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<document id="9116C8C24BEA58F23537CD84CD2F6B99" ID-CLB-Dataset="269786" ID-DOI="10.11646/zootaxa.5346.4.4" ID-GBIF-Dataset="f631dec3-14e8-4dab-b340-df57d7cc8c5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="8390441" ID-ZooBank="3D23DF3F-6B66-4C60-BE3D-A98EA43F01C5" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1695985560601" checkinUser="plazi" docAuthor="Tanasevitch, Andrei V." docDate="2023" docId="03A9879D977DFF9FFF62F6C4D7D87AA9" docLanguage="en" docName="zootaxa.5346.4.4.pdf" docOrigin="Zootaxa 5346 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5346.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Savignia ericola Tanasevitch 2023, new species" docType="treatment" docVersion="6" lastPageNumber="437" masterDocId="FF90FFE59772FF8EFFF5F05FD6617B78" masterDocTitle="Survey of the Ethiopian linyphiid spider fauna. I. Subfamily Erigoninae (Arachnida, Araneae, Linyphiidae)" masterLastPageNumber="442" masterPageNumber="420" pageNumber="435" updateTime="1732480545820" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="960600E398915E3135529153376E6C42">Survey of the Ethiopian linyphiid spider fauna. I. Subfamily Erigoninae (Arachnida, Araneae, Linyphiidae)</mods:title>
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<treatment id="03A9879D977DFF9FFF62F6C4D7D87AA9" ID-DOI="http://doi.org/10.5281/zenodo.8390499" ID-GBIF-Taxon="213625421" ID-Zenodo-Dep="8390499" LSID="urn:lsid:plazi:treatment:03A9879D977DFF9FFF62F6C4D7D87AA9" httpUri="http://treatment.plazi.org/id/03A9879D977DFF9FFF62F6C4D7D87AA9" lastPageId="17" lastPageNumber="437" pageId="15" pageNumber="435" scope_class="Arachnida" scope_family="Linyphiidae" scope_order="Araneae" scope_subFamily="Erigoninae">
<subSubSection id="C31A6500977DFF81FF62F6C4D7927DCD" box="[151,499,1691,1718]" pageId="15" pageNumber="435" type="nomenclature">
<paragraph id="8BBF368B977DFF81FF62F6C4D7927DCD" blockId="15.[151,499,1691,1753]" box="[151,499,1691,1718]" pageId="15" pageNumber="435">
<heading id="D0F781E7977DFF81FF62F6C4D7927DCD" bold="true" box="[151,499,1691,1718]" fontSize="11" level="1" pageId="15" pageNumber="435" reason="1">
<emphasis id="B974EA99977DFF81FF62F6C4D7927DCD" bold="true" box="[151,499,1691,1718]" pageId="15" pageNumber="435">
<taxonomicName id="4C004D08977DFF81FF62F6C4D73A7DCE" ID-CoL="C86TS" authority="Tanasevitch, 2023" authorityName="Tanasevitch" authorityYear="2023" box="[151,347,1691,1718]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="435" phylum="Arthropoda" rank="species" species="ericola" status="sp. nov.">
<emphasis id="B974EA99977DFF81FF62F6C4D73A7DCE" bold="true" box="[151,347,1691,1718]" italics="true" pageId="15" pageNumber="435">Savignia ericola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757E2977DFF81FE96F6C4D7927DCD" box="[355,499,1691,1717]" pageId="15" pageNumber="435" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C31A6500977DFF81FF62F6E0D7727DA1" box="[151,275,1727,1753]" pageId="15" pageNumber="435" type="description">
<paragraph id="8BBF368B977DFF81FF62F6E0D7727DA1" blockId="15.[151,499,1691,1753]" box="[151,275,1727,1753]" pageId="15" pageNumber="435">
