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<document id="CD110E6BF9C9BD3C074CAE3A93BC0F6C" ID-DOI="10.5281/zenodo.176375" ID-GBIF-Dataset="4ae7ea7a-e406-4c68-9777-ec29e3521997" ID-ISSN="1175-5326" ID-Zenodo-Dep="176375" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459832640299" checkinUser="plazi" docAuthor="Williams, Jason D." docDate="2007" docId="03AF87BA6E204D7EFF7AFA822659FE81" docLanguage="en" docName="zt01459p035.pdf" docOrigin="Zootaxa 1459" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Scolelepis hutchingsae Dauer 1985" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FF96FFC26E304D6AFFEDFF9D2452FFD3" masterDocTitle="New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on palp ciliation patterns of the genus Scolelepis" masterLastPageNumber="35" masterPageNumber="1" pageNumber="17" updateTime="1698223090912" updateUser="plazi">
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<mods:title id="5AB62E8441270F3791DAB049049A6BFE">New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on palp ciliation patterns of the genus Scolelepis</mods:title>
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<mods:namePart id="13542DE9416982BF8A06FD5C20A907D2">Williams, Jason D.</mods:namePart>
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<mods:date id="09D12123F0EE2E45D1842BCB366FEE3A">2007</mods:date>
<mods:detail id="947B73789FC65BF58DA54C96CA51C26B" type="volume">
<mods:number id="9357FDF33112D991F175D28893295B72">1459</mods:number>
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<mods:classification id="465BD677A3F6BD2093E2259412564830">journal article</mods:classification>
<mods:identifier id="DCF9EE737D46F0CE5878055343864223" type="DOI">10.5281/zenodo.176375</mods:identifier>
<mods:identifier id="19096D583B56E2318E69B217A3EABDC1" type="GBIF-Dataset">4ae7ea7a-e406-4c68-9777-ec29e3521997</mods:identifier>
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<treatment id="03AF87BA6E204D7EFF7AFA822659FE81" ID-DOI="http://doi.org/10.5281/zenodo.6242421" ID-GBIF-Taxon="119358180" ID-Zenodo-Dep="6242421" LSID="urn:lsid:plazi:treatment:03AF87BA6E204D7EFF7AFA822659FE81" httpUri="http://treatment.plazi.org/id/03AF87BA6E204D7EFF7AFA822659FE81" lastPageId="20" lastPageNumber="21" pageId="16" pageNumber="17">
<subSubSection id="C31C65276E204D7AFF7AFA822566FA8F" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BB936AC6E204D7AFF7AFA822615FAEA" blockId="16.[151,583,1311,1372]" box="[151,583,1311,1337]" pageId="16" pageNumber="17">
<heading id="D0F181C06E204D7AFF7AFA822615FAEA" bold="true" box="[151,583,1311,1337]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C064D2F6E204D7AFF7AFA822615FAEA" ID-CoL="4VRP6" authority="Dauer, 1985" authorityName="Dauer" authorityYear="1985" box="[151,583,1311,1337]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E204D7AFF7AFA822615FAEA" bold="true" box="[151,583,1311,1337]" pageId="16" pageNumber="17">
<emphasis id="B972EABE6E204D7AFF7AFA8225F8FAEA" bold="true" box="[151,426,1311,1337]" italics="true" pageId="16" pageNumber="17">Scolelepis hutchingsae</emphasis>
<bibRefCitation id="EF974B5D6E204D7AFE5FFA822615FAEA" author="Dauer" box="[434,583,1311,1337]" pageId="16" pageNumber="32" refString="Dauer, D. M. (1985) A new species of Scolelepis (Polychaeta: Spionidae) from Lizard Island, Australia. Proceedings of the Biological Society of Washington, 98, 678 - 681." type="journal article" year="1985">Dauer, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB936AC6E204D7AFF7AFADF2566FA8F" blockId="16.[151,583,1311,1372]" box="[151,308,1346,1372]" pageId="16" pageNumber="17">
(
<figureCitation id="133D2A296E204D7AFF72FADF2579FA8F" box="[159,299,1346,1372]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="18.[151,255,1132,1156]" captionStartId-1="19.[151,255,1832,1856]" captionStartId-2="20.[151,255,1094,1118]" captionTargetBox-1="[288,1334,379,1806]" captionTargetBox-2="[288,1311,382,1077]" captionTargetId-1="figure@19.[148,1439,374,1828]" captionTargetId-2="figure@20.[148,1439,375,1090]" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." captionText-1="FIGURE 12. Scolelepis hutchingsae A – G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A – B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm." captionText-2="FIGURE 13. Scolelepis hutchingsae A – E, voucher specimens (USNM 1096812) SEM micrographs. A, setigers 2 – 4 (left to right), lateral view (arrowhead indicates lateral organ); B, setigers 13 – 21 (right to left), lateral view (arrowhead indicates lateral organ); C, setiger 20 neuropodial hooded hooks; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view. Scale bars: A, D = 100 µm; B = 250 µm; C = 25 µm; E = 250 µm." httpUri-1="https://zenodo.org/record/176382/files/figure.png" httpUri-2="https://zenodo.org/record/176383/files/figure.png" pageId="16" pageNumber="17">Figs. 11–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E204D7AFF7AFA0F2700FA7A" box="[151,850,1426,1449]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BB936AC6E204D7AFF7AFA0F2700FA7A" blockId="16.[151,850,1426,1449]" box="[151,850,1426,1449]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB1611826E204D7AFF7AFA0F2700FA7A" box="[151,850,1426,1449]" pageId="16" pageNumber="17">
