197 lines
25 KiB
XML
197 lines
25 KiB
XML
<document id="C8B5FEFBBA0965D6A82A84E09D4B8202" ID-CLB-Dataset="56757" ID-DOI="10.11646/zootaxa.5141.1.2" ID-GBIF-Dataset="f0bcd63f-20f7-438b-b72b-77d49b944722" ID-ISSN="1175-5326" ID-Zenodo-Dep="6577627" ID-ZooBank="F1B8B140-4A7C-4AAA-97C6-A0DAA259C8E3" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653406469686" checkinUser="plazi" docAuthor="Russell, Peter J. C., Lukhtanov, Vladimir A. & Tennent, W. John" docDate="2022" docId="DB6B87B0FFF9FFE7E1FD03ADFB6BF9B6" docLanguage="en" docName="zootaxa.5141.1.2.pdf" docOrigin="Zootaxa 5141 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Melitaea phoebe subsp. caucasica Staudinger 1870" docType="treatment" docVersion="5" lastPageNumber="26" masterDocId="2752FFC8FFF8FFE6E13A005AFFA3FFC3" masterDocTitle="Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis & Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)" masterLastPageNumber="38" masterPageNumber="25" pageNumber="26" updateTime="1699338026890" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="D4682FEA38CFF3E5B026302487963428" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A569F5199BDE15AD87A7F7E313B98C88">
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<mods:title id="730554977E2135C3EB54FB1B5149BC79">Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis & Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)</mods:title>
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<mods:name id="7A068C0ECA853EC2AB6CD492DC19A98F" type="personal">
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<mods:role id="E2535E0E96204F052BB75C95B302BCFB">
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<mods:roleTerm id="298FD0D3068773DC15902668AC4AE8F2">Author</mods:roleTerm>
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<mods:namePart id="694E3EB2A77BC4E0B32B4F6EA7138F6B">Russell, Peter J. C.</mods:namePart>
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<mods:affiliation id="B4B875AB4668A7B5E9ED5D40278E2B09">Oakmeadow, Wessex Avenue, East Wittering, West Sussex PO 20 8 NP, U. K.</mods:affiliation>
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<mods:name id="BF94FD09EAA51F7AE40F953D4FDA2A32" type="personal">
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<mods:roleTerm id="7A064ED624D6BDA9A549328A5A146B0C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0FF53876DDC289E7C47068D85950936F">Lukhtanov, Vladimir A.</mods:namePart>
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<mods:affiliation id="0D8079911FD0602F1AB4B19EE43C9C66">Department of Karyosystematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, 199034 Saint Petersburg, Russia.</mods:affiliation>
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<mods:name id="8BF3FD16F838039BF53F7F9243F65E86" type="personal">
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</mods:role>
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<mods:namePart id="E6418E2455A84E54CBA4C1FBC230BCCF">Tennent, W. John</mods:namePart>
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<mods:affiliation id="7B80F9CCA0F97D85B58E742D4453C43A">Scientific Associate, Division of Insects, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. & Honorary Associate, Oxford University Museum of Natural History, Parks Road, Oxford OX 1 3 PW, U. K</mods:affiliation>
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<mods:typeOfResource id="D7B90407773B1652ECFAFE0A6810AA3E">text</mods:typeOfResource>
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<mods:title id="51926243C93CBD1D448F68A6AAC464A7">Zootaxa</mods:title>
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<mods:part id="0AFA889C292822CED09D2CAAB09DACA7">
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<mods:date id="5ACF35968CFB048C215670D55782B531">2022</mods:date>
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<mods:number id="E8AD5B7C7C789D8EF180093FDCEC9E83">2022-05-24</mods:number>