<figureCitation id="133B2A0E977DFF81FF62F6E0D68A7DA1" box="[151,235,1727,1753]" captionStart="FIGURE 11" captionStartId="16.[151,250,1697,1721]" captionTargetBox="[158,1429,604,1667]" captionTargetId="figure-245@16.[151,1436,597,1673]" captionTargetPageId="16" captionText="FIGURE 11. Photographs of male holotype (A–D) of Savignia ericola n. sp. A, habitus, dorsal view; B, C, same, lateral and anterolateral view, respectively; D, same, frontal view. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.8390477" httpUri="https://zenodo.org/record/8390477/files/figure.png" pageId="15" pageNumber="435">Figs 11</figureCitation>
,
<figureCitation id="133B2A0E977DFF81FF02F6E0D7727DA1" box="[247,275,1727,1753]" captionStart="FIGURE 12" captionStartId="17.[151,250,1440,1464]" captionTargetBox="[167,1433,489,1401]" captionTargetId="figure-134@17.[151,1436,489,1417]" captionTargetPageId="17" captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale." figureDoi="http://doi.org/10.5281/zenodo.8390479" httpUri="https://zenodo.org/record/8390479/files/figure.png" pageId="15" pageNumber="435">12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A6500977DFF81FF62F758D2827C11" pageId="15" pageNumber="435" type="materials_examined">
<paragraph id="8BBF368B977DFF81FF62F758D2827C11" blockId="15.[151,1437,1799,2042]" pageId="15" pageNumber="435">
<emphasis id="B974EA99977DFF81FF62F758D7277C59" bold="true" box="[151,326,1799,1825]" pageId="15" pageNumber="435">Type material.</emphasis>
<materialsCitation id="3B683CD6977DFF81FEBBF758D2827C11" ID-GBIF-Occurrence="4417449301" collectingDate="2022-10-30" collectionCode="ZMMU" collectorName="Arsi Mountains N. P. &amp; A. Tanasevitch" country="Ethiopia" county="Asela Zone" elevation="3503" latitude="7.81944" location="Arsi Mountains" longLatPrecision="1" longitude="39.35429" municipality="Asela" pageId="15" pageNumber="435" specimenCount="1" stateProvince="Oromiya" typeStatus="holotype">
<emphasis id="B974EA99977DFF81FEBBF758D7A57C59" bold="true" box="[334,452,1799,1825]" pageId="15" pageNumber="435">
<typeStatus id="54BB8829977DFF81FEBBF758D7DE7C59" box="[334,447,1799,1825]" pageId="15" pageNumber="435" type="holotype">Holotype</typeStatus>
:
</emphasis>
J (
<collectionCode id="ED11AE4E977DFF81FE04F758D42E7C59" box="[497,591,1799,1825]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="435" type="Museum">ZMMU</collectionCode>
):
<collectingCountry id="F317761B977DFF81FD96F758D4867C59" box="[611,743,1799,1825]" name="Ethiopia" pageId="15" pageNumber="435">ETHIOPIA</collectingCountry>
,
<collectingRegion id="49C4F869977DFF81FD06F758D5C57C59" box="[755,932,1799,1825]" country="Ethiopia" name="Oromiya" pageId="15" pageNumber="435">Oromia Region</collectingRegion>
,
<collectingCounty id="62DE4E07977DFF81FC5AF758D2537C5A" box="[943,1074,1799,1826]" pageId="15" pageNumber="435">Asela Zone</collectingCounty>
, ca 30 air-km SE from
<collectingMunicipality id="6BDBACF1977DFF81FACEF758D31A7C59" box="[1339,1403,1799,1825]" pageId="15" pageNumber="435">Asela</collectingMunicipality>
(= Assela),
<collectorName id="26F5535D977DFF81FF0DF774D7BE7C3E" box="[248,479,1835,1862]" pageId="15" pageNumber="435">
<location id="8EDF6050977DFF81FF0DF774D7C67C3D" LSID="urn:lsid:plazi:treatment:03A9879D977DFF9FFF62F6C4D7D87AA9:8EDF6050977DFF81FF0DF774D7C67C3D" box="[248,423,1835,1861]" country="Ethiopia" county="Asela Zone" latitude="7.81944" longLatPrecision="1" longitude="39.35429" municipality="Asela" name="Arsi Mountains" pageId="15" pageNumber="435" stateProvince="Oromiya">Arsi Mountains</location>
N.P.
