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<document id="1D3B01088E830FAFDE344ADF3451ADBF" ID-DOI="10.11646/zootaxa.3630.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261763" ID-ZooBank="C7529E9C-73F7-4E52-92E7-FCACA8F99A7D" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629846447618" checkinUser="felipe" docAuthor="Liu, Ye, Shi, Hongliang &amp; Liang, Hongbin" docDate="2013" docId="03CF5A007209FFBAFF17FDABF159F937" docLanguage="en" docName="zt03630p518.pdf" docOrigin="Zootaxa 3630 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Chlaenioctenus) eneides Bates 1892" docType="treatment" docVersion="1" lastPageNumber="512" masterDocId="FFF62278720EFFBDFF80FFC0F270FF81" masterDocTitle="Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus" masterLastPageNumber="518" masterPageNumber="505" pageNumber="512" updateTime="1730973406347" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="6FD5E0F1ADE58140B6995FBF3B9A9271">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:namePart id="FA5F83DFF29C10436AED77FFB9F17820">Liu, Ye</mods:namePart>
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<mods:namePart id="7D9A9CBBAA81FCE7619363B369E6685D">Shi, Hongliang</mods:namePart>
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<mods:namePart id="7C5370F5702997D86BE6C39FE176E2D8">Liang, Hongbin</mods:namePart>
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<treatment id="03CF5A007209FFBAFF17FDABF159F937" LSID="urn:lsid:plazi:treatment:03CF5A007209FFBAFF17FDABF159F937" httpUri="http://treatment.plazi.org/id/03CF5A007209FFBAFF17FDABF159F937" lastPageNumber="512" pageId="7" pageNumber="512">
<subSubSection id="C37CB89D7209FFBAFF17FDABF0B6FD05" box="[151,710,619,644]" pageId="7" pageNumber="512" type="nomenclature">
<paragraph id="8BD9EB167209FFBAFF17FDABF0B6FD05" blockId="7.[151,710,619,679]" box="[151,710,619,644]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF17FDABF0B6FD05" ID-CoL="69W7J" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[151,710,619,644]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides" subGenus="Chlaenioctenus">
<emphasis id="B91237047209FFBAFF17FDABF37DFD05" bold="true" box="[151,269,619,644]" italics="true" pageId="7" pageNumber="512">Chlaenius</emphasis>
(
<emphasis id="B91237047209FFBAFE9DFDABF3A1FD05" bold="true" box="[285,465,619,644]" italics="true" pageId="7" pageNumber="512">Chlaenioctenus</emphasis>
)
<emphasis id="B91237047209FFBAFE62FDABF0B6FD05" bold="true" box="[482,710,619,644]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFE62FDABF047FD05" bold="true" box="[482,567,619,644]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
<bibRefCitation id="EFF796E77209FFBAFDBFFDACF0B6FD05" author="Bates H. W." box="[575,710,619,644]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF17FD4FF05CFD26" box="[151,556,655,679]" pageId="7" pageNumber="512" type="description">
<paragraph id="8BD9EB167209FFBAFF17FD4FF05CFD26" blockId="7.[151,710,619,679]" box="[151,556,655,679]" pageId="7" pageNumber="512">
(
<figureCitation id="135DF7937209FFBAFF1FFD4FF358FD26" box="[159,296,655,679]" captionStart="FIGURES 1316" captionStartId="8.[151,263,1909,1932]" captionTargetBox="[192,1336,197,1886]" captionTargetId="figure-51@8.[151,1436,193,1888]" captionTargetPageId="8" captionText="FIGURES 1316. Chlaenius eneides Bates, 1892. 13, Lectotype, male, dorsal view. 14, Lectotype, male, ventral view. 15, Paralectotype, female, dorsal view. 16, Paralectotype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261775" httpUri="https://zenodo.org/record/5261775/files/figure.png" pageId="7" pageNumber="512">Figs. 1316</figureCitation>
,
<figureCitation id="135DF7937209FFBAFEB4FD4FF322FD26" box="[308,338,655,679]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="7" pageNumber="512">24</figureCitation>
,
<figureCitation id="135DF7937209FFBAFEDEFD4FF30CFD26" box="[350,380,655,679]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="7" pageNumber="512">30</figureCitation>
,
<figureCitation id="135DF7937209FFBAFE08FD4FF3D6FD26" box="[392,422,655,679]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="7" pageNumber="512">36</figureCitation>
,
