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<mods:title id="4AEB6EB133089C32A1E3AD7DBFE7CBE9">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:namePart id="4FAD4A21E5955278A7B80A6FEF4477B3">Shi, Hongliang</mods:namePart>
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<mods:namePart id="FD0A3D3DF52A857838D59C88C0879331">Liang, Hongbin</mods:namePart>
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<paragraph id="8BD9EB167209FFBAFF17F8CAF6EDF8A2" blockId="7.[151,1181,1802,1862]" box="[151,1181,1802,1827]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF17F8CAF114F8A3" ID-CoL="5XW7C" authority="Kirschenhofer, 2008" authorityName="Kirschenhofer" authorityYear="2008" box="[151,868,1802,1827]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="langsonensis" status="subgen. nov." subGenus="Chlaenioctenus">
<emphasis id="B91237047209FFBAFF17F8CAF37DF8A2" bold="true" box="[151,269,1802,1827]" italics="true" pageId="7" pageNumber="512">Chlaenius</emphasis>
(
<emphasis id="B91237047209FFBAFE9DF8CAF3A1F8A2" bold="true" box="[285,465,1802,1827]" italics="true" pageId="7" pageNumber="512">Chlaenioctenus</emphasis>
)
<emphasis id="B91237047209FFBAFE62F8CAF007F8A2" bold="true" box="[482,631,1802,1827]" italics="true" pageId="7" pageNumber="512">langsonensis</emphasis>
<bibRefCitation id="EFF796E77209FFBAFDFEF8CAF114F8A3" author="Kirschenhofer, E." box="[638,868,1802,1826]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" pageId="7" pageNumber="512" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer, 2008</bibRefCitation>
</taxonomicName>
<emphasis id="B91237047209FFBAFCEBF8CBF6EDF8A2" bold="true" box="[875,1181,1802,1827]" pageId="7" pageNumber="512">
<taxonomicNameLabel id="A2218A7F7209FFBAFCEBF8CBF661F8A2" box="[875,1041,1803,1827]" pageId="7" pageNumber="512" rank="subGenus">new subgenus</taxonomicNameLabel>
assignment
</emphasis>
</paragraph>
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<subSubSection id="C37CB89D7209FFBAFF17F8EEF3F1F8C7" box="[151,385,1838,1862]" pageId="7" pageNumber="512" type="description">
<paragraph id="8BD9EB167209FFBAFF17F8EEF3F1F8C7" blockId="7.[151,1181,1802,1862]" box="[151,385,1838,1862]" pageId="7" pageNumber="512">(Figs. 19, 25, 31, 37)</paragraph>
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<paragraph id="8BD9EB167209FFBAFF17F8B7F089F80D" blockId="7.[151,761,1910,1933]" box="[151,761,1910,1933]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF17F8B7F089F80D" authority="Kirschenhofer, 2008: 17" authorityName="Kirschenhofer" authorityPageNumber="17" authorityYear="2008" box="[151,761,1910,1933]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="langsonensis" subGenus="Ocybatus">
<emphasis id="B91237047209FFBAFF17F8B7F28EF80C" box="[151,254,1911,1933]" italics="true" pageId="7" pageNumber="512">Chlaenius</emphasis>
(
<emphasis id="B91237047209FFBAFE8CF8B7F31CF80C" box="[268,364,1911,1933]" italics="true" pageId="7" pageNumber="512">Ocybatus</emphasis>
)
<emphasis id="B91237047209FFBAFEFBF8B7F38EF80C" box="[379,510,1911,1933]" italics="true" pageId="7" pageNumber="512">langsonensis</emphasis>
<bibRefCitation id="EFF796E77209FFBAFD84F8B6F089F80D" author="Kirschenhofer, E." box="[516,761,1910,1932]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" page="17" pageId="7" pageNumber="512" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer, 2008: 17</bibRefCitation>
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</paragraph>
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<subSubSection id="C37CB89D7209FFBAFF17F879F6CFF874" pageId="7" pageNumber="512" type="materials_examined">
<paragraph id="8BD9EB167209FFBAFF17F879F6CFF874" blockId="7.[151,1436,1977,2037]" pageId="7" pageNumber="512">
Specimens examined (total
<materialsCitation id="3B0EE14B7209FFBAFE73F879F6CFF874" collectorName="H. Fruhstorfer" country="Vietnam" location="Tonkin" pageId="7" pageNumber="512" specimenCount="1" typeStatus="holotype">
<emphasis id="B91237047209FFBAFE73F879F0FEF850" box="[499,654,1977,2002]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFE73F879F0F6F853" bold="true" box="[499,646,1977,2002]" pageId="7" pageNumber="512">
<specimenCount id="9D60209F7209FFBAFE73F879F00FF853" box="[499,639,1977,2002]" pageId="7" pageNumber="512" type="generic">1 specimen</specimenCount>
)
</emphasis>
.
</emphasis>
<typeStatus id="54DD55B47209FFBAFD1CF879F177F850" box="[668,775,1977,2001]" pageId="7" pageNumber="512" type="holotype">Holotype</typeStatus>
, male (MNHU), “
<collectingCountry id="F371AB867209FFBAFC66F879F64AF850" box="[998,1082,1977,2001]" name="Vietnam" pageId="7" pageNumber="512">Tonkin</collectingCountry>
, Montes Mauson, April. Mai 23000,
<collectorName id="26938EC07209FFBAFF7BF81EF3EDF874" box="[251,413,2013,2037]" pageId="7" pageNumber="512">H. Fruhstorfer</collectorName>
”/ “
<taxonomicName id="4C6690957209FFBAFE42F81DF13EF874" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[450,846,2013,2037]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="langsonensis" status="sp. nov." subGenus="Ocybatus">
<emphasis id="B91237047209FFBAFE42F81DF13EF874" box="[450,846,2013,2037]" italics="true" pageId="7" pageNumber="512">Chlaenius (Ocybatus) langsonensis</emphasis>
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<emphasis id="B91237047209FFBAFCD6F81DF1E5F874" bold="true" box="[854,917,2013,2037]" pageId="7" pageNumber="512">
<taxonomicNameLabel id="A2218A7F7209FFBAFCD6F81DF1E5F874" box="[854,917,2013,2037]" pageId="7" pageNumber="512" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
det.
<bibRefCitation id="EFF796E77209FFBAFC4CF81DF6C0F874" author="Kirschenhofer, E." box="[972,1200,2013,2037]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" pageId="7" pageNumber="512" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer 2008</bibRefCitation>
”.
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</paragraph>
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<caption id="DF19BB9E7206FFB5FF17F8B5F62DF82B" ID-DOI="http://doi.org/10.5281/zenodo.5261775" ID-Zenodo-Dep="5261775" httpUri="https://zenodo.org/record/5261775/files/figure.png" pageId="8" pageNumber="513" startId="8.[151,263,1909,1932]" targetBox="[192,1336,197,1886]" targetPageId="8">
<paragraph id="8BD9EB167206FFB5FF17F8B5F62DF82B" blockId="8.[151,1436,1909,1962]" pageId="8" pageNumber="513">
<emphasis id="B91237047206FFB5FF17F8B5F326F80D" bold="true" box="[151,342,1909,1932]" pageId="8" pageNumber="513">FIGURES 1316.</emphasis>
<taxonomicName id="4C6690957206FFB5FEDFF8B6F0D1F80D" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[351,673,1910,1932]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047206FFB5FEDFF8B6F06AF80D" box="[351,538,1910,1932]" italics="true" pageId="8" pageNumber="513">Chlaenius eneides</emphasis>
<bibRefCitation id="EFF796E77206FFB5FDA4F8B7F0D1F80D" author="Bates H. W." box="[548,673,1910,1932]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="8" pageNumber="513" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
. 13, Lectotype, male, dorsal view. 14, Lectotype, male, ventral view. 15, Paralectotype, female, dorsal view. 16, Paralectotype, female, ventral view. Scale bar = 5.0 mm
</paragraph>
</caption>
<caption id="DF19BB9E7207FFB4FF17FBB7F1AEFB2D" ID-DOI="http://doi.org/10.5281/zenodo.5261779" ID-Zenodo-Dep="5261779" httpUri="https://zenodo.org/record/5261779/files/figure.png" pageId="9" pageNumber="514" startId="9.[151,264,1143,1166]" targetBox="[164,1388,193,1122]" targetPageId="9">
<paragraph id="8BD9EB167207FFB4FF17FBB7F1AEFB2D" blockId="9.[151,1436,1143,1197]" pageId="9" pageNumber="514">
<emphasis id="B91237047207FFB4FF17FBB7F326FB0F" bold="true" box="[151,342,1143,1166]" pageId="9" pageNumber="514">FIGURES 1718.</emphasis>
<taxonomicName id="4C6690957207FFB4FEDEFBB8F076FB0F" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[350,518,1144,1166]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="aodai" status="sp. nov.">
<emphasis id="B91237047207FFB4FEDEFBB8F076FB0F" box="[350,518,1144,1166]" italics="true" pageId="9" pageNumber="514">Chlaenius aodai</emphasis>
</taxonomicName>
<emphasis id="B91237047207FFB4FD90FBB8F03BFB0C" bold="true" box="[528,587,1144,1165]" pageId="9" pageNumber="514">
<taxonomicNameLabel id="A2218A7F7207FFB4FD90FBB8F03BFB0C" box="[528,587,1144,1165]" pageId="9" pageNumber="514" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
17, Holotype, male, dorsal view. 18, Holotype, male, ventral view.
<emphasis id="B91237047207FFB4FA91FBB7F7ECFB0F" bold="true" box="[1297,1436,1143,1166]" pageId="9" pageNumber="514">FIGURE 19.</emphasis>
<taxonomicName id="4C6690957207FFB4FF17FB57F02FFB2D" authority="Kirschenhofer, 2008" authorityName="Kirschenhofer" authorityYear="2008" box="[151,607,1174,1197]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="langsonensis">
<emphasis id="B91237047207FFB4FF17FB57F3F7FB2C" box="[151,391,1175,1197]" italics="true" pageId="9" pageNumber="514">Chlaenius langsonensis</emphasis>
<bibRefCitation id="EFF796E77207FFB4FE0DFB56F02FFB2D" author="Kirschenhofer, E." box="[397,607,1174,1196]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" pageId="9" pageNumber="514" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer, 2008</bibRefCitation>
</taxonomicName>
. Holotype, male. Scale bar = 5.0 mm.
</paragraph>
</caption>
<subSubSection id="C37CB89D7207FFB4FF49FB32F330FA1B" pageId="9" pageNumber="514" type="description">
<paragraph id="8BD9EB167207FFB4FF49FB32F330FA1B" blockId="9.[151,1436,1266,1939]" pageId="9" pageNumber="514">
<emphasis id="B91237047207FFB4FF49FB32F32AFA8B" box="[201,346,1266,1291]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">
<emphasis id="B91237047207FFB4FF49FB32F323FA8A" bold="true" box="[201,339,1266,1291]" pageId="9" pageNumber="514">Description</emphasis>
.
</emphasis>
<bibRefCitation id="EFF796E77207FFB4FEE4FB32F0F5FA8B" author="Kirschenhofer, E." box="[356,645,1266,1290]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" page="17" pageId="9" pageNumber="514" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer (2008: 17)</bibRefCitation>
gave a quite detailed description. We add the following here: Genae glabrous, tumid, as long as half of eyes in dorsal view. Pronotum (
<figureCitation id="135DF7937207FFB4FC28FAD6F18DFAAF" box="[936,1021,1302,1326]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="9" pageNumber="514">Fig. 25</figureCitation>
) with lateral margin straight before posterior angle. Elytral with intervals (
<figureCitation id="135DF7937207FFB4FDC6FAFAF0E8FAD3" box="[582,664,1338,1362]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="9" pageNumber="514">Fig. 31</figureCitation>
) convex, glabrous medially, with a row of pubescence laterally; each elytron with a round yellow spot slightly behind the middle, covering intervals 48. Claws (
<figureCitation id="135DF7937207FFB4FB2CFA9EF68EFAF7" box="[1196,1278,1374,1398]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="9" pageNumber="514">Fig. 37</figureCitation>
) curved, with 67 long teeth.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7207FFB4FF49FA67F137F813" pageId="9" pageNumber="514" type="discussion">
<paragraph id="8BD9EB167207FFB4FF49FA67F619F9CF" blockId="9.[151,1436,1266,1939]" pageId="9" pageNumber="514">
<emphasis id="B91237047207FFB4FF49FA67F34CFA3F" box="[201,316,1447,1471]" inLineHeading="true" pageId="9" pageNumber="514" reason="1">
<emphasis id="B91237047207FFB4FF49FA67F346FA3E" bold="true" box="[201,310,1447,1471]" pageId="9" pageNumber="514">Remarks</emphasis>
.
</emphasis>
<bibRefCitation id="EFF796E77207FFB4FEC5FA67F3A6FA3F" author="Bates H. W." box="[325,470,1446,1470]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="9" pageNumber="514" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates (1892)</bibRefCitation>
erected subgenus
<taxonomicName id="4C6690957207FFB4FD29FA66F127FA3F" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[681,855,1446,1470]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="subGenus" subGenus="Chlaenioctenus">
<emphasis id="B91237047207FFB4FD29FA66F127FA3F" box="[681,855,1446,1470]" italics="true" pageId="9" pageNumber="514">Chlaenioctenus</emphasis>
</taxonomicName>
for claws with teeth, a character unique within the genus
<taxonomicName id="4C6690957207FFB4FF5FFA0BF321FA62" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[223,337,1483,1507]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B91237047207FFB4FF5FFA0BF321FA62" box="[223,337,1483,1507]" italics="true" pageId="9" pageNumber="514">Chlaenius</emphasis>
</taxonomicName>
. This subgenus is similar to subgeneus
<taxonomicName id="4C6690957207FFB4FC8CFA0BF106FA62" box="[780,886,1483,1507]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="subGenus" subGenus="Ocybatus">
<emphasis id="B91237047207FFB4FC8CFA0BF106FA62" box="[780,886,1483,1507]" italics="true" pageId="9" pageNumber="514">Ocybatus</emphasis>
</taxonomicName>
Laferté-Sènectére in having widened labial palpi and two yellow spots on elytra, but differs from the latter in having rounded shoulders (hind wings reduced) in addition to pectinate claws. After examined the
<typeStatus id="54DD55B47207FFB4FD49F9D2F15BF9AB" box="[713,811,1554,1578]" pageId="9" pageNumber="514" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6690957207FFB4FCD7F9D3F67CF9AA" authorityName="Kirschenhofer" authorityYear="2008" box="[855,1036,1555,1579]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="langsonensis">
<emphasis id="B91237047207FFB4FCD7F9D3F67CF9AA" box="[855,1036,1555,1579]" italics="true" pageId="9" pageNumber="514">C. langsonensis</emphasis>
</taxonomicName>
with pectinate claws and rounded shoulders, we assignate this species to subgenus
<taxonomicName id="4C6690957207FFB4FD39F9F6F117F9CF" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[697,871,1590,1614]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="subGenus" subGenus="Chlaenioctenus">
<emphasis id="B91237047207FFB4FD39F9F6F117F9CF" box="[697,871,1590,1614]" italics="true" pageId="9" pageNumber="514">Chlaenioctenus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6690957207FFB4FC2BF9F6F665F9CF" box="[939,1045,1590,1614]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="505" phylum="Arthropoda" rank="subGenus" subGenus="Ocybatus">
<emphasis id="B91237047207FFB4FC2BF9F6F665F9CF" box="[939,1045,1590,1614]" italics="true" pageId="9" pageNumber="514">Ocybatus</emphasis>
</taxonomicName>
herein.
</paragraph>
<paragraph id="8BD9EB167207FFB4FF49F99BF717F8A7" blockId="9.[151,1436,1266,1939]" pageId="9" pageNumber="514">
<taxonomicName id="4C6690957207FFB4FF49F99BF3A7F9F2" authorityName="Kirschenhofer" authorityYear="2008" box="[201,471,1627,1651]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="langsonensis">
<emphasis id="B91237047207FFB4FF49F99BF3A7F9F2" box="[201,471,1627,1651]" italics="true" pageId="9" pageNumber="514">Chlaenius langsonensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C6690957207FFB4FD3EF99BF640F9F3" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="2003" box="[702,1072,1626,1651]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047207FFB4FD3EF99BF1F4F9F2" box="[702,900,1627,1651]" italics="true" pageId="9" pageNumber="514">C. schillhammeri</emphasis>
Kirschenhofer
</taxonomicName>
in having yellow spot slightly behind the middle, femora bicolored and pronotal lateral margin nearly straight before posterior angle (
<figureCitation id="135DF7937207FFB4FA91F9BEF713F917" box="[1297,1379,1662,1686]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="9" pageNumber="514">Fig. 25</figureCitation>
; also see
<figureCitation id="135DF7937207FFB4FF45F962F367F93B" box="[197,279,1698,1722]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="9" pageNumber="514">
Fig.
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</figureCitation>
in
<bibRefCitation id="EFF796E77207FFB4FEC1F962F383F93B" author="Liu, Y. &amp; Liang, H. B. &amp; Kavanaugh, D. &amp; Yang, M. F." box="[321,499,1698,1723]" etAl="et al." firstAuthor="Liu" journalOrPublisher="Zootaxa" pageId="9" pageNumber="514" pagination="15 - 28" part="2397" refId="ref7391" refString="Liu, Y., Liang, H. B., Kavanaugh, D. &amp; Yang, M. F. (2010) Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species. Zootaxa, 2397, 15 - 28." title="Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species" type="journal article" year="2010">
Liu
<emphasis id="B91237047207FFB4FEF2F963F3DEF93A" box="[370,430,1699,1723]" italics="true" pageId="9" pageNumber="514">et al.</emphasis>
2010
</bibRefCitation>
), but differs from the latter in antennomere 1 brown (antennomere 1 black in
<taxonomicName id="4C6690957207FFB4FA02F963F347F95F" authorityName="Kirschenhofer" authorityYear="2003" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047207FFB4FA02F963F347F95F" italics="true" pageId="9" pageNumber="514">C. schillhammeri</emphasis>
</taxonomicName>
), genae strongly tumid (Fig. 19) (genae slightly tumid in
<taxonomicName id="4C6690957207FFB4FC43F906F6F4F95F" authorityName="Kirschenhofer" authorityYear="2003" box="[963,1156,1734,1758]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047207FFB4FC43F906F6F4F95F" box="[963,1156,1734,1758]" italics="true" pageId="9" pageNumber="514">C. schillhammeri</emphasis>
</taxonomicName>
), and pronotal disc (
<figureCitation id="135DF7937207FFB4FAEEF906F2C4F883" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="9" pageNumber="514">Fig. 25</figureCitation>
) more coarsely punctate (less coarsely in
<taxonomicName id="4C6690957207FFB4FD22F92BF116F882" authorityName="Kirschenhofer" authorityYear="2003" box="[674,870,1771,1795]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047207FFB4FD22F92BF116F882" box="[674,870,1771,1795]" italics="true" pageId="9" pageNumber="514">C. schillhammeri</emphasis>
</taxonomicName>
). Their difference of external characters is quite minor, and study on their male genitalia is necessary when male of
<taxonomicName id="4C6690957207FFB4FC09F8CEF639F8A7" authorityName="Kirschenhofer" authorityYear="2003" box="[905,1097,1806,1830]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047207FFB4FC09F8CEF639F8A7" box="[905,1097,1806,1830]" italics="true" pageId="9" pageNumber="514">C. schillhammeri</emphasis>
</taxonomicName>
is available in the future.