</collectorName>
, road from
<location id="8EDF6050977DFF81FD94F774D4CC7C3D" LSID="urn:lsid:plazi:treatment:03A9879D977DFF9FFF62F6C4D7D87AA9:8EDF6050977DFF81FD94F774D4CC7C3D" box="[609,685,1835,1861]" country="Ethiopia" county="Asela Zone" latitude="7.81944" longLatPrecision="1" longitude="39.35429" municipality="Asela" name="Digelu" pageId="15" pageNumber="435" stateProvince="Oromiya">Digelu</location>
to
<location id="8EDF6050977DFF81FD24F774D5757C3D" LSID="urn:lsid:plazi:treatment:03A9879D977DFF9FFF62F6C4D7D87AA9:8EDF6050977DFF81FD24F774D5757C3D" box="[721,788,1835,1861]" country="Ethiopia" county="Asela Zone" latitude="7.81944" longLatPrecision="1" longitude="39.35429" municipality="Asela" name="Ticho" pageId="15" pageNumber="435" stateProvince="Oromiya">Ticho</location>
,
<elevation id="002DD1B8977DFF81FCEAF774D5967C3D" box="[799,1015,1835,1862]" metricMagnitude="3" metricUnit="m" metricValue="3.5025" metricValueMax="3.505" metricValueMin="3.5" pageId="15" pageNumber="435" unit="m" value="3502.5" valueMax="3505.0" valueMin="3500.0">
<quantity id="4CF89B6E977DFF81FCEAF774D5DB7C3E" box="[799,954,1835,1862]" metricMagnitude="3" metricUnit="m" metricValue="3.5025" metricValueMax="3.505" metricValueMin="3.5" pageId="15" pageNumber="435" unit="m" value="3502.5" valueMax="3505.0" valueMin="3500.0">3500–3505 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE34504C977DFF81FBF1F774D21F7C3D" box="[1028,1150,1835,1861]" degrees="7.81944" direction="north" orientation="latitude" pageId="15" pageNumber="435" precision="1" value="7.81944">7.81944°N</geoCoordinate>
<geoCoordinate id="EE34504C977DFF81FB70F774D36F7C3E" box="[1157,1294,1835,1862]" degrees="39.35429" direction="east" orientation="longitude" pageId="15" pageNumber="435" precision="1" value="39.35429">39.35429°E</geoCoordinate>
,
<taxonomicName id="4C004D08977DFF81FAECF773D6DA7C11" class="Arachnida" family="Salticidae" genus="Erica" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="435" phylum="Arthropoda" rank="species" species="arborea">
<emphasis id="B974EA99977DFF81FAECF773D6DA7C11" italics="true" pageId="15" pageNumber="435">Erica arborea</emphasis>
</taxonomicName>
bushes, with grass and green mosses, sifting mosses,
<date id="FFBE104B977DFF81FCE2F710D5F17C11" box="[791,912,1871,1897]" pageId="15" pageNumber="435" value="2022-10-30">
<collectingDate id="EFFAE9A3977DFF81FCE2F710D5F17C11" box="[791,912,1871,1897]" pageId="15" pageNumber="435" value="2022-10-30">30.X.2022</collectingDate>
</date>
, leg.
<collectorName id="26F5535D977DFF81FC3FF710D2107C11" box="[970,1137,1871,1897]" pageId="15" pageNumber="435">A. Tanasevitch</collectorName>
[Eth018].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A6500977DFF81FF32F72CD29D7CF5" box="[199,1276,1907,1934]" pageId="15" pageNumber="435" type="etymology">
<paragraph id="8BBF368B977DFF81FF32F72CD29D7CF5" blockId="15.[151,1437,1799,2042]" box="[199,1276,1907,1934]" pageId="15" pageNumber="435">
<emphasis id="B974EA99977DFF81FF32F72CD72F7CF5" bold="true" box="[199,334,1907,1933]" pageId="15" pageNumber="435">Etymology.</emphasis>
The specific epithet is a Latin adjective meaning a heather-dweller (
<taxonomicName id="4C004D08977DFF81FBBBF72BD28D7CF5" box="[1102,1260,1907,1933]" class="Arachnida" family="Salticidae" genus="Erica" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="435" phylum="Arthropoda" rank="species" species="arborea">