<figureCitation id="135DF7937209FFBAFE32FD4FF38AFD26" box="[434,506,655,679]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="7" pageNumber="512">46, 47</figureCitation>
,
<figureCitation id="135DF7937209FFBAFD86FD4FF053FD26" box="[518,547,655,679]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="7" pageNumber="512">51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF17FD18F0C5FD6F" box="[151,693,728,750]" pageId="7" pageNumber="512" type="reference_group">
<paragraph id="8BD9EB167209FFBAFF17FD18F0C5FD6F" blockId="7.[151,693,728,750]" box="[151,693,728,750]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF17FD18F0C5FD6F" authority="Bates, 1892: 310" authorityName="Bates" authorityPageNumber="310" authorityYear="1892" box="[151,693,728,750]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides" subGenus="Chlaenioctenus">
<emphasis id="B91237047209FFBAFF17FD18F28EFD6F" box="[151,254,728,750]" italics="true" pageId="7" pageNumber="512">Chlaenius</emphasis>
(
<emphasis id="B91237047209FFBAFE8CFD18F3D9FD6F" box="[268,425,728,750]" italics="true" pageId="7" pageNumber="512">Chlaenioctenus</emphasis>
)
<emphasis id="B91237047209FFBAFE38FD18F074FD6F" box="[440,516,728,750]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
<bibRefCitation id="EFF796E77209FFBAFD8AFD18F0C5FD6F" author="Bates H. W." box="[522,693,728,750]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="310" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892: 310</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF17FCDAF7F2FCFB" pageId="7" pageNumber="512" type="materials_examined">
<paragraph id="8BD9EB167209FFBAFF17FCDAF7F2FCFB" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF17FCDAF00BFCB3" box="[151,635,794,819]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF17FCDAF3F6FCB2" bold="true" box="[151,390,794,819]" pageId="7" pageNumber="512">Specimen examined</emphasis>
(
<emphasis id="B91237047209FFBAFE1AFCDBF01BFCB2" bold="true" box="[410,619,794,819]" pageId="7" pageNumber="512">
total
<specimenCount id="9D60209F7209FFBAFE5AFCDAF01BFCB2" box="[474,619,794,819]" pageId="7" pageNumber="512" type="generic">2 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B7209FFBAFD05FCDBF6AEFCD7" collectorName="R. Oberthur" location="Assam" pageId="7" pageNumber="512" specimenCount="1" stateProvince="Assam" typeStatus="lectotype">
<typeStatus id="54DD55B47209FFBAFD05FCDBF085FCB2" box="[645,757,795,819]" pageId="7" pageNumber="512" type="lectotype">Lectotype</typeStatus>
(designated herein): male (MNHN), “
<collectingRegion id="49A225F47209FFBAFB2EFCDAF68CFCB3" box="[1198,1276,794,818]" country="India" name="Assam" pageId="7" pageNumber="512">Assam</collectingRegion>
plains”/ “Ex-Musaeo, H.W.
<bibRefCitation id="EFF796E77209FFBAFEBDFCFFF3B2FCD7" author="Bates H. W." box="[317,450,830,854]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
”/ “Museum Paris, coll.
<collectorName id="26938EC07209FFBAFD4DFCFFF124FCD7" box="[717,852,830,854]" pageId="7" pageNumber="512">R. Oberthür</collectorName>
, 1952”/ “
<taxonomicName id="4C6690957209FFBAFC3DFCFEF6A2FCD7" authority="Bates" authorityName="Bates" authorityYear="1892" box="[957,1234,830,854]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFC3DFCFEF6F9FCD7" box="[957,1161,830,854]" italics="true" pageId="7" pageNumber="512">Chlaenius eneides</emphasis>
Bates
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B0EE14B7209FFBAFB67FCFEF70EFCFB" collectorName="R. Oberthur" location="Assam" pageId="7" pageNumber="512" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="paralectotype">
<typeStatus id="54DD55B47209FFBAFB67FCFEF7F4FCD7" box="[1255,1412,830,854]" pageId="7" pageNumber="512" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="9D60209F7209FFBAFA0EFCFEF293FCFB" pageId="7" pageNumber="512" type="female">1 female</specimenCount>
(MNHN), “
<collectingRegion id="49A225F47209FFBAFEEAFCA2F3C9FCFB" box="[362,441,866,890]" country="India" name="Assam" pageId="7" pageNumber="512">Assam</collectingRegion>
plains / “Ex-Musaeo, H.W.
<bibRefCitation id="EFF796E77209FFBAFC86FCA3F1FFFCFB" author="Bates H. W." box="[774,911,866,890]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
/ “Museum Paris, coll.