</paragraph>
<paragraph id="8BD9EB167207FFB4FF49F8F3F137F813" blockId="9.[151,1436,1266,1939]" pageId="9" pageNumber="514">
<taxonomicName id="4C6690957207FFB4FF49F8F3F30FF8CA" authorityName="Kirschenhofer" authorityYear="2008" box="[201,383,1843,1867]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="langsonensis">
<emphasis id="B91237047207FFB4FF49F8F3F30FF8CA" box="[201,383,1843,1867]" italics="true" pageId="9" pageNumber="514">C. langsonensis</emphasis>
</taxonomicName>
also could be confused with
<taxonomicName id="4C6690957207FFB4FD60F8F3F129F8CA" authorityName="Bates" authorityYear="1892" box="[736,857,1843,1867]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047207FFB4FD60F8F3F129F8CA" box="[736,857,1843,1867]" italics="true" pageId="9" pageNumber="514">C. eneides</emphasis>
</taxonomicName>
in having elytral yellow spot slightly behind the middle and femora bicolored, but differs from the latter in having pronotal lateral margin (
<figureCitation id="135DF7937207FFB4FB16F896F698F8EF" box="[1174,1256,1878,1902]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="9" pageNumber="514">Fig. 25</figureCitation>
) straight before posterior angle (lateral margin sinuate in
<taxonomicName id="4C6690957207FFB4FDE4F8BBF0A9F812" authorityName="Bates" authorityYear="1892" box="[612,729,1915,1939]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047207FFB4FDE4F8BBF0A9F812" box="[612,729,1915,1939]" italics="true" pageId="9" pageNumber="514">C. eneides</emphasis>
</taxonomicName>
,
<figureCitation id="135DF7937207FFB4FD67F8BAF14AF813" box="[743,826,1914,1938]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="9" pageNumber="514">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,242 @@
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<subSubSection id="C37CB89D7209FFBAFF17FDABF0B6FD05" box="[151,710,619,644]" pageId="7" pageNumber="512" type="nomenclature">
<paragraph id="8BD9EB167209FFBAFF17FDABF0B6FD05" blockId="7.[151,710,619,679]" box="[151,710,619,644]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF17FDABF0B6FD05" ID-CoL="69W7J" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[151,710,619,644]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides" subGenus="Chlaenioctenus">
<emphasis id="B91237047209FFBAFF17FDABF37DFD05" bold="true" box="[151,269,619,644]" italics="true" pageId="7" pageNumber="512">Chlaenius</emphasis>
(
<emphasis id="B91237047209FFBAFE9DFDABF3A1FD05" bold="true" box="[285,465,619,644]" italics="true" pageId="7" pageNumber="512">Chlaenioctenus</emphasis>
)
<emphasis id="B91237047209FFBAFE62FDABF0B6FD05" bold="true" box="[482,710,619,644]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFE62FDABF047FD05" bold="true" box="[482,567,619,644]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
<bibRefCitation id="EFF796E77209FFBAFDBFFDACF0B6FD05" author="Bates H. W." box="[575,710,619,644]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF17FD4FF05CFD26" box="[151,556,655,679]" pageId="7" pageNumber="512" type="description">
<paragraph id="8BD9EB167209FFBAFF17FD4FF05CFD26" blockId="7.[151,710,619,679]" box="[151,556,655,679]" pageId="7" pageNumber="512">
(
<figureCitation id="135DF7937209FFBAFF1FFD4FF358FD26" box="[159,296,655,679]" captionStart="FIGURES 1316" captionStartId="8.[151,263,1909,1932]" captionTargetBox="[192,1336,197,1886]" captionTargetId="figure-51@8.[151,1436,193,1888]" captionTargetPageId="8" captionText="FIGURES 1316. Chlaenius eneides Bates, 1892. 13, Lectotype, male, dorsal view. 14, Lectotype, male, ventral view. 15, Paralectotype, female, dorsal view. 16, Paralectotype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261775" httpUri="https://zenodo.org/record/5261775/files/figure.png" pageId="7" pageNumber="512">Figs. 1316</figureCitation>
,
<figureCitation id="135DF7937209FFBAFEB4FD4FF322FD26" box="[308,338,655,679]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="7" pageNumber="512">24</figureCitation>
,
<figureCitation id="135DF7937209FFBAFEDEFD4FF30CFD26" box="[350,380,655,679]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="7" pageNumber="512">30</figureCitation>
,
<figureCitation id="135DF7937209FFBAFE08FD4FF3D6FD26" box="[392,422,655,679]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="7" pageNumber="512">36</figureCitation>
,
<figureCitation id="135DF7937209FFBAFE32FD4FF38AFD26" box="[434,506,655,679]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="7" pageNumber="512">46, 47</figureCitation>
,
<figureCitation id="135DF7937209FFBAFD86FD4FF053FD26" box="[518,547,655,679]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="7" pageNumber="512">51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF17FD18F0C5FD6F" box="[151,693,728,750]" pageId="7" pageNumber="512" type="reference_group">
<paragraph id="8BD9EB167209FFBAFF17FD18F0C5FD6F" blockId="7.[151,693,728,750]" box="[151,693,728,750]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF17FD18F0C5FD6F" authority="Bates, 1892: 310" authorityName="Bates" authorityPageNumber="310" authorityYear="1892" box="[151,693,728,750]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides" subGenus="Chlaenioctenus">
<emphasis id="B91237047209FFBAFF17FD18F28EFD6F" box="[151,254,728,750]" italics="true" pageId="7" pageNumber="512">Chlaenius</emphasis>
(
<emphasis id="B91237047209FFBAFE8CFD18F3D9FD6F" box="[268,425,728,750]" italics="true" pageId="7" pageNumber="512">Chlaenioctenus</emphasis>
)
<emphasis id="B91237047209FFBAFE38FD18F074FD6F" box="[440,516,728,750]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
<bibRefCitation id="EFF796E77209FFBAFD8AFD18F0C5FD6F" author="Bates H. W." box="[522,693,728,750]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="310" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892: 310</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF17FCDAF7F2FCFB" pageId="7" pageNumber="512" type="materials_examined">
<paragraph id="8BD9EB167209FFBAFF17FCDAF7F2FCFB" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF17FCDAF00BFCB3" box="[151,635,794,819]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF17FCDAF3F6FCB2" bold="true" box="[151,390,794,819]" pageId="7" pageNumber="512">Specimen examined</emphasis>
(
<emphasis id="B91237047209FFBAFE1AFCDBF01BFCB2" bold="true" box="[410,619,794,819]" pageId="7" pageNumber="512">
total
<specimenCount id="9D60209F7209FFBAFE5AFCDAF01BFCB2" box="[474,619,794,819]" pageId="7" pageNumber="512" type="generic">2 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B7209FFBAFD05FCDBF6AEFCD7" collectorName="R. Oberthur" location="Assam" pageId="7" pageNumber="512" specimenCount="1" stateProvince="Assam" typeStatus="lectotype">
<typeStatus id="54DD55B47209FFBAFD05FCDBF085FCB2" box="[645,757,795,819]" pageId="7" pageNumber="512" type="lectotype">Lectotype</typeStatus>
(designated herein): male (MNHN), “
<collectingRegion id="49A225F47209FFBAFB2EFCDAF68CFCB3" box="[1198,1276,794,818]" country="India" name="Assam" pageId="7" pageNumber="512">Assam</collectingRegion>
plains”/ “Ex-Musaeo, H.W.
<bibRefCitation id="EFF796E77209FFBAFEBDFCFFF3B2FCD7" author="Bates H. W." box="[317,450,830,854]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
”/ “Museum Paris, coll.
<collectorName id="26938EC07209FFBAFD4DFCFFF124FCD7" box="[717,852,830,854]" pageId="7" pageNumber="512">R. Oberthür</collectorName>
, 1952”/ “
<taxonomicName id="4C6690957209FFBAFC3DFCFEF6A2FCD7" authority="Bates" authorityName="Bates" authorityYear="1892" box="[957,1234,830,854]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFC3DFCFEF6F9FCD7" box="[957,1161,830,854]" italics="true" pageId="7" pageNumber="512">Chlaenius eneides</emphasis>
Bates
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B0EE14B7209FFBAFB67FCFEF70EFCFB" collectorName="R. Oberthur" location="Assam" pageId="7" pageNumber="512" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="paralectotype">
<typeStatus id="54DD55B47209FFBAFB67FCFEF7F4FCD7" box="[1255,1412,830,854]" pageId="7" pageNumber="512" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="9D60209F7209FFBAFA0EFCFEF293FCFB" pageId="7" pageNumber="512" type="female">1 female</specimenCount>
(MNHN), “
<collectingRegion id="49A225F47209FFBAFEEAFCA2F3C9FCFB" box="[362,441,866,890]" country="India" name="Assam" pageId="7" pageNumber="512">Assam</collectingRegion>
plains / “Ex-Musaeo, H.W.
<bibRefCitation id="EFF796E77209FFBAFC86FCA3F1FFFCFB" author="Bates H. W." box="[774,911,866,890]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
/ “Museum Paris, coll.
<collectorName id="26938EC07209FFBAFB23FCA3F75BFCFB" box="[1187,1323,866,890]" pageId="7" pageNumber="512">R. Oberthür</collectorName>
, 1952”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF49FC47F619FAF3" pageId="7" pageNumber="512" type="description">
<paragraph id="8BD9EB167209FFBAFF49FC47F088FC67" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FC47F30FFC1F" box="[201,383,903,927]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FC47F308FC1E" bold="true" box="[201,376,903,927]" pageId="7" pageNumber="512">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F37209FFBAFE47FC46F05AFC1F" box="[455,554,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="7" pageNumber="512" unit="mm" value="14.2">14.2 mm</quantity>
(
<typeStatus id="54DD55B47209FFBAFDB9FC46F0D5FC1F" box="[569,677,902,926]" pageId="7" pageNumber="512" type="lectotype">lectotype</typeStatus>
),
<quantity id="4C9E46F37209FFBAFD38FC46F16BFC1F" box="[696,795,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="7" pageNumber="512" unit="mm" value="15.6">15.6 mm</quantity>
(
<typeStatus id="54DD55B47209FFBAFCAAFC46F1B7FC1F" box="[810,967,902,926]" pageId="7" pageNumber="512" type="paralectotype">paralectotype</typeStatus>
), HW =
<quantity id="4C9E46F37209FFBAFBA5FC46F60DFC1F" box="[1061,1149,902,926]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="512" unit="mm" value="2.3">2.3 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFB09FC46F690FC1F" box="[1161,1248,902,926]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="512" unit="mm" value="2.6">2.6 mm</quantity>
, EYL =
<quantity id="4C9E46F37209FFBAFAC0FC46F7E7FC1F" box="[1344,1431,902,926]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="512" unit="mm" value="0.9">0.9 mm</quantity>
, 1.0 mm, PL = 3.0 mm,
<quantity id="4C9E46F37209FFBAFE1FFC6AF388FC43" box="[415,504,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="512" unit="mm" value="3.5">3.5 mm</quantity>
, PW =
<quantity id="4C9E46F37209FFBAFDCCFC6AF0D5FC43" box="[588,677,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="512" unit="mm" value="3.4">3.4 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFD31FC6AF179FC43" box="[689,777,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="512" unit="mm" value="3.7">3.7 mm</quantity>
, EL =
<quantity id="4C9E46F37209FFBAFCD6FC6AF1DFFC43" box="[854,943,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="512" unit="mm" value="8.3">8.3 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFC3BFC6AF663FC43" box="[955,1043,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="7" pageNumber="512" unit="mm" value="9.2">9.2 mm</quantity>
, EW =
<quantity id="4C9E46F37209FFBAFBE9FC6AF6B2FC43" box="[1129,1218,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="512" unit="mm" value="4.8">4.8 mm</quantity>
,
<quantity id="4C9E46F37209FFBAFB4EFC6AF756FC43" box="[1230,1318,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="512" unit="mm" value="5.8">5.8 mm</quantity>
, PL/PW = 1.10, 1.06, EL/EW = 1.73, 1.59, EW/PW = 1.45, 1.57.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49FC32F17FFBF7" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FC32F32AFB8B" box="[201,346,1010,1035]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FC32F323FB8A" bold="true" box="[201,339,1010,1035]" pageId="7" pageNumber="512">Description</emphasis>
.
</emphasis>
Head and pronotum black, with distinct green metallic shine; elytra black; ventral surface black, with metallic shine; coxae, trochanters, tibiae, antennomeres 2 and 3 black or dark brown; antennomeres 1 and 411, mandibles, palpi and tarsi reddish brown; femora yellow in the middle portion, black at base and apex, basal black areas on hind femora longer than hind trochanters.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49FB42F619FAF3" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
Head convex; vertex and occiput finely punctate; genae glabrous, oblique, as long as one-fifth of eyes; lateral margin of mentum straight. Pronotum (
<figureCitation id="135DF7937209FFBAFDE1FB66F0C6FB3F" box="[609,694,1190,1214]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="7" pageNumber="512">Fig. 24</figureCitation>
) ovoid, widest in the middle; lateral margins slightly bordered, sinuate before posterior angles; disc punctate, median line distinct, base punctate; basal foveae deep, coarsely punctate. Elytra moderately convex, widest in or slightly behind the middle; intervals (
<figureCitation id="135DF7937209FFBAFBF6FB2EF6B9FA87" box="[1142,1225,1262,1286]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="7" pageNumber="512">Fig. 30</figureCitation>
) convex, glabrous medially, with sparse pubescence at each lateral side; yellow spot behind the middle of each elytron, covering intervals 48. Proepisterna very sparsely punctate. Claws (
<figureCitation id="135DF7937209FFBAFCB5FAF6F1F8FACF" box="[821,904,1334,1358]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="7" pageNumber="512">Fig. 36</figureCitation>
) curved, with 47 long teeth. Aedeagus (
<figureCitation id="135DF7937209FFBAFAE4FAF6F2AEFAF3" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="7" pageNumber="512">Figs. 46, 47</figureCitation>
) thin in lateral view. Gonostylus (
<figureCitation id="135DF7937209FFBAFDDBFA9AF0DCFAF3" box="[603,684,1370,1394]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="7" pageNumber="512">Fig. 51</figureCitation>
) with four long spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF49FABFF159F937" pageId="7" pageNumber="512" type="discussion">
<paragraph id="8BD9EB167209FFBAFF49FABFF3FCFA3B" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FABFF34CFA17" box="[201,316,1407,1431]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FABFF346FA16" bold="true" box="[201,310,1407,1431]" pageId="7" pageNumber="512">Remarks</emphasis>
.
</emphasis>
<bibRefCitation id="EFF796E77209FFBAFEC4FABFF07FFA17" author="Bates H. W." box="[324,527,1406,1430]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="310" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates (1892: 310)</bibRefCitation>
gave a very brief description of
<taxonomicName id="4C6690957209FFBAFC03FABEF188FA17" authorityName="Bates" authorityYear="1892" box="[899,1016,1406,1430]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFC03FABEF1E6FA17" box="[899,918,1406,1430]" italics="true" pageId="7" pageNumber="512">C</emphasis>
.
<emphasis id="B91237047209FFBAFC24FABEF188FA17" box="[932,1016,1406,1430]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
</taxonomicName>
. We redescribe this species herein to aid the determination.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49FA06F6BEF9CA" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
According to the original description,
<taxonomicName id="4C6690957209FFBAFD02FA06F089FA5F" authorityName="Bates" authorityYear="1892" box="[642,761,1478,1502]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFD02FA06F0E5FA5F" box="[642,661,1478,1502]" italics="true" pageId="7" pageNumber="512">C</emphasis>
.
<emphasis id="B91237047209FFBAFD25FA06F089FA5F" box="[677,761,1478,1502]" italics="true" pageId="7" pageNumber="512">eneides</emphasis>
</taxonomicName>
was described on a male and a female (
<bibRefCitation id="EFF796E77209FFBAFB4FFA06F7E0FA5F" author="Bates H. W." box="[1231,1424,1478,1502]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="310" pageId="7" pageNumber="512" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates 1892: 310</bibRefCitation>
), without fixing the name-bearing type. In the collection of MNHN,
<specimenCount id="9D60209F7209FFBAFC20FA2BF646F982" box="[928,1078,1515,1539]" pageId="7" pageNumber="512" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
were found collected from the same localities and bearing Bates' determining label. Hence, the male deposited in MNHN is designated as the
<typeStatus id="54DD55B47209FFBAFF17F9F2F28EF9CB" box="[151,254,1586,1610]" pageId="7" pageNumber="512" type="lectotype">lectotype</typeStatus>
herein, for fixing the name to a single specimen and to avoid confusion about the type.