<emphasis id="B974EA99977DFF81FBBBF72BD28D7CF5" box="[1102,1260,1907,1933]" italics="true" pageId="15" pageNumber="435">Erica arborea</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A6500977DFF9EFF32F7C8D5AC7945" lastPageId="16" lastPageNumber="436" pageId="15" pageNumber="435" type="diagnosis">
<paragraph id="8BBF368B977DFF9EFF32F7C8D5AC7945" blockId="15.[151,1437,1799,2042]" lastBlockId="16.[151,1437,151,573]" lastPageId="16" lastPageNumber="436" pageId="15" pageNumber="435">
<emphasis id="B974EA99977DFF81FF32F7C8D7237CC9" bold="true" box="[199,322,1943,1969]" pageId="15" pageNumber="435">Diagnosis.</emphasis>
Based on the formula of chaetotaxy (2.2.1.1) and trichobothriotaxy (Tm I–III present, TmIV absent), the modified carapace, as well as the hypertrophied distal suprategular apophysis, the new species is evidently a member of the
<taxonomicName id="4C004D08977DFF81FEABF7BFD7A17C82" authorityName="Blackwall" authorityYear="1833" box="[350,448,2016,2042]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA99977DFF81FEABF7BFD7A17C82" box="[350,448,2016,2042]" italics="true" pageId="15" pageNumber="435">Savignia</emphasis>
</taxonomicName>
-genus group
<emphasis id="B974EA99977DFF81FDABF7BFD4FD7C81" box="[606,668,2016,2041]" italics="true" pageId="15" pageNumber="435">sensu</emphasis>
<bibRefCitation id="EF914B7A977DFF81FD50F780D53D7C81" author="Millidge, A. F." box="[677,860,2015,2041]" pageId="15" pageNumber="435" pagination="1 - 60" refId="ref11443" refString="Millidge, A. F. (1977) The conformation of the male palpal organs of linyphiid spiders, and its application to the taxonomic and phylogenetic analysis of the family (Araneae: Linyphiidae). Bulletin of the British Arachnological Society, 4, 1 - 60." type="journal article" year="1977">Millidge (1977)</bibRefCitation>
. Among the genera of this group, the new species seems most appropriate to be provisionally assigned to
<taxonomicName id="4C004D089762FF9EFD09F0C8D5947BC9" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[764,1013,151,177]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA999762FF9EFD09F0C8D53F7BC9" box="[764,862,151,177]" italics="true" pageId="16" pageNumber="436">Savignia</emphasis>
<bibRefCitation id="EF914B7A9762FF9EFC91F0C8D5947BC9" author="Simon, E." box="[868,1013,151,177]" pageId="16" pageNumber="436" pagination="180 - 885" refId="ref11731" refString="Simon, E. (1884) Les arachnides de France. Tome cinquieme, deuxieme et troisieme partie. Roret, Paris, pp. 180 - 885, pls. 26 - 27." type="book chapter" year="1884">Simon, 1884</bibRefCitation>
</taxonomicName>
, as certain resemble is observed in the structure of the embolic division, namely, the peculiar shape of the embolus, as well as by the presence of a narrow process at its base (MRP in
<figureCitation id="133B2A0E9762FF9EFE24F080D4547B81" box="[465,565,223,249]" captionStart="FIGURE 12" captionStartId="17.[151,250,1440,1464]" captionTargetBox="[167,1433,489,1401]" captionTargetId="figure-134@17.[151,1436,489,1417]" captionTargetPageId="17" captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale." figureDoi="http://doi.org/10.5281/zenodo.8390479" httpUri="https://zenodo.org/record/8390479/files/figure.png" pageId="16" pageNumber="436">Fig. 12E</figureCitation>
). The shape of the embolus is similar to that in many Asian congeners, e.g.