<taxonomicName id="4C064D2F6E204D7AFF7AFA0F2683FA7A" ID-CoL="4VRP6" authority="Dauer, 1985: 678" authorityName="Dauer" authorityPageNumber="678" authorityYear="1985" box="[151,721,1426,1449]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="hutchingsae" subGenus="Scolelepis">
<emphasis id="B972EABE6E204D7AFF7AFA0F265DFA7A" box="[151,527,1426,1449]" italics="true" pageId="16" pageNumber="17">Scolelepis (Scolelepis) hutchingsae</emphasis>
<treatmentCitation id="0AA710BD6E204D7AFDF8FA0F2683FA7A" author="Dauer" box="[533,721,1426,1449]" page="678" pageId="16" pageNumber="17" year="1985">
<bibRefCitation id="EF974B5D6E204D7AFDF8FA0F26CEFA7A" author="Dauer" box="[533,668,1426,1449]" pageId="16" pageNumber="32" refString="Dauer, D. M. (1985) A new species of Scolelepis (Polychaeta: Spionidae) from Lizard Island, Australia. Proceedings of the Biological Society of Washington, 98, 678 - 681." type="journal article" year="1985">Dauer, 1985</bibRefCitation>
: 678
</treatmentCitation>
</taxonomicName>
-681,
<figureCitation id="133D2A296E204D7AFCE1FA0F2719FA7A" box="[780,843,1426,1449]" captionStart="FIGURE 1" captionStartId="2.[151,255,1919,1943]" captionText="FIGURE 1. Map of the Philippines with three collecting sites indicated in the text: 1) Morong, Bataan, 2) Bacoor, Cavite (Manila Bay), 3) Boracay, Aklan." pageId="16" pageNumber="17">Fig. 1</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E204D7AFF7AFA472068F947" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BB936AC6E204D7AFF7AFA472068F947" blockId="16.[151,1436,1498,2004]" pageId="16" pageNumber="17">
<emphasis id="B972EABE6E204D7AFF7AFA47269BFA27" bold="true" box="[151,713,1498,1524]" pageId="16" pageNumber="17">Material examined. Diniwid Beach, Boracay</emphasis>
,
<geoCoordinate id="EE32506B6E204D7AFD36FA472713FA27" box="[731,833,1498,1524]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="12.0">11°60´N</geoCoordinate>
,
<geoCoordinate id="EE32506B6E204D7AFCBCFA472796FA27" box="[849,964,1498,1524]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="121.9">121°54´E</geoCoordinate>
, sandy beach,
<date id="FFB8106C6E204D7AFB9AFA472143FA27" box="[1143,1297,1498,1524]" pageId="16" pageNumber="17" value="1999-04-13">13 Apr 1999</date>
(four complete specimens, four anterior ends (two with regenerating posterior ends), three posterior ends in alcohol,
<collectionCode id="ED17AE696E204D7AFF7AF9B724BEF997" LSID="urn:lsid:biocol.org:col:34871" box="[151,236,1578,1604]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1096811).—
<emphasis id="B972EABE6E204D7AFE66F9B7271BF997" bold="true" box="[395,841,1578,1604]" pageId="16" pageNumber="17">
<collectingCountry id="F311763C6E204D7AFE66F9B7264BF997" box="[395,537,1578,1604]" name="Philippines" pageId="16" pageNumber="17">Philippines</collectingCountry>
, Boracay, White Beach
</emphasis>
,
<geoCoordinate id="EE32506B6E204D7AFCB1F9B72790F997" box="[860,962,1578,1604]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="11.983334">11°59´N</geoCoordinate>
,
<geoCoordinate id="EE32506B6E204D7AFC3EF9B72014F997" box="[979,1094,1578,1604]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="121.916664">121°55´E</geoCoordinate>
, sandy beach,
<date id="FFB8106C6E204D7AFB13F9B721CEF997" box="[1278,1436,1578,1604]" pageId="16" pageNumber="17" value="1999-04-14">14 Apr 1999</date>
(three complete specimens, three anterior ends, four posterior ends in alcohol,
<collectionCode id="ED17AE696E204D7AFBDCF9CF2038F9BF" box="[1073,1130,1618,1644]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="16" pageNumber="17">ZRC</collectionCode>
2006.0224; one complete specimen on two SEM stubs, one pair of palps on SEM stub,
<collectionCode id="ED17AE696E204D7AFC8BF9E727E9F947" LSID="urn:lsid:biocol.org:col:34871" box="[870,955,1658,1684]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17">USNM</collectionCode>
1096812).
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E204D7BFF2AF93F212CFB92" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BB936AC6E204D7AFF2AF93F2634F8DF" blockId="16.[151,1436,1498,2004]" pageId="16" pageNumber="17">
<emphasis id="B972EABE6E204D7AFF2AF93F250DF96F" bold="true" box="[199,351,1698,1724]" pageId="16" pageNumber="17">Description.</emphasis>
Largest specimen of 73 setigers, 16.3-mm long, 0.8-mm wide at setiger 16. Body widest anteriorly, gradually tapering to posterior end; body nearly rectangular in cross section. Color in alcohol opaque white, no pigmentation present.