</paragraph>
<paragraph id="8BD9EB167209FFBAFF49F996F159F937" blockId="7.[151,1436,794,1718]" pageId="7" pageNumber="512">
<taxonomicName id="4C6690957209FFBAFF49F996F331F9EF" authorityName="Bates" authorityYear="1892" box="[201,321,1622,1646]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B91237047209FFBAFF49F996F331F9EF" box="[201,321,1622,1646]" italics="true" pageId="7" pageNumber="512">C. eneides</emphasis>
</taxonomicName>
is similar with
<taxonomicName id="4C6690957209FFBAFE7FF996F0B4F9EF" authorityName="Kirschenhofer" authorityYear="2003" box="[511,708,1622,1646]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047209FFBAFE7FF996F0B4F9EF" box="[511,708,1622,1646]" italics="true" pageId="7" pageNumber="512">C. schillhammeri</emphasis>
</taxonomicName>
in having elytral spots slightly behind the middle and femora bicolored, but differs from the latter by pronotal lateral margin sinuate before posterior angle in contrast to lateral margin straight before posterior angle in
<taxonomicName id="4C6690957209FFBAFDE1F95EF152F937" authorityName="Kirschenhofer" authorityYear="2003" box="[609,802,1694,1718]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B91237047209FFBAFDE1F95EF152F937" box="[609,802,1694,1718]" italics="true" pageId="7" pageNumber="512">C. schillhammeri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,371 @@
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<mods:title id="A84DB5B91638534E6EC69AF59B748207">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:namePart id="9C4B521AD36565C4E2E91290EC26617F">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="1628737F79ED8EC040D0C14E730F2D5C" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<treatment id="03CF5A00720BFFB8FF17FF57F3E0FA1A" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFB8FF17FF57F3E0FA1A" httpUri="http://treatment.plazi.org/id/03CF5A00720BFFB8FF17FF57F3E0FA1A" lastPageNumber="510" pageId="5" pageNumber="510">
<subSubSection id="C37CB89D720BFFB8FF17FF57F0A4FF31" box="[151,724,151,176]" pageId="5" pageNumber="510" type="nomenclature">
<paragraph id="8BD9EB16720BFFB8FF17FF57F0A4FF31" blockId="5.[151,724,151,212]" box="[151,724,151,176]" pageId="5" pageNumber="510">
<taxonomicName id="4C669095720BFFB8FF17FF57F0FEFF31" ID-CoL="5Y7VC" authority="Liu &amp; Liang, 2013" authorityName="Liu &amp; Liang" authorityYear="2013" box="[151,654,151,176]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="li" subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FF17FF57F37DFF31" bold="true" box="[151,269,151,176]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FE9DFF57F3A1FF31" bold="true" box="[285,465,151,176]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720BFFB8FE62FF57F0A4FF31" bold="true" box="[482,724,151,176]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FE62FF57F381FF31" bold="true" box="[482,497,151,176]" italics="true" pageId="5" pageNumber="510">li</emphasis>
Liu &amp; Liang
<taxonomicNameLabel id="A2218A7F720BFFB8FD15FF58F0A4FF31" box="[661,724,152,176]" pageId="5" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF17FF7CF06DFF55" box="[151,541,188,212]" pageId="5" pageNumber="510" type="description">
<paragraph id="8BD9EB16720BFFB8FF17FF7CF06DFF55" blockId="5.[151,724,151,212]" box="[151,541,188,212]" pageId="5" pageNumber="510">
(
<figureCitation id="135DF793720BFFB8FF1FFF7CF36BFF55" box="[159,283,188,212]" captionStart="FIGURES 912" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[282,1273,195,1883]" captionTargetId="figure-51@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURES 912. Chlaenius li sp. n. 9, Holotype, male, dorsal view. 10, Holotype, male, ventral view. 11, Paratype, female, dorsal view. 12, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261773" httpUri="https://zenodo.org/record/5261773/files/figure.png" pageId="5" pageNumber="510">Figs. 912</figureCitation>
,
<figureCitation id="135DF793720BFFB8FEA6FF7CF335FF55" box="[294,325,188,212]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="5" pageNumber="510">22</figureCitation>
,
<figureCitation id="135DF793720BFFB8FED0FF7CF31EFF55" box="[336,366,188,212]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="5" pageNumber="510">28</figureCitation>
,
<figureCitation id="135DF793720BFFB8FEFAFF7CF3E8FF55" box="[378,408,188,212]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="5" pageNumber="510">34</figureCitation>
,
<figureCitation id="135DF793720BFFB8FE24FF7CF39CFF55" box="[420,492,188,212]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="5" pageNumber="510">42, 43</figureCitation>
,
<figureCitation id="135DF793720BFFB8FE78FF7CF065FF55" box="[504,533,188,212]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="5" pageNumber="510">49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF17FEC3F66AFD05" pageId="5" pageNumber="510" type="materials_examined">
<paragraph id="8BD9EB16720BFFB8FF17FEC3F66AFD05" blockId="5.[151,1436,259,1436]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF17FEC3F00BFE9A" box="[151,635,259,284]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF17FEC3F3F7FE9D" bold="true" box="[151,391,259,284]" pageId="5" pageNumber="510">Specimen examined</emphasis>
(
<emphasis id="B9123704720BFFB8FE1AFEC4F01BFE9D" bold="true" box="[410,619,259,284]" pageId="5" pageNumber="510">
total
<specimenCount id="9D60209F720BFFB8FE5AFEC3F01BFE9D" box="[474,619,259,284]" pageId="5" pageNumber="510" type="generic">4 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B720BFFB8FD05FEC4F375FEE5" collectingDate="2012-04-18" collectionCode="IZCAS" collectorName="Liu Y. &amp; Shi H. L." country="China" elevation="921" latitude="18.7251" location="Diaoluoshan Nature Reserve" longLatPrecision="1" longitude="109.86861" pageId="5" pageNumber="510" specimenCount="1" specimenCount-male="1" stateProvince="Hainan Prov." typeStatus="holotype">
<typeStatus id="54DD55B4720BFFB8FD05FEC4F082FE9D" box="[645,754,260,284]" pageId="5" pageNumber="510" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60209F720BFFB8FD7FFEC4F145FE9D" box="[767,821,260,284]" pageId="5" pageNumber="510" type="male">male</specimenCount>
(
<collectionCode id="ED7773D3720BFFB8FCC7FEC4F1EDFE9D" box="[839,925,260,284]" pageId="5" pageNumber="510">IZCAS</collectionCode>
), “
<collectingCountry id="F371AB86720BFFB8FC3EFEC4F675FE9D" box="[958,1029,260,284]" name="China" pageId="5" pageNumber="510">China</collectingCountry>
,
<collectingRegion id="49A225F4720BFFB8FB93FEC4F6DBFE9A" box="[1043,1195,260,284]" country="China" name="Hainan" pageId="5" pageNumber="510">Hainan Prov.</collectingRegion>
,
<location id="8EB9BDCD720BFFB8FB38FEC4F284FEBE" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFB8FF17FF57F3E0FA1A:8EB9BDCD720BFFB8FB38FEC4F284FEBE" country="China" latitude="18.7251" longLatPrecision="1" longitude="109.86861" name="Diaoluoshan Nature Reserve" pageId="5" pageNumber="510" stateProvince="Hainan Prov.">Diaoluoshan Nature Reserve</location>
, on tree trunk,
<geoCoordinate id="EE528DD1720BFFB8FE30FEE7F05FFEBE" box="[432,559,295,319]" degrees="18.72510" direction="north" orientation="latitude" pageId="5" pageNumber="510" precision="1" value="18.7251">N18.72510</geoCoordinate>
,
<geoCoordinate id="EE528DD1720BFFB8FDC0FEE7F0BEFEBE" box="[576,718,295,319]" degrees="109.86861" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="1" value="109.86861">E109.86861</geoCoordinate>
/ “
<quantity id="4C9E46F3720BFFB8FC82FEE7F136FEBE" box="[770,838,295,319]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="5" pageNumber="510" unit="m" value="921.0">
<elevation id="004B0C25720BFFB8FC82FEE7F136FEBE" box="[770,838,295,319]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="5" pageNumber="510" unit="m" value="921.0">921m</elevation>
</quantity>
,
<date id="FFD8CDD6720BFFB8FCD7FEE7F1B7FEBE" box="[855,967,295,319]" pageId="5" pageNumber="510" value="2012-04-18">
<collectingDate id="EF9C343E720BFFB8FCD7FEE7F1B7FEBE" box="[855,967,295,319]" pageId="5" pageNumber="510" value="2012-04-18">2012.4.18</collectingDate>
</date>
N,
<collectorName id="26938EC0720BFFB8FC79FEE7F634FEBE" box="[1017,1092,295,319]" pageId="5" pageNumber="510">Liu Y.</collectorName>
&amp;
<collectorName id="26938EC0720BFFB8FBF1FEE7F6A5FEBE" box="[1137,1237,295,319]" pageId="5" pageNumber="510">Shi H.L.</collectorName>
leg., Institute of Zoology
</materialsCitation>
/ “
<typeStatus id="54DD55B4720BFFB8FEA6FE8CF3CFFEE5" box="[294,447,332,356]" pageId="5" pageNumber="510" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C669095720BFFB8FE4BFE8CF174FEE5" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[459,772,332,356]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FE4BFE8CF04DFEE5" box="[459,573,332,356]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FDCDFE8CF08BFEE5" box="[589,763,332,356]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
</taxonomicName>
<emphasis id="B9123704720BFFB8FC8CFE8CF16BFEE5" box="[780,795,332,356]" italics="true" pageId="5" pageNumber="510">li</emphasis>
<emphasis id="B9123704720BFFB8FCA2FE8CF111FEE5" bold="true" box="[802,865,332,356]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A2218A7F720BFFB8FCA2FE8CF111FEE5" box="[802,865,332,356]" pageId="5" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [red label].
<typeStatus id="54DD55B4720BFFB8FAAFFE8CF7ECFEE5" box="[1327,1436,332,356]" pageId="5" pageNumber="510" type="paratype">Paratypes</typeStatus>
(total
<specimenCount id="9D60209F720BFFB8FF63FEAFF30DFE06" box="[227,381,367,391]" pageId="5" pageNumber="510" type="generic">3 specimens</specimenCount>
):
<materialsCitation id="3B0EE14B720BFFB8FE19FEAFF777FE2D" collectingDate="2012-04-16" collectionCode="IZCAS" collectorName="Liu Y. &amp; Shi H. L." country="China" elevation="921" latitude="18.7251" location="Diaoluoshan Nature Reserve" longLatPrecision="1" longitude="109.86861" municipality="Diaoluoshan Nature Reserve" pageId="5" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Prov." typeStatus="paratype">
<specimenCount id="9D60209F720BFFB8FE19FEAFF074FE06" box="[409,516,367,391]" pageId="5" pageNumber="510" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720BFFB8FD9DFEAFF003FE06" box="[541,627,367,391]" pageId="5" pageNumber="510">IZCAS</collectionCode>
), “
<collectingCountry id="F371AB86720BFFB8FD1BFEAFF093FE06" box="[667,739,367,391]" name="China" pageId="5" pageNumber="510">China</collectingCountry>
,
<collectingRegion id="49A225F4720BFFB8FD77FEAFF1E6FE06" box="[759,918,367,391]" country="China" name="Hainan" pageId="5" pageNumber="510">Hainan Prov.</collectingRegion>
,
<location id="8EB9BDCD720BFFB8FC2BFEAFF773FE06" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFB8FF17FF57F3E0FA1A:8EB9BDCD720BFFB8FC2BFEAFF773FE06" box="[939,1283,367,391]" country="China" latitude="18.7251" longLatPrecision="1" longitude="109.86861" municipality="Diaoluoshan Nature Reserve" name="Diaoluoshan Nature Reserve" pageId="5" pageNumber="510" stateProvince="Hainan Prov.">Diaoluoshan Nature Reserve</location>
, on ground,
<geoCoordinate id="EE528DD1720BFFB8FF17FE54F366FE2D" box="[151,278,404,428]" degrees="18.72510" direction="north" orientation="latitude" pageId="5" pageNumber="510" precision="1" value="18.7251">N18.72510</geoCoordinate>
,
<geoCoordinate id="EE528DD1720BFFB8FEA5FE54F3C0FE2D" box="[293,432,404,428]" degrees="109.86861" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="1" value="109.86861">E109.86861</geoCoordinate>
” / “
<quantity id="4C9E46F3720BFFB8FE60FE54F054FE2D" box="[480,548,404,428]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="5" pageNumber="510" unit="m" value="921.0">
<elevation id="004B0C25720BFFB8FE60FE54F054FE2D" box="[480,548,404,428]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="5" pageNumber="510" unit="m" value="921.0">921m</elevation>
</quantity>
,
<date id="FFD8CDD6720BFFB8FDB3FE54F0D2FE2D" box="[563,674,404,428]" pageId="5" pageNumber="510" value="2012-04-16">
<collectingDate id="EF9C343E720BFFB8FDB3FE54F0D2FE2D" box="[563,674,404,428]" pageId="5" pageNumber="510" value="2012-04-16">2012.4.16</collectingDate>
</date>
N,
<collectorName id="26938EC0720BFFB8FD52FE54F16BFE2A" box="[722,795,404,428]" pageId="5" pageNumber="510">Liu Y.</collectorName>
&amp;
<collectorName id="26938EC0720BFFB8FCC4FE54F1D7FE2A" box="[836,935,404,428]" pageId="5" pageNumber="510">Shi H.L.</collectorName>
leg., Institute of Zoology / “
</materialsCitation>
<typeStatus id="54DD55B4720BFFB8FA87FE54F7E8FE2D" box="[1287,1432,404,428]" pageId="5" pageNumber="510" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C669095720BFFB8FF17FE78F3A5FE4E" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,469,439,464]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FF17FE78F378FE51" box="[151,264,440,464]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FE9DFE78F3BBFE51" box="[285,459,440,464]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
</taxonomicName>
<emphasis id="B9123704720BFFB8FE60FE78F39FFE51" box="[480,495,440,464]" italics="true" pageId="5" pageNumber="510">li</emphasis>
<emphasis id="B9123704720BFFB8FE7CFE78F030FE51" bold="true" box="[508,576,440,464]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A2218A7F720BFFB8FE7CFE78F030FE51" box="[508,576,440,464]" pageId="5" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by
<collectorName id="26938EC0720BFFB8FD32FE77F0A9FE4E" box="[690,729,439,463]" pageId="5" pageNumber="510">Liu</collectorName>
&amp;
<collectorName id="26938EC0720BFFB8FC86FE77F13AFE4E" box="[774,842,439,463]" pageId="5" pageNumber="510">Liang</collectorName>
, 2012” [yellow label];
<materialsCitation id="3B0EE14B720BFFB8FBDEFE77F056FD96" collectingDate="2012-03-20" collectionCode="IZCAS" collectorName="Zhu Xiaoyu" country="China" elevation="921" latitude="18.7251" location="Diaoluoshan Nature Reserve" longLatPrecision="1" longitude="109.86861" pageId="5" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Prov." typeStatus="paratype">
<specimenCount id="9D60209F720BFFB8FBDEFE77F6B4FE4E" box="[1118,1220,439,463]" pageId="5" pageNumber="510" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720BFFB8FB57FE77F75DFE4E" box="[1239,1325,439,463]" pageId="5" pageNumber="510">IZCAS</collectionCode>
), “
<collectingCountry id="F371AB86720BFFB8FAD0FE77F7E8FE4E" box="[1360,1432,439,463]" name="China" pageId="5" pageNumber="510">China</collectingCountry>
,
<collectingRegion id="49A225F4720BFFB8FF17FE1CF341FE72" box="[151,305,476,500]" country="China" name="Hainan" pageId="5" pageNumber="510">Hainan Prov.</collectingRegion>
,
<location id="8EB9BDCD720BFFB8FEC0FE1CF0FFFE72" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFB8FF17FF57F3E0FA1A:8EB9BDCD720BFFB8FEC0FE1CF0FFFE72" box="[320,655,476,500]" country="China" latitude="18.7251" longLatPrecision="1" longitude="109.86861" name="Diaoluoshan Nature Reserve" pageId="5" pageNumber="510" stateProvince="Hainan Prov.">Diaoluoshan Nature Reserve</location>
, on ground,
<geoCoordinate id="EE528DD1720BFFB8FCA9FE1CF1D9FE75" box="[809,937,476,500]" degrees="18.72510" direction="north" orientation="latitude" pageId="5" pageNumber="510" precision="1" value="18.7251">N18.72510</geoCoordinate>
,
<geoCoordinate id="EE528DD1720BFFB8FC39FE1CF634FE75" box="[953,1092,476,500]" degrees="109.86861" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="1" value="109.86861">E109.86861</geoCoordinate>
” / “
<quantity id="4C9E46F3720BFFB8FBF8FE1CF6CBFE75" box="[1144,1211,476,500]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="5" pageNumber="510" unit="m" value="921.0">
<elevation id="004B0C25720BFFB8FBF8FE1CF6CBFE75" box="[1144,1211,476,500]" metricMagnitude="2" metricUnit="m" metricValue="9.209999999999999" pageId="5" pageNumber="510" unit="m" value="921.0">921m</elevation>
</quantity>
,
<date id="FFD8CDD6720BFFB8FB4CFE1CF74CFE75" box="[1228,1340,476,500]" pageId="5" pageNumber="510" value="2012-03-20">
<collectingDate id="EF9C343E720BFFB8FB4CFE1CF74CFE75" box="[1228,1340,476,500]" pageId="5" pageNumber="510" value="2012-03-20">2012.3.20</collectingDate>
</date>
N,
<collectorName id="26938EC0720BFFB8FAEFFE1CF299FD96" pageId="5" pageNumber="510">Zhu Xiaoyu</collectorName>
leg., Institute of Zoology
</materialsCitation>
/ “
<typeStatus id="54DD55B4720BFFB8FDCEFE3FF0AFFD96" box="[590,735,511,535]" pageId="5" pageNumber="510" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C669095720BFFB8FD6DFDC0F65AFD96" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[749,1066,511,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FD6DFDC0F12FFD99" box="[749,863,512,536]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FCF3FDC0F651FD99" box="[883,1057,512,536]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
</taxonomicName>
<emphasis id="B9123704720BFFB8FBB5FDC0F634FD99" box="[1077,1092,512,536]" italics="true" pageId="5" pageNumber="510">li</emphasis>
<emphasis id="B9123704720BFFB8FBCFFDC0F6E2FD99" bold="true" box="[1103,1170,512,536]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A2218A7F720BFFB8FBCFFDC0F6E2FD99" box="[1103,1170,512,536]" pageId="5" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by
<collectorName id="26938EC0720BFFB8FA82FE3FF759FD96" box="[1282,1321,511,535]" pageId="5" pageNumber="510">Liu</collectorName>
&amp;
<collectorName id="26938EC0720BFFB8FAD3FE3FF7E7FD96" box="[1363,1431,511,535]" pageId="5" pageNumber="510">Liang</collectorName>
, 2012” [yellow label];
<materialsCitation id="3B0EE14B720BFFB8FE23FDE4F6AAFDDE" collectingDate="2009-04-06" collectionCode="IZCAS" collectorName="Zhu Xiaoyu" country="China" elevation="982" latitude="18.74363" location="Jianfengling Nature Reserve" longLatPrecision="1" longitude="108.84311" pageId="5" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Prov." typeStatus="paratype">
<specimenCount id="9D60209F720BFFB8FE23FDE4F07CFDBD" box="[419,524,548,572]" pageId="5" pageNumber="510" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720BFFB8FDA3FDE4F009FDBD" box="[547,633,548,572]" pageId="5" pageNumber="510">IZCAS</collectionCode>
), “
<collectingCountry id="F371AB86720BFFB8FD1EFDE4F095FDBD" box="[670,741,548,572]" name="China" pageId="5" pageNumber="510">China</collectingCountry>
,
<collectingRegion id="49A225F4720BFFB8FD78FDE4F1E5FDBA" box="[760,917,548,572]" country="China" name="Hainan" pageId="5" pageNumber="510">Hainan Prov.</collectingRegion>
,
<location id="8EB9BDCD720BFFB8FC27FDE4F688FDBA" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFB8FF17FF57F3E0FA1A:8EB9BDCD720BFFB8FC27FDE4F688FDBA" box="[935,1272,548,572]" country="China" latitude="18.74363" longLatPrecision="1" longitude="108.84311" name="Jianfengling Nature Reserve" pageId="5" pageNumber="510" stateProvince="Hainan Prov.">Jianfengling Nature Reserve</location>
,
<location id="8EB9BDCD720BFFB8FA8BFDE4F7E8FDBD" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFB8FF17FF57F3E0FA1A:8EB9BDCD720BFFB8FA8BFDE4F7E8FDBD" box="[1291,1432,548,572]" country="China" latitude="18.74363" longLatPrecision="1" longitude="108.84311" municipality="Jianfengling Nature Reserve" name="Mingfenggu" pageId="5" pageNumber="510" stateProvince="Hainan Prov.">Mingfenggu</location>
, “
<geoCoordinate id="EE528DD1720BFFB8FF21FD87F353FDDE" box="[161,291,583,607]" degrees="18.74363" direction="north" orientation="latitude" pageId="5" pageNumber="510" precision="1" value="18.74363">N18.74363</geoCoordinate>
,
<geoCoordinate id="EE528DD1720BFFB8FEB5FD87F3CFFDDE" box="[309,447,583,607]" degrees="108.84311" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="1" value="108.84311">E108.84311</geoCoordinate>
” / “
<quantity id="4C9E46F3720BFFB8FE76FD87F04BFDDE" box="[502,571,583,607]" metricMagnitude="2" metricUnit="m" metricValue="9.82" pageId="5" pageNumber="510" unit="m" value="982.0">
<elevation id="004B0C25720BFFB8FE76FD87F04BFDDE" box="[502,571,583,607]" metricMagnitude="2" metricUnit="m" metricValue="9.82" pageId="5" pageNumber="510" unit="m" value="982.0">982m</elevation>
</quantity>
,
<date id="FFD8CDD6720BFFB8FDCDFD87F0C3FDDE" box="[589,691,583,607]" pageId="5" pageNumber="510" value="2009-04-06">
<collectingDate id="EF9C343E720BFFB8FDCDFD87F0C3FDDE" box="[589,691,583,607]" pageId="5" pageNumber="510" value="2009-04-06">2009.4.6</collectingDate>
</date>
6.15,
<collectorName id="26938EC0720BFFB8FC84FD87F1E0FDDE" box="[772,912,583,607]" pageId="5" pageNumber="510">Zhu Xiaoyu</collectorName>
leg., Institute of Zoology
</materialsCitation>
/ “
<typeStatus id="54DD55B4720BFFB8FA87FD87F7E8FDDE" box="[1287,1432,583,607]" pageId="5" pageNumber="510" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C669095720BFFB8FF17FDACF3A0FD05" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,464,620,644]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FF17FDACF378FD05" box="[151,264,620,644]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FE99FDACF3B7FD05" box="[281,455,620,644]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
</taxonomicName>
<emphasis id="B9123704720BFFB8FE57FDACF396FD05" box="[471,486,620,644]" italics="true" pageId="5" pageNumber="510">li</emphasis>
<emphasis id="B9123704720BFFB8FE6EFDACF05CFD05" bold="true" box="[494,556,620,644]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A2218A7F720BFFB8FE6EFDACF05CFD05" box="[494,556,620,644]" pageId="5" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label].
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF49FD50F691FB45" pageId="5" pageNumber="510" type="description">
<paragraph id="8BD9EB16720BFFB8FF49FD50F125FC95" blockId="5.[151,1436,259,1436]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF49FD50F30FFD26" box="[201,383,656,680]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF49FD50F308FD29" bold="true" box="[201,376,656,680]" pageId="5" pageNumber="510">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F3720BFFB8FE46FD4FF019FD26" box="[454,617,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.56" metricValueMin="1.51" pageId="5" pageNumber="510" unit="mm" value="15.35" valueMax="15.6" valueMin="15.1">15.115.6 mm</quantity>
(mean =
<quantity id="4C9E46F3720BFFB8FD52FD4FF14AFD26" box="[722,826,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="5" pageNumber="510" unit="mm" value="15.4">15.4 mm</quantity>
), HW =
<quantity id="4C9E46F3720BFFB8FC17FD4FF66EFD26" box="[919,1054,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="5" pageNumber="510" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.52.6 mm</quantity>
(mean =
<quantity id="4C9E46F3720BFFB8FB08FD4FF691FD26" box="[1160,1249,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="510" unit="mm" value="2.5">2.5 mm</quantity>
), EYL = 0.91.0 mm (mean =
<quantity id="4C9E46F3720BFFB8FEABFD74F3F5FD4D" box="[299,389,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="510" unit="mm" value="0.9">0.9 mm</quantity>
), PL =
<quantity id="4C9E46F3720BFFB8FE55FD74F02CFD4A" box="[469,604,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="5" pageNumber="510" unit="mm" value="3.3" valueMax="3.4" valueMin="3.2">3.23.4 mm</quantity>
(mean =
<quantity id="4C9E46F3720BFFB8FD46FD74F16FFD4D" box="[710,799,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="510" unit="mm" value="3.3">3.3 mm</quantity>
), PW =
<quantity id="4C9E46F3720BFFB8FCF8FD74F18FFD4A" box="[888,1023,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="5" pageNumber="510" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.53.8 mm</quantity>
(mean =
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), EL = 8.79.0 mm (mean =
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), EW = 5.0
<quantity id="4C9E46F3720BFFB8FE7BFD17F024FD6E" box="[507,596,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="510" unit="mm" value="5.6">5.6 mm</quantity>
(mean =
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), ratio PW/PL = 1.031.15 (mean = 1.10), EL/EW = 1.581.78 (mean = 1.66), EW/PW = 1.431.53 (mean = 1.48).