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<mods:number id="2B75E81E58D25920F3D585B7A4330458">5141</mods:number>
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<mods:detail id="35B5BEEC37D0D3E9E8697C96E38D52F8" type="issue">
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<mods:number id="D8678DDA6B5E56777FC5DDE3B2A115DE">1</mods:number>
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<mods:identifier id="A47C5E3C5EA11AC05F14B8161E416251" type="DOI">10.11646/zootaxa.5141.1.2</mods:identifier>
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<treatment id="DB6B87B0FFF9FFE7E1FD03ADFB6BF9B6" ID-DOI="http://doi.org/10.5281/zenodo.6580814" ID-GBIF-Taxon="195626720" ID-Zenodo-Dep="6580814" LSID="urn:lsid:plazi:treatment:DB6B87B0FFF9FFE7E1FD03ADFB6BF9B6" httpUri="http://treatment.plazi.org/id/DB6B87B0FFF9FFE7E1FD03ADFB6BF9B6" lastPageNumber="26" pageId="1" pageNumber="26">
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<subSubSection id="1BD8652DFFF9FFE7E1FD03ADFD3FFBD2" box="[199,668,1014,1041]" pageId="1" pageNumber="26" type="nomenclature">
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<paragraph id="537D36A6FFF9FFE7E1FD03ADFD3FFBD2" blockId="1.[151,1437,870,2013]" box="[199,668,1014,1041]" pageId="1" pageNumber="26">
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<taxonomicName id="94C24D25FFF9FFE7E1FD03ADFD3FFBD2" ID-CoL="3ZK6S" authority="Staudinger, 1870" authorityName="Staudinger" authorityYear="1870" box="[199,668,1014,1041]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="caucasica">
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<emphasis id="61B6EAB4FFF9FFE7E1FD03ADFE6BFBD2" bold="true" box="[199,456,1014,1041]" italics="true" pageId="1" pageNumber="26">M. phoebe caucasica</emphasis>
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<bibRefCitation id="37534B57FFF9FFE7E0EC03ADFD3FFBD2" author="Staudinger, O." box="[470,668,1015,1041]" pageId="1" pageNumber="26" pagination="3" refId="ref11169" refString="Staudinger, O. (1870) Beitrag zur Lepidopterenfauna Griechenlands. Horae Societatis Entomologicae Rossicae, 7, [3] - 304." type="journal article" year="1870">Staudinger, 1870</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="1BD8652DFFF9FFE7E39003ADFB6BF9B6" pageId="1" pageNumber="26" type="discussion">
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<paragraph id="537D36A6FFF9FFE7E39003ADFB6BF9B6" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="26">
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[TL: “Kindermann ganz ähnliche Stücke im Caucasus fing (?- Helenendorf; Kindermann leg.)”]. The name
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<emphasis id="61B6EAB4FFF9FFE7E3B50441FD5CFBF7" box="[655,767,1051,1076]" italics="true" pageId="1" pageNumber="26">caucasica</emphasis>
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was preoccupied by
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<taxonomicName id="94C24D25FFF9FFE7E2D00441FA3FFBF6" authority="Staudinger, 1861" authorityName="Staudinger" authorityYear="1861" box="[1002,1436,1051,1077]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="didyma" subSpecies="caucasica">
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<emphasis id="61B6EAB4FFF9FFE7E2D00441FB7BFBF7" box="[1002,1240,1051,1077]" italics="true" pageId="1" pageNumber="26">M. didyma caucasica</emphasis>
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<bibRefCitation id="37534B57FFF9FFE7E5E40441FA3FFBF6" author="Staudinger, O." box="[1246,1436,1051,1077]" pageId="1" pageNumber="26" refId="ref11128" refString="Staudinger, O. (1861) In Staudinger, O. & Wocke, M. F. Catalog der Lepidopteren Europa's und der angrenzenden Lander, Staudinger & Burdach, Dresden, Germany, XVI + 192 pp." type="book" year="1861">Staudinger, 1861</bibRefCitation>
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</taxonomicName>
