greenland</collectingCountry>
from Zackenberg, all based on females. However, a few doubts were expressed about the assignment of these females to
<taxonomicName id="4C534D62FFDAFF16798898B4FD46FB20" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[496,708,1184,1210]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF16798898B4FD46FB20" box="[496,708,1184,1210]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
: (1) the type locality lies in the warmer maritime climatic zone on the south coast of Alaska; (2) Huckett (1932, 1965) emphasized the presence of abundant setulae on the declivities of the scutellum in the male
<typeStatus id="54E88843FFDAFF1679AE98FCFDB9FA98" box="[470,571,1256,1282]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
, but these declivities are practically bare in these females.
</paragraph>
<paragraph id="8BEC36E1FFDAFF1678BF9918FE45FA2C" blockId="20.[151,1437,1112,1714]" pageId="20" pageNumber="21">
Later, male and female specimens similar to
<taxonomicName id="4C534D62FFDAFF167ACC9918FC0AFABC" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[692,904,1292,1318]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF167ACC9918FC0AFABC" box="[692,904,1292,1318]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
from Wrangel Island were found in Gorodkov’s collections in the ZISP, and additional material of the same species has been collected on Wrangel Island. Further material similar to
<taxonomicName id="4C534D62FFDAFF1679109940FDBFFAF4" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[360,573,1364,1390]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF1679109940FDBFFAF4" box="[360,573,1364,1390]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
from the
<collectingRegion id="4997F803FFDAFF167AD09940FCDCFAF4" box="[680,862,1364,1390]" country="Russia" name="Magadan" pageId="20" pageNumber="21">Magadan region</collectingRegion>
and
<collectingRegion id="4997F803FFDAFF167BE89940FC4AFAF4" box="[912,968,1364,1390]" country="Russia" name="Altay" pageId="20" pageNumber="21">Altai</collectingRegion>
Mts of
<collectingCountry id="F3447671FFDAFF167C659940FBEAFAF4" box="[1053,1128,1364,1390]" name="Russia" pageId="20" pageNumber="21">Russia</collectingCountry>
was also collected recently. The good condition of these flies and the presence of males made it possible to study more carefully all the characters of the collected specimens.
</paragraph>
<paragraph id="8BEC36E1FFDAFF1678BF99D4FE3FF914" blockId="20.[151,1437,1112,1714]" pageId="20" pageNumber="21">
Specimens from the
<collectingRegion id="4997F803FFDAFF1679CD99D4FDECFA40" box="[437,622,1472,1498]" country="Russia" name="Magadan" pageId="20" pageNumber="21">Magadan region</collectingRegion>
and
<collectingRegion id="4997F803FFDAFF167ADF99D4FD5DFA40" box="[679,735,1472,1498]" country="Russia" name="Altay" pageId="20" pageNumber="21">Altai</collectingRegion>
Mts agreed fully with the description of
<taxonomicName id="4C534D62FFDAFF167CC499D4FA16FA40" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[1212,1428,1472,1498]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF167CC499D4FA16FA40" box="[1212,1428,1472,1498]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
; these localities are not further north than the
<typeStatus id="54E88843FFDAFF167AEF99F0FD45FA64" box="[663,711,1508,1534]" pageId="20" pageNumber="21">type</typeStatus>
locality of
<taxonomicName id="4C534D62FFDAFF167B3399F0FBA3FA64" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[843,1057,1508,1534]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF167B3399F0FBA3FA64" box="[843,1057,1508,1534]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
. Specimens from Wrangel Island are very similar to
<taxonomicName id="4C534D62FFDAFF16791D9A1CFDB8F9B8" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[357,570,1544,1570]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF16791D9A1CFDB8F9B8" box="[357,570,1544,1570]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
, but there are not only some morphological differences between these species, including the genitalia, but they are also genetically distinct. The specimens from Wrangel Island, including the females listed in
<bibRefCitation id="EFC24B10FFDAFF16792E9A44FD2BF9F0" author="Sorokina, V. S. &amp; Michelsen, V." box="[342,681,1616,1642]" pageId="20" pageNumber="21" pagination="512 - 520" refId="ref19692" refString="Sorokina, V. S. &amp; Michelsen, V. (2014) Contributions to the taxonomy and faunistics of some arctic species of Spilogona Schnabl (Diptera: Muscidae). Zootaxa, 3814 (4), 512-520. https://doi.org/10.11646/zootaxa.3814.4.4" type="journal article" year="2014">Sorokina &amp; Michelsen (2014)</bibRefCitation>
, thus proved to be a species new for science,
<taxonomicName id="4C534D62FFDAFF167CD19A44FA1EF9F0" authorityName="Sorokina" authorityYear="2025" box="[1193,1436,1616,1642]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latizygoma" status="sp. nov.">
<emphasis id="B927EAF3FFDAFF167CD19A44FA1EF9F0" box="[1193,1436,1616,1642]" italics="true" pageId="20" pageNumber="21">Spilogona latizygoma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2145788FFDAFF1678EF9A60FF73F914" box="[151,241,1652,1678]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B927EAF3FFDAFF1678EF9A60FF6EF914" bold="true" box="[151,236,1652,1678]" pageId="20" pageNumber="21">sp. nov</emphasis>
.
</taxonomicNameLabel>
, described above.
</paragraph>
<paragraph id="8BEC36E1FFDAFF1678BF9A8CFB72F928" blockId="20.[151,1437,1112,1714]" box="[199,1264,1688,1714]" pageId="20" pageNumber="21">
<taxonomicName id="4C534D62FFDAFF1678BF9A8CFE1FF928" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[199,413,1688,1714]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF1678BF9A8CFE1FF928" box="[199,413,1688,1714]" italics="true" pageId="20" pageNumber="21">Spilogona tendipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D62FFDAFF1679AB9A8CFD45F928" authorityName="Sorokina" authorityYear="2025" box="[467,711,1688,1714]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latizygoma" status="sp. nov.">
<emphasis id="B927EAF3FFDAFF1679AB9A8CFD45F928" box="[467,711,1688,1714]" italics="true" pageId="20" pageNumber="21">Spilogona latizygoma</emphasis>
</taxonomicName>
<emphasis id="B927EAF3FFDAFF167AB69A8CFCA5F928" bold="true" box="[718,807,1688,1714]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A2145788FFDAFF167AB69A8CFCA5F928" box="[718,807,1688,1714]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated as in following couple:
</paragraph>
<paragraph id="8BEC36E1FFDAFF1678EF9AC9FA1EF8B7" blockId="20.[151,1436,1757,1925]" pageId="20" pageNumber="21">
<emphasis id="B927EAF3FFDAFF1678EF9AC9FF1DF968" box="[151,159,1757,1778]" italics="true" pageId="20" pageNumber="21">-</emphasis>
Mid tibia without
<emphasis id="B927EAF3FFDAFF1679EA9AC9FE1FF968" box="[402,413,1757,1778]" italics="true" pageId="20" pageNumber="21">v</emphasis>
setae; hind femur with distinct long
<emphasis id="B927EAF3FFDAFF167B7B9AC9FC98F968" box="[771,794,1757,1778]" italics="true" pageId="20" pageNumber="21">pv</emphasis>
in basal half; all femora with numerous hair-like
<emphasis id="B927EAF3FFDAFF167C859AC9FA96F968" box="[1277,1300,1757,1778]" italics="true" pageId="20" pageNumber="21">av</emphasis>
setae in basal half; scutum with long ground-setulae; sternites 3–5 with long setae (
<figureCitation id="13682A64FFDAFF167BF89AEEFC49F895" box="[896,971,1786,1808]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="20" pageNumber="21">Fig. 3A</figureCitation>
); eye with inconspicuous hairs; male terminalia as in
<figureCitation id="13682A64FFDAFF16796D9B03FEFCF8B7" box="[277,382,1815,1837]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="20" pageNumber="21">Fig. 3G, H</figureCitation>
; female hypoproct with weak individual spinules (
<figureCitation id="13682A64FFDAFF167B1B9B03FC28F8B7" box="[867,938,1815,1837]" captionStart="FIGURE 3" captionStartId="9.[151,250,1560,1584]" captionTargetBox="[166,1422,181,1525]" captionTargetId="figure-17@9.[166,1422,181,1536]" captionTargetPageId="9" captionText="FIGURE 3. Spilogona latizygoma sp. nov. A. Holotype, male, lateral view. B. Holotype, frons and scutum, dorsal view. C. Paratype, female, lateral view. D. Terminalia, lateral view, digital photo, paratype. E. Sternite 5, digital photo, paratype. F. Ovipositor, lateral, dorsal and ventral views, digital photo. G. Terminalia, lateral view, paratype. H. Cercal plate, dorsal view. I. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812240" httpUri="https://zenodo.org/record/14812240/files/figure.png" pageId="20" pageNumber="21">Fig. 3F</figureCitation>
)......................
<taxonomicName id="4C534D62FFDAFF167CB09B0CFACBF8B6" authorityName="Sorokina" authorityYear="2025" box="[1224,1353,1815,1837]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latizygoma" status="sp. nov.">
<emphasis id="B927EAF3FFDAFF167CB09B0CFB56F8B7" box="[1224,1236,1816,1837]" italics="true" pageId="20" pageNumber="21">S</emphasis>
.