</paragraph>
<paragraph id="8BD9EB16720BFFB8FF49FCDFF730FCFE" blockId="5.[151,1436,259,1436]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF49FCDFF32AFCB9" bold="true" box="[201,346,799,824]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">Description.</emphasis>
Head and pronotum black, vertex with weak coppery shine, pronotum with strong coppery metallic shine laterally; elytra black, with a round yellow spot at posterior third of each elytron; ventral surface, coxae, trochanters, femora, tibiae and mandibles black, tarsi brown; antennomeres 13 black, 411 brown.
</paragraph>
<paragraph id="8BD9EB16720BFFB8FF49FC4CF691FB45" blockId="5.[151,1436,259,1436]" pageId="5" pageNumber="510">
Head with vertex convex, finely and sparsely punctate; occiput densely punctate; genae glabrous, oblique, as long as one-fifth of eyes in dorsal view; lateral margins of mentum rounded. Pronotum (
<figureCitation id="135DF793720BFFB8FB03FC6FF6A5FC46" box="[1155,1237,943,967]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="5" pageNumber="510">Fig. 22</figureCitation>
) ovoid, widest in the middle; lateral margins distinctly bordered, sinuate before posterior angles; disc glabrous, finely and sparsely punctate, median line distinct, shallow; posterior angles nearly rectangular, rounded at apex; base coarsely punctate; basal foveae deep, coarsely punctate. Elytra moderately convex, widest in the middle; intervals (
<figureCitation id="135DF793720BFFB8FAC2FBDCF7E3FBB5" box="[1346,1427,1052,1076]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="5" pageNumber="510">Fig. 28</figureCitation>
) convex, intervals 28 smooth and glabrous medially, with a row of punctures and pubescence laterally (near to striae), intervals 1 and 9 punctate and pubescent throughout; yellow spot covering intervals 48. Proepisterna very sparsely punctate. Claws (
<figureCitation id="135DF793720BFFB8FE3FFB48F061FB21" box="[447,529,1160,1184]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="5" pageNumber="510">Fig. 34</figureCitation>
) curved, with 68 very short teeth. Aedeagus (
<figureCitation id="135DF793720BFFB8FBA9FB48F6C1FB21" box="[1065,1201,1160,1184]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="5" pageNumber="510">Figs. 42, 43</figureCitation>
) thin in lateral view, apical lamella rounded and very short. Gonostylus (
<figureCitation id="135DF793720BFFB8FD5AFB6CF15BFB45" box="[730,811,1196,1220]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="5" pageNumber="510">Fig. 50</figureCitation>
) with six short spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF49FB10F608FB69" box="[201,1144,1232,1256]" pageId="5" pageNumber="510" type="etymology">
<paragraph id="8BD9EB16720BFFB8FF49FB10F608FB69" blockId="5.[151,1436,259,1436]" box="[201,1144,1232,1256]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF49FB10F33FFB66" box="[201,335,1232,1256]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF49FB10F338FB69" bold="true" box="[201,328,1232,1256]" pageId="5" pageNumber="510">Etymology</emphasis>
.
</emphasis>
Latinized name
<emphasis id="B9123704720BFFB8FD8BFB10F06AFB69" box="[523,538,1232,1256]" italics="true" pageId="5" pageNumber="510">li</emphasis>
refers to Li ethnic nationality living in
<collectingRegion id="49A225F4720BFFB8FC54FB10F655FB69" box="[980,1061,1232,1256]" country="China" name="Hainan" pageId="5" pageNumber="510">Hainan</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF49FB34F3E0FA1A" pageId="5" pageNumber="510" type="discussion">
<paragraph id="8BD9EB16720BFFB8FF49FB34F3E0FA1A" blockId="5.[151,1436,259,1436]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF49FB34F34CFA8A" box="[201,316,1268,1292]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF49FB34F346FA8D" bold="true" box="[201,310,1268,1292]" pageId="5" pageNumber="510">Remarks</emphasis>
.
</emphasis>
<emphasis id="B9123704720BFFB8FEC7FB34F3A2FA8D" box="[327,466,1268,1292]" italics="true" pageId="5" pageNumber="510">
<taxonomicName id="4C669095720BFFB8FEC7FB34F3C9FA8D" box="[327,441,1268,1292]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">Chlaenius</taxonomicName>
li
</emphasis>
is an endemic species to
<collectingRegion id="49A225F4720BFFB8FC81FB34F122FA8D" box="[769,850,1268,1292]" country="China" name="Hainan" pageId="5" pageNumber="510">Hainan</collectingRegion>
Island (
<figureCitation id="135DF793720BFFB8FC33FB34F67AFA8D" box="[947,1034,1268,1292]" captionStart="FIGURE 52" captionStartId="12.[151,250,1844,1867]" captionTargetBox="[151,1436,935,1822]" captionTargetId="figure-69@12.[151,1436,935,1822]" captionTargetPageId="12" captionText="FIGURE 52. Geographical distribution of Chlaenius (Chlaenioctenus) species." figureDoi="http://doi.org/10.5281/zenodo.5261797" httpUri="https://zenodo.org/record/5261797/files/figure.png" pageId="5" pageNumber="510">Fig. 52</figureCitation>
). It is similar to
<taxonomicName id="4C669095720BFFB8FB50FB34F756FA8D" authorityName="Jedlicka" authorityYear="1960" box="[1232,1318,1268,1292]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720BFFB8FB50FB34F756FA8D" box="[1232,1318,1268,1292]" italics="true" pageId="5" pageNumber="510">C. freyi</emphasis>
</taxonomicName>
in having wholly black femora, and elytron with yellow spot at posterior third, but differs from the latter in head, pronotum glabrous and each of intervals 28 glabrous medially (
<figureCitation id="135DF793720BFFB8FC9BFAFCF103FAD5" box="[795,883,1340,1364]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="5" pageNumber="510">Fig. 28</figureCitation>
), gonostylus (
<figureCitation id="135DF793720BFFB8FB98FAFCF61EFAD5" box="[1048,1134,1340,1364]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="5" pageNumber="510">Fig. 50</figureCitation>
) with six spines on outer margin in contrast to head and pronotum pubescent, all elytral intervals regularly pubescent, gonostylus with only four spines in
<taxonomicName id="4C669095720BFFB8FEB6FA44F3F9FA1D" authorityName="Jedlicka" authorityYear="1960" box="[310,393,1412,1436]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720BFFB8FEB6FA44F3F9FA1D" box="[310,393,1412,1436]" italics="true" pageId="5" pageNumber="510">C. freyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,231 @@
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<mods:title id="871A3D076742CA679AE9B77FCECE5DA9">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<treatment id="03CF5A00720BFFBAFF17FA25F602FD96" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFBAFF17FA25F602FD96" httpUri="http://treatment.plazi.org/id/03CF5A00720BFFBAFF17FA25F602FD96" lastPageId="7" lastPageNumber="512" pageId="5" pageNumber="510">
<subSubSection id="C37CB89D720BFFB8FF17FA25F175FA7C" box="[151,773,1509,1534]" pageId="5" pageNumber="510" type="nomenclature">
<paragraph id="8BD9EB16720BFFB8FF17FA25F175FA7C" blockId="5.[151,773,1509,1569]" box="[151,773,1509,1534]" pageId="5" pageNumber="510">
<taxonomicName id="4C669095720BFFB8FF17FA25F0CEFA7F" ID-CoL="5XW7J" authority="Liu &amp; Liang" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,702,1509,1534]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="aodai" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FF17FA25F37DFA7F" bold="true" box="[151,269,1509,1534]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FE9DFA25F3A1FA7F" bold="true" box="[285,465,1509,1534]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720BFFB8FE62FA25F0CEFA7F" bold="true" box="[482,702,1509,1534]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FE62FA25F051FA7F" bold="true" box="[482,545,1509,1534]" italics="true" pageId="5" pageNumber="510">aodai</emphasis>
Liu &amp; Liang
</emphasis>
</taxonomicName>
<emphasis id="B9123704720BFFB8FD45FA25F175FA7C" bold="true" box="[709,773,1509,1533]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A2218A7F720BFFB8FD45FA25F175FA7C" box="[709,773,1509,1533]" pageId="5" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF17F9C9F070F9A0" box="[151,512,1545,1569]" pageId="5" pageNumber="510" type="description">
<paragraph id="8BD9EB16720BFFB8FF17F9C9F070F9A0" blockId="5.[151,773,1509,1569]" box="[151,512,1545,1569]" pageId="5" pageNumber="510">
(
<figureCitation id="135DF793720BFFB8FF1FF9C9F356F9A0" box="[159,294,1545,1569]" captionStart="FIGURES 1718" captionStartId="9.[151,264,1143,1166]" captionTargetBox="[164,1388,193,1122]" captionTargetId="figure-349@9.[151,1436,193,1122]" captionTargetPageId="9" captionText="FIGURES 1718. Chlaenius aodai sp. n. 17, Holotype, male, dorsal view. 18, Holotype, male, ventral view. FIGURE 19. Chlaenius langsonensis Kirschenhofer, 2008. Holotype, male. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261779" httpUri="https://zenodo.org/record/5261779/files/figure.png" pageId="5" pageNumber="510">Figs. 17, 18</figureCitation>
,
<figureCitation id="135DF793720BFFB8FEB2F9C9F320F9A0" box="[306,336,1545,1569]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="5" pageNumber="510">23</figureCitation>
,
<figureCitation id="135DF793720BFFB8FEDCF9C9F30AF9A0" box="[348,378,1545,1569]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="5" pageNumber="510">29</figureCitation>
,
<figureCitation id="135DF793720BFFB8FE06F9C9F3D4F9A0" box="[390,420,1545,1569]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="5" pageNumber="510">35</figureCitation>
,
<figureCitation id="135DF793720BFFB8FE30F9C9F387F9A0" box="[432,503,1545,1569]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="5" pageNumber="510">44, 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFB8FF17F991F1DDF930" pageId="5" pageNumber="510" type="materials_examined">
<paragraph id="8BD9EB16720BFFB8FF17F991F1DDF930" blockId="5.[151,1436,1617,2037]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF17F991F0D7F9E8" box="[151,679,1617,1642]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF17F991F3E2F9EB" bold="true" box="[151,402,1617,1642]" pageId="5" pageNumber="510">Specimen examined</emphasis>
(
<emphasis id="B9123704720BFFB8FE30F991F0E6F9EB" bold="true" box="[432,662,1617,1642]" pageId="5" pageNumber="510">
total
<specimenCount id="9D60209F720BFFB8FE7BF991F0E6F9EB" box="[507,662,1617,1642]" pageId="5" pageNumber="510" type="generic">1 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B720BFFB8FD3CF991F635F90C" collectingDate="1989-05-24" collectorName="Pacholatko" country="Vietnam" location="Tamdao" pageId="5" pageNumber="510" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DD55B4720BFFB8FD3CF991F158F9E8" box="[700,808,1617,1641]" pageId="5" pageNumber="510" type="holotype">Holotype</typeStatus>
: male (NHMB), “N.
<collectingCountry id="F371AB86720BFFB8FBCAF991F6D9F9E8" box="[1098,1193,1617,1641]" name="Vietnam" pageId="5" pageNumber="510">Vietnam</collectingCountry>
(Tonkin),
<location id="8EB9BDCD720BFFB8FABBF991F7E8F9E8" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFBAFF17FA25F602FD96:8EB9BDCD720BFFB8FABBF991F7E8F9E8" box="[1339,1432,1617,1641]" country="Vietnam" name="Tamdao" pageId="5" pageNumber="510">Tamdao</location>
, 12
<date id="FFD8CDD6720BFFB8FF40F9B5F346F90C" box="[192,310,1653,1677]" pageId="5" pageNumber="510" value="1989-05-24">
<collectingDate id="EF9C343E720BFFB8FF40F9B5F346F90C" box="[192,310,1653,1677]" pageId="5" pageNumber="510" value="1989-05-24">24.5.1989</collectingDate>
</date>
,
<collectorName id="26938EC0720BFFB8FECCF9B5F3B8F90C" box="[332,456,1653,1677]" pageId="5" pageNumber="510">Pacholatko</collectorName>
leg.” / “Freiwlliger, Museumsverein, Basel, 1989”
</materialsCitation>
/ “
<typeStatus id="54DD55B4720BFFB8FBFBF9B5F764F90C" box="[1147,1300,1653,1677]" pageId="5" pageNumber="510" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C669095720BFFB8FAAAF9B5F3EFF930" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="aodai" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720BFFB8FAAAF9B5F7EBF90C" box="[1322,1435,1653,1677]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
(
<emphasis id="B9123704720BFFB8FF20F959F33EF930" box="[160,334,1689,1713]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720BFFB8FEDFF959F3EFF930" box="[351,415,1689,1713]" italics="true" pageId="5" pageNumber="510">aodai</emphasis>
</taxonomicName>
<emphasis id="B9123704720BFFB8FE25F959F394F930" bold="true" box="[421,484,1689,1713]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A2218A7F720BFFB8FE25F959F394F930" box="[421,484,1689,1713]" pageId="5" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012 [red label].
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720BFFBAFF49F97DF7FCFEBE" lastPageId="7" lastPageNumber="512" pageId="5" pageNumber="510" type="description">
<paragraph id="8BD9EB16720BFFB8FF49F97DF1F5F978" blockId="5.[151,1436,1617,2037]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF49F97DF30FF954" box="[201,383,1725,1749]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF49F97DF308F954" bold="true" box="[201,376,1725,1749]" pageId="5" pageNumber="510">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F3720BFFB8FE4AF97DF040F954" box="[458,560,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="5" pageNumber="510" unit="mm" value="16.6">16.6 mm</quantity>
, HW =
<quantity id="4C9E46F3720BFFB8FD0BF97DF093F954" box="[651,739,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="510" unit="mm" value="2.6">2.6 mm</quantity>
, EYL = 1.0 mm, PL =
<quantity id="4C9E46F3720BFFB8FC6AF97DF633F954" box="[1002,1091,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="510" unit="mm" value="3.5">3.5 mm</quantity>
, PW =
<quantity id="4C9E46F3720BFFB8FB18F97DF681F954" box="[1176,1265,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="510" unit="mm" value="3.5">3.5 mm</quantity>
, EL =
<quantity id="4C9E46F3720BFFB8FABFF97DF7E7F954" box="[1343,1431,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="510" unit="mm" value="9.5">9.5 mm</quantity>
, EW =
<quantity id="4C9E46F3720BFFB8FF60F921F348F978" box="[224,312,1761,1785]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="510" unit="mm" value="5.2">5.2 mm</quantity>
, ratio PL/PW = 1.0, EL/EW = 1.83, EW/PW = 1.49.
</paragraph>
<paragraph id="8BD9EB16720BFFB8FF49F8C5F3A9F808" blockId="5.[151,1436,1617,2037]" pageId="5" pageNumber="510">
<emphasis id="B9123704720BFFB8FF49F8C5F32AF89C" box="[201,346,1797,1822]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
<emphasis id="B9123704720BFFB8FF49F8C5F323F89F" bold="true" box="[201,339,1797,1822]" pageId="5" pageNumber="510">Description</emphasis>
.
</emphasis>
Head and pronotum black, with distinct metallic shine; elytra black, with weak shine; coxae, trochanters, tibiae, antennomeres 2 and 3 black or dark brown; antennomeres 1 and 411, mandibles, palpi and tarsi reddish brown; femora yellow in the middle portion, black at base and apex, basal black areas on hind femora longer than hind trochanters.
</paragraph>
<paragraph id="8BD9EB16720BFFBAFF49F855F7FCFEBE" blockId="5.[151,1436,1617,2037]" lastBlockId="7.[151,1436,151,536]" lastPageId="7" lastPageNumber="512" pageId="5" pageNumber="510">
Head convex, vertex and occiput punctate; genae glabrous, oblique, as long as one-fourth of eyes in dorsal view; lateral margins of mentum rounded. Pronotum (
<figureCitation id="135DF793720BFFB8FC84F879F128F850" box="[772,856,1977,2001]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="5" pageNumber="510">Fig. 23</figureCitation>
) long-ovoid, widest in the middle; lateral margins indistinctly bordered, sinuate before posterior angles; disc finely punctate, glabrous, median line distinct; posterior angles nearly rectangular, rounded at apex; base punctate, basal foveae deep, sparsely punctate. Elytra moderately convex, widest near posterior third; intervals (
<figureCitation id="135DF7937209FFBAFD2DFF7CF172FF55" box="[685,770,188,212]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="7" pageNumber="512">Fig. 29</figureCitation>
) slightly convex, with regular punctures and pubescence; yellow spot at posterior third of each elytron, small, covering half interval 4 to interval 8. Proepisterna very sparsely punctate. Hind femora with basal black areas slightly longer than hind trochanters, claws (
<figureCitation id="135DF7937209FFBAFB69FEC4F749FE9D" box="[1257,1337,260,284]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="7" pageNumber="512">Fig. 35</figureCitation>
) curved, with 56 short teeth. Aedeagus (
<figureCitation id="135DF7937209FFBAFD83FEE7F0FAFEBE" box="[515,650,295,319]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="7" pageNumber="512">Figs. 44, 45</figureCitation>
) moderately robust in lateral view, apical lamella rounded and short.
</paragraph>
</subSubSection>
<caption id="DF19BB9E7208FFBBFF17F8B5F132F82B" ID-DOI="http://doi.org/10.5281/zenodo.5261773" ID-Zenodo-Dep="5261773" httpUri="https://zenodo.org/record/5261773/files/figure.png" pageId="6" pageNumber="511" startId="6.[151,264,1909,1932]" targetBox="[282,1273,195,1883]" targetPageId="6">
<paragraph id="8BD9EB167208FFBBFF17F8B5F132F82B" blockId="6.[151,1436,1909,1962]" pageId="6" pageNumber="511">
<emphasis id="B91237047208FFBBFF17F8B5F339F80D" bold="true" box="[151,329,1909,1932]" pageId="6" pageNumber="511">FIGURES 912.</emphasis>
<emphasis id="B91237047208FFBBFED0F8B6F3BDF80D" box="[336,461,1910,1932]" italics="true" pageId="6" pageNumber="511">
<taxonomicName id="4C6690957208FFBBFED0F8B6F3C7F80D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[336,439,1910,1932]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="genus" status="sp. nov.">Chlaenius</taxonomicName>
li
</emphasis>
<emphasis id="B91237047208FFBBFE55F8B6F060F80A" bold="true" box="[469,528,1910,1931]" pageId="6" pageNumber="511">
<taxonomicNameLabel id="A2218A7F7208FFBBFE55F8B6F060F80A" box="[469,528,1910,1931]" pageId="6" pageNumber="511" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
9, Holotype, male, dorsal view. 10, Holotype, male, ventral view. 11, Paratype, female, dorsal view. 12, Paratype, female, ventral view. Scale bar = 5.0 mm
</paragraph>
</caption>
<subSubSection id="C37CB89D7209FFBAFF49FE8CF0FBFE06" pageId="7" pageNumber="512" type="etymology">
<paragraph id="8BD9EB167209FFBAFF49FE8CF0FBFE06" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FE8CF33FFEE2" box="[201,335,332,356]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FE8CF338FEE5" bold="true" box="[201,328,332,356]" pageId="7" pageNumber="512">Etymology</emphasis>
.