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and the name was replaced first by
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<taxonomicName id="94C24D25FFF9FFE7E30F0465FC6CFB9A" authority="Fruhstorfer 1917" authorityName="Fruhstorfer" authorityYear="1917" box="[565,975,1087,1113]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="ottonis">
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<emphasis id="61B6EAB4FFF9FFE7E30F0465FCA0FB9B" box="[565,771,1087,1113]" italics="true" pageId="1" pageNumber="26">M. phoebe ottonis</emphasis>
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<bibRefCitation id="37534B57FFF9FFE7E2370465FC6CFB9A" author="Fruhstorfer, H." box="[781,975,1087,1113]" pageId="1" pageNumber="26" pagination="19" refId="ref9078" refString="Fruhstorfer, H. (1917) Neue Rhopaloceren aus der Sammlung Leonhard. Societas Entomologica, 32 (5), 19." type="journal article" year="1917">Fruhstorfer 1917</bibRefCitation>
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</taxonomicName>
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. A
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<typeStatus id="8C798804FFF9FFE7E2C20465FBFCFB9A" box="[1016,1119,1087,1113]" pageId="1" pageNumber="26" type="lectotype">lectotype</typeStatus>
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female and a
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<typeStatus id="8C798804FFF9FFE7E43F0465FA3FFB9A" box="[1285,1436,1087,1113]" pageId="1" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
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male were designated by Nekrutenko (
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<bibRefCitation id="37534B57FFF9FFE7E36F0439FC27FBBE" author="Hesselbarth, G. & van Oorschot, H. & Wagener, S." box="[597,900,1122,1149]" pageId="1" pageNumber="26" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. & Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
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Hesselbarth
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<emphasis id="61B6EAB4FFF9FFE7E3DF0439FCBAFBBE" box="[741,793,1123,1149]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
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. 1995: 2
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</bibRefCitation>
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: 1028) from the Staudinger collection, housed at Zoologisches Museum der Humboldt Universität,
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<collectingRegion id="9106F844FFF9FFE7E23904DDFCEAFB62" box="[771,841,1159,1185]" country="Germany" name="Berlin" pageId="1" pageNumber="26">Berlin</collectingRegion>
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(figs 5A, B, C & 6A, B, C). Verity subsequently also proposed a replacement name,
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<emphasis id="61B6EAB4FFF9FFE7E30F04F1FD18FB06" box="[565,699,1195,1221]" italics="true" pageId="1" pageNumber="26">caucasicola</emphasis>
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<bibRefCitation id="37534B57FFF9FFE7E3FE04F1FCF1FB06" author="Verity, R." box="[708,850,1195,1221]" pageId="1" pageNumber="26" pagination="26 - 31" refId="ref11898" refString="Verity, R. (1919) Seasonal Polymorphism and Races of some European Grypocera and Rhopalocera. Entomologist's Record and Journal of Variation 31 (2), 26 - 31, (3), 43 - 48, (5), 87 - 89, (7), 121 - 129, (10), 178 - 184, (11), 193 - 201." type="journal article" year="1919">Verity, 1919</bibRefCitation>
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, this being a synonym of
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<emphasis id="61B6EAB4FFF9FFE7E5B804F1FB6DFB07" box="[1154,1230,1195,1220]" italics="true" pageId="1" pageNumber="26">ottonis</emphasis>
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.