<emphasis id="B927EAF3FFDAFF167C989B03FACBF8B6" box="[1248,1353,1815,1836]" italics="true" pageId="20" pageNumber="21">latizygoma</emphasis>
</taxonomicName>
<emphasis id="B927EAF3FFDAFF167D289B0CFA1EF8B7" bold="true" box="[1360,1436,1816,1837]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A2145788FFDAFF167D289B0CFA1EF8B7" box="[1360,1436,1816,1837]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEC36E1FFDAFF1678EF9B21FA1EF81E" blockId="20.[151,1436,1757,1925]" pageId="20" pageNumber="21">
<emphasis id="B927EAF3FFDAFF1678EF9B21FF1DF8D0" box="[151,159,1845,1866]" italics="true" pageId="20" pageNumber="21">-</emphasis>
Mid tibia with a short
<emphasis id="B927EAF3FFDAFF1679C19B21FE46F8D0" box="[441,452,1845,1866]" italics="true" pageId="20" pageNumber="21">v</emphasis>
seta; hind femur without distinct long
<emphasis id="B927EAF3FFDAFF167B439B21FCD0F8D0" box="[827,850,1845,1866]" italics="true" pageId="20" pageNumber="21">pv</emphasis>
in basal half; all femora without numerous hair-like
<emphasis id="B927EAF3FFDAFF167D379B21FAE4F8D0" box="[1359,1382,1845,1866]" italics="true" pageId="20" pageNumber="21">av</emphasis>
setae in basal half; scutum with short ground-setulae; sternites 3–5 with short setae (
<figureCitation id="13682A64FFDAFF167BB59B46FB9BF8FD" box="[973,1049,1874,1896]" captionStart="FIGURE 7" captionStartId="21.[151,250,1845,1869]" captionTargetBox="[153,1434,525,1814]" captionTargetId="figure-149@21.[153,1434,525,1822]" captionTargetPageId="21" captionText="FIGURE 7. Spilogona tendipes. A. Female, lateral view. B. Female, lateral view. C. Terminalia, lateral view, digital photo. D. Sternite 5, digital photo. E. Ovipositor, lateral, dorsal and ventral views, digital photo. F. Terminalia, lateral view. H. Cercal plate and surstylus, dorsal view. H. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812253" httpUri="https://zenodo.org/record/14812253/files/figure.png" pageId="20" pageNumber="21">Fig. 7B</figureCitation>
); eye with distinct hairs; male terminalia as in
<figureCitation id="13682A64FFDAFF16796D9B7BFEFAF81F" box="[277,376,1903,1925]" captionStart="FIGURE 7" captionStartId="21.[151,250,1845,1869]" captionTargetBox="[153,1434,525,1814]" captionTargetId="figure-149@21.[153,1434,525,1822]" captionTargetPageId="21" captionText="FIGURE 7. Spilogona tendipes. A. Female, lateral view. B. Female, lateral view. C. Terminalia, lateral view, digital photo. D. Sternite 5, digital photo. E. Ovipositor, lateral, dorsal and ventral views, digital photo. F. Terminalia, lateral view. H. Cercal plate and surstylus, dorsal view. H. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812253" httpUri="https://zenodo.org/record/14812253/files/figure.png" pageId="20" pageNumber="21">Fig. 7F, G</figureCitation>
; female hypoproct with numerous spinules (
<figureCitation id="13682A64FFDAFF167B5A9B7BFCE8F81F" box="[802,874,1903,1925]" captionStart="FIGURE 7" captionStartId="21.[151,250,1845,1869]" captionTargetBox="[153,1434,525,1814]" captionTargetId="figure-149@21.[153,1434,525,1822]" captionTargetPageId="21" captionText="FIGURE 7. Spilogona tendipes. A. Female, lateral view. B. Female, lateral view. C. Terminalia, lateral view, digital photo. D. Sternite 5, digital photo. E. Ovipositor, lateral, dorsal and ventral views, digital photo. F. Terminalia, lateral view. H. Cercal plate and surstylus, dorsal view. H. Sternite 5. Bare: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14812253" httpUri="https://zenodo.org/record/14812253/files/figure.png" pageId="20" pageNumber="21">Fig. 7E</figureCitation>
).....................................
<taxonomicName id="4C534D62FFDAFF167D4C9B64FA1EF81E" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[1332,1436,1903,1925]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDAFF167D4C9B64FAC2F81F" box="[1332,1344,1904,1925]" italics="true" pageId="20" pageNumber="21">S</emphasis>
.
<emphasis id="B927EAF3FFDAFF167D349B7BFA1EF81E" box="[1356,1436,1903,1924]" italics="true" pageId="20" pageNumber="21">tendipes</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDAFF1778BF9BA0FCC1FE87" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BEC36E1FFDAFF1678BF9BA0FA1EF854" blockId="20.[151,1437,1972,2035]" box="[199,1436,1972,1998]" pageId="20" pageNumber="21">
<emphasis id="B927EAF3FFDAFF1678BF9BA0FE31F854" bold="true" box="[199,435,1972,1998]" pageId="20" pageNumber="21">Material examined.</emphasis>
<materialsCitation id="3B3B3CBCFFDAFF1679C59BA0FB48F854" box="[445,1226,1972,1998]" country="Russia" location="Magadan region" pageId="20" pageNumber="21" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Magadan">
<collectingCountry id="F3447671FFDAFF1679C59BA0FDA7F854" box="[445,549,1972,1998]" name="Russia" pageId="20" pageNumber="21">RUSSIA</collectingCountry>
,
<emphasis id="B927EAF3FFDAFF167A4E9BA0FD71F854" box="[566,755,1972,1998]" italics="true" pageId="20" pageNumber="21">
<collectingRegion id="4997F803FFDAFF167A4E9BA0FD71F854" box="[566,755,1972,1998]" country="Russia" name="Magadan" pageId="20" pageNumber="21">Magadan region</collectingRegion>
</emphasis>
:
<specimenCount id="9D55FD68FFDAFF167B7D9BA0FCAAF854" box="[773,808,1972,1998]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
<specimenCount id="9D55FD68FFDAFF167B499BA0FCD4F854" box="[817,854,1972,1998]" count="2" pageId="20" pageNumber="21" type="female">2♀</specimenCount>
, Yablonevyi Pass,
<quantity id="4CAB9B04FFDAFF167C579BA0FB05F854" box="[1071,1159,1972,1998]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="21" unit="km" value="120.0">120 km</quantity>
NNE
</materialsCitation>
<materialsCitation id="3B3B3CBCFFDAFF167CAB9BA0FA15F854" box="[1235,1431,1972,1998]" country="Russia" elevation="900" location="Magadan" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Magadan">
<collectingRegion id="4997F803FFDAFF167CAB9BA0FABCF854" box="[1235,1342,1972,1998]" country="Russia" name="Magadan" pageId="20" pageNumber="21">Magadan</collectingRegion>
,
<quantity id="4CAB9B04FFDAFF167D349BA0FA15F854" box="[1356,1431,1972,1998]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="21" unit="m" value="900.0">
<elevation id="007ED1D2FFDAFF167D349BA0FA15F854" box="[1356,1431,1972,1998]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="21" unit="m" value="900.0">900 m</elevation>
</quantity>
</materialsCitation>
,
</paragraph>
<paragraph id="8BEC36E1FFDAFF1778EF9BCCFCC1FE87" blockId="20.[151,1437,1972,2035]" lastBlockId="21.[151,1438,150,465]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<materialsCitation id="3B3B3CBCFFDAFF1778EF9BCCFD37FF2B" collectingDate="2014-07-17" collectionCode="SZMN, ZMUM" collectorName="N. Vikhrev" lastPageId="21" lastPageNumber="22" latitude="60.983334" location="Yablonevyi Pass" pageId="20" pageNumber="21" specimenCode="BUIC-DIP1759" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<geoCoordinate id="EE675026FFDAFF1678EF9BCCFF75F869" box="[151,247,2008,2035]" degrees="60" direction="north" minutes="59" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="60.983334">60°59’N</geoCoordinate>
151°53 Е,
<date id="FFED1021FFDAFF1679F99BCCFD81F868" box="[385,515,2008,2034]" pageId="20" pageNumber="21" value="2014-07-17">
<collectingDate id="EFA9E9C9FFDAFF1679F99BCCFD81F868" box="[385,515,2008,2034]" pageId="20" pageNumber="21" value="2014-07-17">17.vii.2014</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFDAFF167A3E9BCCFD44F868" box="[582,710,2008,2034]" pageId="20" pageNumber="21">N. Vikhrev</collectorName>
(
<specimenCount id="9D55FD68FFDAFF167AA49BCCFD7FF868" box="[732,765,2008,2034]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
(
<specimenCode id="DBF59E9AFFDAFF167B779BCCFC46F869" box="[783,964,2008,2035]" collectionCode="BUIC-DIP" pageId="20" pageNumber="21">BUIC-DIP1759</specimenCode>
) in
<collectionCode id="ED42AE24FFDAFF167B8E9BCCFBC5F868" box="[1014,1095,2008,2034]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="20" pageNumber="21">SZMN</collectionCode>
,
<specimenCount id="9D55FD68FFDAFF167C2E9BCCFBFBF868" box="[1110,1145,2008,2034]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
<specimenCount id="9D55FD68FFDAFF167CFC9BCCFB25F868" box="[1156,1191,2008,2034]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
in
<collectionCode id="ED42AE24FFDAFF167CAA9BCCFAACF868" box="[1234,1326,2008,2034]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="20" pageNumber="21">ZMUM</collectionCode>
);
<specimenCount id="9D55FD68FFDAFF167D3D9BCCFAE5F868" box="[1349,1383,2008,2034]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
<specimenCount id="9D55FD68FFDAFF167D0A9BCCFA15F868" box="[1394,1431,2008,2034]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<location id="8E8C603AFFDBFF1778EF9C83FED0FF2B" LSID="urn:lsid:plazi:treatment:03FA87F7FFDAFF1778EF9FFFFBD5FE4B:8E8C603AFFDBFF1778EF9C83FED0FF2B" box="[151,338,151,177]" latitude="60.983334" name="Yablonevyi Pass" pageId="21" pageNumber="22">Yablonevyi Pass</location>
,
<locationDeviation id="9A5E2FC3FFDBFF1779269C83FDE7FF2B" box="[350,613,151,177]" location="Magadan" pageId="21" pageNumber="22">
<quantity id="4CAB9B04FFDBFF1779269C83FE31FF2B" box="[350,435,151,177]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="km" value="120.0">120 km</quantity>
NNE Magadan
</locationDeviation>
, 783 м
</materialsCitation>
,
<materialsCitation id="3B3B3CBCFFDBFF177AB99C82FF2DFF63" collectingDate="2017-07-11" collectionCode="V, SZMN" collectorName="Altai" county="Sorokina" elevation="2075" latitude="60.583332" location="Akbul River" municipality="Kosh-Agach district" pageId="21" pageNumber="22" specimenCode="BUIC-DIP1758" specimenCount="1" specimenCount-male="1">
<geoCoordinate id="EE675026FFDBFF177AB99C82FCA0FF2B" box="[705,802,150,177]" degrees="60" direction="north" minutes="35" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="60.583332">60°35’N</geoCoordinate>
151°32 Е,
<collectingMethod id="52124EF6FFDBFF177BD89C83FBD3FF2B" box="[928,1105,151,177]" pageId="21" pageNumber="22">yellow pan trap</collectingMethod>
,
<date id="FFED1021FFDBFF177C249C83FB5FFF2B" box="[1116,1245,151,177]" pageId="21" pageNumber="22" value="2017-07-11">
<collectingDate id="EFA9E9C9FFDBFF177C249C83FB5FFF2B" box="[1116,1245,151,177]" pageId="21" pageNumber="22" value="2017-07-11">11.vii.2017</collectingDate>
</date>
, leg.
<collectionCode id="ED42AE24FFDBFF177D619C83FAAFFF2B" box="[1305,1325,151,177]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="21" pageNumber="22" type="Museum">V</collectionCode>
.
<collectingCounty id="628D4E6DFFDBFF177D4F9C83FA1EFF2B" box="[1335,1436,151,177]" pageId="21" pageNumber="22">Sorokina</collectingCounty>
(
<specimenCode id="DBF59E9AFFDBFF1778E79CAEFED7FF4F" box="[159,341,186,213]" collectionCode="BUIC-DIP" pageId="21" pageNumber="22">BUIC-DIP1758</specimenCode>
,
<collectionCode id="ED42AE24FFDBFF17791A9CAFFE37FF4F" box="[354,437,187,213]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="21" pageNumber="22">SZMN</collectionCode>
).