</emphasis>
Latinized name
<taxonomicName id="4C6690957209FFBAFD89FE8CF039FEE5" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[521,585,332,356]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="aodai">
<emphasis id="B91237047209FFBAFD89FE8CF039FEE5" box="[521,585,332,356]" italics="true" pageId="7" pageNumber="512">aodai</emphasis>
</taxonomicName>
refers to Ao Dai, a traditional costume of Vietnamese women, implying that the new species is discovered from
<collectingCountry id="F371AB867209FFBAFDA5FEAFF0F7FE06" box="[549,647,367,391]" name="Vietnam" pageId="7" pageNumber="512">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D7209FFBAFF49FE54F602FD96" pageId="7" pageNumber="512" type="discussion">
<paragraph id="8BD9EB167209FFBAFF49FE54F602FD96" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="512">
<emphasis id="B91237047209FFBAFF49FE54F34CFE2A" box="[201,316,404,428]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
<emphasis id="B91237047209FFBAFF49FE54F346FE2D" bold="true" box="[201,310,404,428]" pageId="7" pageNumber="512">Remarks</emphasis>
.
</emphasis>
<taxonomicName id="4C6690957209FFBAFEC3FE54F38BFE2D" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[323,507,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="aodai">
<emphasis id="B91237047209FFBAFEC3FE54F38BFE2D" box="[323,507,404,428]" italics="true" pageId="7" pageNumber="512">Chlaenius aodai</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C6690957209FFBAFD0BFE54F097FE2D" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[651,743,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B91237047209FFBAFD0BFE54F097FE2D" box="[651,743,404,428]" italics="true" pageId="7" pageNumber="512">C. cheni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6690957209FFBAFC9DFE54F1FEFE2D" authorityName="Kirschenhofer" authorityYear="2008" box="[797,910,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B91237047209FFBAFC9DFE54F1FEFE2D" box="[797,910,404,428]" italics="true" pageId="7" pageNumber="512">C. sehnali</emphasis>
</taxonomicName>
in having yellow spot at posterior third of each elytron, but differs from
<taxonomicName id="4C6690957209FFBAFE30FE78F07FFE51" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[432,527,440,464]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B91237047209FFBAFE30FE78F07FFE51" box="[432,527,440,464]" italics="true" pageId="7" pageNumber="512">C. cheni</emphasis>
</taxonomicName>
in having pronotum wider than long contrast to pronotum wider than long in
<taxonomicName id="4C6690957209FFBAFA02FE78F2A4FE75" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B91237047209FFBAFA02FE78F2A4FE75" italics="true" pageId="7" pageNumber="512">C. cheni</emphasis>
</taxonomicName>
, and from
<taxonomicName id="4C6690957209FFBAFED4FE1CF3B7FE75" authorityName="Kirschenhofer" authorityYear="2008" box="[340,455,476,500]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B91237047209FFBAFED4FE1CF3B7FE75" box="[340,455,476,500]" italics="true" pageId="7" pageNumber="512">C. sehnali</emphasis>
</taxonomicName>
in having all elytral intervals regularly pubescent and pronotal disc finely punctate in contrast to intervals glabrous medially and pronotal disc coarsely punctate in
<taxonomicName id="4C6690957209FFBAFC7AFDC0F61BFD99" authorityName="Kirschenhofer" authorityYear="2008" box="[1018,1131,512,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B91237047209FFBAFC7AFDC0F61BFD99" box="[1018,1131,512,536]" italics="true" pageId="7" pageNumber="512">C. sehnali</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="63E91D6D84DE29F496E4585EC129C0CC" ID-DOI="10.11646/zootaxa.3630.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261763" ID-ZooBank="C7529E9C-73F7-4E52-92E7-FCACA8F99A7D" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629846447618" checkinUser="felipe" docAuthor="Liu, Ye, Shi, Hongliang &amp; Liang, Hongbin" docDate="2013" docId="03CF5A00720DFFBEFF17FC83F1FEF85D" docLanguage="en" docName="zt03630p518.pdf" docOrigin="Zootaxa 3630 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Chlaenioctenus) bodhidharma Liu &amp; Shi &amp; Liang 2013, sp. n." docType="treatment" docVersion="1" lastPageNumber="508" masterDocId="FFF62278720EFFBDFF80FFC0F270FF81" masterDocTitle="Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus" masterLastPageNumber="518" masterPageNumber="505" pageNumber="508" updateTime="1730973406347" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="9D571337E0115D9B2BBA849E0E7FDB1F">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:namePart id="64B58B08F5B519001F25D79F616B8A23">Liu, Ye</mods:namePart>
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<mods:namePart id="A4B931F6E3A8C2A7AE6FE68E07580C65">Shi, Hongliang</mods:namePart>
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<mods:namePart id="501D1ACD7E3C389BDA49A088880C18D8">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="A05E257E18054389A1E7F626B50CDDA3" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:date id="DDB16C82E1918F05A48532AAE2CAFC95">2013</mods:date>
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<mods:number id="A2F5EFD6900EB450D8F7335EA22510A5">2013-03-21</mods:number>
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<treatment id="03CF5A00720DFFBEFF17FC83F1FEF85D" LSID="urn:lsid:plazi:treatment:03CF5A00720DFFBEFF17FC83F1FEF85D" httpUri="http://treatment.plazi.org/id/03CF5A00720DFFBEFF17FC83F1FEF85D" lastPageNumber="508" pageId="3" pageNumber="508">
<subSubSection id="C37CB89D720DFFBEFF17FC83F111FCDD" box="[151,865,835,860]" pageId="3" pageNumber="508" type="nomenclature">
<paragraph id="8BD9EB16720DFFBEFF17FC83F111FCDD" blockId="3.[151,865,835,895]" box="[151,865,835,860]" pageId="3" pageNumber="508">
<taxonomicName id="4C669095720DFFBEFF17FC83F16AFCDD" ID-CoL="5Y87J" authority="Liu &amp; Liang" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,794,835,860]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFF17FC83F37DFCDD" bold="true" box="[151,269,835,860]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFE9DFC83F3A1FCDD" bold="true" box="[285,465,835,860]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720DFFBEFE62FC83F16AFCDD" bold="true" box="[482,794,835,860]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFE62FC83F00DFCDD" bold="true" box="[482,637,835,860]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
Liu &amp; Liang
</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFCA1FC84F111FCDD" bold="true" box="[801,865,836,860]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFCA1FC84F111FCDD" box="[801,865,836,860]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF17FCA7F07FFCFE" box="[151,527,871,895]" pageId="3" pageNumber="508" type="description">
<paragraph id="8BD9EB16720DFFBEFF17FCA7F07FFCFE" blockId="3.[151,865,835,895]" box="[151,527,871,895]" pageId="3" pageNumber="508">
(
<figureCitation id="135DF793720DFFBEFF1FFCA7F37DFCFE" box="[159,269,871,895]" captionStart="FIGURES 58" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 58. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="3" pageNumber="508">Figs. 58</figureCitation>
,
<figureCitation id="135DF793720DFFBEFE98FCA7F346FCFE" box="[280,310,871,895]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="3" pageNumber="508">21</figureCitation>
,
<figureCitation id="135DF793720DFFBEFEC2FCA7F310FCFE" box="[322,352,871,895]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="3" pageNumber="508">27</figureCitation>
,
<figureCitation id="135DF793720DFFBEFEECFCA7F3FAFCFE" box="[364,394,871,895]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="3" pageNumber="508">33</figureCitation>
,
<figureCitation id="135DF793720DFFBEFE16FCA7F3AEFCFE" box="[406,478,871,895]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">40, 41</figureCitation>
,
<figureCitation id="135DF793720DFFBEFE6AFCA7F077FCFE" box="[490,519,871,895]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF17FC6FF382FB1E" pageId="3" pageNumber="508" type="materials_examined">
<paragraph id="8BD9EB16720DFFBEFF17FC6FF382FB1E" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF17FC6FF0F4FC46" box="[151,644,943,968]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF17FC6FF3F9FC49" bold="true" box="[151,393,943,968]" pageId="3" pageNumber="508">Specimen examined</emphasis>
(
<emphasis id="B9123704720DFFBEFE1EFC70F004FC49" bold="true" box="[414,628,943,968]" pageId="3" pageNumber="508">
total
<specimenCount id="9D60209F720DFFBEFE61FC6FF004FC49" box="[481,628,943,968]" pageId="3" pageNumber="508" type="generic">3 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B720DFFBEFD10FC6FF3EDFC6D" collectingDate="1982-04-25" collectorName="Ch. J. Rai'" elevation="2900" location="Alghera" pageId="3" pageNumber="508" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DD55B4720DFFBEFD10FC6FF08DFC46" box="[656,765,943,967]" pageId="3" pageNumber="508" type="holotype">Holotype</typeStatus>
: male (NHMB), “Darjeeling D.,
<collectorName id="26938EC0720DFFBEFB05FC6FF77AFC46" box="[1157,1290,943,967]" pageId="3" pageNumber="508">Ch. J. Rai</collectorName>
/ “
<location id="8EB9BDCD720DFFBEFAB9FC6FF7E8FC46" LSID="urn:lsid:plazi:treatment:03CF5A00720DFFBEFF17FC83F1FEF85D:8EB9BDCD720DFFBEFAB9FC6FF7E8FC46" box="[1337,1432,943,967]" name="Alghera" pageId="3" pageNumber="508">Alghera</location>
,
<quantity id="4C9E46F3720DFFBEFF17FC14F298FC6D" box="[151,232,980,1004]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">
<elevation id="004B0C25720DFFBEFF17FC14F298FC6D" box="[151,232,980,1004]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">2900m</elevation>
</quantity>
,
<date id="FFD8CDD6720DFFBEFF72FC14F303FC6D" box="[242,371,980,1004]" pageId="3" pageNumber="508" value="1982-04-25">
<collectingDate id="EF9C343E720DFFBEFF72FC14F303FC6D" box="[242,371,980,1004]" pageId="3" pageNumber="508" value="1982-04-25">25.IV.1982</collectingDate>
</date>
/ “
</materialsCitation>
<typeStatus id="54DD55B4720DFFBEFE1DFC14F046FC6D" box="[413,566,980,1004]" pageId="3" pageNumber="508" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C669095720DFFBEFDC1FC14F667FC6D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[577,1047,980,1004]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFDC1FC14F0C3FC6D" box="[577,691,980,1004]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFD42FC14F100FC6D" box="[706,880,980,1004]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720DFFBEFC00FC14F667FC6D" box="[896,1047,980,1004]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFB9EFC14F62CFC6D" bold="true" box="[1054,1116,980,1004]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFB9EFC14F62CFC6D" box="[1054,1116,980,1004]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [red label].
<materialsCitation id="3B0EE14B720DFFBEFEA3FC38F333FBB5" collectingDate="1982-04-25" elevation="2900" pageId="3" pageNumber="508" specimenCount="3" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54DD55B4720DFFBEFEA3FC38F3E0FB91" box="[291,400,1016,1040]" pageId="3" pageNumber="508" type="paratype">Paratypes</typeStatus>
(total
<specimenCount id="9D60209F720DFFBEFE63FC37F008FB8E" box="[483,632,1015,1039]" pageId="3" pageNumber="508" type="generic">2 specimens</specimenCount>
):
<specimenCount id="9D60209F720DFFBEFD0FFC37F090FB8E" box="[655,736,1015,1039]" pageId="3" pageNumber="508" type="male">1 male</specimenCount>
(NHMB), “Darjeeling D., Ch. J. Rai / “Alghera,
<quantity id="4C9E46F3720DFFBEFAC7FC37F7E8FB8E" box="[1351,1432,1015,1039]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">
<elevation id="004B0C25720DFFBEFAC7FC37F7E8FB8E" box="[1351,1432,1015,1039]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="508" unit="m" value="2900.0">2900m</elevation>
</quantity>
,
<date id="FFD8CDD6720DFFBEFF17FBDCF368FBB5" box="[151,280,1052,1076]" pageId="3" pageNumber="508" value="1982-04-25">
<collectingDate id="EF9C343E720DFFBEFF17FBDCF368FBB5" box="[151,280,1052,1076]" pageId="3" pageNumber="508" value="1982-04-25">25.IV.1982</collectingDate>
</date>
/ “
</materialsCitation>
<typeStatus id="54DD55B4720DFFBEFEC3FBDCF3A4FBB5" box="[323,468,1052,1076]" pageId="3" pageNumber="508" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C669095720DFFBEFE5FFBDCF1C6FBB5" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[479,950,1052,1076]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFE5FFBDCF021FBB5" box="[479,593,1052,1076]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFDE1FBDCF17FFBB5" box="[609,783,1052,1076]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720DFFBEFC9FFBDCF1C6FBB5" box="[799,950,1052,1076]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFC3DFBDCF18CFBB5" bold="true" box="[957,1020,1052,1076]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFC3DFBDCF18CFBB5" box="[957,1020,1052,1076]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by
<collectorName id="26938EC0720DFFBEFBE1FBDCF6F8FBB5" box="[1121,1160,1052,1076]" pageId="3" pageNumber="508">Liu</collectorName>
&amp;
<collectorName id="26938EC0720DFFBEFB2BFBDCF69FFBB5" box="[1195,1263,1052,1076]" pageId="3" pageNumber="508">Liang</collectorName>
, 2012” [yellow label];
<materialsCitation id="3B0EE14B720DFFBEFF67FBFFF7EBFBD6" box="[231,1435,1087,1111]" elevation="1800.0" pageId="3" pageNumber="508">
<specimenCount id="9D60209F720DFFBEFF67FBFFF33AFBD6" box="[231,330,1087,1111]" pageId="3" pageNumber="508" type="female">1 female</specimenCount>
(NHMB), “
<collectingCountry id="F371AB86720DFFBEFE53FBFFF06FFBD6" box="[467,543,1087,1111]" name="India" pageId="3" pageNumber="508">Indien</collectingCountry>
, Darjeeling D., B. Bhakta / “Sukhay Simana,
<quantity id="4C9E46F3720DFFBEFBC0FBFFF6A8FBD6" box="[1088,1240,1087,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="3" pageNumber="508" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">
<elevation id="004B0C25720DFFBEFBC0FBFFF6A8FBD6" box="[1088,1240,1087,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="3" pageNumber="508" unit="m" value="1800.0" valueMax="2000.0" valueMin="1600.0">16002000m</elevation>
</quantity>
, 31.IV11.V.86
</materialsCitation>
[date 31.IV is wrong, possibly 30.IV] / “
<typeStatus id="54DD55B4720DFFBEFDD9FBA4F09AFBFD" box="[601,746,1124,1148]" pageId="3" pageNumber="508" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C669095720DFFBEFD74FBA4F6BBFBFD" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[756,1227,1124,1148]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720DFFBEFD74FBA4F115FBFD" box="[756,869,1124,1148]" italics="true" pageId="3" pageNumber="508">Chlaenius</emphasis>
(
<emphasis id="B9123704720DFFBEFCF6FBA4F654FBFD" box="[886,1060,1124,1148]" italics="true" pageId="3" pageNumber="508">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720DFFBEFBB4FBA4F6BBFBFD" box="[1076,1227,1124,1148]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720DFFBEFB52FBA4F761FBFD" bold="true" box="[1234,1297,1124,1148]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A2218A7F720DFFBEFB52FBA4F761FBFD" box="[1234,1297,1124,1148]" pageId="3" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label].
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49FB6CF74AF885" pageId="3" pageNumber="508" type="description">
<paragraph id="8BD9EB16720DFFBEFF49FB6CF008FAAE" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FB6CF30FFB42" box="[201,383,1196,1220]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49FB6CF308FB45" bold="true" box="[201,376,1196,1220]" pageId="3" pageNumber="508">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F3720DFFBEFE4EFB6CF004FB42" box="[462,628,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" metricValueMax="1.69" metricValueMin="1.53" pageId="3" pageNumber="508" unit="mm" value="16.1" valueMax="16.9" valueMin="15.3">15.316.9 mm</quantity>
(mean =
<quantity id="4C9E46F3720DFFBEFD65FB6CF13FFB45" box="[741,847,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="3" pageNumber="508" unit="mm" value="15.9">15.9 mm</quantity>
), HW =
<quantity id="4C9E46F3720DFFBEFC35FB6CF64FFB42" box="[949,1087,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="3" pageNumber="508" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.42.9 mm</quantity>
(mean =
<quantity id="4C9E46F3720DFFBEFB30FB6CF77CFB45" box="[1200,1292,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="508" unit="mm" value="2.7">2.7 mm</quantity>
), EYL = 1.0 mm, PL =
<quantity id="4C9E46F3720DFFBEFE95FB0FF3EFFB66" box="[277,415,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.6" metricValueMin="3.3" pageId="3" pageNumber="508" unit="mm" value="3.45" valueMax="3.6" valueMin="3.3">3.33.6 mm</quantity>
(mean =
<quantity id="4C9E46F3720DFFBEFD90FB0FF01CFB66" box="[528,620,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="508" unit="mm" value="3.4">3.4 mm</quantity>
), PW =
<quantity id="4C9E46F3720DFFBEFD4DFB0FF128FB66" box="[717,856,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="3" pageNumber="508" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.74.3 mm</quantity>
(mean = 4.0 mm), EL =
<quantity id="4C9E46F3720DFFBEFBFDFB0FF778FB66" box="[1149,1288,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="9.149999999999999" metricValueMax="9.6" metricValueMin="8.7" pageId="3" pageNumber="508" unit="mm" value="9.149999999999999" valueMax="9.6" valueMin="8.7">8.79.6 mm</quantity>
(mean = 9.0 mm), EW =
<quantity id="4C9E46F3720DFFBEFEA3FB34F3DBFA8A" box="[291,427,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="6.1" metricValueMin="5.4" pageId="3" pageNumber="508" unit="mm" value="5.75" valueMax="6.1" valueMin="5.4">5.46.1 mm</quantity>
(mean =
<quantity id="4C9E46F3720DFFBEFD96FB34F000FA8D" box="[534,624,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="508" unit="mm" value="5.6">5.6 mm</quantity>
), ratio PW/PL = 1.121.19 (mean = 1.15), EL/EW = 1.561.63 (mean = 1.60), EW/PW = 1.381.46 (mean = 1.42).
</paragraph>
<paragraph id="8BD9EB16720DFFBEFF49FAFBF36DFA3E" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FAFBF32AFAD5" bold="true" box="[201,346,1339,1364]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">Description.</emphasis>
Head and pronotum black, pronotum with green metallic shine laterally; elytra black, with a yellow spot in the middle of each elytron; ventral surface, coxae, trochanters, tibiae black, femora yellow in the middle, black at base and apex (less so in female,
<figureCitation id="135DF793720DFFBEFD42FA44F174FA1D" box="[706,772,1412,1436]" captionStart="FIGURES 58" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 58. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="3" pageNumber="508">Fig. 8</figureCitation>
); antennomeres yellow or reddish brown; mandibles, palpi, tarsi brown.
</paragraph>
<paragraph id="8BD9EB16720DFFBEFF49FA0CF74AF885" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
Head convex, vertex and occiput finely punctate; genae glabrous, oblique, as long as one-fifth of eyes in dorsal view; lateral margins of mentum nearly straight. Pronotum (
<figureCitation id="135DF793720DFFBEFCC3FA2FF1E5F986" box="[835,917,1519,1543]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="3" pageNumber="508">Fig. 21</figureCitation>
) moderately ovoid, widest slightly before the middle; lateral margins distinctly bordered, slightly sinuate before posterior angles; disc finely punctate, glabrous, median line distinct; posterior angles rounded; base punctate; basal foveae deep, coarsely punctate. Elytra moderately convex, widest slightly behind the middle; intervals 28 convex, glabrous medially, with a row of punctures laterally (near striae) (
<figureCitation id="135DF793720DFFBEFD91F9BFF018F916" box="[529,616,1663,1687]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="3" pageNumber="508">Fig. 27</figureCitation>
), intervals 1 and 9 irregularly pubescent; yellow spot covering half of interval 4 to interval 8. Proepisterna sparsely punctate. Hind femora with basal black areas slightly longer than hind trochanters; claws (
<figureCitation id="135DF793720DFFBEFEF8F908F3BCF961" box="[376,460,1736,1760]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="3" pageNumber="508">Fig. 33</figureCitation>
) curved, with 79 long teeth. Aedeagus (
<figureCitation id="135DF793720DFFBEFC33F908F630F961" box="[947,1088,1736,1760]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Figs. 40, 41</figureCitation>
) robust in lateral view, apical lamella narrow and long. Gonostylus (
<figureCitation id="135DF793720DFFBEFDC5F92CF0E6F885" box="[581,662,1772,1796]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">Fig. 49</figureCitation>
) of female ovipositor with five long spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49F8D0F6B7F8A9" box="[201,1223,1808,1832]" pageId="3" pageNumber="508" type="etymology">
<paragraph id="8BD9EB16720DFFBEFF49F8D0F6B7F8A9" blockId="3.[151,1436,943,2012]" box="[201,1223,1808,1832]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49F8D0F33FF8A6" box="[201,335,1808,1832]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49F8D0F338F8A9" bold="true" box="[201,328,1808,1832]" pageId="3" pageNumber="508">Etymology</emphasis>
.