<taxonomicName id="4C004D089762FF9EFA72F080D7FF7A65" authority="Eskov, 1991" authorityName="Eskov" authorityYear="1991" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="B974EA999762FF9EFA72F080D7687A64" italics="true" pageId="16" pageNumber="436">S. amurensis</emphasis>
<bibRefCitation id="EF914B7A9762FF9EFEE4F15CD7FF7A65" author="Eskov, K. Y." box="[273,414,259,285]" pageId="16" pageNumber="436" pagination="140 - 144" refId="ref10791" refString="Eskov, K. Y. (1991) A spider genus Savignya (s. str.) (Aranei, Linyphiidae) in the fauna of the Far East and central Asia. Zoologicheskii Zhurnal, 70 (5), 140 - 144." type="journal article" year="1991">Eskov, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D089762FF9EFE5EF15CD5CE7A65" authority="Marusik, Koponen &amp; Danilov, 2001" authorityName="Marusik, Koponen &amp; Danilov" authorityYear="2001" box="[427,943,259,285]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="species" species="eskovi">
<emphasis id="B974EA999762FF9EFE5EF15CD46C7A65" box="[427,525,259,285]" italics="true" pageId="16" pageNumber="436">S. eskovi</emphasis>
<bibRefCitation id="EF914B7A9762FF9EFDE0F15CD5CE7A65" author="Marusik, Y. M. &amp; Koponen, S. &amp; Danilov, S. N." box="[533,943,259,285]" pageId="16" pageNumber="436" pagination="83 - 92" refId="ref11279" refString="Marusik, Y. M., Koponen, S. &amp; Danilov, S. N. (2001) Taxonomic and faunistic notes on linyphiids of Transbaikalia and south Siberia (Araneae, Linyphiidae). Bulletin of the British Arachnological Society, 12, 83 - 92." type="journal article" year="2001">Marusik, Koponen &amp; Danilov, 2001</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D089762FF9EFC4EF15CD2CA7A65" authority="Eskov, 1988" authorityName="Eskov" authorityYear="1988" box="[955,1195,259,285]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="species" species="saitoi">
<emphasis id="B974EA999762FF9EFC4EF15CD2747A64" box="[955,1045,259,285]" italics="true" pageId="16" pageNumber="436">S. saitoi</emphasis>
<bibRefCitation id="EF914B7A9762FF9EFBE8F15CD2CA7A65" author="Eskov, K. Y." box="[1053,1195,259,285]" pageId="16" pageNumber="436" pagination="13 - 39" refId="ref10745" refString="Eskov, K. Y. (1988) The spider genera Savignya Blackwall, Diplocephalus Bertkau and Archaraeoncus Tanasevitch (Aranei, Linyphiidae) in the fauna of Siberia and the Soviet Far East. Folia Entomologica Hungarica, 49, 13 - 39." type="journal article" year="1988">Eskov, 1988</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D089762FF9EFB42F15CD3F97A65" authority="Eskov, 1988" authorityName="Eskov" authorityYear="1988" box="[1207,1432,259,285]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="species" species="zero">
<emphasis id="B974EA999762FF9EFB42F15CD3637A64" box="[1207,1282,259,285]" italics="true" pageId="16" pageNumber="436">S. zero</emphasis>
<bibRefCitation id="EF914B7A9762FF9EFAFFF15CD3F97A65" author="Eskov, K. Y." box="[1290,1432,259,285]" pageId="16" pageNumber="436" pagination="13 - 39" refId="ref10745" refString="Eskov, K. Y. (1988) The spider genera Savignya Blackwall, Diplocephalus Bertkau and Archaraeoncus Tanasevitch (Aranei, Linyphiidae) in the fauna of Siberia and the Soviet Far East. Folia Entomologica Hungarica, 49, 13 - 39." type="journal article" year="1988">Eskov, 1988</bibRefCitation>
</taxonomicName>
, etc., all of which also show a median radical process in the embolic division. However, the new species stands apart among other congeners in lacking of a mesal outgrowth on the distal suprategular apophysis (MOSA in
<bibRefCitation id="EF914B7A9762FF9EFAE2F114D71E7AF1" author="Tanasevitch, A. V. &amp; Trilikauskas, L. A." pageId="16" pageNumber="436" pagination="269 - 270" refId="ref11767" refString="Tanasevitch, A. V. &amp; Trilikauskas, L. A. (2006) A new species of the genus Savignia Blackwall from the Khabarovsk Province, Russian Far East (Aranei: Linyphiidae: Erigoninae). Arthropoda Selecta, 14 (3), 269 - 270. [for 2005]" type="journal article" year="2006">Tanasevitch &amp; Trilikauskas 2006</bibRefCitation>
, figs 2, 4), a significant generic synapomorphy. Besides this,
<taxonomicName id="4C004D089762FF9EFBDBF130D2877AF1" authority="Tanasevitch, 2023" authorityName="Tanasevitch" authorityYear="2023" box="[1070,1254,367,393]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="species" species="ericola" status="sp. nov.">