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<bibRefCitation id="37534B57FFF9FFE7E5E504F1FEAFFB2A" author="Kemal, M. & Kocak, A. O." pageId="1" pageNumber="26" refId="ref9502" refString="Kemal, M. & Kocak, A. O. (2011) A synonymical and distributional checklist of the Papilionoidea and Hesperioidea of East Mediterranean countries, including Turkey (Lepidoptera). Priamus, 25 (Supplement), 1 - 162, Plates 1 - 42." type="book" year="2011">Kemal & Koçak (2011: 44)</bibRefCitation>
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used the name ‘
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<taxonomicName id="94C24D25FFF9FFE7E0FF0495FD09FB2A" box="[453,682,1231,1257]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subGenus" subGenus="Cinclidia">
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<emphasis id="61B6EAB4FFF9FFE7E0FF0495FD8AFB2A" box="[453,553,1231,1257]" italics="true" pageId="1" pageNumber="26">Melitaea</emphasis>
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(
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<emphasis id="61B6EAB4FFF9FFE7E3030495FD02FB2A" box="[569,673,1231,1257]" italics="true" pageId="1" pageNumber="26">Cinclidia</emphasis>
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)
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</taxonomicName>
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(
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<taxonomicName id="94C24D25FFF9FFE7E3860495FCAEFB2A" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[700,781,1231,1257]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="61B6EAB4FFF9FFE7E3860495FCAEFB2A" box="[700,781,1231,1257]" italics="true" pageId="1" pageNumber="26">phoebe</emphasis>
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</taxonomicName>
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)
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<emphasis id="61B6EAB4FFF9FFE7E2270495FCC8FB2A" box="[797,875,1231,1257]" italics="true" pageId="1" pageNumber="26">sextilis</emphasis>
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<bibRefCitation id="37534B57FFF9FFE7E2480495FB88FB2A" author="Jachontov, A. A." box="[882,1067,1231,1257]" pageId="1" pageNumber="26" pagination="282 - 292" refId="ref9463" refString="Jachontov, A. A. (1909) Notice sur les Lepidopteres Rhopaloceres du Caucase. Russkoe entomologicheskoe obozreniie (Revue Russe d'Entomologie), 8 (1908), 282 - 292. [In Russian]" type="journal article" year="1909">Jachontov, 1909</bibRefCitation>
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’ as a replacement name giving it subspecific(?) status; however,
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<bibRefCitation id="37534B57FFF9FFE7E0CD04A9FD50FACE" author="Jachontov, A. A." box="[503,755,1266,1293]" pageId="1" pageNumber="26" pagination="282 - 292" refId="ref9463" refString="Jachontov, A. A. (1909) Notice sur les Lepidopteres Rhopaloceres du Caucase. Russkoe entomologicheskoe obozreniie (Revue Russe d'Entomologie), 8 (1908), 282 - 292. [In Russian]" type="journal article" year="1909">Jachontov (1909: 285)</bibRefCitation>
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used this name for a variety of second generation
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<taxonomicName id="94C24D25FFF9FFE7E41C04A9FA3FFACE" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[1318,1436,1267,1293]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="61B6EAB4FFF9FFE7E41C04A9FA3FFACE" box="[1318,1436,1267,1293]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
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</taxonomicName>
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and, so far as the authors are aware, no author since has used the name
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<emphasis id="61B6EAB4FFF9FFE7E2E3054DFB84FAF2" box="[985,1063,1303,1329]" italics="true" pageId="1" pageNumber="26">sextilis</emphasis>
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in favour of
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<emphasis id="61B6EAB4FFF9FFE7E5F9054DFAACFAF3" box="[1219,1295,1303,1328]" italics="true" pageId="1" pageNumber="26">ottonis</emphasis>
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<bibRefCitation id="37534B57FFF9FFE7E422054DFF71FA96" author="Fruhstorfer, H." pageId="1" pageNumber="26" pagination="19" refId="ref9078" refString="Fruhstorfer, H. (1917) Neue Rhopaloceren aus der Sammlung Leonhard. Societas Entomologica, 32 (5), 19." type="journal article" year="1917">Fruhstorfer, 1917</bibRefCitation>
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. In fact the
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<taxonomicName id="94C24D25FFF9FFE7E0580561FE7FFA96" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[354,476,1339,1365]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
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<emphasis id="61B6EAB4FFF9FFE7E0580561FE7FFA96" box="[354,476,1339,1365]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
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</taxonomicName>
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species group portrayed by
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<bibRefCitation id="37534B57FFF9FFE7E2120561FBC9FA96" author="Kemal, M. & Kocak, A. O." box="[808,1130,1339,1365]" pageId="1" pageNumber="26" refId="ref9502" refString="Kemal, M. & Kocak, A. O. (2011) A synonymical and distributional checklist of the Papilionoidea and Hesperioidea of East Mediterranean countries, including Turkey (Lepidoptera). Priamus, 25 (Supplement), 1 - 162, Plates 1 - 42." type="book" year="2011">Kemal & Koçak (2011: 44)</bibRefCitation>
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, in their article on eastern Mediterranean butterflies, included
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<taxonomicName id="94C24D25FFF9FFE7E3170505FD03FABB" authorityName="Oberthur" authorityYear="1876" box="[557,672,1375,1400]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="punica">
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<emphasis id="61B6EAB4FFF9FFE7E3170505FD03FABB" box="[557,672,1375,1400]" italics="true" pageId="1" pageNumber="26">M. punica</emphasis>
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</taxonomicName>
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, a species absent from the eastern Mediterranean. This perpetuates confusion, which the first author with others has been trying to resolve.