</emphasis>
Latinized name
<taxonomicName id="4C669095720DFFBEFD8AF8D0F0D1F8A9" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[522,673,1808,1832]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma">
<emphasis id="B9123704720DFFBEFD8AF8D0F0D1F8A9" box="[522,673,1808,1832]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
refers to the great Buddhist monk Bodhidharma.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49F8F4F1FEF85D" pageId="3" pageNumber="508" type="discussion">
<paragraph id="8BD9EB16720DFFBEFF49F8F4F1FEF85D" blockId="3.[151,1436,943,2012]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49F8F4F34CF8CA" box="[201,316,1844,1868]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49F8F4F346F8CD" bold="true" box="[201,310,1844,1868]" pageId="3" pageNumber="508">Remarks</emphasis>
.
</emphasis>
<taxonomicName id="4C669095720DFFBEFEC6F8F4F028F8CD" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[326,600,1844,1868]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma">
<emphasis id="B9123704720DFFBEFEC6F8F4F028F8CD" box="[326,600,1844,1868]" italics="true" pageId="3" pageNumber="508">Chlaenius bodhidharma</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C669095720DFFBEFD75F8F4F1C8F8CD" authorityName="Kirschenhofer" authorityYear="2003" box="[757,952,1844,1868]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B9123704720DFFBEFD75F8F4F1C8F8CD" box="[757,952,1844,1868]" italics="true" pageId="3" pageNumber="508">C. schillhammeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C669095720DFFBEFC74F8F4F61CF8CD" authorityName="Bates" authorityYear="1892" box="[1012,1132,1844,1868]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720DFFBEFC74F8F4F61CF8CD" box="[1012,1132,1844,1868]" italics="true" pageId="3" pageNumber="508">C. eneides</emphasis>
</taxonomicName>
in having elytral intervals glabrous in the middle, but differs from the latter in having yellow spot at the middle of each elytron and covering only half of interval
<quantity id="4C9E46F3720DFFBEFE06F8BCF3C3F815" box="[390,435,1916,1940]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="508" unit="in" value="4.0">4 in</quantity>
contrast to yellow spot placed slightly behind the middle and covering whole width of interval
<quantity id="4C9E46F3720DFFBEFF74F860F36FF839" box="[244,287,1952,1976]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="508" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C669095720DFFBEFEA6F860F397F839" authorityName="Kirschenhofer" authorityYear="2003" box="[294,487,1952,1976]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B9123704720DFFBEFEA6F860F397F839" box="[294,487,1952,1976]" italics="true" pageId="3" pageNumber="508">C. schillhammeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C669095720DFFBEFD9FF860F0E5F839" authorityName="Bates" authorityYear="1892" box="[543,661,1952,1976]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720DFFBEFD9FF860F0E5F839" box="[543,661,1952,1976]" italics="true" pageId="3" pageNumber="508">C. eneides</emphasis>
</taxonomicName>
. Additionally, aedeagus of
<taxonomicName id="4C669095720DFFBEFC4AF860F6F2F839" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[970,1154,1952,1976]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="bodhidharma">
<emphasis id="B9123704720DFFBEFC4AF860F1ADF839" box="[970,989,1952,1976]" italics="true" pageId="3" pageNumber="508">C</emphasis>
.
<emphasis id="B9123704720DFFBEFC6BF860F6F2F839" box="[1003,1154,1952,1976]" italics="true" pageId="3" pageNumber="508">bodhidharma</emphasis>
</taxonomicName>
is more robust (
<figureCitation id="135DF793720DFFBEFABBF860F7FFF839" box="[1339,1423,1952,1976]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Fig. 40</figureCitation>
), and apical lamella (
<figureCitation id="135DF793720DFFBEFEF2F804F3B3F85D" box="[370,451,1988,2012]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Fig. 41</figureCitation>
) longer than in
<taxonomicName id="4C669095720DFFBEFDF5F804F09AF85D" authorityName="Bates" authorityYear="1892" box="[629,746,1988,2012]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720DFFBEFDF5F804F09AF85D" box="[629,746,1988,2012]" italics="true" pageId="3" pageNumber="508">C. eneides</emphasis>
</taxonomicName>
(
<figureCitation id="135DF793720DFFBEFD79F804F10FF85D" box="[761,895,1988,2012]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Figs. 46, 47</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,361 @@
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<mods:title id="B02D70509BC31C6A741828740D856D3E">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:namePart id="B1A74F9C40E525E898C02DD78EBD3C6A">Liu, Ye</mods:namePart>
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<mods:namePart id="F2F28ED85DAC9AA7CD1FEAE57FE61D7D">Shi, Hongliang</mods:namePart>
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<mods:namePart id="7017A4734DB064B28FDCE8FE25A59285">Liang, Hongbin</mods:namePart>
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<mods:date id="EBCC050E968745249E7E9BB52468CD40">2013</mods:date>
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Key to species of subgenus
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<emphasis id="B9123704720FFFBCFE62FEC4F0EFFE9F" bold="true" box="[482,671,260,286]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFD26FEC3F144FE9C" author="Bates H. W." box="[678,820,259,285]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="1" pageNumber="506" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
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(Modified from
<bibRefCitation id="EFF796E7720FFFBCFECDFEE6F384FEBF" author="Liu, Y. &amp; Liang, H. B. &amp; Kavanaugh, D. &amp; Yang, M. F." box="[333,500,294,318]" etAl="et al." firstAuthor="Liu" journalOrPublisher="Zootaxa" pageId="1" pageNumber="506" pagination="15 - 28" part="2397" refId="ref7391" refString="Liu, Y., Liang, H. B., Kavanaugh, D. &amp; Yang, M. F. (2010) Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species. Zootaxa, 2397, 15 - 28." title="Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species" type="journal article" year="2010">
Liu
<emphasis id="B9123704720FFFBCFEFBFEE7F3DCFEBF" box="[379,428,294,318]" italics="true" pageId="1" pageNumber="506">et al</emphasis>
. 2010
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1. Femora wholly black (
<figureCitation id="135DF793720FFFBCFE3AFEAEF05CFE02" box="[442,556,366,387]" captionStart="FIGURES 912" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[282,1273,195,1883]" captionTargetId="figure-51@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURES 912. Chlaenius li sp. n. 9, Holotype, male, dorsal view. 10, Holotype, male, ventral view. 11, Paratype, female, dorsal view. 12, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261773" httpUri="https://zenodo.org/record/5261773/files/figure.png" pageId="1" pageNumber="506">Figs. 10, 12</figureCitation>
)....................................................................... 2
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- Femora more or less yellow in the middle (
<figureCitation id="135DF793720FFFBCFDFEFE4BF0A9FE21" box="[638,729,395,416]" captionStart="FIGURES 14" captionStartId="2.[151,263,1909,1932]" captionTargetBox="[224,1309,194,1888]" captionTargetId="figure-51@2.[151,1436,193,1888]" captionTargetPageId="2" captionText="FIGURES 14. Chlaenius cheni sp. n. 1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="1" pageNumber="506">Figs. 2, 3</figureCitation>
,
<figureCitation id="135DF793720FFFBCFD64FE4BF179FE21" box="[740,777,395,416]" captionStart="FIGURES 58" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 58. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="1" pageNumber="506">6, 8</figureCitation>
,
<figureCitation id="135DF793720FFFBCFC94FE4BF122FE21" box="[788,850,395,416]" captionStart="FIGURES 1316" captionStartId="8.[151,263,1909,1932]" captionTargetBox="[192,1336,197,1886]" captionTargetId="figure-51@8.[151,1436,193,1888]" captionTargetPageId="8" captionText="FIGURES 1316. Chlaenius eneides Bates, 1892. 13, Lectotype, male, dorsal view. 14, Lectotype, male, ventral view. 15, Paralectotype, female, dorsal view. 16, Paralectotype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261775" httpUri="https://zenodo.org/record/5261775/files/figure.png" pageId="1" pageNumber="506">14, 16</figureCitation>
,
<figureCitation id="135DF793720FFFBCFCDCFE4BF106FE21" box="[860,886,395,416]" captionStart="FIGURES 1718" captionStartId="9.[151,264,1143,1166]" captionTargetBox="[164,1388,193,1122]" captionTargetId="figure-349@9.[151,1436,193,1122]" captionTargetPageId="9" captionText="FIGURES 1718. Chlaenius aodai sp. n. 17, Holotype, male, dorsal view. 18, Holotype, male, ventral view. FIGURE 19. Chlaenius langsonensis Kirschenhofer, 2008. Holotype, male. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261779" httpUri="https://zenodo.org/record/5261779/files/figure.png" pageId="1" pageNumber="506">18</figureCitation>
, 19)......................................... 5
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2. Elytron with a yellow spot at posterior third (
<figureCitation id="135DF793720FFFBCFD12FE69F09DFE3F" box="[658,749,425,446]" captionStart="FIGURES 14" captionStartId="2.[151,263,1909,1932]" captionTargetBox="[224,1309,194,1888]" captionTargetId="figure-51@2.[151,1436,193,1888]" captionTargetPageId="2" captionText="FIGURES 14. Chlaenius cheni sp. n. 1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="1" pageNumber="506">Figs. 1, 3</figureCitation>
,
<figureCitation id="135DF793720FFFBCFD78FE69F159FE3F" box="[760,809,425,446]" captionStart="FIGURES 912" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[282,1273,195,1883]" captionTargetId="figure-51@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURES 912. Chlaenius li sp. n. 9, Holotype, male, dorsal view. 10, Holotype, male, ventral view. 11, Paratype, female, dorsal view. 12, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261773" httpUri="https://zenodo.org/record/5261773/files/figure.png" pageId="1" pageNumber="506">9, 11</figureCitation>
,
<figureCitation id="135DF793720FFFBCFCB3FE69F13CFE3F" box="[819,844,425,446]" captionStart="FIGURES 1718" captionStartId="9.[151,264,1143,1166]" captionTargetBox="[164,1388,193,1122]" captionTargetId="figure-349@9.[151,1436,193,1122]" captionTargetPageId="9" captionText="FIGURES 1718. Chlaenius aodai sp. n. 17, Holotype, male, dorsal view. 18, Holotype, male, ventral view. FIGURE 19. Chlaenius langsonensis Kirschenhofer, 2008. Holotype, male. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261779" httpUri="https://zenodo.org/record/5261779/files/figure.png" pageId="1" pageNumber="506">17</figureCitation>
)............................................... 3
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- Elytron with a yellow spot near the middle (
<figureCitation id="135DF793720FFFBCFD07FE06F092FE5A" box="[647,738,454,475]" captionStart="FIGURES 58" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 58. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="1" pageNumber="506">Figs. 5, 7</figureCitation>
,
<figureCitation id="135DF793720FFFBCFD6DFE06F15BFE5A" box="[749,811,454,475]" captionStart="FIGURES 1316" captionStartId="8.[151,263,1909,1932]" captionTargetBox="[192,1336,197,1886]" captionTargetId="figure-51@8.[151,1436,193,1888]" captionTargetPageId="8" captionText="FIGURES 1316. Chlaenius eneides Bates, 1892. 13, Lectotype, male, dorsal view. 14, Lectotype, male, ventral view. 15, Paralectotype, female, dorsal view. 16, Paralectotype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261775" httpUri="https://zenodo.org/record/5261775/files/figure.png" pageId="1" pageNumber="506">13, 15</figureCitation>
, 19)............................................... 4
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3. Each of elytral intervals 28 glabrous medially, with a row of pubescence laterally (near striae); pronotum glabrous, with coppery metallic shine laterally.
<collectingCountry id="F371AB86720FFFBCFE76FDC1F05FFD97" box="[502,559,513,534]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFDBCFDC1F0F4FD97" box="[572,644,513,534]" country="China" name="Hainan" pageId="1" pageNumber="506">Hainan</collectingRegion>
) (
<figureCitation id="135DF793720FFFBCFD18FDC1F08FFD97" box="[664,767,513,534]" captionStart="FIGURES 912" captionStartId="6.[151,264,1909,1932]" captionTargetBox="[282,1273,195,1883]" captionTargetId="figure-51@6.[151,1436,193,1888]" captionTargetPageId="6" captionText="FIGURES 912. Chlaenius li sp. n. 9, Holotype, male, dorsal view. 10, Holotype, male, ventral view. 11, Paratype, female, dorsal view. 12, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261773" httpUri="https://zenodo.org/record/5261773/files/figure.png" pageId="1" pageNumber="506">Figs. 9, 11</figureCitation>
,
<figureCitation id="135DF793720FFFBCFC89FDC1F153FD97" box="[777,803,513,534]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">22</figureCitation>
,
<figureCitation id="135DF793720FFFBCFCADFDC1F136FD97" box="[813,838,513,534]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="1" pageNumber="506">28</figureCitation>
)........................................
<emphasis id="B9123704720FFFBCFAB6FDC1F72FFD97" box="[1334,1375,513,534]" italics="true" pageId="1" pageNumber="506">C. li</emphasis>
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- All elytral intervals regularly pubescent; pronotum pubescent, with green metallic shine laterally.
<collectingCountry id="F371AB86720FFFBCFB0AFDDEF6B3FDB2" box="[1162,1219,542,563]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
(
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,
<collectingRegion id="49A225F4720FFFBCFAA3FDDEF7E4FDB2" box="[1315,1428,542,563]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong</collectingRegion>
)...................................................................................
<taxonomicName id="4C669095720FFFBCFB44FDFCF7ECFDD1" authority="Jedlicka, 1960" authorityName="Jedlicka" authorityYear="1960" box="[1220,1436,571,593]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720FFFBCFB44FDFCF77BFDD0" box="[1220,1291,572,593]" italics="true" pageId="1" pageNumber="506">C. freyi</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFA91FDFBF7ECFDD1" author="Jedlicka A." box="[1297,1436,571,592]" firstAuthor="Jedlicka" journalOrPublisher="Entomologischen Arbeiten aus dem Museum G. Frey" pageId="1" pageNumber="506" pagination="587 - 598" part="11" refId="ref7233" refString="Jedlicka A. (1960) Neue Carabiden aus den Sammlungen des Museums Frey (Col.). Entomologischen Arbeiten aus dem Museum G. Frey, 11, 587 - 598." title="Neue Carabiden aus den Sammlungen des Museums Frey (Col.)" type="journal article" year="1960">Jedlička, 1960</bibRefCitation>
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4. Each of elytral intervals 2-8 with a row of regular punctures laterally (near striae); pronotum finely punctate.
<collectingCountry id="F371AB86720FFFBCFB78FD99F728FDEF" box="[1272,1368,601,622]" name="Myanmar" pageId="1" pageNumber="506">Myanmar</collectingCountry>
......................................................................................
<taxonomicName id="4C669095720FFFBCFB27FDB6F7ECFD0A" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[1191,1436,630,651]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="B9123704720FFFBCFB27FDB6F754FD0A" box="[1191,1316,630,651]" italics="true" pageId="1" pageNumber="506">C. pectinipes</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFAABFDB6F7ECFD0A" author="Bates H. W." box="[1323,1436,630,651]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="1" pageNumber="506" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FD54F7ECFD47" blockId="1.[151,1437,366,1267]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FD54F7ECFD47" pageId="1" pageNumber="506">
- Each of elytral intervals 2-8 with irregular punctures laterally (near striae), not displayed in a row; pronotum coarsely punctate.
<collectingCountry id="F371AB86720FFFBCFF61FD71F346FD47" box="[225,310,689,710]" name="Thailand" pageId="1" pageNumber="506">Thailand</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFEC2FD71F3C7FD47" box="[322,439,689,710]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="506">Chiang Mai</collectingRegion>
)...........................................................
<taxonomicName id="4C669095720FFFBCFB0BFD71F7ECFD47" authority="Kirschenhofer, 2009" authorityName="Kirschenhofer" authorityYear="2009" box="[1163,1436,689,710]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="bilyi">
<emphasis id="B9123704720FFFBCFB0BFD71F6A2FD47" box="[1163,1234,689,710]" italics="true" pageId="1" pageNumber="506">C. bilyi</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFB58FD71F7ECFD47" author="Kirschenhofer, E." box="[1240,1436,689,710]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" pageId="1" pageNumber="506" pagination="3 - 18" part="25" refId="ref7353" refString="Kirschenhofer, E. (2009) Neue Arten der Gattung Chlaenius Bonelli 1810 aus der orientalischen und pazifischen Region (Coleoptera: Carabidae). Acta Coleopterologica, 25 (1), 3 - 18." title="Neue Arten der Gattung Chlaenius Bonelli 1810 aus der orientalischen und pazifischen Region (Coleoptera: Carabidae)" type="journal article" year="2009">Kirschenhofer, 2009</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="3092F3B3720FFFBCFF17FD0EF7EBFC80" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FD0EF7EBFD62" blockId="1.[151,1437,366,1267]" box="[151,1435,718,739]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FD0EF7EBFD62" box="[151,1435,718,739]" pageId="1" pageNumber="506">5. Elytron with a yellow spot in or slightly behind the middle.....................................................6</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FD2CF7EBFC80" blockId="1.[151,1437,366,1267]" box="[151,1435,748,769]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FD2CF7EBFC80" box="[151,1435,748,769]" pageId="1" pageNumber="506">- Elytron with a yellow spot at posterior third................................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="3092F3B3720FFFBCFF17FCC9F7ECFCBA" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FCC9F7EBFC9F" blockId="1.[151,1437,366,1267]" box="[151,1435,777,798]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FCC9F7EBFC9F" box="[151,1435,777,798]" pageId="1" pageNumber="506">
6. Elytral intervals regularly pubescent.
<collectingCountry id="F371AB86720FFFBCFDC9FCC9F0F2FC9F" box="[585,642,777,798]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFD0EFCC9F0ABFC9F" box="[654,731,777,798]" country="China" name="Yunnan" pageId="1" pageNumber="506">Yunnan</collectingRegion>
).........................
<taxonomicName id="4C669095720FFFBCFB8AFCC9F7EBFC9F" authority="Liu &amp; Kavanaugh, 2010" authorityName="Liu &amp; Kavanaugh" authorityYear="2010" box="[1034,1435,777,798]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="rotundithorax">
<emphasis id="B9123704720FFFBCFB8AFCC9F6DCFC9F" box="[1034,1196,777,798]" italics="true" pageId="1" pageNumber="506">C. rotundithorax</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFB32FCC9F7EBFC9F" author="Liu, Y. &amp; Liang, H. B. &amp; Kavanaugh, D. &amp; Yang, M. F." box="[1202,1435,777,798]" firstAuthor="Liu" journalOrPublisher="Zootaxa" pageId="1" pageNumber="506" pagination="15 - 28" part="2397" refId="ref7391" refString="Liu, Y., Liang, H. B., Kavanaugh, D. &amp; Yang, M. F. (2010) Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species. Zootaxa, 2397, 15 - 28." title="Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species" type="journal article" year="2010">Liu &amp; Kavanaugh, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FCE6F7ECFCBA" blockId="1.[151,1437,366,1267]" box="[151,1436,806,827]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FCE6F7ECFCBA" box="[151,1436,806,827]" pageId="1" pageNumber="506">- Elytral intervals pubescent laterally, glabrous medially........................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="3092F3B3720FFFBCFF17FC84F7ECFCF7" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FC84F7ECFCD9" blockId="1.[151,1437,366,1267]" box="[151,1436,835,857]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FC84F7ECFCD9" box="[151,1436,835,857]" pageId="1" pageNumber="506">
7. Elytron with a yellow spot in the middle; antennomeres 13 brown (
<figureCitation id="135DF793720FFFBCFCF1FC84F1BFFCD8" box="[881,975,836,857]" captionStart="FIGURES 58" captionStartId="4.[151,264,1909,1932]" captionTargetBox="[287,1297,195,1886]" captionTargetId="figure-51@4.[151,1436,193,1888]" captionTargetPageId="4" captionText="FIGURES 58. Chlaenius bodhidharma sp. n. 5, Holotype, male, dorsal view. 6, Holotype, male, ventral view. 7, Paratype, female, dorsal view. 8, Paratype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261769" httpUri="https://zenodo.org/record/5261769/files/figure.png" pageId="1" pageNumber="506">Figs. 58</figureCitation>
).