<emphasis id="B974EA999762FF9EFBDBF130D2877AF1" box="[1070,1254,367,393]" italics="true" pageId="16" pageNumber="436">Savignia ericola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757E29762FF9EFB1BF130D34B7AF1" box="[1262,1322,367,393]" pageId="16" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
possesses a characteristic “
<taxonomicName id="4C004D089762FF9EFEA7F1CCD7D07AD5" authorityName="Tanasevitch" authorityYear="1987" box="[338,433,403,429]" class="Arachnida" family="Linyphiidae" genus="Bisetifer" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA999762FF9EFEA7F1CCD7D07AD5" box="[338,433,403,429]" italics="true" pageId="16" pageNumber="436">Bisetifer</emphasis>
</taxonomicName>
-like” palpal tibia which bears three long, strong, lateral setae absent from other congeners of
<taxonomicName id="4C004D089762FF9EFF43F1E8D7797AA9" authorityName="Simon" authorityYear="1884" box="[182,280,439,465]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA999762FF9EFF43F1E8D7797AA9" box="[182,280,439,465]" italics="true" pageId="16" pageNumber="436">Savignia</emphasis>
</taxonomicName>
-genus group. This new species is only conditionally to be placed in
<taxonomicName id="4C004D089762FF9EFBEEF1E8D21C7AA9" authorityName="Simon" authorityYear="1884" box="[1051,1149,439,465]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA999762FF9EFBEEF1E8D21C7AA9" box="[1051,1149,439,465]" italics="true" pageId="16" pageNumber="436">Savignia</emphasis>
</taxonomicName>
, so at least until a female is found. If the structure of the epigynum turns out to be similar to that of
<taxonomicName id="4C004D089762FF9EFC25F184D2537A8D" authorityName="Simon" authorityYear="1884" box="[976,1074,475,501]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA999762FF9EFC25F184D2537A8D" box="[976,1074,475,501]" italics="true" pageId="16" pageNumber="436">Savignia</emphasis>
</taxonomicName>
(namely, with a bisected ventral plate), then the placement of the new species into the genus will become justified. At the same time, it is possible that the new species represents a new genus of the
<taxonomicName id="4C004D089762FF9EFD25F27CD5537945" authorityName="Simon" authorityYear="1884" box="[720,818,547,573]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA999762FF9EFD25F27CD5537945" box="[720,818,547,573]" italics="true" pageId="16" pageNumber="436">Savignia</emphasis>
</taxonomicName>
-genus group.
</paragraph>
</subSubSection>
<caption id="DF7F66039762FF9EFF62F6FED51E7DA5" ID-DOI="http://doi.org/10.5281/zenodo.8390477" ID-Zenodo-Dep="8390477" httpUri="https://zenodo.org/record/8390477/files/figure.png" pageId="16" pageNumber="436" startId="16.[151,250,1697,1721]" targetBox="[158,1429,604,1667]" targetPageId="16" targetType="figure">
<paragraph id="8BBF368B9762FF9EFF62F6FED51E7DA5" blockId="16.[151,1437,1697,1757]" pageId="16" pageNumber="436">
<emphasis id="B974EA999762FF9EFF62F6FED77E7DC1" bold="true" box="[151,287,1697,1721]" pageId="16" pageNumber="436">FIGURE 11.</emphasis>
Photographs of male holotype (
<emphasis id="B974EA999762FF9EFD9CF6FED4FB7DC1" bold="true" box="[617,666,1697,1721]" pageId="16" pageNumber="436">A–D</emphasis>
) of
<taxonomicName id="4C004D089762FF9EFD33F6FED50F7DC1" authority="Tanasevitch, 2023" authorityName="Tanasevitch" authorityYear="2023" box="[710,878,1697,1721]" class="Arachnida" family="Linyphiidae" genus="Savignia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="436" phylum="Arthropoda" rank="species" species="ericola" status="sp. nov.">
<emphasis id="B974EA999762FF9EFD33F6FED50F7DC1" box="[710,878,1697,1721]" italics="true" pageId="16" pageNumber="436">Savignia ericola</emphasis>
</taxonomicName>
<emphasis id="B974EA999762FF9EFC80F6FED5A97DC1" bold="true" box="[885,968,1697,1721]" pageId="16" pageNumber="436">
<taxonomicNameLabel id="A24757E29762FF9EFC80F6FED5CE7DC1" box="[885,943,1697,1721]" pageId="16" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
A
</emphasis>
, habitus, dorsal view;
<emphasis id="B974EA999762FF9EFB44F6FED2A37DC1" bold="true" box="[1201,1218,1697,1721]" pageId="16" pageNumber="436">B</emphasis>
,
<emphasis id="B974EA999762FF9EFB25F6FED2837DC1" bold="true" box="[1232,1250,1697,1721]" pageId="16" pageNumber="436">C</emphasis>
, same, lateral and anterolateral view, respectively;
<emphasis id="B974EA999762FF9EFE15F69AD7937DA5" bold="true" box="[480,498,1733,1757]" pageId="16" pageNumber="436">D</emphasis>
, same, frontal view. Scale bar: 0.5mm.