</paragraph>
<paragraph id="8BB936AC6E204D7BFF2AF88720BFFCB0" blockId="16.[151,1436,1498,2004]" lastBlockId="17.[124,1875,771,1414]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Prostomium conical, extending anteriorly to sharp point, posteriorly continuing as short, pointed caruncle, to middle of setiger 1; peristomial wings surrounding base of palps (
<figureCitation id="133D2A296E204D7AFC58F8DF2041F88F" box="[949,1043,1858,1884]" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="16" pageNumber="17">Figs. 11</figureCitation>
A–B, 12A–E). Eyes and occipital tentacle absent. Palps extending to about setiger 15, with two distinctly separated transverse rows of cilia along ventral surfaces, with long and short rows in approximately 2:1 ratio, longer rows of cilia approximately 35-µm long, shorter rows approximately 10-µm long and elevated on 5–10 µm lobes (oriented medially) Species Author Occipital Neuropodial Neuropodial Notopodial Notosetae Branchial fusion Notes tentacle hooks, hooks, # of hooks setiger 1
</paragraph>
<caption id="DF7966246E214D7BFF91FC9E208FFCC9" box="[124,1245,771,794]" pageId="17" pageNumber="18">
<paragraph id="8BB936AC6E214D7BFF91FC9E208FFCC9" blockId="17.[124,1875,771,1414]" box="[124,1245,771,794]" pageId="17" pageNumber="18">
<emphasis id="B972EABE6E214D7BFF91FC9E24BCFCC9" bold="true" box="[124,238,771,794]" pageId="17" pageNumber="18">TABLE 1.</emphasis>
Taxonomic characters of species in the genus
<taxonomicName id="4C064D2F6E214D7BFD3FFC9E2769FCC9" box="[722,827,771,794]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="genus">
<emphasis id="B972EABE6E214D7BFD3FFC9E2769FCC9" box="[722,827,771,794]" italics="true" pageId="17" pageNumber="18">Scolelepis</emphasis>
</taxonomicName>
described since Hutchings et al. (1998).
</paragraph>
</caption>
<paragraph id="8BB936AC6E214D7BFD04FCF42797FCAD" blockId="17.[124,1875,771,1414]" box="[745,965,873,894]" pageId="17" pageNumber="18">begin on setiger teeth</paragraph>
<paragraph id="8BB936AC6E214D7BFF79FC1A231AFC3F" blockId="17.[124,1875,771,1414]" pageId="17" pageNumber="18">
<taxonomicName id="4C064D2F6E214D7BFF79FC1A24B1FC4F" box="[148,227,903,924]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E214D7BFF79FC1A24B1FC4F" box="[148,227,903,924]" italics="true" pageId="17" pageNumber="18">alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E214D7BFF05FC1A254FFC4F" bold="true" box="[232,285,903,924]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A24157C56E214D7BFF05FC1A254FFC4F" box="[232,285,903,924]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
present study Absent
<date id="FFB8106C6E214D7BFD04FC1A27F6FC4F" box="[745,932,903,924]" pageId="17" pageNumber="18" value="1932-02-29">29-32 2</date>
Present Present Partially fused (tips free) notopodial postsetal lamellae with up to 9 lobes by setiger 5; neuropodial postsetal lamellae with up to 3 lobes by setiger 5
</paragraph>
<paragraph id="8BB936AC6E214D7BFF79FC682365FBD9" blockId="17.[124,1875,771,1414]" box="[148,1847,1012,1034]" pageId="17" pageNumber="18">
<emphasis id="B972EABE6E214D7BFF79FC6824BCFBD9" box="[148,238,1013,1034]" italics="true" pageId="17" pageNumber="18">anakenae</emphasis>
Rozbaczylo &amp; Castilla 1988 Absent 35-37 2 Absent Present Partially fused (tips free) Not cited in Hutchings
<emphasis id="B972EABE6E214D7BF8E5FC682365FBD9" box="[1800,1847,1013,1034]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
</paragraph>
<paragraph id="8BB936AC6E214D7BF9C3FB92220FFBF7" blockId="17.[124,1875,771,1414]" box="[1582,1629,1039,1060]" pageId="17" pageNumber="18">1998</paragraph>
<paragraph id="8BB936AC6E214D7BFF79FBB1212CFB92" blockId="17.[124,1875,771,1414]" box="[148,1406,1068,1089]" pageId="17" pageNumber="18">
<emphasis id="B972EABE6E214D7BFF79FBB1249AFB92" box="[148,200,1068,1089]" italics="true" pageId="17" pageNumber="18">dicha</emphasis>
Hutchings, Frouin &amp; Hily, 1998 Absent
<date id="FFB8106C6E214D7BFD04FBB127F6FB92" box="[745,932,1068,1089]" pageId="17" pageNumber="18" value="1932-02-26">26-32 2</date>
Absent Absent Fused basally