<emphasis id="B927EAF3FFDBFF1779B19CAFFDEEFF4F" box="[457,620,187,213]" italics="true" pageId="21" pageNumber="22">
<collectorName id="26A65337FFDBFF1779B19CAFFE7DFF4F" box="[457,511,187,213]" pageId="21" pageNumber="22">Altai</collectorName>
<taxonomicName id="4C534D62FFDBFF177A709CAFFDEEFF4F" box="[520,620,187,213]" class="Repensiviricetes" family="Geminiviridae" genus="Republic" higherTaxonomySource="CoL" kingdom="Shotokuvirae" order="Geplafuvirales" pageId="21" pageNumber="22" phylum="Cressdnaviricota" rank="genus">Republic</taxonomicName>
</emphasis>
:
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,
<collectingMunicipality id="6B88AC9BFFDBFF177AC89CAFFC15FF4F" box="[688,919,187,213]" pageId="21" pageNumber="22">Kosh-Agach district</collectingMunicipality>
,
<location id="8E8C603AFFDBFF177BDB9CAFFBB0FF4F" LSID="urn:lsid:plazi:treatment:03FA87F7FFDAFF1778EF9FFFFBD5FE4B:8E8C603AFFDBFF177BDB9CAFFBB0FF4F" box="[931,1074,187,213]" county="Sorokina" latitude="60.583332" municipality="Kosh-Agach district" name="Akbul River" pageId="21" pageNumber="22">Akbul River</location>
, on stones near the water,
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</quantity>
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,
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88º1’Е,
<date id="FFED1021FFDBFF1779F19CCBFD8EFF63" box="[393,524,223,249]" pageId="21" pageNumber="22" value="2009-07-12">
<collectingDate id="EFA9E9C9FFDBFF1779F19CCBFD8EFF63" box="[393,524,223,249]" pageId="21" pageNumber="22" value="2009-07-12">12.vii.2009</collectingDate>
</date>
, leg.
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.
<collectorName id="26A65337FFDBFF177A0B9CCBFD5AFF63" box="[627,728,223,249]" pageId="21" pageNumber="22">Sorokina</collectorName>
(
<collectionCode id="ED42AE24FFDBFF177A949CCBFCC2FF63" box="[748,832,223,249]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="21" pageNumber="22">SZMN</collectionCode>
).
<collectingCountry id="F3447671FFDBFF177B209CCBFC57FF63" box="[856,981,223,249]" name="Canada" pageId="21" pageNumber="22">CANADA</collectingCountry>
,
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<collectingRegion id="4997F803FFDBFF177B909CCBFBA9FF63" box="[1000,1067,223,249]" country="Canada" name="Yukon" pageId="21" pageNumber="22">Yukon</collectingRegion>
</emphasis>
:
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,
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</materialsCitation>
,
<materialsCitation id="3B3B3CBCFFDBFF177D469CCAFCBDFE87" collectingDate="1987-07-01" collectingDateMax="1987-07-05" collectingDateMin="1987-07-01" collectionCode="SZMN" collectorName="M. Polak &amp; M. Wood" country="Canada" latitude="67.96667" longLatPrecision="988" longitude="-136.48334" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Yukon">
<geoCoordinate id="EE675026FFDBFF177D469CCAFA1EFF63" box="[1342,1436,222,249]" degrees="67" direction="north" minutes="58" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="67.96667">67º58’N</geoCoordinate>
<geoCoordinate id="EE675026FFDBFF1778EF9D17FE88FE87" box="[151,266,259,285]" degrees="136" direction="west" minutes="29" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-136.48334">136º29’W</geoCoordinate>
,
<date id="FFED1021FFDBFF17796D9D16FE27FE87" box="[277,421,258,285]" pageId="21" pageNumber="22" value="1987-07-01" valueMax="1987-07-05" valueMin="1987-07-01">
<collectingDate id="EFA9E9C9FFDBFF17796D9D16FE27FE87" box="[277,421,258,285]" pageId="21" pageNumber="22" value="1987-07-01" valueMax="1987-07-05" valueMin="1987-07-01">1–5.vii.1987</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFDBFF1779999D17FDCAFE87" box="[481,584,259,285]" pageId="21" pageNumber="22">M. Polak</collectorName>
&amp;
<collectorName id="26A65337FFDBFF177A139D17FD56FE87" box="[619,724,259,285]" pageId="21" pageNumber="22">M. Wood</collectorName>
(
<collectionCode id="ED42AE24FFDBFF177A9B9D17FCB5FE87" box="[739,823,259,285]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="21" pageNumber="22">SZMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDBFF1778BF9D32FB5BFE13" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BEC36E1FFDBFF1778BF9D32FF6AFEFF" blockId="21.[151,1438,150,465]" pageId="21" pageNumber="22">
<emphasis id="B927EAF3FFDBFF1778BF9D32FEE3FEDA" bold="true" box="[199,353,294,320]" pageId="21" pageNumber="22">Distribution.</emphasis>
PALAEARCTIC:?
<collectingCountry id="F3447671FFDBFF177A339D33FD40FEDB" box="[587,706,295,321]" name="Greenland" pageId="21" pageNumber="22">Greenland</collectingCountry>
(NE),
<collectingCountry id="F3447671FFDBFF177B639D33FCE4FEDB" box="[795,870,295,321]" name="Russia" pageId="21" pageNumber="22">Russia</collectingCountry>
(
<collectingRegion id="4997F803FFDBFF177B039D33FBBEFEDB" box="[891,1084,295,321]" country="Russia" name="Magadan" pageId="21" pageNumber="22">Magadan region</collectingRegion>
,
<collectingRegion id="4997F803FFDBFF177C349D33FB0DFEDB" box="[1100,1167,295,321]" country="Russia" name="Altay" pageId="21" pageNumber="22">Altay</collectingRegion>
). NEARCTIC: Alaska, Yukon.
</paragraph>
<paragraph id="8BEC36E1FFDBFF1778BF9D7AFB5BFE13" blockId="21.[151,1438,150,465]" box="[199,1241,366,393]" pageId="21" pageNumber="22">
<emphasis id="B927EAF3FFDBFF1778BF9D7AFEEBFE13" bold="true" box="[199,361,366,393]" pageId="21" pageNumber="22">DNA barcode</emphasis>
. BOLD BIN: BOLD:ADZ7217. See
<tableCitation id="C6D1035AFFDBFF177B749D7BFCDDFE13" box="[780,863,367,393]" captionStart="TABLE 1" captionStartId="2.[152,245,657,681]" captionText="TABLE 1. List of Russian Spilogona species with barcode index numbers (BIN) and GenBank accession numbers." pageId="21" pageNumber="22">Table 1</tableCitation>
for GenBank accession numbers.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDBFF1778BF9D87FBD5FE4B" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BEC36E1FFDBFF1778BF9D87FBD5FE4B" blockId="21.[151,1438,150,465]" pageId="21" pageNumber="22">
<emphasis id="B927EAF3FFDBFF1778BF9D87FEB9FE37" bold="true" box="[199,315,403,429]" pageId="21" pageNumber="22">Remarks.</emphasis>
<taxonomicName id="4C534D62FFDBFF17793D9D87FD99FE37" authorityName="F" baseAuthorityName="Malloch" baseAuthorityYear="1920" box="[325,539,403,429]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tendipes">
<emphasis id="B927EAF3FFDBFF17793D9D87FD99FE37" box="[325,539,403,429]" italics="true" pageId="21" pageNumber="22">Spilogona tendipes</emphasis>
</taxonomicName>
is morphologically very similar to
<taxonomicName id="4C534D62FFDBFF177BC29D87FB32FE37" authorityName="Sorokina" authorityYear="2025" box="[954,1200,403,429]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="latizygoma">
<emphasis id="B927EAF3FFDBFF177BC29D87FB32FE37" box="[954,1200,403,429]" italics="true" pageId="21" pageNumber="22">Spilogona latizygoma</emphasis>
</taxonomicName>
. But based on DNA barcodes, these species are genetically different; the p-distance between them is 3.36%.
</paragraph>
</subSubSection>
</treatment>
</document>

200
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(
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<emphasis id="B927EAF3FFDCFF1078EF9EE8FE96FC8E" box="[151,276,764,788]" italics="true" pageId="18" pageNumber="19">Limnophora</emphasis>
(
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)
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in
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. Type-locality: “Vier
<specimenCount id="9D55FD68FFDCFF107CF79EE9FB20FC8E" box="[1167,1186,765,788]" count="1" pageId="18" pageNumber="19" type="male">♂</specimenCount>
Exemplare in Warschau, Vorstadt Park Praga. Juli” [
<collectingCountry id="F3447671FFDCFF1079A09F0FFDC6FCA9" box="[472,580,795,819]" name="Poland" pageId="18" pageNumber="19">POLAND</collectingCountry>
,
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<treatmentCitationGroup id="AB4311CFFFDCFF1078EF9F2EFDF8FCEB" pageId="18" pageNumber="19">
<treatmentCitation id="0AF210F0FFDCFF1078EF9F2EFDD7FCC8" author="Malloch, J. R." box="[151,597,826,850]" page="226" pageId="18" pageNumber="19" year="1917">
<taxonomicName id="4C534D62FFDCFF1078EF9F2EFDD7FCC8" ID-CoL="898XJ" authority="Malloch, 1917: 226" authorityName="Malloch" authorityPageNumber="226" authorityYear="1917" box="[151,597,826,850]" class="Insecta" family="Muscidae" genus="Tetramerinx" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B927EAF3FFDCFF1078EF9F2EFE0BFCC8" box="[151,393,826,850]" italics="true" pageId="18" pageNumber="19">Tetramerinx brevicornis</emphasis>
<materialsCitation id="3B3B3CBCFFDCFF1079F69F2EFDD7FCC8" box="[398,597,826,850]" collectingDate="1906-08-23" country="United States of America" location="Taken" municipality="Lake" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Michigan">
<bibRefCitation id="EFC24B10FFDCFF1079F69F2EFDD7FCC8" author="Malloch, J. R." box="[398,597,826,850]" pageId="18" pageNumber="19" pagination="225 - 226" refId="ref18873" refString="Malloch, J. R. (1917) A new North American species of the genus Tetramerinx (Diptera, Anthomyiidae). Canadian Entomologist, 49 (7), 225-226. https://doi.org/10.4039/Ent49225-7" type="journal article" year="1917">Malloch, 1917: 226</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. Type-locality: “Waukegan, Ill.,
<date id="FFED1021FFDCFF107BEF9F2EFBC2FCC8" box="[919,1088,826,850]" pageId="18" pageNumber="19" value="1906-08-23">
<collectingDate id="EFA9E9C9FFDCFF107BEF9F2EFBC2FCC8" box="[919,1088,826,850]" pageId="18" pageNumber="19" value="1906-08-23">August 23, 1906</collectingDate>
</date>
.
<location id="8E8C603AFFDCFF107C3F9F2EFB07FCC8" LSID="urn:lsid:plazi:treatment:03FA87F7FFDCFF1078EF9E87FC25FAD1:8E8C603AFFDCFF107C3F9F2EFB07FCC8" box="[1095,1157,826,850]" country="United States of America" municipality="Lake" name="Taken" pageId="18" pageNumber="19" stateProvince="Michigan">Taken</location>
on shore of
<collectingMunicipality id="6B88AC9BFFDCFF107C859F2EFAB2FCC8" box="[1277,1328,826,850]" pageId="18" pageNumber="19">Lake</collectingMunicipality>
<collectingRegion id="4997F803FFDCFF107D4D9F2EFA1BFCC8" box="[1333,1433,826,850]" country="United States of America" name="Michigan" pageId="18" pageNumber="19">Michigan</collectingRegion>
.