<collectorName id="26938EC07209FFBAFB23FCA3F75BFCFB" box="[1187,1323,866,890]" pageId="7" pageNumber="512">R. Oberthür</collectorName>
, 1952”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF49FC47F619FAF3" pageId="7" pageNumber="512" type="description">
<paragraph id="8BD9EB167209FFBAFF49FC47F088FC67" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FC47F30FFC1F" box="[201,383,903,927]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FC47F308FC1E" bold="true" box="[201,376,903,927]" pageId="7" pageNumber="512">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F37209FFBAFE47FC46F05AFC1F" box="[455,554,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="7" pageNumber="512" unit="mm" value="14.2">14.2 mm</quantity>
(
<typeStatus id="54DD55B47209FFBAFDB9FC46F0D5FC1F" box="[569,677,902,926]" pageId="7" pageNumber="512" type="lectotype">lectotype</typeStatus>
),
<quantity id="4C9E46F37209FFBAFD38FC46F16BFC1F" box="[696,795,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="7" pageNumber="512" unit="mm" value="15.6">15.6 mm</quantity>
(
<typeStatus id="54DD55B47209FFBAFCAAFC46F1B7FC1F" box="[810,967,902,926]" pageId="7" pageNumber="512" type="paralectotype">paralectotype</typeStatus>
), HW =
<quantity id="4C9E46F37209FFBAFBA5FC46F60DFC1F" box="[1061,1149,902,926]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="512" unit="mm" value="2.3">2.3 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFB09FC46F690FC1F" box="[1161,1248,902,926]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="512" unit="mm" value="2.6">2.6 mm</quantity>
, EYL =
<quantity id="4C9E46F37209FFBAFAC0FC46F7E7FC1F" box="[1344,1431,902,926]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="512" unit="mm" value="0.9">0.9 mm</quantity>
, 1.0 mm, PL = 3.0 mm,
<quantity id="4C9E46F37209FFBAFE1FFC6AF388FC43" box="[415,504,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="512" unit="mm" value="3.5">3.5 mm</quantity>
, PW =
<quantity id="4C9E46F37209FFBAFDCCFC6AF0D5FC43" box="[588,677,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="512" unit="mm" value="3.4">3.4 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFD31FC6AF179FC43" box="[689,777,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="512" unit="mm" value="3.7">3.7 mm</quantity>
, EL =
<quantity id="4C9E46F37209FFBAFCD6FC6AF1DFFC43" box="[854,943,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="512" unit="mm" value="8.3">8.3 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFC3BFC6AF663FC43" box="[955,1043,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="7" pageNumber="512" unit="mm" value="9.2">9.2 mm</quantity>
, EW =
<quantity id="4C9E46F37209FFBAFBE9FC6AF6B2FC43" box="[1129,1218,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="512" unit="mm" value="4.8">4.8 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFB4EFC6AF756FC43" box="[1230,1318,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="512" unit="mm" value="5.8">5.8 mm</quantity>
, PL/PW = 1.10, 1.06, EL/EW = 1.73, 1.59, EW/PW = 1.45, 1.57.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49FC32F17FFBF7" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FC32F32AFB8B" box="[201,346,1010,1035]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FC32F323FB8A" bold="true" box="[201,339,1010,1035]" pageId="7" pageNumber="512">Description</emphasis>
.