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E214D7BFF79FBD32301FB61" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BB936AC6E214D7BFF79FBD32365FBB0" blockId="17.[124,1875,771,1414]" box="[148,1847,1101,1123]" pageId="17" pageNumber="18">
<taxonomicName id="4C064D2F6E214D7BFF79FBD3269FFBB1" ID-CoL="4VRMZ" authority="Eibye-Jacobsen, 1997 Absent" authorityName="Eibye-Jacobsen, 1997 Absent" box="[148,717,1101,1123]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="laciniata">
<emphasis id="B972EABE6E214D7BFF79FBD324B5FBB0" box="[148,231,1102,1123]" italics="true" pageId="17" pageNumber="18">laciniata</emphasis>
<bibRefCitation id="EF974B5D6E214D7BFEB6FBD0267AFBB1" author="Eibye-Jacobsen" box="[347,552,1101,1122]" pageId="17" pageNumber="32" refString="Eibye-Jacobsen, D. (1997) A new species of Scolelepis (Polychaeta: Spionidae), highly abundant on the sand beaches of western Phuket Island, Thailand. Bulletin of Marine Science, 60, 240 - 251." type="journal article" year="1997">Eibye-Jacobsen, 1997</bibRefCitation>
Absent
</taxonomicName>
<date id="FFB8106C6E214D7BFD04FBD027E5FBB1" box="[745,951,1101,1122]" pageId="17" pageNumber="18" value="1924-02-26" valueMax="1924-03-26" valueMin="1924-02-26">24-26 2-3</date>
Absent Absent Partially fused (tips free) Not cited in Hutchings
<emphasis id="B972EABE6E214D7BF8E5FBD32365FBB0" box="[1800,1847,1102,1123]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
</paragraph>
<paragraph id="8BB936AC6E214D7BF9C3FBF52301FB61" blockId="17.[124,1875,771,1414]" pageId="17" pageNumber="18">1998; notopodial postsetal lamellae with up to 5 lobes by setiger 18</paragraph>
</subSubSection>
<subSubSection id="C31C65276E214D7BFF79FB2621B5FA99" pageId="17" pageNumber="18" type="description">
<paragraph id="8BB936AC6E214D7BFF79FB26218DFB1C" blockId="17.[124,1875,771,1414]" box="[148,1503,1210,1232]" pageId="17" pageNumber="18">
<taxonomicName id="4C064D2F6E214D7BFF79FB262545FB03" box="[148,279,1211,1232]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="magnicornuta" status="sp. nov.">
<emphasis id="B972EABE6E214D7BFF79FB262545FB03" box="[148,279,1211,1232]" italics="true" pageId="17" pageNumber="18">magnicornuta</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E214D7BFEF1FB262503FB03" bold="true" box="[284,337,1211,1232]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A24157C56E214D7BFEF1FB262503FB03" box="[284,337,1211,1232]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
present study Present
<date id="FFB8106C6E214D7BFD05FB2727F6FB1C" box="[744,932,1210,1231]" pageId="17" pageNumber="18" value="1925-02-30">25-30 2</date>
Present Present Partially fused (tips free
</paragraph>
<paragraph id="8BB936AC6E214D7BFB13FB48224AFB39" blockId="17.[124,1875,771,1414]" box="[1278,1560,1237,1258]" pageId="17" pageNumber="18">anteriorly, at base posteriorly)</paragraph>
<paragraph id="8BB936AC6E214D7BFF79FB6E2253FAD4" blockId="17.[124,1875,771,1414]" box="[148,1537,1266,1288]" pageId="17" pageNumber="18">
<taxonomicName id="4C064D2F6E214D7BFF79FB6E248FFADB" box="[148,221,1267,1288]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="magnus">
<emphasis id="B972EABE6E214D7BFF79FB6E248FFADB" box="[148,221,1267,1288]" italics="true" pageId="17" pageNumber="18">magnus</emphasis>
</taxonomicName>
Ozolin'sh, 1990 Present 34 2 Present Present Completely fused (anterior)
</paragraph>
<paragraph id="8BB936AC6E214D7BFF79FA8921B5FAFB" blockId="17.[124,1875,771,1414]" box="[148,1511,1299,1321]" pageId="17" pageNumber="18">
<emphasis id="B972EABE6E214D7BFF79FA8924B5FAFA" box="[148,231,1300,1321]" italics="true" pageId="17" pageNumber="18">marionis</emphasis>
Branch, 1998 Absent 42 2 Absent Present Partially fused (tips free)
</paragraph>
<paragraph id="8BB936AC6E214D7BFF79FAA821B5FA99" blockId="17.[124,1875,771,1414]" box="[148,1511,1333,1354]" pageId="17" pageNumber="18">
<taxonomicName id="4C064D2F6E214D7BFF79FAA82553FA99" box="[148,257,1333,1354]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="villosivaina" status="sp. nov.">