<specimenCount id="9D55FD68FFDCFF1078BF9F4DFEE4FCEA" box="[199,358,856,880]" count="2" pageId="18" pageNumber="19" type="generic">Two specimens</specimenCount>
” [
<collectingCountry id="F3447671FFDCFF1079039F4DFE32FCEB" box="[379,432,857,881]" name="United States of America" pageId="18" pageNumber="19">USA</collectingCountry>
,
<collectingRegion id="4997F803FFDCFF1079C59F4CFD88FCEA" box="[445,522,856,880]" country="United States of America" name="Illinois" pageId="18" pageNumber="19">Illinois</collectingRegion>
].
<emphasis id="B927EAF3FFDCFF107A629F4CFDF8FCEB" bold="true" box="[538,634,856,881]" pageId="18" pageNumber="19">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFDCFF1078EF9F8BFE89FB0F" blockId="18.[151,1437,927,1356]" pageId="18" pageNumber="19">
<emphasis id="B927EAF3FFDCFF1078EF9F8BFF63FC23" bold="true" box="[151,225,927,953]" pageId="18" pageNumber="19">Notes.</emphasis>
The series of Nearctic specimens of
<taxonomicName id="4C534D62FFDCFF107AF99F8BFCFBFC23" ID-CoL="4Z3SW" baseAuthorityName="Malloch" baseAuthorityYear="1917" box="[641,889,927,953]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B927EAF3FFDCFF107AF99F8BFCFBFC23" box="[641,889,927,953]" italics="true" pageId="18" pageNumber="19">Spilogona brevicornis</emphasis>
</taxonomicName>
and European specimens of
<taxonomicName id="4C534D62FFDCFF107CC59F8BFA1FFC23" ID-CoL="4Z3YH" baseAuthorityName="Schnabl" baseAuthorityYear="1911" box="[1213,1437,927,953]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="scutulata">
<emphasis id="B927EAF3FFDCFF107CC59F8BFA1FFC23" box="[1213,1437,927,953]" italics="true" pageId="18" pageNumber="19">Spilogona scutulata</emphasis>
</taxonomicName>
were examined. The descriptions of both species and the male terminalia are the same. These species are very distinctive in both sexes, with strong presutural
<emphasis id="B927EAF3FFDCFF107ABC9FFDFD6BFB98" box="[708,745,1001,1026]" italics="true" pageId="18" pageNumber="19">acr</emphasis>
setae in 2 even rows which are strikingly stronger than the ground hairs (
<figureCitation id="13682A64FFDCFF10794E9819FE35FBBD" box="[310,439,1037,1064]" captionStart="FIGURE 5" captionStartId="16.[151,250,1785,1809]" captionTargetBox="[151,1429,181,1761]" captionTargetId="figure-41@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURE 5. Spilogona ssp. A. Spilogona fulvibasis, lateral view, holotype (photo: Garry Ouellette). B–G. Spilogona lapponica (photos of paratypes of Spilogona compacta). B. Sternite 5. C. Terminalia, lateral view. D. Male, lateral view. E. Female, lateral view. F. Male abdomen apex, ventral view. G. Cercal plate, dorsal view. H–J. Spilogona padlei. H. Terminalia, lateral view. I. Sternite 5. J. Female abdomen apex, lateral view. K–M. Spilogona scutulata. K. Male, lateral view. L. Female, lateral view. M. Sternite 5." figureDoi="http://doi.org/10.5281/zenodo.14812247" httpUri="https://zenodo.org/record/14812247/files/figure.png" pageId="18" pageNumber="19">Figs 5K, L</figureCitation>
), and hind femur without
<emphasis id="B927EAF3FFDCFF107A9B981AFD7FFBBD" box="[739,765,1038,1063]" italics="true" pageId="18" pageNumber="19">pv</emphasis>
setae; males with rather broad fronto-orbital plates, which touch and are very silvery, shining; very light grey scutum with a brown spot between
<emphasis id="B927EAF3FFDCFF107C1B9826FB31FBD1" box="[1123,1203,1073,1099]" italics="true" pageId="18" pageNumber="19">post dc</emphasis>
; light abdomen with small spots, and a brown dusted scutellum; sternite 5 with with a broad median notch and a pointed caudal margins (
<figureCitation id="13682A64FFDCFF1078E7986EFF7DFB0F" box="[159,255,1146,1173]" captionStart="FIGURE 5" captionStartId="16.[151,250,1785,1809]" captionTargetBox="[151,1429,181,1761]" captionTargetId="figure-41@16.[151,1435,181,1761]" captionTargetPageId="16" captionText="FIGURE 5. Spilogona ssp. A. Spilogona fulvibasis, lateral view, holotype (photo: Garry Ouellette). B–G. Spilogona lapponica (photos of paratypes of Spilogona compacta). B. Sternite 5. C. Terminalia, lateral view. D. Male, lateral view. E. Female, lateral view. F. Male abdomen apex, ventral view. G. Cercal plate, dorsal view. H–J. Spilogona padlei. H. Terminalia, lateral view. I. Sternite 5. J. Female abdomen apex, lateral view. K–M. Spilogona scutulata. K. Male, lateral view. L. Female, lateral view. M. Sternite 5." figureDoi="http://doi.org/10.5281/zenodo.14812247" httpUri="https://zenodo.org/record/14812247/files/figure.png" pageId="18" pageNumber="19">Fig. 5M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDCFF1078BF988BFDB5FB44" pageId="18" pageNumber="19" type="description">
<paragraph id="8BEC36E1FFDCFF1078BF988BFDB5FB44" blockId="18.[151,1437,927,1356]" pageId="18" pageNumber="19">
I therefore conclude that
<taxonomicName id="4C534D62FFDCFF10798998B4FC37FB23" authority="Malloch, 1917" authorityName="Malloch" authorityYear="1917" box="[497,949,1183,1209]" class="Insecta" family="Muscidae" genus="Tetramerinx" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B927EAF3FFDCFF10798998B4FC81FB23" box="[497,771,1183,1209]" italics="true" pageId="18" pageNumber="19">Tetramerinx brevicornis</emphasis>
<bibRefCitation id="EFC24B10FFDCFF107B76988BFC37FB23" author="Malloch, J. R." box="[782,949,1183,1209]" pageId="18" pageNumber="19" pagination="225 - 226" refId="ref18873" refString="Malloch, J. R. (1917) A new North American species of the genus Tetramerinx (Diptera, Anthomyiidae). Canadian Entomologist, 49 (7), 225-226. https://doi.org/10.4039/Ent49225-7" type="journal article" year="1917">Malloch, 1917</bibRefCitation>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="4C534D62FFDCFF107D6A988BFDB0FB44" authority="Schnabl, 1911" authorityName="Schnabl" authorityYear="1911" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="scutulata" subGenus="Spilogona">
<emphasis id="B927EAF3FFDCFF107D6A988BFA1EFB23" box="[1298,1436,1183,1209]" italics="true" pageId="18" pageNumber="19">Limnophora</emphasis>
(
<emphasis id="B927EAF3FFDCFF1078D998D7FE91FB47" box="[161,275,1219,1245]" italics="true" pageId="18" pageNumber="19">Spilogona</emphasis>
)
<emphasis id="B927EAF3FFDCFF10795A98D7FE0BFB47" box="[290,393,1219,1245]" italics="true" pageId="18" pageNumber="19">scutulata</emphasis>
<bibRefCitation id="EFC24B10FFDCFF1079E898D7FDB0FB44" author="Schnabl, J. &amp; Dziedzicki, H." box="[400,562,1219,1246]" pageId="18" pageNumber="19" pagination="55 - 358" refId="ref19071" refString="Schnabl, J. &amp; Dziedzicki, H. (1911) Die Anthomyiden. Nova Acta Academiae Caesarea Leopoldina-Carolinae Germanicum Naturae Curiosorum, 95, 55-358." type="journal article" year="1911">Schnabl, 1911</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDCFF1078BF98FCFC25FAD1" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BEC36E1FFDCFF1078BF98FCFC25FAD1" blockId="18.[151,1437,927,1356]" pageId="18" pageNumber="19">
<emphasis id="B927EAF3FFDCFF1078BF98FCFEE3FA98" bold="true" box="[199,353,1256,1282]" pageId="18" pageNumber="19">Distribution.</emphasis>
PALAEARCTIC:
<collectingCountry id="F3447671FFDCFF107A4A98FCFD05FA98" box="[562,647,1256,1282]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107AEA98FCFD77FA98" box="[658,757,1256,1282]" name="Belgium" pageId="18" pageNumber="19">Belgium</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107A8798FCFCC0FA98" box="[767,834,1256,1282]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
(
<collectingRegion id="4997F803FFDCFF107B2898FCFC20FA98" box="[848,930,1256,1282]" country="China" name="Shanxi" pageId="18" pageNumber="19">Shanxi</collectingRegion>
,
<collectingRegion id="4997F803FFDCFF107BD598FCFC7BFA98" box="[941,1017,1256,1282]" country="China" name="Gansu" pageId="18" pageNumber="19">Gansu</collectingRegion>
),
<collectingCountry id="F3447671FFDCFF107C7398FCFB42FA98" box="[1035,1216,1256,1282]" name="Czech Republic" pageId="18" pageNumber="19">
Czech
<taxonomicName id="4C534D62FFDCFF107C2098FCFB42FA98" box="[1112,1216,1256,1282]" class="Repensiviricetes" family="Geminiviridae" genus="Republic" higherTaxonomySource="CoL" kingdom="Shotokuvirae" order="Geplafuvirales" pageId="18" pageNumber="22" phylum="Cressdnaviricota" rank="genus">Republic</taxonomicName>