<collectingCountry id="F371AB86720FFFBCFC60FC84F661FCD8" box="[992,1041,836,857]" name="India" pageId="1" pageNumber="506">India</collectingCountry>
(Darjeeling)....
<taxonomicName id="4C669095720FFFBCFB43FC84F710FCD8" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[1219,1376,836,857]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="bodhidharma" status="sp. nov.">
<emphasis id="B9123704720FFFBCFB43FC84F710FCD8" box="[1219,1376,836,857]" italics="true" pageId="1" pageNumber="506">C. bodhidharma</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFAE6FC83F7ECFCD9" bold="true" box="[1382,1436,835,856]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFAE6FC83F7ECFCD9" box="[1382,1436,835,856]" pageId="1" pageNumber="506" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FCA1F7ECFCF7" blockId="1.[151,1437,366,1267]" box="[151,1436,865,886]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FCA1F7ECFCF7" box="[151,1436,865,886]" pageId="1" pageNumber="506">
- Elytron with a yellow spot slightly behind the middle (
<figureCitation id="135DF793720FFFBCFD6DFCA1F110FCF7" box="[749,864,865,886]" captionStart="FIGURES 1316" captionStartId="8.[151,263,1909,1932]" captionTargetBox="[192,1336,197,1886]" captionTargetId="figure-51@8.[151,1436,193,1888]" captionTargetPageId="8" captionText="FIGURES 1316. Chlaenius eneides Bates, 1892. 13, Lectotype, male, dorsal view. 14, Lectotype, male, ventral view. 15, Paralectotype, female, dorsal view. 16, Paralectotype, female, ventral view. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.5261775" httpUri="https://zenodo.org/record/5261775/files/figure.png" pageId="1" pageNumber="506">Figs. 13, 15</figureCitation>
, 19); antennomeres 13 totally or partly black......... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3092F3B3720FFFBCFF17FCBEF7ECFC30" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FCBEF7ECFC12" blockId="1.[151,1437,366,1267]" box="[151,1436,894,915]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FCBEF7ECFC12" box="[151,1436,894,915]" pageId="1" pageNumber="506">
8. Lateral margin of pronotum sinuate before posterior angle (
<figureCitation id="135DF793720FFFBCFC9BFCBEF110FC12" box="[795,864,894,915]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">Fig. 24</figureCitation>
).............................
<taxonomicName id="4C669095720FFFBCFB3DFCBEF7ECFC12" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[1213,1436,894,915]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="eneides">
<emphasis id="B9123704720FFFBCFB3DFCBEF755FC12" box="[1213,1317,894,915]" italics="true" pageId="1" pageNumber="506">C. eneides</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFAABFCBEF7ECFC12" author="Bates H. W." box="[1323,1436,894,915]" firstAuthor="Bates" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="1" pageNumber="506" pagination="267 - 428" part="12" refId="ref7190" refString="Bates H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 12, 267 - 428." title="Viaggio di Leonardo Fea in Birmania e regioni vicini XLIV. List of the Carabidae" type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FC5CF7ECFC30" blockId="1.[151,1437,366,1267]" box="[151,1436,924,945]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FC5CF7ECFC30" box="[151,1436,924,945]" pageId="1" pageNumber="506">
- Lateral margin of pronotum straight before posterior angle (
<figureCitation id="135DF793720FFFBCFC9EFC5CF114FC30" box="[798,868,924,945]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">Fig. 25</figureCitation>
)............................................. 9
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BD9EB16720FFFBCFF17FC79F7ECFC6A" blockId="1.[151,1437,366,1267]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FC79F7ECFC6A" pageId="1" pageNumber="506">
<keyStep id="3092F3B3720FFFBCFF17FC79F7ECFC6A" pageId="1" pageNumber="506">
9. Antennomere 1 yellow or light brown, genae strongly tumid, as long as half of eye in dorsal view (Fig. 19)
<collectingCountry id="F371AB86720FFFBCFB5FFC79F740FC4F" box="[1247,1328,953,974]" name="Vietnam" pageId="1" pageNumber="506">Vietnam</collectingCountry>
(Mau Son).......................................................................
<taxonomicName id="4C669095720FFFBCFBBAFC16F7ECFC6A" authority="Kirschenhofer, 2008" authorityName="Kirschenhofer" authorityYear="2008" box="[1082,1436,982,1003]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="langsonensis">
<emphasis id="B9123704720FFFBCFBBAFC16F6A1FC6A" box="[1082,1233,982,1003]" italics="true" pageId="1" pageNumber="506">C. langsonensis</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFB58FC16F7ECFC6A" author="Kirschenhofer, E." box="[1240,1436,982,1003]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" pageId="1" pageNumber="506" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer, 2008</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FC34F7ECFBA7" blockId="1.[151,1437,366,1267]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FC34F7ECFBA7" pageId="1" pageNumber="506">
<keyStep id="3092F3B3720FFFBCFF17FC34F7ECFBA7" pageId="1" pageNumber="506">
10. Antennomere 1 dark brown; genae slightly tumid, as long as one-third of eye in dorsal view.
<collectingCountry id="F371AB86720FFFBCFBDBFC34F6C8FB88" box="[1115,1208,1012,1033]" name="Myanmar" pageId="1" pageNumber="506">Myanmar</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFB45FC34F77DFB88" box="[1221,1293,1012,1033]" country="Myanmar" name="Kachin" pageId="1" pageNumber="506">Kachin</collectingRegion>
).................................................................................
<taxonomicName id="4C669095720FFFBCFBACFBD1F7ECFBA7" authority="Kirschenhofer, 2003" authorityName="Kirschenhofer" authorityYear="2003" box="[1068,1436,1041,1062]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="schillhammeri">
<emphasis id="B9123704720FFFBCFBACFBD1F6A1FBA7" box="[1068,1233,1041,1062]" italics="true" pageId="1" pageNumber="506">C. schillhammeri</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFB58FBD1F7ECFBA7" author="Kirschenhofer, E." box="[1240,1436,1041,1062]" firstAuthor="Kirschenhofer" journalOrPublisher="Entomofauna" pageId="1" pageNumber="506" pagination="29 - 60" part="24" refId="ref7267" refString="Kirschenhofer, E. (2003) Uber neue und wenig bekannte Carabidae aus der athiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae). Entomofauna, 24 (3), 29 - 60." title="Uber neue und wenig bekannte Carabidae aus der athiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae)" type="journal article" year="2003">Kirschenhofer, 2003</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<keyStep id="3092F3B3720FFFBCFF17FBEEF7EBFBE0" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FBEEF7ECFBC2" blockId="1.[151,1437,366,1267]" box="[151,1436,1070,1091]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FBEEF7ECFBC2" box="[151,1436,1070,1091]" pageId="1" pageNumber="506">
11. Genae pubescent.
<collectingCountry id="F371AB86720FFFBCFE10FBEEF3B9FBC2" box="[400,457,1070,1091]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFE56FBEEF055FBC2" box="[470,549,1070,1091]" country="China" name="Taiwan)" pageId="1" pageNumber="506">Taiwan)</collectingRegion>
.................................................
<taxonomicName id="4C669095720FFFBCFBEFFBEEF7ECFBC2" authority="Liu &amp; Liang, 2010" authorityName="Liu &amp; Liang" authorityYear="2010" box="[1135,1436,1070,1091]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="piligenys">
<emphasis id="B9123704720FFFBCFBEFFBEEF691FBC2" box="[1135,1249,1070,1091]" italics="true" pageId="1" pageNumber="506">C. piligenys</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFB68FBEEF7ECFBC2" author="Liu, Y. &amp; Liang, H. B. &amp; Kavanaugh, D. &amp; Yang, M. F." box="[1256,1436,1070,1091]" firstAuthor="Liu" journalOrPublisher="Zootaxa" pageId="1" pageNumber="506" pagination="15 - 28" part="2397" refId="ref7391" refString="Liu, Y., Liang, H. B., Kavanaugh, D. &amp; Yang, M. F. (2010) Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species. Zootaxa, 2397, 15 - 28." title="Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species" type="journal article" year="2010">Liu &amp; Liang, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FB8CF7EBFBE0" blockId="1.[151,1437,366,1267]" box="[151,1435,1100,1121]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FB8CF7EBFBE0" box="[151,1435,1100,1121]" pageId="1" pageNumber="506">- Genae glabrous...................................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="3092F3B3720FFFBCFF17FBA9F7ECFB38" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FBA9F7ECFB1A" blockId="1.[151,1437,366,1267]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FBA9F7ECFB1A" pageId="1" pageNumber="506">
12. Pronotum coarsely punctate; elytral intervals pubescent laterally, glabrous medially.
<collectingCountry id="F371AB86720FFFBCFB8AFBA9F633FBFF" box="[1034,1091,1129,1150]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFBD0FBA9F6D9FBFF" box="[1104,1193,1129,1150]" country="China" name="Guangxi" pageId="1" pageNumber="506">Guangxi</collectingRegion>
)................................................................................................
<taxonomicName id="4C669095720FFFBCFBF0FB46F7ECFB1A" authority="Kirschenhofer, 2008" authorityName="Kirschenhofer" authorityYear="2008" box="[1136,1436,1158,1179]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B9123704720FFFBCFBF0FB46F6A1FB1A" box="[1136,1233,1158,1179]" italics="true" pageId="1" pageNumber="506">C. sehnali</emphasis>
<bibRefCitation id="EFF796E7720FFFBCFB58FB46F7ECFB1A" author="Kirschenhofer, E." box="[1240,1436,1158,1179]" firstAuthor="Kirschenhofer" journalOrPublisher="Acta Coleopterologica" pageId="1" pageNumber="506" pagination="3 - 34" part="24" refId="ref7307" refString="Kirschenhofer, E. (2008) Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae). Acta Coleopterologica, 24 (3), 3 - 34." title="Neue und wenig bekannte Arten sowie drei neue Synonyme (Tribus Chlaeniini) der palaarktischen, orientalischen und himalayanischen Region. (Coleoptera: Carabidae)" type="journal article" year="2008">Kirschenhofer, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FB64F7ECFB38" blockId="1.[151,1437,366,1267]" box="[151,1436,1188,1209]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FB64F7ECFB38" box="[151,1436,1188,1209]" pageId="1" pageNumber="506">- Pronotum finely punctate; elytral intervals regularly pubescent................................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="3092F3B3720FFFBCFF17FB01F7ECFB72" pageId="1" pageNumber="506">
<paragraph id="8BD9EB16720FFFBCFF17FB01F7ECFB54" blockId="1.[151,1437,366,1267]" box="[151,1436,1216,1238]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FB01F7ECFB54" box="[151,1436,1216,1238]" pageId="1" pageNumber="506">
13. Antennomeres 13 yellow; pronotum wider than long (
<figureCitation id="135DF793720FFFBCFD72FB01F13EFB57" box="[754,846,1217,1238]" captionStart="FIGURES 14" captionStartId="2.[151,263,1909,1932]" captionTargetBox="[224,1309,194,1888]" captionTargetId="figure-51@2.[151,1436,193,1888]" captionTargetPageId="2" captionText="FIGURES 14. Chlaenius cheni sp. n. 1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="1" pageNumber="506">Figs. 1, 3</figureCitation>
,
<figureCitation id="135DF793720FFFBCFCD8FB01F104FB57" box="[856,884,1217,1238]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">20</figureCitation>
).
<collectingCountry id="F371AB86720FFFBCFC04FB01F1CDFB57" box="[900,957,1217,1238]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
(
<collectingRegion id="49A225F4720FFFBCFC4AFB01F64DFB57" box="[970,1085,1217,1238]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong</collectingRegion>
)................
<taxonomicName id="4C669095720FFFBCFA90FB01F710FB57" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[1296,1376,1217,1238]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov.">
<emphasis id="B9123704720FFFBCFA90FB01F710FB57" box="[1296,1376,1217,1238]" italics="true" pageId="1" pageNumber="506">C. cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFAE6FB00F7ECFB54" bold="true" box="[1382,1436,1216,1237]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFAE6FB00F7ECFB54" box="[1382,1436,1216,1237]" pageId="1" pageNumber="506" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF17FB1EF7ECFB72" blockId="1.[151,1437,366,1267]" box="[151,1436,1246,1267]" pageId="1" pageNumber="506">
<keyLead id="30974823720FFFBCFF17FB1EF7ECFB72" box="[151,1436,1246,1267]" pageId="1" pageNumber="506">
- Antennomeres 13 black or brown; pronotum longer than wide (
<figureCitation id="135DF793720FFFBCFCCDFB1EF1B1FB72" box="[845,961,1246,1267]" captionStart="FIGURES 1718" captionStartId="9.[151,264,1143,1166]" captionTargetBox="[164,1388,193,1122]" captionTargetId="figure-349@9.[151,1436,193,1122]" captionTargetPageId="9" captionText="FIGURES 1718. Chlaenius aodai sp. n. 17, Holotype, male, dorsal view. 18, Holotype, male, ventral view. FIGURE 19. Chlaenius langsonensis Kirschenhofer, 2008. Holotype, male. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261779" httpUri="https://zenodo.org/record/5261779/files/figure.png" pageId="1" pageNumber="506">Figs. 17, 18</figureCitation>
,
<figureCitation id="135DF793720FFFBCFC4BFB1EF196FB72" box="[971,998,1246,1267]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">23</figureCitation>
).
<collectingCountry id="F371AB86720FFFBCFC77FB1EF638FB72" box="[1015,1096,1246,1267]" name="Vietnam" pageId="1" pageNumber="506">Vietnam</collectingCountry>
(Tam Dao).......
<taxonomicName id="4C669095720FFFBCFA8DFB1EF72FFB72" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[1293,1375,1246,1267]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="aodai" status="sp. nov.">
<emphasis id="B9123704720FFFBCFA8DFB1EF72FFB72" box="[1293,1375,1246,1267]" italics="true" pageId="1" pageNumber="506">C. aodai</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFAE6FB1EF7ECFB72" bold="true" box="[1382,1436,1246,1267]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFAE6FB1EF7ECFB72" box="[1382,1436,1246,1267]" pageId="1" pageNumber="506" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,319 @@
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<mods:title id="A9D8EAA55F7BB0BB33247996B8F33C89">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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<mods:namePart id="B0CE30528BBE0F015694208F4C3EC6D0">Liu, Ye</mods:namePart>
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<mods:namePart id="FB6877F21D5D1869CDE885FF1D88B006">Shi, Hongliang</mods:namePart>
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<mods:namePart id="6185BE2EFA0C54DC3D9E0527736A900F">Liang, Hongbin</mods:namePart>
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<treatment id="03CF5A00720FFFBEFF17FA83F6C5FD6E" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E" httpUri="http://treatment.plazi.org/id/03CF5A00720FFFBEFF17FA83F6C5FD6E" lastPageId="3" lastPageNumber="508" pageId="1" pageNumber="506">
<subSubSection id="C37CB89D720FFFBCFF17FA83F174FADA" box="[151,772,1347,1372]" pageId="1" pageNumber="506" type="nomenclature">
<paragraph id="8BD9EB16720FFFBCFF17FA83F174FADA" blockId="1.[150,772,1347,1407]" box="[151,772,1347,1372]" pageId="1" pageNumber="506">
<taxonomicName id="4C669095720FFFBCFF17FA83F0CEFADD" ID-CoL="69VVJ" authority="Liu &amp; Liang" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,702,1347,1372]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFF17FA83F37DFADD" bold="true" box="[151,269,1347,1372]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFE9DFA83F3A1FADD" bold="true" box="[285,465,1347,1372]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFE62FA83F0CEFADD" bold="true" box="[482,702,1347,1372]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFE62FA83F051FADD" bold="true" box="[482,545,1347,1372]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
Liu &amp; Liang
</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFD45FA84F174FADA" bold="true" box="[709,772,1348,1371]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFD45FA84F174FADA" box="[709,772,1348,1371]" pageId="1" pageNumber="506" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720FFFBCFF16FAA7F07FFAFE" box="[150,527,1383,1407]" pageId="1" pageNumber="506" type="description">
<paragraph id="8BD9EB16720FFFBCFF16FAA7F07FFAFE" blockId="1.[150,772,1347,1407]" box="[150,527,1383,1407]" pageId="1" pageNumber="506">
(
<figureCitation id="135DF793720FFFBCFF1FFAA7F37CFAFE" box="[159,268,1383,1407]" captionStart="FIGURES 14" captionStartId="2.[151,263,1909,1932]" captionTargetBox="[224,1309,194,1888]" captionTargetId="figure-51@2.[151,1436,193,1888]" captionTargetPageId="2" captionText="FIGURES 14. Chlaenius cheni sp. n. 1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="1" pageNumber="506">Figs. 14</figureCitation>
,
<figureCitation id="135DF793720FFFBCFE98FAA7F347FAFE" box="[280,311,1383,1407]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="1" pageNumber="506">20</figureCitation>
,
<figureCitation id="135DF793720FFFBCFEC2FAA7F310FAFE" box="[322,352,1383,1407]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="1" pageNumber="506">26</figureCitation>
,
<figureCitation id="135DF793720FFFBCFEECFAA7F3FAFAFE" box="[364,394,1383,1407]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="1" pageNumber="506">32</figureCitation>
,
<figureCitation id="135DF793720FFFBCFE16FAA7F3AEFAFE" box="[406,478,1383,1407]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="1" pageNumber="506">38, 39</figureCitation>
,
<figureCitation id="135DF793720FFFBCFE6AFAA7F077FAFE" box="[490,519,1383,1407]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="1" pageNumber="506">48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720FFFBCFF17FA6FF1BFF91E" pageId="1" pageNumber="506" type="materials_examined">
<paragraph id="8BD9EB16720FFFBCFF17FA6FF1BFF91E" blockId="1.[151,1438,1455,1948]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFF17FA6FF005FA46" box="[151,629,1455,1480]" inLineHeading="true" pageId="1" pageNumber="506" reason="1">
<emphasis id="B9123704720FFFBCFF17FA6FF3F5FA49" bold="true" box="[151,389,1455,1480]" pageId="1" pageNumber="506">Specimen examined</emphasis>
(
<emphasis id="B9123704720FFFBCFE17FA70F015FA49" bold="true" box="[407,613,1455,1480]" pageId="1" pageNumber="506">
total
<specimenCount id="9D60209F720FFFBCFE56FA6FF015FA49" box="[470,613,1455,1480]" pageId="1" pageNumber="506" type="generic">3 specimens</specimenCount>
</emphasis>
).
</emphasis>
<materialsCitation id="3B0EE14B720FFFBCFDFEFA6FF3DFF98E" collectingDate="1989-03-21" collectionCode="SCAU" collectorName="Chen Shoujian" country="China" location="Chebaling Nature Reserve" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Guangdong Prov." typeStatus="holotype">
<typeStatus id="54DD55B4720FFFBCFDFEFA6FF09BFA46" box="[638,747,1455,1479]" pageId="1" pageNumber="506" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60209F720FFFBCFD76FA6FF15CFA46" box="[758,812,1455,1479]" pageId="1" pageNumber="506" type="male">male</specimenCount>
(
<collectionCode id="ED7773D3720FFFBCFCBDFA6FF1FDFA46" box="[829,909,1455,1479]" pageId="1" pageNumber="506">SCAU</collectionCode>
), “
<collectingCountry id="F371AB86720FFFBCFC2CFA6FF184FA46" box="[940,1012,1455,1479]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
,
<collectingRegion id="49A225F4720FFFBCFB80FA6FF6B8FA46" box="[1024,1224,1455,1479]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong Prov.</collectingRegion>
,
<location id="8EB9BDCD720FFFBCFB54FA6FF285FA6D" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E:8EB9BDCD720FFFBCFB54FA6FF285FA6D" country="China" name="Chebaling Nature Reserve" pageId="1" pageNumber="506" stateProvince="Guangdong Prov.">Chebaling Nature Reserve</location>
,
<date id="FFD8CDD6720FFFBCFF79FA14F31BFA6D" box="[249,363,1492,1516]" pageId="1" pageNumber="506" value="1989-03-21">
<collectingDate id="EF9C343E720FFFBCFF79FA14F31BFA6D" box="[249,363,1492,1516]" pageId="1" pageNumber="506" value="1989-03-21">1989.3.21</collectingDate>
</date>
,
<collectorName id="26938EC0720FFFBCFEF7FA14F06EFA6D" box="[375,542,1492,1516]" pageId="1" pageNumber="506">Chen Shoujian</collectorName>
leg. / “HOLOTYPE,
<taxonomicName id="4C669095720FFFBCFCA7FA14F6D6FA6D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[807,1190,1492,1516]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFCA7FA14F1E8FA6D" box="[807,920,1492,1516]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFC2BFA14F629FA6D" box="[939,1113,1492,1516]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFBEAFA14F6D6FA6D" box="[1130,1190,1492,1516]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFB2FFA14F69FFA6D" bold="true" box="[1199,1263,1492,1516]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFB2FFA14F69FFA6D" box="[1199,1263,1492,1516]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [red label]
</materialsCitation>
.