<emphasis id="B972EABE6E214D7BFF79FAA82553FA99" box="[148,257,1333,1354]" italics="true" pageId="17" pageNumber="18">villosivaina</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E214D7BFEE1FAA82513FA99" bold="true" box="[268,321,1333,1354]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A24157C56E214D7BFEE1FAA82513FA99" box="[268,321,1333,1354]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
present study Absent
<date id="FFB8106C6E214D7BFD04FAA827E5FA99" box="[745,951,1333,1354]" pageId="17" pageNumber="18" value="1925-01-29" valueMax="1925-02-29" valueMin="1925-01-29">25-29 1-2</date>
Present Present Partially fused (tips free)
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E214D7BFF79FACB222CFA55" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BB936AC6E214D7BFF79FACB2371FAB8" blockId="17.[124,1875,771,1414]" box="[148,1827,1366,1387]" pageId="17" pageNumber="18">
<taxonomicName id="4C064D2F6E214D7BFF79FACB269FFAB8" ID-CoL="4VRQ2" authority="Eibye-Jacobsen &amp; Soares, 2000 Absent" authorityName="Eibye-Jacobsen &amp; Soares, 2000 Absent" box="[148,717,1366,1387]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="vazaha">
<emphasis id="B972EABE6E214D7BFF79FACB2484FAB8" box="[148,214,1366,1387]" italics="true" pageId="17" pageNumber="18">vazaha</emphasis>
<bibRefCitation id="EF974B5D6E214D7BFEB6FACB26D6FAB8" author="Eibye-Jacobsen" box="[347,644,1366,1387]" pageId="17" pageNumber="32" refString="Eibye-Jacobsen, D. &amp; Soares, A. G. (2000) New records of Scolelepis (Polychaeta: Spionidae) from the sandy beaches of Madagascar, with the description of a new species. Bulletin of Marine Science, 67, 571 - 586." type="journal article" year="2000">Eibye-Jacobsen &amp; Soares, 2000</bibRefCitation>
Absent
</taxonomicName>
<date id="FFB8106C6E214D7BFD05FACB27F1FAB8" box="[744,931,1366,1387]" pageId="17" pageNumber="18" value="1925-03-31">25-31 3</date>
Present Present Partially fused (at base) large notopodial spines on
</paragraph>
<paragraph id="8BB936AC6E214D7BF9C3FAEC222CFA55" blockId="17.[124,1875,771,1414]" box="[1582,1662,1393,1414]" pageId="17" pageNumber="18">setiger 4</paragraph>
</subSubSection>
<subSubSection id="C31C65276E224D78FF7AFF0524A9F8B7" pageId="18" pageNumber="19" type="description">
<paragraph id="8BB936AC6E224D78FF7AFF0520C3FED1" blockId="18.[151,1436,152,498]" pageId="18" pageNumber="19">
(
<figureCitation id="133D2A296E224D78FF72FF0524ADFF61" box="[159,255,152,178]" captionStart="FIGURE 12" captionStartId="19.[151,255,1832,1856]" captionTargetBox="[288,1334,379,1806]" captionTargetId="figure@19.[148,1439,374,1828]" captionTargetPageId="19" captionText="FIGURE 12. Scolelepis hutchingsae A – G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A – B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm." httpUri="https://zenodo.org/record/176382/files/figure.png" pageId="18" pageNumber="19">Figs. 12</figureCitation>
G–H, 18D); rows of mucus secreting cells (represented by tubular necks) present proximal to transverse ciliary rows, median ciliated groove absent; base of palps with ciliary patches (
<figureCitation id="133D2A296E224D78FB9FFF5D2097FF09" box="[1138,1221,192,218]" captionStart="FIGURE 12" captionStartId="19.[151,255,1832,1856]" captionTargetBox="[288,1334,379,1806]" captionTargetId="figure@19.[148,1439,374,1828]" captionTargetPageId="19" captionText="FIGURE 12. Scolelepis hutchingsae A – G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A – B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm." httpUri="https://zenodo.org/record/176382/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
E–F). Nuchal cilia in U-shaped pattern on both sides of caruncle, posterior to the base of palps (
<figureCitation id="133D2A296E224D78FBF7FF752022FED1" box="[1050,1136,232,258]" captionStart="FIGURE 12" captionStartId="19.[151,255,1832,1856]" captionTargetBox="[288,1334,379,1806]" captionTargetId="figure@19.[148,1439,374,1828]" captionTargetPageId="19" captionText="FIGURE 12. Scolelepis hutchingsae A – G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A – B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm." httpUri="https://zenodo.org/record/176382/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
E).