</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107CB298FCFAB7FA98" box="[1226,1333,1256,1282]" name="Denmark" pageId="18" pageNumber="19">Denmark</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107D3898FCFA1AFA98" box="[1344,1432,1256,1282]" name="Finland" pageId="18" pageNumber="19">Finland</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF1078EF9919FE82FABD" box="[151,256,1293,1319]" name="Germany" pageId="18" pageNumber="19">Germany</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF1079689919FE28FABC" box="[272,426,1292,1319]" name="United Kingdom" pageId="18" pageNumber="19">Great Britain</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF1079C29919FD9FFABD" box="[442,541,1293,1319]" name="Hungary" pageId="18" pageNumber="19">Hungary</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107A549919FD05FABD" box="[556,647,1293,1319]" name="Norway" pageId="18" pageNumber="19">Norway</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107AEF9918FD6AFABC" box="[663,744,1292,1318]" name="Poland" pageId="18" pageNumber="19">Poland</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107A8F9918FCC0FABC" box="[759,834,1292,1318]" name="Russia" pageId="18" pageNumber="19">Russia</collectingCountry>
(Europe),
<collectingCountry id="F3447671FFDCFF107BBB9918FBAAFABC" box="[963,1064,1292,1318]" name="Slovakia" pageId="18" pageNumber="19">Slovakia</collectingCountry>
,
<collectingCountry id="F3447671FFDCFF107C4F9918FB11FABC" box="[1079,1171,1292,1318]" name="Sweden" pageId="18" pageNumber="19">Sweden</collectingCountry>
. NEARCTIC: Oregon, NWT to Colorado &amp; S Dakota, New York, Manitoba, Quebec, Maine.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="03FA87F7FFDFFF1078EF9A97FB63FDD8" ID-DOI="http://doi.org/10.5281/zenodo.14812269" ID-Zenodo-Dep="14812269" LSID="urn:lsid:plazi:treatment:03FA87F7FFDFFF1078EF9A97FB63FDD8" httpUri="http://treatment.plazi.org/id/03FA87F7FFDFFF1078EF9A97FB63FDD8" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<paragraph id="8BEC36E1FFDFFF1378EF9A97FD0DF907" blockId="17.[151,655,1667,1729]" box="[151,655,1667,1694]" pageId="17" pageNumber="18">
<heading id="D0A4818DFFDFFF1378EF9A97FD0DF907" bold="true" box="[151,655,1667,1694]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="4C534D62FFDFFF1378EF9A97FD0DF907" ID-CoL="4Z3Y9" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,655,1667,1694]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFDFFF1378EF9A97FD0DF907" bold="true" box="[151,655,1667,1694]" pageId="17" pageNumber="18">
<emphasis id="B927EAF3FFDFFF1378EF9A97FE44F907" bold="true" box="[151,454,1667,1694]" italics="true" pageId="17" pageNumber="18">Spilogona quinquesetosa</emphasis>
(Schnabl, 1915)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<figureCitation id="13682A64FFDFFF1378EF9AB3FF6EF95B" box="[151,236,1703,1729]" captionStart="FIGURE 4" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[161,1436,502,1903]" captionTargetId="figure-177@12.[151,1436,489,1913]" captionTargetPageId="12" captionText="FIGURE 4. Spilogona ssp. A–C. Spilogona churchillensis. A. Terminalia, lateral view. B. Cercal plate, dorsal view. C. Sternite 5. D. Spilogona quinquesetosa, sternite 5. E–K. Spilogona cordyluraeformis, scutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14812242" httpUri="https://zenodo.org/record/14812242/files/figure.png" pageId="17" pageNumber="18">Fig. 4D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDFFF1378EF9AF8FAB8F8E4" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BEC36E1FFDFFF1378EF9AF8FEDBF8B8" blockId="17.[151,1436,1771,1918]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB4311CFFFDFFF1378EF9AF8FEDBF8B8" pageId="17" pageNumber="18">
<taxonomicName id="4C534D62FFDFFF1378EF9AF8FD05F89E" ID-CoL="8CKTD" authority="Schnabl" authorityName="Schnabl" authorityYear="1915" box="[151,647,1772,1796]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quinquesetosa" subGenus="Spilogona">
<emphasis id="B927EAF3FFDFFF1378EF9AF8FE96F89E" box="[151,276,1772,1796]" italics="true" pageId="17" pageNumber="18">Limnophora</emphasis>
(
<emphasis id="B927EAF3FFDFFF13795D9AF8FE0EF89E" box="[293,396,1772,1796]" italics="true" pageId="17" pageNumber="18">Spilogona</emphasis>
)
<emphasis id="B927EAF3FFDFFF1379E49AF8FDAFF899" box="[412,557,1772,1795]" italics="true" pageId="17" pageNumber="18">quinquesetosa</emphasis>
Schnabl
</taxonomicName>
in
<treatmentCitation id="0AF210F0FFDFFF137AD29AF8FC18F89E" author="Becker, T. &amp; Dziedzicki, H. &amp; Schnabl, J. &amp; Villeneuve, J." box="[682,922,1771,1796]" page="33" pageId="17" pageNumber="18" year="1915">
<bibRefCitation id="EFC24B10FFDFFF137AD29AF8FC18F89E" author="Becker, T. &amp; Dziedzicki, H. &amp; Schnabl, J. &amp; Villeneuve, J." box="[682,922,1771,1796]" pageId="17" pageNumber="18" pagination="1 - 67" refId="ref18056" refString="Becker, T., Dziedzicki, H., Schnabl, J. &amp; Villeneuve, J. (1915) Resultats scientifiques de l'Expedition des freres Kuznecov (Kouznetzov) a l'Oural Arctique en 1909, sous la direction de H. Backlund. Diptera. Zapiski Imperatorskoi Akademii Nauk (VIII), Fiziko-Matematicheskoe Otdeleniye, 28 (7), 1-67." type="journal article" year="1915">
Becker
<emphasis id="B927EAF3FFDFFF137A829AF8FCADF89E" box="[762,815,1772,1796]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
, 1915: 33
</bibRefCitation>
</treatmentCitation>
. Type-locality:
<collectingCountry id="F3447671FFDFFF137C439AF8FB03F89E" box="[1083,1153,1772,1796]" name="Russia" pageId="17" pageNumber="18">Russia</collectingCountry>
, Yugorskiy peninsula, Kara district [seen].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFDFFF1378EF9B3DFDC6F8FA" blockId="17.[151,1436,1771,1918]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB4311CFFFDFFF1378EF9B3DFDC6F8FA" pageId="17" pageNumber="18">
<treatmentCitation id="0AF210F0FFDFFF1378EF9B3DFDD5F8DB" author="Huckett, H. C." box="[151,599,1833,1857]" page="212" pageId="17" pageNumber="18" year="1965">
<taxonomicName id="4C534D62FFDFFF1378EF9B3DFDD5F8DB" ID-CoL="4Z3SN" authority="Huckett, 1965: 212" authorityName="Huckett" authorityPageNumber="212" authorityYear="1965" box="[151,599,1833,1857]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="barrowensis">
<emphasis id="B927EAF3FFDFFF1378EF9B3DFE01F8DB" box="[151,387,1833,1857]" italics="true" pageId="17" pageNumber="18">Spilogona barrowensis</emphasis>
<materialsCitation id="3B3B3CBCFFDFFF1379F49B3DFDD5F8DB" box="[396,599,1833,1857]" collectingDate="1952-07-20" collectorName="P. D. Hurd &amp; Syn." country="United States of America" location="Point Barrow" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Alaska">
<bibRefCitation id="EFC24B10FFDFFF1379F49B3DFDD5F8DB" author="Huckett, H. C." box="[396,599,1833,1857]" pageId="17" pageNumber="18" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965: 212</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54E88843FFDFFF137A1B9B3DFD17F8DB" box="[611,661,1833,1857]" pageId="17" pageNumber="18">Type</typeStatus>
locality: “
<location id="8E8C603AFFDFFF137B7A9B3DFC12F8DA" LSID="urn:lsid:plazi:treatment:03FA87F7FFDFFF1078EF9A97FB63FDD8:8E8C603AFFDFFF137B7A9B3DFC12F8DA" box="[770,912,1833,1857]" country="United States of America" name="Point Barrow" pageId="17" pageNumber="18" stateProvince="Alaska">Point Barrow</location>
,
<collectingRegion id="4997F803FFDFFF137BE39B3DFC66F8DB" box="[923,996,1833,1857]" country="United States of America" name="Alaska" pageId="17" pageNumber="18">Alaska</collectingRegion>
, 7.20.52 [i.e.
<date id="FFED1021FFDFFF137C0E9B3DFB6FF8DB" box="[1142,1261,1833,1857]" pageId="17" pageNumber="18" value="1952-07-20">
<collectingDate id="EFA9E9C9FFDFFF137C0E9B3DFB6FF8DB" box="[1142,1261,1833,1857]" pageId="17" pageNumber="18" value="1952-07-20">20.vii.1952</collectingDate>
</date>
]” [
<collectingCountry id="F3447671FFDFFF137D749B3DFAC3F8DB" box="[1292,1345,1833,1857]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
,
<collectingRegion id="4997F803FFDFFF137D379B3DFA1BF8DB" box="[1359,1433,1833,1857]" country="United States of America" name="Alaska" pageId="17" pageNumber="18">Alaska</collectingRegion>
; collector
<collectorName id="26A65337FFDFFF13795E9B5CFE0CF8FA" box="[294,398,1864,1888]" pageId="17" pageNumber="18">P.D. Hurd</collectorName>
] [seen].
<emphasis id="B927EAF3FFDFFF13799C9B53FDC6F8FA" bold="true" box="[484,580,1863,1888]" pageId="17" pageNumber="18">
<collectorName id="26A65337FFDFFF13799C9B53FD91F8C5" box="[484,531,1863,1887]" pageId="17" pageNumber="18">Syn.</collectorName>
nov.
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEC36E1FFDFFF1378EF9B72FAB8F8E4" blockId="17.[151,1436,1771,1918]" box="[151,1338,1894,1918]" pageId="17" pageNumber="18">
<taxonomicName id="4C534D62FFDFFF1378EF9B72FDC3F8E4" ID-CoL="4Z3Y9" authority="(Schnabl, 1915)" baseAuthorityName="Schnabl" baseAuthorityYear="1915" box="[151,577,1894,1918]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFDFFF1378EF9B72FE17F8E4" box="[151,405,1894,1918]" italics="true" pageId="17" pageNumber="18">Spilogona quinquesetosa</emphasis>
(Schnabl, 1915)
</taxonomicName>
; Sorokina &amp; Thidrikh, 2021: 226 (
<collectingRegion id="4997F803FFDFFF137BE09B72FC79F8E4" box="[920,1019,1894,1918]" country="Russia" name="Chukotka" pageId="17" pageNumber="18">Chukotka</collectingRegion>
AO, including Wrangel Island).