<materialsCitation id="3B0EE14B720FFFBCFE3DFA38F62DF9D6" collectingDate="1989-03-21" collectionCode="SCAU" collectorName="Chen Shoujian" country="China" location="Chebaling Nature Reserve" pageId="1" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong Prov." typeStatus="paratype">
<typeStatus id="54DD55B4720FFFBCFE3DFA38F05AF991" box="[445,554,1528,1552]" pageId="1" pageNumber="506" type="paratype">Paratypes</typeStatus>
(total 2 specimens):
<specimenCount id="9D60209F720FFFBCFC9DFA37F1F0F98E" box="[797,896,1527,1551]" pageId="1" pageNumber="506" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720FFFBCFC11FA37F191F98E" box="[913,993,1527,1551]" pageId="1" pageNumber="506">SCAU</collectionCode>
), “
<collectingCountry id="F371AB86720FFFBCFB80FA37F638F98E" box="[1024,1096,1527,1551]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
,
<collectingRegion id="49A225F4720FFFBCFBD5FA37F76EF98E" box="[1109,1310,1527,1551]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong Prov.</collectingRegion>
,
<location id="8EB9BDCD720FFFBCFAABFA37F339F9B2" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E:8EB9BDCD720FFFBCFAABFA37F339F9B2" country="China" name="Chebaling Nature Reserve" pageId="1" pageNumber="506" stateProvince="Guangdong Prov.">Chebaling Nature Reserve</location>
,
<date id="FFD8CDD6720FFFBCFED6F9DCF3B9F9B5" box="[342,457,1564,1588]" pageId="1" pageNumber="506" value="1989-03-21">
<collectingDate id="EF9C343E720FFFBCFED6F9DCF3B9F9B5" box="[342,457,1564,1588]" pageId="1" pageNumber="506" value="1989-03-21">1989.3.21</collectingDate>
</date>
,
<collectorName id="26938EC0720FFFBCFE56F9DCF00DF9B5" box="[470,637,1564,1588]" pageId="1" pageNumber="506">Chen Shoujian</collectorName>
leg. / “
<taxonomicName id="4C669095720FFFBCFD66F9DCF1BCF9B5" authorityName="Motschulsky" authorityYear="1854" box="[742,972,1564,1588]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="posticalis">
<emphasis id="B9123704720FFFBCFD66F9DCF1BCF9B5" box="[742,972,1564,1588]" italics="true" pageId="1" pageNumber="506">Chlaenius posticalis</emphasis>
</taxonomicName>
M. det. Chen Shoujian” / “PARATYPE,
<taxonomicName id="4C669095720FFFBCFF17F980F067F9D9" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[151,535,1599,1624]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFF17F980F378F9D9" box="[151,264,1600,1624]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFE9BF980F3B9F9D9" box="[283,457,1600,1624]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFE5BF980F067F9D9" box="[475,535,1600,1624]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFDA2F980F011F9D9" bold="true" box="[546,609,1600,1624]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFDA2F980F011F9D9" box="[546,609,1600,1624]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label];
</materialsCitation>
<materialsCitation id="3B0EE14B720FFFBCFBE6F9FFF1BFF91E" collectionCode="IZCAS" collectorName="An Shuwen" country="China" location="Chebaling Nature Reserve" pageId="1" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong Prov." typeStatus="paratype">
<specimenCount id="9D60209F720FFFBCFBE6F9FFF6B9F9D6" box="[1126,1225,1599,1623]" pageId="1" pageNumber="506" type="female">1 female</specimenCount>
(
<collectionCode id="ED7773D3720FFFBCFB5AF9FFF740F9D6" box="[1242,1328,1599,1623]" pageId="1" pageNumber="506">IZCAS</collectionCode>
), “
<collectingCountry id="F371AB86720FFFBCFAD0F9FFF7E8F9D6" box="[1360,1432,1599,1623]" name="China" pageId="1" pageNumber="506">China</collectingCountry>
,
<collectingRegion id="49A225F4720FFFBCFF17F9A4F31EF9FA" box="[151,366,1636,1660]" country="China" name="Guangdong" pageId="1" pageNumber="506">Guangdong Prov.</collectingRegion>
,
<location id="8EB9BDCD720FFFBCFE09F9A4F0A3F9FA" LSID="urn:lsid:plazi:treatment:03CF5A00720FFFBEFF17FA83F6C5FD6E:8EB9BDCD720FFFBCFE09F9A4F0A3F9FA" box="[393,723,1636,1660]" country="China" name="Chebaling Nature Reserve" pageId="1" pageNumber="506" stateProvince="Guangdong Prov.">Chebaling Nature Reserve</location>
, 2003.7,
<collectorName id="26938EC0720FFFBCFCDAF9A4F19DF9FD" box="[858,1005,1636,1660]" pageId="1" pageNumber="506">An Shuwen</collectorName>
leg. / “PARATYPE,
<taxonomicName id="4C669095720FFFBCFAAAF9A4F3EBF921" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov." subGenus="Chlaenioctenus">
<emphasis id="B9123704720FFFBCFAAAF9A4F7EBF9FD" box="[1322,1435,1636,1660]" italics="true" pageId="1" pageNumber="506">Chlaenius</emphasis>
(
<emphasis id="B9123704720FFFBCFF20F948F33EF921" box="[160,334,1672,1696]" italics="true" pageId="1" pageNumber="506">Chlaenioctenus</emphasis>
)
<emphasis id="B9123704720FFFBCFEDFF948F3EBF921" box="[351,411,1672,1696]" italics="true" pageId="1" pageNumber="506">cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720FFFBCFE22F948F391F921" bold="true" box="[418,481,1672,1696]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="A2218A7F720FFFBCFE22F948F391F921" box="[418,481,1672,1696]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, des. by Liu &amp; Liang, 2012” [yellow label].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720FFFBEFF49F96CF1E5FE4E" lastPageId="3" lastPageNumber="508" pageId="1" pageNumber="506" type="description">
<paragraph id="8BD9EB16720FFFBCFF49F96CF040F8AE" blockId="1.[151,1438,1455,1948]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFF49F96CF30FF942" box="[201,383,1708,1732]" inLineHeading="true" pageId="1" pageNumber="506" reason="1">
<emphasis id="B9123704720FFFBCFF49F96CF308F945" bold="true" box="[201,376,1708,1732]" pageId="1" pageNumber="506">Measurements</emphasis>
.
</emphasis>
BL =
<quantity id="4C9E46F3720FFFBCFE4EF96CF004F942" box="[462,628,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6199999999999999" metricValueMin="1.48" pageId="1" pageNumber="506" unit="mm" value="15.5" valueMax="16.2" valueMin="14.8">14.816.2 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFD65F96CF13FF945" box="[741,847,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="1" pageNumber="506" unit="mm" value="15.4">15.4 mm</quantity>
), HW =
<quantity id="4C9E46F3720FFFBCFC35F96CF64FF942" box="[949,1087,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="1" pageNumber="506" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.62.7 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFB30F96CF77CF945" box="[1200,1292,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="506" unit="mm" value="2.6">2.6 mm</quantity>
), EYL = 1.0 mm, PL =
<quantity id="4C9E46F3720FFFBCFE8EF90FF3E5F966" box="[270,405,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.4" metricValueMin="3.3" pageId="1" pageNumber="506" unit="mm" value="3.3499999999999996" valueMax="3.4" valueMin="3.3">3.33.4 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFD80F90FF02AF966" box="[512,602,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="506" unit="mm" value="3.4">3.4 mm</quantity>
), PW =
<quantity id="4C9E46F3720FFFBCFD34F90FF14BF966" box="[692,827,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="1" pageNumber="506" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.53.7 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFC26F90FF189F966" box="[934,1017,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="506" unit="mm" value="3.6">3.6mm</quantity>
), EL =
<quantity id="4C9E46F3720FFFBCFBCBF90FF6A3F966" box="[1099,1235,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.5" metricValueMin="8.7" pageId="1" pageNumber="506" unit="mm" value="9.1" valueMax="9.5" valueMin="8.7">8.79.5 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFABDF90FF7E0F966" box="[1341,1424,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="506" unit="mm" value="9.1">9.1mm</quantity>
), EW =
<quantity id="4C9E46F3720FFFBCFF5FF934F317F88A" box="[223,359,1780,1804]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="5.7" metricValueMin="5.2" pageId="1" pageNumber="506" unit="mm" value="5.45" valueMax="5.7" valueMin="5.2">5.25.7 mm</quantity>
(mean =
<quantity id="4C9E46F3720FFFBCFE52F934F05BF88D" box="[466,555,1780,1804]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="506" unit="mm" value="5.5">5.5 mm</quantity>
), ratio PW/PL = 1.061.09 (mean = 1.07), EL/EW = 1.581.70 (mean = 1.65), EW/PW = 1.491.58 (mean = 1.53).
</paragraph>
<paragraph id="8BD9EB16720FFFBCFF49F8FBF6E2F81A" blockId="1.[151,1438,1455,1948]" pageId="1" pageNumber="506">
<emphasis id="B9123704720FFFBCFF49F8FBF32AF8D5" bold="true" box="[201,346,1851,1876]" inLineHeading="true" pageId="1" pageNumber="506" reason="1">Description.</emphasis>
Head and pronotum black, with green metallic shine; elytra black, with a round yellow spot at posterior third of each elytron; ventral surface, coxae, trochanters, and tibiae black; antennae brown; palpi, tarsi reddish brown or dark brown; femora yellow in the middle portion, black at base and apex.
</paragraph>
<caption id="DF19BB9E720CFFBFFF17F8B5F14DF82B" ID-DOI="http://doi.org/10.5281/zenodo.5261767" ID-Zenodo-Dep="5261767" httpUri="https://zenodo.org/record/5261767/files/figure.png" pageId="2" pageNumber="507" startId="2.[151,263,1909,1932]" targetBox="[224,1309,194,1888]" targetPageId="2">
<paragraph id="8BD9EB16720CFFBFFF17F8B5F14DF82B" blockId="2.[151,1436,1909,1962]" pageId="2" pageNumber="507">
<emphasis id="B9123704720CFFBFFF17F8B5F34BF80D" bold="true" box="[151,315,1909,1932]" pageId="2" pageNumber="507">FIGURES 14.</emphasis>
<taxonomicName id="4C669095720CFFBFFEC3F8B6F399F80D" authority="Liu &amp; Shi &amp; Liang, 2013" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[323,489,1910,1932]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov.">
<emphasis id="B9123704720CFFBFFEC3F8B6F399F80D" box="[323,489,1910,1932]" italics="true" pageId="2" pageNumber="507">Chlaenius cheni</emphasis>
</taxonomicName>
<emphasis id="B9123704720CFFBFFE70F8B6F05BF80A" bold="true" box="[496,555,1910,1931]" pageId="2" pageNumber="507">
<taxonomicNameLabel id="A2218A7F720CFFBFFE70F8B6F05BF80A" box="[496,555,1910,1931]" pageId="2" pageNumber="507" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
1, Holotype, male, dorsal view. 2, Holotype, male, ventral view. 3, Paratype, female, dorsal view. 4, Paratype, female, ventral view. Scale bar = 5.0 mm.
</paragraph>
</caption>
<paragraph id="8BD9EB16720DFFBEFF49FF57F1E5FE4E" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="508">
Head with vertex convex, finely punctate; occiput coarsely punctate and sparsely pubescent; genae glabrous, oblique, as long as one-fifth of eyes in dorsal view; lateral margins of mentum rounded. Pronotum (
<figureCitation id="135DF793720DFFBEFB74FF7CF736FF55" box="[1268,1350,188,212]" captionStart="FIGURES 2025" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 2025. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="3" pageNumber="508">Fig. 20</figureCitation>
) ovoid, widest in the middle; lateral margins distinctly bordered, sinuate before posterior angles; disc finely punctate, pubescent, median line distinct, shallow; posterior angles obtuse (&gt;90 degrees), rounded at apex; base rugosepunctate, pubescent; basal foveae deep, coarsely punctate. Elytra moderately convex, widest near posterior third; intervals (
<figureCitation id="135DF793720DFFBEFE8CFE8CF313FEE5" box="[268,355,332,356]" captionStart="FIGURES 2631" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 2631. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="3" pageNumber="508">Fig. 26</figureCitation>
) convex, densely punctate and pubescent; yellow spot covering intervals 48. Proepisterna sparsely punctate. Hind femora with basal black areas slightly shorter than hind trochanters, claws (
<figureCitation id="135DF793720DFFBEFB69FEAFF74AFE06" box="[1257,1338,367,391]" captionStart="FIGURES 3237" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 3237. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="3" pageNumber="508">Fig. 32</figureCitation>
) curved, with 67 short teeth. Aedeagus (
<figureCitation id="135DF793720DFFBEFD90FE54F0EBFE2D" box="[528,667,404,428]" captionStart="FIGURES 3847" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 3847. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="3" pageNumber="508">Figs. 38, 39</figureCitation>
) thin in lateral view, apical lamella rounded and short. Gonostylus (
<figureCitation id="135DF793720DFFBEFF1FFE77F281FE4E" box="[159,241,439,463]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">Fig. 48</figureCitation>
) of female ovipositor with five long spines on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49FE1CF7FAFE75" box="[201,1418,476,500]" pageId="3" pageNumber="508" type="etymology">
<paragraph id="8BD9EB16720DFFBEFF49FE1CF7FAFE75" blockId="3.[151,1436,151,752]" box="[201,1418,476,500]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FE1CF33FFE72" box="[201,335,476,500]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49FE1CF338FE75" bold="true" box="[201,328,476,500]" pageId="3" pageNumber="508">Etymology</emphasis>
.
</emphasis>
Latinized name
<taxonomicName id="4C669095720DFFBEFD8AFE1CF037FE75" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[522,583,476,500]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B9123704720DFFBEFD8AFE1CF037FE75" box="[522,583,476,500]" italics="true" pageId="3" pageNumber="508">cheni</emphasis>
</taxonomicName>
refers to Professor Chen Shoujian, a respected carabid specialist in
<collectingCountry id="F371AB86720DFFBEFAC0FE1CF7F6FE75" box="[1344,1414,476,500]" name="China" pageId="3" pageNumber="508">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37CB89D720DFFBEFF49FDC0F6C5FD6E" pageId="3" pageNumber="508" type="discussion">
<paragraph id="8BD9EB16720DFFBEFF49FDC0F35CFD02" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="508">
<emphasis id="B9123704720DFFBEFF49FDC0F34CFD96" box="[201,316,512,536]" inLineHeading="true" pageId="3" pageNumber="508" reason="1">
<emphasis id="B9123704720DFFBEFF49FDC0F346FD99" bold="true" box="[201,310,512,536]" pageId="3" pageNumber="508">Remarks</emphasis>
.
</emphasis>
<taxonomicName id="4C669095720DFFBEFEC7FDC0F070FD99" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[327,512,512,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B9123704720DFFBEFEC7FDC0F070FD99" box="[327,512,512,536]" italics="true" pageId="3" pageNumber="508">Chlaenius cheni</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C669095720DFFBEFD1FFDC0F164FD99" authorityName="Kirschenhofer" authorityYear="2008" box="[671,788,512,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B9123704720DFFBEFD1FFDC0F164FD99" box="[671,788,512,536]" italics="true" pageId="3" pageNumber="508">C. sehnali</emphasis>
</taxonomicName>
in having yellow spot on posterior third of elytron and femora yellow in the middle portion, but differs from the latter in having pronotal disc finely punctate and elytral intervals regularly pubescent in contrast to pronotal disc coarsely punctate and elytral intervals glabrous medially in
<taxonomicName id="4C669095720DFFBEFF34FDACF355FD05" authorityName="Kirschenhofer" authorityYear="2008" box="[180,293,620,644]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="sehnali">
<emphasis id="B9123704720DFFBEFF34FDACF355FD05" box="[180,293,620,644]" italics="true" pageId="3" pageNumber="508">C. sehnali</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD9EB16720DFFBEFF49FD50F6C5FD6E" blockId="3.[151,1436,151,752]" pageId="3" pageNumber="508">
<taxonomicName id="4C669095720DFFBEFF49FD50F36EFD29" authorityName="Jedlicka" authorityYear="1960" box="[201,286,656,680]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFF49FD50F36EFD29" box="[201,286,656,680]" italics="true" pageId="3" pageNumber="508">C. freyi</emphasis>
</taxonomicName>
is the nearest species with
<taxonomicName id="4C669095720DFFBEFDDEFD50F0CFFD29" authorityName="Liu &amp; Shi &amp; Liang" authorityYear="2013" box="[606,703,656,680]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cheni">
<emphasis id="B9123704720DFFBEFDDEFD50F0CFFD29" box="[606,703,656,680]" italics="true" pageId="3" pageNumber="508">C. cheni</emphasis>
</taxonomicName>
in distribution (
<figureCitation id="135DF793720DFFBEFCFAFD4FF1A0FD26" box="[890,976,655,679]" captionStart="FIGURE 52" captionStartId="12.[151,250,1844,1867]" captionTargetBox="[151,1436,935,1822]" captionTargetId="figure-69@12.[151,1436,935,1822]" captionTargetPageId="12" captionText="FIGURE 52. Geographical distribution of Chlaenius (Chlaenioctenus) species." figureDoi="http://doi.org/10.5281/zenodo.5261797" httpUri="https://zenodo.org/record/5261797/files/figure.png" pageId="3" pageNumber="508">Fig. 52</figureCitation>
), and their position of yellow spots and interval pubescence on elytra are also similar. However, the new species differs from
<taxonomicName id="4C669095720DFFBEFBE3FD74F6C7FD4D" authorityName="Jedlicka" authorityYear="1960" box="[1123,1207,692,716]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFBE3FD74F606FD4D" box="[1123,1142,692,716]" italics="true" pageId="3" pageNumber="508">C</emphasis>
.
<emphasis id="B9123704720DFFBEFB05FD74F6C7FD4D" box="[1157,1207,692,716]" italics="true" pageId="3" pageNumber="508">freyi</emphasis>
</taxonomicName>
in having bicolored femora (wholly black in
<taxonomicName id="4C669095720DFFBEFE2CFD18F38EFD71" authorityName="Jedlicka" authorityYear="1960" box="[428,510,728,752]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFE2CFD18F38EFD71" box="[428,510,728,752]" italics="true" pageId="3" pageNumber="508">C. freyi</emphasis>
</taxonomicName>
), and long gonostylus spines (
<figureCitation id="135DF793720DFFBEFCD4FD17F1D5FD6E" box="[852,933,727,751]" captionStart="FIGURES 4851" captionStartId="12.[151,264,823,846]" captionTargetBox="[151,1436,193,765]" captionTargetId="figure-65@12.[151,1436,193,802]" captionTargetPageId="12" captionText="FIGURES 4851. Female gonostylus of Chlaenius (Chlaenioctenus) species. 48, C. cheni, paratype. 49, C. bodhidharma, paratype. 50, C. li, paratype. 51, C. eneides, paralectotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261793" httpUri="https://zenodo.org/record/5261793/files/figure.png" pageId="3" pageNumber="508">Fig. 48</figureCitation>
) (very short in
<taxonomicName id="4C669095720DFFBEFBD2FD18F6D5FD71" authorityName="Jedlicka" authorityYear="1960" box="[1106,1189,728,752]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B9123704720DFFBEFBD2FD18F6D5FD71" box="[1106,1189,728,752]" italics="true" pageId="3" pageNumber="508">C. freyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>