</paragraph>
<paragraph id="8BB936AC6E224D78FF2AFE8D25CEFE21" blockId="18.[151,1436,152,498]" pageId="18" pageNumber="19">
Setiger 1 well developed with rounded notopodial and neuropodial postsetal lamellae, notosetae and neurosetae present. Postsetal neuropodial lamellae of setigers 1–18 rounded, becoming broadest by setiger 10 (
<figureCitation id="133D2A296E224D78FF72FEFD24ACFEA9" box="[159,254,352,378]" captionStart="FIGURE 12" captionStartId="19.[151,255,1832,1856]" captionTargetBox="[288,1334,379,1806]" captionTargetId="figure@19.[148,1439,374,1828]" captionTargetPageId="19" captionText="FIGURE 12. Scolelepis hutchingsae A – G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A – B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm." httpUri="https://zenodo.org/record/176382/files/figure.png" pageId="18" pageNumber="19">Figs. 12</figureCitation>
A–D; 13A–B), by setiger 20 lamellae forming low, broad lobe (
<figureCitation id="133D2A296E224D78FC0FFEFD2012FEA9" box="[994,1088,352,378]" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="18" pageNumber="19">Figs. 11</figureCitation>
C, 13B), lamellae of posterior setigers broadly triangular on dorsal end, rounded on ventral end with neurosetae (
<figureCitation id="133D2A296E224D78FB71FE1520A6FE71" box="[1180,1268,392,418]" captionStart="FIGURE 13" captionStartId="20.[151,255,1094,1118]" captionTargetBox="[288,1311,382,1077]" captionTargetId="figure@20.[148,1439,375,1090]" captionTargetPageId="20" captionText="FIGURE 13. Scolelepis hutchingsae A – E, voucher specimens (USNM 1096812) SEM micrographs. A, setigers 2 – 4 (left to right), lateral view (arrowhead indicates lateral organ); B, setigers 13 – 21 (right to left), lateral view (arrowhead indicates lateral organ); C, setiger 20 neuropodial hooded hooks; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view. Scale bars: A, D = 100 µm; B = 250 µm; C = 25 µm; E = 250 µm." httpUri="https://zenodo.org/record/176383/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
D–E). Lateral organs present from setiger 1 between notopodial and neuropodial postsetal lamellae continuing to posterior setigers (
<figureCitation id="133D2A296E224D78FEEDFE452506FE21" box="[256,340,472,498]" captionStart="FIGURE 13" captionStartId="20.[151,255,1094,1118]" captionTargetBox="[288,1311,382,1077]" captionTargetId="figure@20.[148,1439,375,1090]" captionTargetPageId="20" captionText="FIGURE 13. Scolelepis hutchingsae A – E, voucher specimens (USNM 1096812) SEM micrographs. A, setigers 2 – 4 (left to right), lateral view (arrowhead indicates lateral organ); B, setigers 13 – 21 (right to left), lateral view (arrowhead indicates lateral organ); C, setiger 20 neuropodial hooded hooks; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view. Scale bars: A, D = 100 µm; B = 250 µm; C = 25 µm; E = 250 µm." httpUri="https://zenodo.org/record/176383/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
A, B).
</paragraph>
<paragraph id="8BB936AC6E224D78FED4FDB2248BF9A7" blockId="18.[313,1044,559,598]" lastBlockId="18.[151,1436,1346,2012]" pageId="18" pageNumber="19">
A B C D E F G H I F-I Notosetae of setiger 1 and subsequent setigers arranged in two vertical rows of limbate capillaries (
<figureCitation id="133D2A296E224D78FA80FADF24E6FA57" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="18" pageNumber="19">
Fig.
<date id="FFB8106C6E224D78FF7AFAF724E6FA57" box="[151,180,1386,1412]" pageId="18" pageNumber="19" value="1911-01">11</date>
</figureCitation>
I), dorsalmost notosetae of these rows longer than ventral ones; 1–3 notopodial hooded hooks from setiger 18–20, with up to four hooks per fascicle in posterior setigers; notopodial hooded hooks bidentate with acute main fang and single accessory tooth (
<figureCitation id="133D2A296E224D78FDB5FA2726FEFA07" box="[600,684,1466,1492]" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="18" pageNumber="19">Fig. 11</figureCitation>
H). Neurosetae of setiger 1 and subsequent setigers in two vertical rows of limbate capillaries (
<figureCitation id="133D2A296E224D78FDE3FA7F2630FA2F" box="[526,610,1506,1532]" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="18" pageNumber="19">Fig. 11</figureCitation>
G), 1–3 neuropodial hooded hooks from setiger 17–20, typically with 5–8 hooks in middle to posterior setigers; neuropodial hooded hooks bidentate with acute main fang and single accessory tooth, neuropodial hooded hooks with longer shafts than notopodial hooded hooks (
<figureCitation id="133D2A296E224D78FACBF9AF21D7F99F" box="[1318,1413,1586,1612]" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="18" pageNumber="19">Figs. 11</figureCitation>
F, 13C).
</paragraph>
<caption id="DF7966246E224D78FF7AFBF126F3FADD" pageId="18" pageNumber="19">
<paragraph id="8BB936AC6E224D78FF7AFBF126F3FADD" blockId="18.[151,1437,1132,1294]" pageId="18" pageNumber="19">
<emphasis id="B972EABE6E224D78FF7AFBF1257AFB57" bold="true" box="[151,296,1132,1156]" pageId="18" pageNumber="19">FIGURE 11.</emphasis>
<taxonomicName id="4C064D2F6E224D78FEDFFBF1267BFB50" box="[306,553,1132,1155]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E224D78FEDFFBF1267BFB50" box="[306,553,1132,1155]" italics="true" pageId="18" pageNumber="19">Scolelepis hutchingsae</emphasis>
</taxonomicName>
A–I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A–B, D–E = 200 µm; C = 125 µm; F–I = 25 µm.