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDFFF1078EF9BB9FB5BFD87" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BEC36E1FFDFFF1078EF9BB9FCEFFED9" blockId="17.[151,1437,1964,2027]" lastBlockId="18.[151,1437,151,578]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B927EAF3FFDFFF1378EF9BB9FF58F85D" bold="true" box="[151,218,1965,1991]" pageId="17" pageNumber="18">Notes</emphasis>
. The type material of
<taxonomicName id="4C534D62FFDFFF1379AD9BB8FCEBF85C" authority="Schnabl" authorityName="Schnabl" authorityYear="1915" box="[469,873,1964,1990]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFDFFF1379AD9BB8FC8AF85C" box="[469,776,1964,1990]" italics="true" pageId="17" pageNumber="18">Limnophora quinquesetosa</emphasis>
Schnabl
</taxonomicName>
(in
<collectionCode id="ED42AE24FFDFFF137BE09BB9FC51F85D" box="[920,979,1965,1991]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="17" pageNumber="18">ZISP</collectionCode>
) and
<taxonomicName id="4C534D62FFDFFF137C6B9BB8FAF4F85C" authority="Huckett" authorityName="Huckett" authorityYear="1965" box="[1043,1398,1964,1990]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="barrowensis">
<emphasis id="B927EAF3FFDFFF137C6B9BB8FA97F85C" box="[1043,1301,1964,1990]" italics="true" pageId="17" pageNumber="18">Spilogona barrowensis</emphasis>
Huckett
</taxonomicName>
(in Canadian National Collection) was examined. The descriptions of both species and the male terminalia are the same. This is black species with brown dust, without marks on scutum and abdomen, a shining prementum, with preapical setulae on upper border of declivities of scutellum, facial edge not projecting beyond level of profrons, strong
<emphasis id="B927EAF3FFDCFF107D199CA8FAF9FF4F" box="[1377,1403,188,213]" italics="true" pageId="18" pageNumber="19">av</emphasis>
,
<emphasis id="B927EAF3FFDCFF107DF19CA8FA17FF4F" box="[1417,1429,188,213]" italics="true" pageId="18" pageNumber="19">v</emphasis>
,
<emphasis id="B927EAF3FFDCFF1078EF9CF4FF33FF63" box="[151,177,224,249]" italics="true" pageId="18" pageNumber="19">pv</emphasis>
setae on mid tibia, and hind femur without
<emphasis id="B927EAF3FFDCFF107AE39CF4FD37FF63" box="[667,693,224,249]" italics="true" pageId="18" pageNumber="19">pv</emphasis>
setae. According to Huckett’s description (1965), the posthumeral and notopleural regions are pale in
<taxonomicName id="4C534D62FFDCFF107A489D10FCB6FE84" authorityName="Huckett" authorityYear="1965" box="[560,820,260,286]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="barrowensis">
<emphasis id="B927EAF3FFDCFF107A489D10FCB6FE84" box="[560,820,260,286]" italics="true" pageId="18" pageNumber="19">Spilogona barrowensis</emphasis>
</taxonomicName>
, but in the
<typeStatus id="54E88843FFDCFF107BC49D10FB9CFE84" box="[956,1054,260,286]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
and the
<typeStatus id="54E88843FFDCFF107CFC9D11FB6DFE85" box="[1156,1263,261,287]" pageId="18" pageNumber="19" type="paratype">paratypes</typeStatus>
of this species these regions are brown dusted as in
<taxonomicName id="4C534D62FFDCFF107A4C9D3DFCE4FED9" authorityName="Schnabl" authorityYear="1915" box="[564,870,297,323]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quinquesetosa">
<emphasis id="B927EAF3FFDCFF107A4C9D3DFCE4FED9" box="[564,870,297,323]" italics="true" pageId="18" pageNumber="19">Limnophora quinquesetosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFDCFF1078BF9D59FDF3FE16" blockId="18.[151,1437,151,578]" pageId="18" pageNumber="19">
I therefore conclude that
<taxonomicName id="4C534D62FFDCFF10798E9D59FC2FFEF2" authority="Huckett, 1965" authorityName="Huckett" authorityYear="1965" box="[502,941,333,360]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="barrowensis">
<emphasis id="B927EAF3FFDCFF10798E9D59FD7FFEFD" box="[502,765,333,359]" italics="true" pageId="18" pageNumber="19">Spilogona barrowensis</emphasis>
<bibRefCitation id="EFC24B10FFDCFF107B719D59FC2FFEF2" author="Huckett, H. C." box="[777,941,333,360]" pageId="18" pageNumber="19" pagination="1 - 370" refId="ref18822" refString="Huckett, H. C. (1965) The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Memoirs of the Entomological Society of Canada, 42, 1-370. [Ottawa] https://doi.org/10.4039/entm9742fv" type="journal article" year="1965">Huckett, 1965</bibRefCitation>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="4C534D62FFDCFF107D6A9D59FDEFFE16" authority="Schnabl, 1915" authorityName="Schnabl" authorityYear="1915" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quinquesetosa" subGenus="Spilogona">
<emphasis id="B927EAF3FFDCFF107D6A9D59FA1EFEFD" box="[1298,1436,333,359]" italics="true" pageId="18" pageNumber="19">Limnophora</emphasis>
(
<emphasis id="B927EAF3FFDCFF1078D99D65FE91FE11" box="[161,275,369,395]" italics="true" pageId="18" pageNumber="19">Spilogona</emphasis>
)
<emphasis id="B927EAF3FFDCFF10795A9D66FE40FE11" box="[290,450,370,395]" italics="true" pageId="18" pageNumber="19">quinquesetosa</emphasis>
Schnabl, 1915
</taxonomicName>
.
</paragraph>
<paragraph id="8BEC36E1FFDCFF1078BF9D81FC95FE62" blockId="18.[151,1437,151,578]" pageId="18" pageNumber="19">
<emphasis id="B927EAF3FFDCFF1078BF9D81FDBBFE35" bold="true" box="[199,569,405,431]" pageId="18" pageNumber="19">Additional material examined</emphasis>
.
<materialsCitation id="3B3B3CBCFFDCFF107A369D81FD07FE62" collectingDate="2016-08-04" collectingDateMax="2016-08-25" collectingDateMin="2016-08-04" collectorName="A. Nikhaeva &amp; M. Bizin" country="Russia" latitude="72.96667" location="Shokalsky Island" longLatPrecision="964" longitude="72.45" municipality="Gydanskiy Proliv" pageId="18" pageNumber="19" specimenCode="BUIC-DIP2175, BUIC-DIP2176" specimenCount="6" specimenCount-female="6" stateProvince="Yamalo-Nenets">
<collectingCountry id="F3447671FFDCFF107A369D81FD34FE35" box="[590,694,405,431]" name="Russia" pageId="18" pageNumber="19">RUSSIA</collectingCountry>
,
<emphasis id="B927EAF3FFDCFF107AB39D81FC27FE2A" box="[715,933,405,432]" italics="true" pageId="18" pageNumber="19">
<collectingRegion id="4997F803FFDCFF107AB39D81FCF6FE35" box="[715,884,405,431]" country="Russia" name="Yamalo-Nenets" pageId="18" pageNumber="19">Yamalo-Nenets</collectingRegion>
AO
</emphasis>
:
<specimenCount id="9D55FD68FFDCFF107BC39D81FC5DFE35" box="[955,991,405,431]" count="3" pageId="18" pageNumber="19" type="female">3♀</specimenCount>
,
<collectingMunicipality id="6B88AC9BFFDCFF107B8A9D81FB3CFE35" box="[1010,1214,405,431]" pageId="18" pageNumber="19">Gydanskiy Proliv</collectingMunicipality>
,
<location id="8E8C603AFFDCFF107CB79D81FA1AFE35" LSID="urn:lsid:plazi:treatment:03FA87F7FFDFFF1078EF9A97FB63FDD8:8E8C603AFFDCFF107CB79D81FA1AFE35" box="[1231,1432,405,431]" country="Russia" latitude="72.96667" longLatPrecision="964" longitude="72.45" municipality="Gydanskiy Proliv" name="Shokalsky Island" pageId="18" pageNumber="19" stateProvince="Yamalo-Nenets">Shokalsky Island</location>
,
<geoCoordinate id="EE675026FFDCFF1078EF9DAEFF7AFE4F" box="[151,248,442,469]" degrees="72" direction="north" minutes="58" orientation="latitude" pageId="18" pageNumber="19" precision="925" value="72.96667">72°58’N</geoCoordinate>
<geoCoordinate id="EE675026FFDCFF1078879DAEFEDDFE4E" box="[255,351,442,468]" degrees="72" direction="east" minutes="27" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="72.45">72°27’E</geoCoordinate>
, swamp by a stream (
<specimenCount id="9D55FD68FFDCFF107A2F9DAEFDF4FE4E" box="[599,630,442,468]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
), arctic fox hole (
<specimenCount id="9D55FD68FFDCFF107B3D9DAEFCE6FE4E" box="[837,868,442,468]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
), tundra (
<specimenCount id="9D55FD68FFDCFF107BA09DAEFC75FE4E" box="[984,1015,442,468]" count="1" pageId="18" pageNumber="19" type="female">1♀</specimenCount>
),
<date id="FFED1021FFDCFF107C689DAEFB35FE4F" box="[1040,1207,442,469]" pageId="18" pageNumber="19" value="2016-08-04" valueMax="2016-08-25" valueMin="2016-08-04">
<collectingDate id="EFA9E9C9FFDCFF107C689DAEFB35FE4F" box="[1040,1207,442,469]" pageId="18" pageNumber="19" value="2016-08-04" valueMax="2016-08-25" valueMin="2016-08-04">4–25.viii.2016</collectingDate>
</date>
, leg.
<collectorName id="26A65337FFDCFF107C899DAEFAFDFE4E" box="[1265,1407,442,468]" pageId="18" pageNumber="19">A. Nikhaeva</collectorName>
&amp;
<collectorName id="26A65337FFDCFF1078EF9DCBFF79FE62" box="[151,251,478,505]" pageId="18" pageNumber="19">M. Bizin</collectorName>
(
<specimenCode id="DBF59E9AFFDCFF1079719DCAFE3DFE63" box="[265,447,478,505]" collectionCode="BUIC-DIP" pageId="18" pageNumber="19">BUIC-DIP2175</specimenCode>
,
<specimenCode id="DBF59E9AFFDCFF1079B29DCAFDFCFE62" box="[458,638,478,504]" collectionCode="BUIC-DIP" pageId="18" pageNumber="19">BUIC-DIP2176</specimenCode>
)
</materialsCitation>
(all
<collectionCode id="ED42AE24FFDCFF107AC09DCAFC89FE62" box="[696,779,478,504]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="18" pageNumber="19">SZMN</collectionCode>
).
</paragraph>
<paragraph id="8BEC36E1FFDCFF1078BF9E17FB5BFD87" blockId="18.[151,1437,151,578]" box="[199,1241,515,542]" pageId="18" pageNumber="19">
<emphasis id="B927EAF3FFDCFF1078BF9E17FEEBFD87" bold="true" box="[199,361,515,541]" pageId="18" pageNumber="19">DNA barcode</emphasis>
.
<collectionCode id="ED42AE24FFDCFF10790F9E17FE7CFD87" box="[375,510,515,541]" pageId="18" pageNumber="19">BOLD BIN</collectionCode>
:
<collectionCode id="ED42AE24FFDCFF107A719E17FDD8FD87" box="[521,602,515,541]" pageId="18" pageNumber="19">BOLD</collectionCode>
:AEK7965. See
<tableCitation id="C6D1035AFFDCFF107B749E17FCDDFD87" box="[780,863,515,541]" captionStart="TABLE 1" captionStartId="2.[152,245,657,681]" captionText="TABLE 1. List of Russian Spilogona species with barcode index numbers (BIN) and GenBank accession numbers." pageId="18" pageNumber="19">Table 1</tableCitation>
for GenBank accession numbers.
</paragraph>
</subSubSection>
<subSubSection id="C349656AFFDCFF1078BF9E33FB63FDD8" box="[199,1249,551,578]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BEC36E1FFDCFF1078BF9E33FB63FDD8" blockId="18.[151,1437,151,578]" box="[199,1249,551,578]" pageId="18" pageNumber="19">
<emphasis id="B927EAF3FFDCFF1078BF9E33FEE3FDDB" bold="true" box="[199,353,551,577]" pageId="18" pageNumber="19">Distribution.</emphasis>
PALAEARCTIC:
<collectingCountry id="F3447671FFDCFF107A4D9E33FD02FDDB" box="[565,640,551,577]" name="Russia" pageId="18" pageNumber="19">Russia</collectingCountry>
(Arctic Siberia). NEARCTIC: Alaska (Point Barrow).