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<bibRefCitation id="37534B57FFF9FFE7E28905D9FB7AFA5E" author="Hesselbarth, G. & van Oorschot, H. & Wagener, S." box="[947,1241,1410,1437]" pageId="1" pageNumber="26" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. & Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
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Hesselbarth
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<emphasis id="61B6EAB4FFF9FFE7E50405D9FBCDFA5E" box="[1086,1134,1411,1437]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
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. (1995: 3
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</bibRefCitation>
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, Tafel 80/81: figs 30–
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<specimenCount id="45C4FD2FFFF9FFE7E1F805FDFF5EFA02" box="[194,253,1447,1473]" pageId="1" pageNumber="26" type="male">33 ♂</specimenCount>
|
||
; Tafel 82/83: figs 1–
|
||
<specimenCount id="45C4FD2FFFF9FFE7E0DF05FDFDACFA02" box="[485,527,1447,1473]" pageId="1" pageNumber="26" type="female">4 ♀</specimenCount>
|
||
) placed
|
||
<emphasis id="61B6EAB4FFF9FFE7E34905FDFD1CFA03" box="[627,703,1447,1472]" italics="true" pageId="1" pageNumber="26">ottonis</emphasis>
|
||
as a synonym of
|
||
<taxonomicName id="94C24D25FFF9FFE7E2BD05FDFC5DFA02" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[903,1022,1447,1473]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
|
||
<emphasis id="61B6EAB4FFF9FFE7E2BD05FDFC5DFA02" box="[903,1022,1447,1473]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
|
||
</taxonomicName>
|
||
. Although the
|
||
<typeStatus id="8C798804FFF9FFE7E59D05FDFAADFA02" box="[1191,1294,1447,1473]" pageId="1" pageNumber="26" type="lectotype">lectotype</typeStatus>
|
||
female does not show all the characters typical of
|
||
<taxonomicName id="94C24D25FFF9FFE7E3070591FD17FA26" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[573,692,1483,1509]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
|
||
<emphasis id="61B6EAB4FFF9FFE7E3070591FD17FA26" box="[573,692,1483,1509]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
|
||
</taxonomicName>
|
||
, for instance the underside submarginal black arches do not touch the intervening veins (see
|
||
<figureCitation id="CBF92A23FFF9FFE7E08405B5FDB6F9CA" box="[446,533,1518,1545]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[153,1436,200,1692]" captionTargetId="figure-26@2.[151,1436,181,1693]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, Melitaea ornata male underside, Russian Federation, southern Urals, Orenburg Province, Guberli, ca. 190 m, 14 May 2014. 2, Melitaea phoebe male underside, Russian Federation, southern Urals, Orenburg Province, Krasnoznamenka, ca. 210 m, 16 May 2014. 3, Melitaea ornata female underside, Slovenia, Koper, ca. 2km NW of Rakitovec ca. 525 m, 17 May 2011. 4, Melitaea phoebe female underside, Slovenia, Koper, ca. 2km S. of Rakitovec ca. 525 m, 17 May 2011. 5, Lectotype female of Melitaea phoebe caucasica Staudinger, 1870. 6, Paralectotype male of Melitaea phoebe caucasica Staudinger, 1870. Figs 1–4 © P. Russell. Figs 5–6 © T. Léger. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577631" httpUri="https://zenodo.org/record/6577631/files/figure.png" pageId="1" pageNumber="26">Fig. 5B</figureCitation>
|
||
), the
|
||
<typeStatus id="8C798804FFF9FFE7E36E05B5FD48F9CA" box="[596,747,1519,1545]" pageId="1" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
|
||
underside (
|
||