</paragraph>
</caption>
<paragraph id="8BB936AC6E224D78FF2AF91F270AF8C7" blockId="18.[151,1436,1346,2012]" pageId="18" pageNumber="19">
Branchiae from setiger 2, fused to postsetal notopodial lamellae for about half of branchial length in anterior setigers, fusion reduced to about one-third of branchial length in posterior setigers; with band of cilia along inner edge of each branchia, joined to corresponding branchiae on opposite side by two dorsal ciliary bands, anterior band broader than posterior band (
<figureCitation id="133D2A296E224D78FD32F9672764F8C7" box="[735,822,1786,1812]" captionStart="FIGURE 12" captionStartId="19.[151,255,1832,1856]" captionTargetBox="[288,1334,379,1806]" captionTargetId="figure@19.[148,1439,374,1828]" captionTargetPageId="19" captionText="FIGURE 12. Scolelepis hutchingsae A – G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A – B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm." httpUri="https://zenodo.org/record/176382/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
B).
</paragraph>
<paragraph id="8BB936AC6E224D78FF2AF8BF24A9F8B7" blockId="18.[151,1436,1346,2012]" pageId="18" pageNumber="19">
Pygidium broadly rounded, with short conical papillae surrounding dorsal anal opening (
<figureCitation id="133D2A296E224D78FB10F8BF210EF8EF" box="[1277,1372,1826,1852]" captionStart="FIGURE 11" captionStartId="18.[151,255,1132,1156]" captionText="FIGURE 11. Scolelepis hutchingsae A – I, voucher specimens (USNM 1096811). A, anterior end, dorsal view (palps missing); B, anterior end, lateral view; C, setiger 48, anterior lateral view; D, posterior setigers and pygidium, dorsal view; E, posterior setigers and pygidium, lateral view; F, neuropodial hooded hook from setiger 48, lateral view; G, neuroseta from setiger 48; H, notopodial hooded hook from setiger 48, lateral view; I, notoseta from setiger 48. Scale bars: A – B, D – E = 200 µm; C = 125 µm; F – I = 25 µm." pageId="18" pageNumber="19">Figs. 11</figureCitation>
D–E, 13D–E).
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E224D7EFF2AF8EC279CFED1" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BB936AC6E224D79FF2AF8EC206DFE81" blockId="18.[151,1436,1346,2012]" lastBlockId="19.[151,1437,152,338]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B972EABE6E224D78FF2AF8EC2512F858" bold="true" box="[199,320,1905,1931]" pageId="18" pageNumber="19">Remarks.</emphasis>
The
<collectingCountry id="F311763C6E224D78FE6DF8EF25ABF85F" box="[384,505,1906,1932]" name="Philippines" pageId="18" pageNumber="19">Philippine</collectingCountry>
specimens agree with the original description of
<taxonomicName id="4C064D2F6E224D78FBA1F8EF2108F858" box="[1100,1370,1906,1931]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E224D78FBA1F8EF2108F858" box="[1100,1370,1906,1931]" italics="true" pageId="18" pageNumber="19">Scolelepis hutchingsae</emphasis>
</taxonomicName>
from Lizard
<collectingCountry id="F311763C6E224D78FF00F807256AF867" box="[237,312,1946,1972]" name="Iceland" pageId="18" pageNumber="19">Island</collectingCountry>
,
<collectingCountry id="F311763C6E224D78FEABF80725E3F867" box="[326,433,1946,1972]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
by
<bibRefCitation id="EF974B5D6E224D78FE0EF80726D1F867" author="Dauer" box="[483,643,1946,1972]" pageId="18" pageNumber="32" refString="Dauer, D. M. (1985) A new species of Scolelepis (Polychaeta: Spionidae) from Lizard Island, Australia. Proceedings of the Biological Society of Washington, 98, 678 - 681." type="journal article" year="1985">Dauer (1985)</bibRefCitation>
including the palp morphology (
<bibRefCitation id="EF974B5D6E224D78FBFCF80720F3F867" author="Dauer" box="[1041,1185,1946,1972]" pageId="18" pageNumber="32" refString="Dauer, D. M. (1987) Systematic significance of the morphology of spionid polychaete palps. Bulletin of the Biological Society of Washington No. 7, 41 - 45." type="book chapter" year="1987">Dauer 1987</bibRefCitation>
,
<bibRefCitation id="EF974B5D6E224D78FB5DF80720BDF867" author="Dauer" box="[1200,1263,1946,1972]" pageId="18" pageNumber="32" refString="Dauer, D. M. (1994) Functional ciliary groups of the feeding palps of Spionid polychaetes. In: Dauvin, J. - C., Laubier, L. &amp; Reish, D. J. (Eds.) Actes de la 4 eme Conference internationale des Polychetes. Memoires du Museum national d'Histoire naturelle, 162, 81 - 84." type="journal article" year="1994">1994</bibRefCitation>
).
<collectingCountry id="F311763C6E224D78FAEBF8072175F867" box="[1286,1319,1946,1972]" name="American Samoa" pageId="18" pageNumber="19">As</collectingCountry>
indicated previously for
<taxonomicName id="4C064D2F6E224D78FEABF85F259AF808" box="[326,456,1986,2011]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E224D78FEABF85F259AF808" box="[326,456,1986,2011]" italics="true" pageId="18" pageNumber="19">S. alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E224D78FE3DF85C2646F808" bold="true" box="[464,532,1985,2011]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A24157C56E224D78FE3DF85C2646F808" box="[464,532,1985,2011]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C064D2F6E224D78FDC9F85F2682F808" box="[548,720,1986,2011]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E224D78FDC9F85F2682F808" box="[548,720,1986,2011]" italics="true" pageId="18" pageNumber="19">S. hutchingsae</emphasis>
</taxonomicName>
belongs to a group of five species that possess notosetae on setiger 1, notopodial hooded hooks, bidentate neuropodial hooded hooks, and lack an occipital tentacle.