</emphasis>
Head and pronotum black, with distinct green metallic shine; elytra black; ventral surface black, with metallic shine; coxae, trochanters, tibiae, antennomeres 2 and 3 black or dark brown; antennomeres 1 and 411, mandibles, palpi and tarsi reddish brown; femora yellow in the middle portion, black at base and apex, basal black areas on hind femora longer than hind trochanters.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49FB42F619FAF3" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
Head convex; vertex and occiput finely punctate; genae glabrous, oblique, as long as one-fifth of eyes; lateral margin of mentum straight. Pronotum (
<figureCitation id="135DF7937209FFBAFDE1FB66F0C6FB3F" box="[609,694,1190,1214]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="7" pageNumber="512">Fig. 24</figureCitation>
) ovoid, widest in the middle; lateral margins slightly bordered, sinuate before posterior angles; disc punctate, median line distinct, base punctate; basal foveae deep, coarsely punctate. Elytra moderately convex, widest in or slightly behind the middle; intervals (
<figureCitation id="135DF7937209FFBAFBF6FB2EF6B9FA87" box="[1142,1225,1262,1286]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="7" pageNumber="512">Fig. 30</figureCitation>
) convex, glabrous medially, with sparse pubescence at each lateral side; yellow spot behind the middle of each elytron, covering intervals 48. Proepisterna very sparsely punctate. Claws (
<figureCitation id="135DF7937209FFBAFCB5FAF6F1F8FACF" box="[821,904,1334,1358]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="7" pageNumber="512">Fig. 36</figureCitation>
) curved, with 47 long teeth. Aedeagus (
<figureCitation id="135DF7937209FFBAFAE4FAF6F2AEFAF3" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="7" pageNumber="512">Figs. 46, 47</figureCitation>
) thin in lateral view. Gonostylus (
<figureCitation id="135DF7937209FFBAFDDBFA9AF0DCFAF3" box="[603,684,1370,1394]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="7" pageNumber="512">Fig. 51</figureCitation>
) with four long spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF49FABFF159F937" pageId="7" pageNumber="512" type="discussion">
<paragraph id="8BD9EB167209FFBAFF49FABFF3FCFA3B" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FABFF34CFA17" box="[201,316,1407,1431]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FABFF346FA16" bold="true" box="[201,310,1407,1431]" pageId="7" pageNumber="512">Remarks</emphasis>
.
</emphasis>
<bibRefCitation id="EFF796E77209FFBAFEC4FABFF07FFA17" author="Bates H. W." box="[324,527,1406,1430]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="310" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates (1892: 310)</bibRefCitation>
gave a very brief description of
<taxonomicName id="4C6690957209FFBAFC03FABEF188FA17" authorityName="Bates" authorityYear="1892" box="[899,1016,1406,1430]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFC03FABEF1E6FA17" box="[899,918,1406,1430]" italics="true" pageId="7" pageNumber="512">C</emphasis>
.
<emphasis id="B91237047209FFBAFC24FABEF188FA17" box="[932,1016,1406,1430]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
</taxonomicName>
. We redescribe this species herein to aid the determination.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49FA06F6BEF9CA" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
According to the original description,
<taxonomicName id="4C6690957209FFBAFD02FA06F089FA5F" authorityName="Bates" authorityYear="1892" box="[642,761,1478,1502]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFD02FA06F0E5FA5F" box="[642,661,1478,1502]" italics="true" pageId="7" pageNumber="512">C</emphasis>
.
<emphasis id="B91237047209FFBAFD25FA06F089FA5F" box="[677,761,1478,1502]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
</taxonomicName>
was described on a male and a female (
<bibRefCitation id="EFF796E77209FFBAFB4FFA06F7E0FA5F" author="Bates H. W." box="[1231,1424,1478,1502]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="310" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates 1892: 310</bibRefCitation>
), without fixing the name-bearing type. In the collection of MNHN,
<specimenCount id="9D60209F7209FFBAFC20FA2BF646F982" box="[928,1078,1515,1539]" pageId="7" pageNumber="512" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
were found collected from the same localities and bearing Bates' determining label. Hence, the male deposited in MNHN is designated as the
<typeStatus id="54DD55B47209FFBAFF17F9F2F28EF9CB" box="[151,254,1586,1610]" pageId="7" pageNumber="512" type="lectotype">lectotype</typeStatus>
herein, for fixing the name to a single specimen and to avoid confusion about the type.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49F996F159F937" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF49F996F331F9EF" authorityName="Bates" authorityYear="1892" box="[201,321,1622,1646]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFF49F996F331F9EF" box="[201,321,1622,1646]" italics="true" pageId="7" pageNumber="512">C. eneides</emphasis>
</taxonomicName>
is similar with
<taxonomicName id="4C6690957209FFBAFE7FF996F0B4F9EF" authorityName="Kirschenhofer" authorityYear="2003" box="[511,708,1622,1646]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047209FFBAFE7FF996F0B4F9EF" box="[511,708,1622,1646]" italics="true" pageId="7" pageNumber="512">C. schillhammeri</emphasis>
</taxonomicName>
in having elytral spots slightly behind the middle and femora bicolored, but differs from the latter by pronotal lateral margin sinuate before posterior angle in contrast to lateral margin straight before posterior angle in
<taxonomicName id="4C6690957209FFBAFDE1F95EF152F937" authorityName="Kirschenhofer" authorityYear="2003" box="[609,802,1694,1718]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047209FFBAFDE1F95EF152F937" box="[609,802,1694,1718]" italics="true" pageId="7" pageNumber="512">C. schillhammeri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>