</paragraph>
</subSubSection>
</treatment>
</document>

373
data/7D/57/F1/7D57F10BFFC5FF8403CCFC19FAD05811.xml Normal file
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@ -0,0 +1,373 @@
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<mods:title id="1C4B30B6CA04BFBA9B543CC0DC87D80E">Cephisus siccifolia (Walker) (Hemiptera: Aphrophoridae): description of the immature and adult stages, including genital morphology, with notes on species delimitation challenges in the genus</mods:title>
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<mods:name id="8432B18E84685F864F25479274A39475" type="personal">
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<emphasis id="C78A9C0FFFC5FF8403CCFC19FEDB5DE7" bold="true" box="[151,378,943,970]" italics="true" pageId="3" pageNumber="50">Cephisus siccifolia</emphasis>
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<emphasis id="C78A9C0FFFC5FF8403CCFC42FED45A21" box="[151,373,1012,1036]" italics="true" pageId="3" pageNumber="50">Aphrophora siccifolia</emphasis>
Walker, 1851: 698
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<emphasis id="C78A9C0FFFC5FF8403CCFBA4FEC05A07" box="[151,353,1042,1066]" italics="true" pageId="3" pageNumber="50">Cephisus siccifolius</emphasis>
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[sic]
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<emphasis id="C78A9C0FFFC5FF8403CCFB87FE2E5A64" box="[151,399,1073,1097]" italics="true" pageId="3" pageNumber="50">Sphodroscarta siccifolia</emphasis>
Stål, 1869: 18
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<emphasis id="C78A9C0FFFC5FF8403CCFB23FED85A82" bold="true" box="[151,377,1173,1199]" pageId="3" pageNumber="50">Material examined</emphasis>
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:
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, L. Alasmar, leg.”
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leg”
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<collectionCode id="93EFD8D8FFC5FF840736FA90FB3C5B6D" box="[1133,1181,1318,1344]" country="Australia" name="Australian Museum" pageId="3" pageNumber="50" type="Museum">AM</collectionCode>
,
<collectingMunicipality id="1525DA67FFC5FF8407F2FA90FB705B6D" box="[1193,1233,1318,1344]" pageId="3" pageNumber="50">Rio</collectingMunicipality>
\
<location id="F02116C6FFC5FF8407B1FA90FAF25B6D" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF8407B1FA90FAF25B6D" box="[1258,1363,1318,1344]" country="Brazil" county="Ipixuna" latitude="-7.168333" longLatPrecision="21" longitude="-70.81833" municipality="Rio" name="Gregorio" pageId="3" pageNumber="50" stateProvince="Acre">Gregório</location>
;
<location id="F02116C6FFC5FF840604FA90FF705B49" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF840604FA90FF705B49" country="Brazil" county="Ipixuna" latitude="-7.168333" longLatPrecision="21" longitude="-70.81833" municipality="Rio" name="Com. Lago" pageId="3" pageNumber="50" stateProvince="Acre">Com. Lago</location>
\
<location id="F02116C6FFC5FF8403BDFAFCFE9A5B49" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF8403BDFAFCFE9A5B49" box="[230,315,1354,1380]" country="Brazil" county="Ipixuna" latitude="-7.168333" longLatPrecision="21" longitude="-70.81833" municipality="Rio" name="Grande" pageId="3" pageNumber="50" stateProvince="Acre">Grande</location>
,
<location id="F02116C6FFC5FF84021DFAFCFE025B49" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF84021DFAFCFE025B49" box="[326,419,1354,1380]" country="Brazil" county="Ipixuna" latitude="-7.168333" longLatPrecision="21" longitude="-70.81833" municipality="Rio" name="Seringal" pageId="3" pageNumber="50" stateProvince="Acre">Seringal</location>
\
<location id="F02116C6FFC5FF8402E2FAFCFDB15B49" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF8402E2FAFCFDB15B49" box="[441,528,1354,1380]" country="Brazil" county="Ipixuna" latitude="-7.168333" longLatPrecision="21" longitude="-70.81833" municipality="Rio" name="Recreio" pageId="3" pageNumber="50" stateProvince="Acre">Recreio</location>
\
<geoCoordinate id="90CA26DAFFC5FF84017DFAFCFD285B49" box="[550,649,1354,1380]" degrees="7" direction="south" minutes="10" orientation="latitude" pageId="3" pageNumber="50" precision="15" seconds="6" value="-7.168333">7°10'6&quot;S</geoCoordinate>
<geoCoordinate id="90CA26DAFFC5FF8401CBFAFCFCAF5B49" box="[656,782,1354,1380]" degrees="70" direction="west" minutes="49" orientation="longitude" pageId="3" pageNumber="50" precision="15" seconds="6" value="-70.81833">70°49'6&quot;W</geoCoordinate>
” “
<quantity id="3206EDF8FFC5FF840071FAFCFCCD5B48" box="[810,876,1354,1381]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="50" unit="m" value="145.0">
<elevation id="7ED3A72EFFC5FF840071FAFCFCCD5B48" box="[810,876,1354,1381]" metricMagnitude="2" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="50" unit="m" value="145.0">145m</elevation>
</quantity>
-
<collectingMethod id="2CBF380AFFC5FF8400D8FAFCFC485B49" box="[899,1001,1354,1380]" pageId="3" pageNumber="50">light trap</collectingMethod>
\
<date id="814066DDFFC5FF8400A5FAFCFB2F5B49" box="[1022,1166,1354,1380]" pageId="3" pageNumber="50" value="2011-05-18" valueMax="2011-05-23" valueMin="2011-05-18">
<collectingDate id="91049F35FFC5FF8400A5FAFCFB2F5B49" box="[1022,1166,1354,1380]" pageId="3" pageNumber="50" value="2011-05-18" valueMax="2011-05-23" valueMin="2011-05-18">18-23V.2011</collectingDate>
</date>
\ Cavichioli, Gonçalves &amp; \ Takiya”
</materialsCitation>
;
<materialsCitation id="45964A40FFC5FF840277FAD8FB9E5BA5" box="[300,1087,1390,1417]" collectorName="M. Alvarenga" country="Brazil" elevation="580" location="Encruzilhada" pageId="3" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<specimenCount id="E3F88B94FFC5FF840277FAD8FEEE5BA5" box="[300,335,1390,1416]" count="1" pageId="3" pageNumber="50" type="male">1♂</specimenCount>
“
<location id="F02116C6FFC5FF84023AFAD8FE565BA5" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF84023AFAD8FE565BA5" box="[353,503,1390,1416]" country="Brazil" name="Encruzilhada" pageId="3" pageNumber="50" stateProvince="Bahia">Encruzilhada</location>
-
<collectingRegion id="373A8EFFFFC5FF84014BFAD8FDF05BA5" box="[528,593,1390,1416]" country="Brazil" name="Bahia" pageId="3" pageNumber="50">Bahia</collectingRegion>
\
<collectingCountry id="8DE9008DFFC5FF840132FAD8FD0E5BA5" box="[617,687,1390,1416]" name="Brazil" pageId="3" pageNumber="50">Brasil</collectingCountry>
,
<quantity id="3206EDF8FFC5FF8401E0FAD8FD5F5BA4" box="[699,766,1390,1417]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="50" unit="m" value="580.0">
<elevation id="7ED3A72EFFC5FF8401E0FAD8FD5F5BA4" box="[699,766,1390,1417]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="50" unit="m" value="580.0">580m</elevation>
</quantity>
, XI/74 \
<collectorName id="580B25CBFFC5FF840038FAD8FC5F5BA5" box="[867,1022,1390,1416]" pageId="3" pageNumber="50">M. Alvarenga</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="45964A40FFC5FF840710FAD8FAB55B81" collectingDate="2016-10-01" collectingDateMax="2016-10-31" collectingDateMin="2016-10-01" collectionCode="MT, R" collectorName="O. A. Correa" country="Brazil" county="Chapada" elevation="698" latitude="-15.4372225" location="Aldeia Velha" longLatPrecision="21" longitude="-55.758335" municipality="Guimaraes" pageId="3" pageNumber="50" specimenCount="1" specimenCount-male="1">
<specimenCount id="E3F88B94FFC5FF840710FAD8FBCF5BA5" box="[1099,1134,1390,1416]" count="1" pageId="3" pageNumber="50" type="male">1♂</specimenCount>
“
<collectingCountry id="8DE9008DFFC5FF8407DAFAD8FB685BA5" box="[1153,1225,1390,1416]" name="Brazil" pageId="3" pageNumber="50">Brasil</collectingCountry>
,
<collectionCode id="93EFD8D8FFC5FF84078FFAD8FB5E5BA5" box="[1236,1279,1390,1416]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="3" pageNumber="50">MT</collectionCode>
,
<collectingCounty id="1C203891FFC5FF840650FAD8FACC5BA5" box="[1291,1389,1390,1416]" pageId="3" pageNumber="50">Chapada</collectingCounty>
dos \
<collectingMunicipality id="1525DA67FFC5FF8403F3FA24FE855B81" box="[168,292,1426,1452]" pageId="3" pageNumber="50">Guimarães</collectingMunicipality>
,
<location id="F02116C6FFC5FF84026BFA24FE655B81" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF84026BFA24FE655B81" box="[304,452,1426,1452]" country="Brazil" county="Chapada" latitude="-15.4372225" longLatPrecision="21" longitude="-55.758335" municipality="Guimaraes" name="Aldeia Velha" pageId="3" pageNumber="50">Aldeia Velha</location>
\
<geoCoordinate id="90CA26DAFFC5FF840285FA24FDFC5B80" box="[478,605,1426,1453]" degrees="15" direction="south" minutes="26" orientation="latitude" pageId="3" pageNumber="50" precision="15" seconds="14" value="-15.4372225">15°26'14&quot;S</geoCoordinate>
<geoCoordinate id="90CA26DAFFC5FF84013CFA24FD505B80" box="[615,753,1426,1453]" degrees="55" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="50" precision="15" seconds="30" value="-55.758335">55°45'30&quot;W</geoCoordinate>
\
<quantity id="3206EDF8FFC5FF840050FA24FCEB5B81" box="[779,842,1426,1452]" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" pageId="3" pageNumber="50" unit="m" value="698.0">
<elevation id="7ED3A72EFFC5FF840050FA24FCEB5B81" box="[779,842,1426,1452]" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" pageId="3" pageNumber="50" unit="m" value="698.0">698m</elevation>
</quantity>
<date id="814066DDFFC5FF84000FFA24FC435B81" box="[852,994,1426,1452]" pageId="3" pageNumber="50" value="2016-10-01" valueMax="2016-10-31" valueMin="2016-10-01">
<collectingDate id="91049F35FFC5FF84000FFA24FC435B81" box="[852,994,1426,1452]" pageId="3" pageNumber="50" value="2016-10-01" valueMax="2016-10-31" valueMin="2016-10-01">1-31.X.2016</collectingDate>
</date>
Luz \ P.