</paragraph>
</caption>
<subSubSection id="C31A65009762FF9FFF32F754D7F47AF1" lastPageId="17" lastPageNumber="437" pageId="16" pageNumber="436" type="description">
<paragraph id="8BBF368B9762FF9FFF32F754D7597A1D" blockId="16.[151,1437,1803,2010]" lastBlockId="17.[151,1437,151,465]" lastPageId="17" lastPageNumber="437" pageId="16" pageNumber="436">
<emphasis id="B974EA999762FF9EFF32F754D7387C5D" bold="true" box="[199,345,1803,1829]" pageId="16" pageNumber="436">Description.</emphasis>
Male
<typeStatus id="54BB88299762FF9EFE5DF754D46F7C5D" box="[424,526,1803,1829]" pageId="16" pageNumber="436" type="holotype">holotype</typeStatus>
. Total length 1.83. Carapace slightly modified as shown in
<figureCitation id="133B2A0E9762FF9EFB3BF754D33A7C5D" box="[1230,1371,1803,1829]" captionStart="FIGURE 11" captionStartId="16.[151,250,1697,1721]" captionTargetBox="[158,1429,604,1667]" captionTargetId="figure-245@16.[151,1436,597,1673]" captionTargetPageId="16" captionText="FIGURE 11. Photographs of male holotype (A–D) of Savignia ericola n. sp. A, habitus, dorsal view; B, C, same, lateral and anterolateral view, respectively; D, same, frontal view. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.8390477" httpUri="https://zenodo.org/record/8390477/files/figure.png" pageId="16" pageNumber="436">Figs 11A–C</figureCitation>
, 1.03 long, 0.63 wide, pale greyish brown to greyish yellow. Anterior part of carapace somewhat protruded anteriorly, eyes relatively small. Chelicerae 0.22 long, a mastidion absent. Legs yellow. Leg I 2.06 long (0.63 + 0.22 + 0.48 + 0.40 + 0.33), IV 2.19 long (0.63 + 0.20 +0.58 + 0.48 + 0.30). Chaetotaxy 2.2.1.1, spines about as long as 0.5–1 diameter of corresponding leg segment. Metatarsi I–III with a trichobothrium each. TmI 0.41. Palp (
<figureCitation id="133B2A0E9762FF9EFB0AF7C4D3F07CCD" box="[1279,1425,1947,1973]" captionStart="FIGURE 12" captionStartId="17.[151,250,1440,1464]" captionTargetBox="[167,1433,489,1401]" captionTargetId="figure-134@17.[151,1436,489,1417]" captionTargetPageId="17" captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale." figureDoi="http://doi.org/10.5281/zenodo.8390479" httpUri="https://zenodo.org/record/8390479/files/figure.png" pageId="16" pageNumber="436">Figs 12A–G</figureCitation>
): Patella slightly elongated. Tibia with a dorsal, conical, apically rounded outgrowth and a small projection bearing three long, stout and slightly curved setae. Paracymbium J-shaped. Distal suprategular apophysis very large, massive, ending with a strong, straight spike. Radix rounded proximally, with a membranous tissue near middle, a flat, subtrapeziform, anterior apophysis (ARA in
<figureCitation id="133B2A0E9763FF9FFD61F080D49B7B81" box="[660,762,223,249]" captionStart="FIGURE 12" captionStartId="17.[151,250,1440,1464]" captionTargetBox="[167,1433,489,1401]" captionTargetId="figure-134@17.[151,1436,489,1417]" captionTargetPageId="17" captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale." figureDoi="http://doi.org/10.5281/zenodo.8390479" httpUri="https://zenodo.org/record/8390479/files/figure.png" pageId="17" pageNumber="437">Fig. 12E</figureCitation>
), and an relatively long, narrow, apically unciform process (