<taxonomicName id="4C064D2F6E234D79FF7AFF5D25F6FF0A" box="[151,420,192,217]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E234D79FF7AFF5D25F6FF0A" box="[151,420,192,217]" italics="true" pageId="19" pageNumber="20">Scolelepis hutchingsae</emphasis>
</taxonomicName>
is unique in the genus in possessing notopodial hooded hooks that begin in the same setiger or one setiger posterior to the beginning of the neuropodial hooded hooks. In addition,
<taxonomicName id="4C064D2F6E234D79FB1DFF7521CEFED2" box="[1264,1436,232,257]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E234D79FB1DFF7521CEFED2" box="[1264,1436,232,257]" italics="true" pageId="19" pageNumber="20">S. hutchingsae</emphasis>
</taxonomicName>
has patches of cilia at the base of the palps on the dorsal side near the caruncle and nuchal organ, although these patches of cilia were not noted in specimens from
<collectingCountry id="F311763C6E234D79FCC5FEA527C1FE81" box="[808,915,312,338]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
(
<bibRefCitation id="EF974B5D6E234D79FC4EFEA52060FE81" author="Dauer" box="[931,1074,312,338]" pageId="19" pageNumber="32" refString="Dauer, D. M. (1985) A new species of Scolelepis (Polychaeta: Spionidae) from Lizard Island, Australia. Proceedings of the Biological Society of Washington, 98, 678 - 681." type="journal article" year="1985">Dauer 1985</bibRefCitation>
).
</paragraph>
<caption id="DF7966246E234D79FF7AF8B525A5F83F" httpUri="https://zenodo.org/record/176382/files/figure.png" pageId="19" pageNumber="20" targetBox="[288,1334,379,1806]" targetPageId="19">
<paragraph id="8BB936AC6E234D79FF7AF8B525A5F83F" blockId="19.[151,1436,1832,2028]" pageId="19" pageNumber="20">
<emphasis id="B972EABE6E234D79FF7AF8B52574F893" bold="true" box="[151,294,1832,1856]" pageId="19" pageNumber="20">FIGURE 12.</emphasis>
<taxonomicName id="4C064D2F6E234D79FEC1F8B4264DF893" box="[300,543,1833,1856]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E234D79FEC1F8B4264DF893" box="[300,543,1833,1856]" italics="true" pageId="19" pageNumber="20">Scolelepis hutchingsae</emphasis>
</taxonomicName>
A–G, voucher specimens (USNM 1096812) SEM micrographs. A, anterior end, lateral view (palp missing on right side); B, anterior end, dorsal view; C, anterior end, oblique en-face view; D, anterior end, lateral view; E, caruncle and nuchal organ (base of palp at top, arrowhead indicates patches of cilia on palp base shown in F); F, base of palp with patches of cilia; G, lateral view of palp; H, middle region of palp, oblique frontal view (arrowheads show two short ciliary rows on elevated lobes). Scale bars: A–B, D = 250 µm; C = 125 µm; E = 50 µm; F = 25 µm; G = 100 µm; H = 20 µm.
</paragraph>
</caption>
<paragraph id="8BB936AC6E244D7EFF2AFF05279CFED1" blockId="20.[151,1436,152,338]" pageId="20" pageNumber="21">
Two of the specimens of
<taxonomicName id="4C064D2F6E244D7EFE1CFF0526CFFF62" box="[497,669,152,177]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E244D7EFE1CFF0526CFFF62" box="[497,669,152,177]" italics="true" pageId="20" pageNumber="21">S. hutchingsae</emphasis>
</taxonomicName>
exhibited posterior ends that were regenerating. Two incomplete specimens (posterior end fragments) collected in April had eggs within the body. One specimen had five bivalve larvae in the digestive tract, each approximately 480-µm long.
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E244D7EFF2AFE922659FE81" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BB936AC6E244D7EFF2AFE922659FE81" blockId="20.[151,1436,152,338]" pageId="20" pageNumber="21">
<emphasis id="B972EABE6E244D7EFF2AFE922535FEFA" bold="true" box="[199,359,271,297]" pageId="20" pageNumber="21">Distribution.</emphasis>
Sandy beach in Boracay of the Aklan province in the
<collectingCountry id="F311763C6E244D7EFC0FFE8D2038FEF9" box="[994,1130,272,298]" name="Philippines" pageId="20" pageNumber="21">Philippines</collectingCountry>
; shallow subtidal (&lt;
<quantity id="4CFE9B496E244D7EFAB0FE8D21DCFEF9" box="[1373,1422,272,298]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="5.0">5 m</quantity>
);
<collectingCountry id="F311763C6E244D7EFF7AFEA52550FE81" box="[151,258,312,338]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
(Lizard
<collectingCountry id="F311763C6E244D7EFE8AFEA525E0FE81" box="[359,434,312,338]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
, GBR).
</paragraph>
</subSubSection>
</treatment>
</document>
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