<collectionCode id="93EFD8D8FFC5FF840715FA24FBC55B81" box="[1102,1124,1426,1452]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="50">R</collectionCode>
.
<collectorName id="580B25CBFFC5FF84072AFA24FAAB5B81" box="[1137,1290,1426,1452]" pageId="3" pageNumber="50">O. A. Correa</collectorName>
”
</materialsCitation>
;
<materialsCitation id="45964A40FFC5FF840679FA24FD6E5BFD" collectingDate="1975-10" collectorName="M. Alvarenga" country="Brazil" location="Sinop" pageId="3" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso">
<specimenCount id="E3F88B94FFC5FF840679FA24FAE45B81" box="[1314,1349,1426,1452]" count="1" pageId="3" pageNumber="50" type="male">1♂</specimenCount>
“
<location id="F02116C6FFC5FF840602FA24FA3C5B81" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF840602FA24FA3C5B81" box="[1369,1437,1426,1452]" country="Brazil" name="Sinop" pageId="3" pageNumber="50" stateProvince="Mato Grosso">Sinop</location>
<collectingRegion id="373A8EFFFFC5FF8403CCFA00FE885BFD" box="[151,297,1462,1488]" country="Brazil" name="Mato Grosso" pageId="3" pageNumber="50">Mato Grosso</collectingRegion>
\
<collectingCountry id="8DE9008DFFC5FF840264FA00FE235BFD" box="[319,386,1462,1488]" name="Brazil" pageId="3" pageNumber="50">Brasil</collectingCountry>
<date id="814066DDFFC5FF8402D2FA00FE7F5BFC" box="[393,478,1462,1489]" pageId="3" pageNumber="50" value="1975-10">
<collectingDate id="91049F35FFC5FF8402D2FA00FE7F5BFC" box="[393,478,1462,1489]" pageId="3" pageNumber="50" value="1975-10">X-1975</collectingDate>
</date>
\
<collectorName id="580B25CBFFC5FF8402AEFA00FD315BFD" box="[501,656,1462,1488]" pageId="3" pageNumber="50">M. Alvarenga</collectorName>
leg.”
</materialsCitation>
;
<materialsCitation id="45964A40FFC5FF840180FA00FB1E5BD9" collectingDate="2009-02-19" collectionCode="R" collectorName="Grossi" country="Brazil" county="Dourados" latitude="-22.209446" location="Coqueiro" longLatPrecision="21" longitude="-54.91278" municipality="Reserva Florestal Faz." pageId="3" pageNumber="50" specimenCount="1" specimenCount-male="1">
<specimenCount id="E3F88B94FFC5FF840180FA00FD5F5BFD" box="[731,766,1462,1488]" count="1" pageId="3" pageNumber="50" type="male">1♂</specimenCount>
“
<collectingCountry id="8DE9008DFFC5FF84004BFA00FCF95BFD" box="[784,856,1462,1488]" name="Brazil" pageId="3" pageNumber="50">Brasil</collectingCountry>
, MS,
<collectingCounty id="1C203891FFC5FF8400C1FA00FBAB5BFD" box="[922,1034,1462,1488]" pageId="3" pageNumber="50">Dourados</collectingCounty>
, \
<collectingMunicipality id="1525DA67FFC5FF84077FFA00FABF5BFD" box="[1060,1310,1462,1488]" pageId="3" pageNumber="50">Reserva Florestal Faz.</collectingMunicipality>
\
<location id="F02116C6FFC5FF84066FFA00FA3D5BFD" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF84066FFA00FA3D5BFD" box="[1332,1436,1462,1488]" country="Brazil" county="Dourados" latitude="-22.209446" longLatPrecision="21" longitude="-54.91278" municipality="Reserva Florestal Faz." name="Coqueiro" pageId="3" pageNumber="50">Coqueiro</location>
<geoCoordinate id="90CA26DAFFC5FF8403CCFA6CFEB75BD9" box="[151,278,1498,1524]" degrees="22" direction="south" minutes="12" orientation="latitude" pageId="3" pageNumber="50" precision="15" seconds="34" value="-22.209446">22°12'34&quot;S</geoCoordinate>
\
<geoCoordinate id="90CA26DAFFC5FF84026FFA6CFE1F5BD8" box="[308,446,1498,1525]" degrees="54" direction="west" minutes="54" orientation="longitude" pageId="3" pageNumber="50" precision="15" seconds="46" value="-54.91278">54°54'46&quot;W</geoCoordinate>
<date id="814066DDFFC5FF840292FA6CFD9F5BD9" box="[457,574,1498,1524]" pageId="3" pageNumber="50" value="2009-02-19">
<collectingDate id="91049F35FFC5FF840292FA6CFD9F5BD9" box="[457,574,1498,1524]" pageId="3" pageNumber="50" value="2009-02-19">19.II.2009</collectingDate>
</date>
A. \ Paladini, D.
<collectionCode id="93EFD8D8FFC5FF840052FA6CFCBE5BD9" box="[777,799,1498,1524]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="50">R</collectionCode>
. Parizotto, P. C. \
<collectorName id="580B25CBFFC5FF8400A0FA6CFBE55BD9" box="[1019,1092,1498,1524]" pageId="3" pageNumber="50">Grossi</collectorName>
leg. Luz”
</materialsCitation>
;
<materialsCitation id="45964A40FFC5FF840795FA6CFB155835" collectingDate="2013-04-01" collectingDateMax="2013-04-30" collectingDateMin="2013-04-01" collectorName="M. Savaris &amp; S. Lampert" country="Brazil" location="Monte Belo" municipality="Chapeco" pageId="3" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina">
<specimenCount id="E3F88B94FFC5FF840795FA6CFB505BD9" box="[1230,1265,1498,1524]" count="1" pageId="3" pageNumber="50" type="male">1♂</specimenCount>
“
<collectingCountry id="8DE9008DFFC5FF84065CFA6CFAEE5BD9" box="[1287,1359,1498,1524]" name="Brazil" pageId="3" pageNumber="50">Brasil</collectingCountry>
,
<collectingRegion id="373A8EFFFFC5FF840605FA6CFF585835" country="Brazil" name="Santa Catarina" pageId="3" pageNumber="50">Santa Catarina</collectingRegion>
, \
<collectingMunicipality id="1525DA67FFC5FF84024FFA48FED85835" box="[276,377,1534,1560]" pageId="3" pageNumber="50">Chapecó</collectingMunicipality>
,
<location id="F02116C6FFC5FF8402DEFA48FDAA5835" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF8402DEFA48FDAA5835" box="[389,523,1534,1560]" country="Brazil" municipality="Chapeco" name="Monte Belo" pageId="3" pageNumber="50" stateProvince="Santa Catarina">Monte Belo</location>
\
<date id="814066DDFFC5FF840179FA48FD1C5835" box="[546,701,1534,1560]" pageId="3" pageNumber="50" value="2013-04-01" valueMax="2013-04-30" valueMin="2013-04-01">
<collectingDate id="91049F35FFC5FF840179FA48FD1C5835" box="[546,701,1534,1560]" pageId="3" pageNumber="50" value="2013-04-01" valueMax="2013-04-30" valueMin="2013-04-01">01-30.iv.2013</collectingDate>
</date>
\ (Malaise)
<collectorName id="580B25CBFFC5FF84001CFA48FC655835" box="[839,964,1534,1560]" pageId="3" pageNumber="50">M. Savaris</collectorName>
, &amp; \
<collectorName id="580B25CBFFC5FF8400A6FA48FBDA5835" box="[1021,1147,1534,1560]" pageId="3" pageNumber="50">S. Lampert</collectorName>
leg”
</materialsCitation>
;
<materialsCitation id="45964A40FFC5FF84079BFA48FACC5811" collectingDate="2016-11-21" collectingDateMax="2016-11-24" collectingDateMin="2016-11-21" collectorName="A. C. Domahovski" country="Brazil" elevation="1000" latitude="-24.561388" location="Guartela" longLatPrecision="21" longitude="-50.25722" municipality="Tibagi" pageId="3" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
<specimenCount id="E3F88B94FFC5FF84079BFA48FB425835" box="[1216,1251,1534,1560]" count="1" pageId="3" pageNumber="50" type="male">1♂</specimenCount>
“
<collectingCountry id="8DE9008DFFC5FF8407AEFA48FA9C5835" box="[1269,1341,1534,1560]" name="Brazil" pageId="3" pageNumber="50">Brasil</collectingCountry>
,
<collectingRegion id="373A8EFFFFC5FF840612FA48FA395835" box="[1353,1432,1534,1560]" country="Brazil" name="Parana" pageId="3" pageNumber="50">Paraná</collectingRegion>
,
<collectingMunicipality id="1525DA67FFC5FF8403CCF994FF435811" box="[151,226,1570,1596]" pageId="3" pageNumber="50">Tibagi</collectingMunicipality>
, \ P. E. do
<location id="F02116C6FFC5FF840202F994FE1D5811" LSID="urn:lsid:plazi:treatment:7D57F10BFFC5FF8403CCFC19FAD05811:F02116C6FFC5FF840202F994FE1D5811" box="[345,444,1570,1596]" country="Brazil" latitude="-24.561388" longLatPrecision="21" longitude="-50.25722" municipality="Tibagi" name="Guartela" pageId="3" pageNumber="50" stateProvince="Parana">Guartelá</location>
,
<quantity id="3206EDF8FFC5FF84029CF994FDB55811" box="[455,532,1570,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="50" unit="m" value="1000.0">
<elevation id="7ED3A72EFFC5FF84029CF994FDB55811" box="[455,532,1570,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="50" unit="m" value="1000.0">1000m</elevation>
</quantity>
\
<geoCoordinate id="90CA26DAFFC5FF840170F994FD0B5811" box="[555,682,1570,1596]" degrees="24" direction="south" minutes="33" orientation="latitude" pageId="3" pageNumber="50" precision="15" seconds="41" value="-24.561388">24°33'41&quot;S</geoCoordinate>
<geoCoordinate id="90CA26DAFFC5FF8401EAF994FC9B5810" box="[689,826,1570,1597]" degrees="50" direction="west" minutes="15" orientation="longitude" pageId="3" pageNumber="50" precision="15" seconds="26" value="-50.25722">50°15'26&quot;W</geoCoordinate>
\
<date id="814066DDFFC5FF84000BF994FC545811" box="[848,1013,1570,1596]" pageId="3" pageNumber="50" value="2016-11-21" valueMax="2016-11-24" valueMin="2016-11-21">
<collectingDate id="91049F35FFC5FF84000BF994FC545811" box="[848,1013,1570,1596]" pageId="3" pageNumber="50" value="2016-11-21" valueMax="2016-11-24" valueMin="2016-11-21">21-24.XI.2016</collectingDate>
</date>
Sweep \
<collectorName id="580B25CBFFC5FF840700F994FA8C5811" box="[1115,1325,1570,1596]" pageId="3" pageNumber="50">A. C. Domahovski</collectorName>
leg.”
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>