<figureCitation id="133B2A0E9763FF9FFF6AF15CD7427A65" box="[159,291,259,285]" captionStart="FIGURE 12" captionStartId="17.[151,250,1440,1464]" captionTargetBox="[167,1433,489,1401]" captionTargetId="figure-134@17.[151,1436,489,1417]" captionTargetPageId="17" captionText="FIGURE 12. Details of male palpal structure (A–G) of holotype of Savignia ericola n. sp. A, left palp, retrolateral view; B, tibia, dorsal view; C, tegulum, distal suprategular apophysis and embolic division, prolateral view; D, distal suprategular apophysis, lateral view; E, embolic division, mesal view; F, apex of median radical process; G, apex of embolus. Scale bars: A–E, 0.1mm; G, F, not to scale." figureDoi="http://doi.org/10.5281/zenodo.8390479" httpUri="https://zenodo.org/record/8390479/files/figure.png" pageId="17" pageNumber="437">Figs 12E, F</figureCitation>
) situated at base of embolus. Embolus starting from middle of radix as a narrow stripe, sharply curved in middle part to gradually taper thereafter into a thin and curved tip. Opisthosoma 0.90 long, 0.58 wide, pale grey (
<figureCitation id="133B2A0E9763FF9FFF6AF114D74D7A1D" box="[159,300,331,357]" captionStart="FIGURE 11" captionStartId="16.[151,250,1697,1721]" captionTargetBox="[158,1429,604,1667]" captionTargetId="figure-245@16.[151,1436,597,1673]" captionTargetPageId="16" captionText="FIGURE 11. Photographs of male holotype (A–D) of Savignia ericola n. sp. A, habitus, dorsal view; B, C, same, lateral and anterolateral view, respectively; D, same, frontal view. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.8390477" httpUri="https://zenodo.org/record/8390477/files/figure.png" pageId="17" pageNumber="437">Figs 11A–C</figureCitation>
).
</paragraph>
<paragraph id="8BBF368B9763FF9FFF32F130D7F47AF1" blockId="17.[151,1437,151,465]" box="[199,405,367,393]" pageId="17" pageNumber="437">
<emphasis id="B974EA999763FF9FFF32F130D77F7AF1" bold="true" box="[199,286,367,393]" pageId="17" pageNumber="437">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31A65009763FF9FFF32F1CDD7D87AA9" pageId="17" pageNumber="437" type="distribution">
<paragraph id="8BBF368B9763FF9FFF32F1CDD7D87AA9" blockId="17.[151,1437,151,465]" pageId="17" pageNumber="437">
<emphasis id="B974EA999763FF9FFF32F1CDD7007AD4" bold="true" box="[199,353,402,428]" pageId="17" pageNumber="437">Distribution.</emphasis>
Known only from the
<typeStatus id="54BB88299763FF9FFDABF1CCD4EF7AD5" box="[606,654,403,429]" pageId="17" pageNumber="437">type</typeStatus>
locality in the highlands of Arsi Mountains N.P. (
<quantity id="4CF89B6E9763FF9FFB40F1CDD3337AD5" box="[1205,1362,402,429]" metricMagnitude="3" metricUnit="m" metricValue="3.5025" metricValueMax="3.505" metricValueMin="3.5" pageId="17" pageNumber="437" unit="m" value="3502.5" valueMax="3505.0" valueMin="3500.0">3500–3505 m</quantity>
a.s.l.),
<collectingRegion id="49C4F8699763FF9FFF62F1E8D7297AA9" box="[151,328,439,465]" country="Ethiopia" name="Oromiya" pageId="17" pageNumber="437">Oromia Region</collectingRegion>
,
<collectingCountry id="F317761B9763FF9FFEA6F1E8D7D77AA9" box="[339,438,439,465]" name="Ethiopia" pageId="17" pageNumber="437">Ethiopia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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