<figureCitation id="CBF92A23FFF9FFE7E25405B5FC67F9C9" box="[878,964,1519,1546]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[153,1436,200,1692]" captionTargetId="figure-26@2.[151,1436,181,1693]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, Melitaea ornata male underside, Russian Federation, southern Urals, Orenburg Province, Guberli, ca. 190 m, 14 May 2014. 2, Melitaea phoebe male underside, Russian Federation, southern Urals, Orenburg Province, Krasnoznamenka, ca. 210 m, 16 May 2014. 3, Melitaea ornata female underside, Slovenia, Koper, ca. 2km NW of Rakitovec ca. 525 m, 17 May 2011. 4, Melitaea phoebe female underside, Slovenia, Koper, ca. 2km S. of Rakitovec ca. 525 m, 17 May 2011. 5, Lectotype female of Melitaea phoebe caucasica Staudinger, 1870. 6, Paralectotype male of Melitaea phoebe caucasica Staudinger, 1870. Figs 1–4 © P. Russell. Figs 5–6 © T. Léger. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577631" httpUri="https://zenodo.org/record/6577631/files/figure.png" pageId="1" pageNumber="26">Fig. 6B</figureCitation>
|
||
) certainly shows all the characters typical of
|
||
<taxonomicName id="94C24D25FFF9FFE7E18C0649FE8EF9EE" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[182,301,1555,1581]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
|
||
<emphasis id="61B6EAB4FFF9FFE7E18C0649FE8EF9EE" box="[182,301,1555,1581]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
|
||
</taxonomicName>
|
||
. Recent authors, such
|
||
<bibRefCitation id="37534B57FFF9FFE7E3160649FC9CF9EE" author="Tshikolovets, V. V." box="[556,831,1555,1581]" pageId="1" pageNumber="26" refId="ref11624" refString="Tshikolovets, V. V. (2011) Butterflies of Europe & the Mediterranean area. Tshikolovets Publications, Pardubice, Czech Republic, 544 pp." type="book" year="2011">Tshikolovets (2011: 497</bibRefCitation>
|
||
; 2003: plate 24: figs
|
||
<specimenCount id="45C4FD2FFFF9FFE7E5140649FBCCF9EE" box="[1070,1135,1555,1581]" pageId="1" pageNumber="26" type="generic">
|
||
<quantity id="943A9B43FFF9FFE7E5140649FBC8F9EE" box="[1070,1131,1555,1581]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="26" unit="m" value="16.0">16 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
and
|
||
<specimenCount id="45C4FD2FFFF9FFE7E59C0649FB7FF9ED" box="[1190,1244,1555,1582]" pageId="1" pageNumber="26" type="generic">17 f.</specimenCount>
|
||
),
|
||
<bibRefCitation id="37534B57FFF9FFE7E5C80649FEDAF992" author="Tshikolovets, V. V. & Naderi A. & Eckweiler, W." pageId="1" pageNumber="26" refId="ref11652" refString="Tshikolovets, V. V., Naderi A. & Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, Czech Republic, 366 pp. + 65 colour plates." type="book" year="2014">
|
||
Tshikolovets
|
||
<emphasis id="61B6EAB4FFF9FFE7E4B20649FF17F992" italics="true" pageId="1" pageNumber="26">et al.</emphasis>
|
||
(2014: 318–319)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="37534B57FFF9FFE7E0B9066DFCAFF991" author="Van Oorschot, H. & Coutsis, J. G." box="[387,780,1591,1618]" pageId="1" pageNumber="26" refId="ref11693" refString="Van Oorschot, H. & Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot & Coutsis (2014: 60)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="37534B57FFF9FFE7E278066DFB26F992" author="Russell, PJ & Tennent, W. J." box="[834,1157,1590,1617]" pageId="1" pageNumber="26" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. & Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis & Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell & Tennent (2016: 45</bibRefCitation>
|
||
, note 22) have all agreed that this is a subspecies of
|
||
<taxonomicName id="94C24D25FFF9FFE7E0F80601FD9BF9B6" baseAuthorityName="Denis & Schiffermuller" baseAuthorityYear="1775" box="[450,568,1627,1653]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
|
||
<emphasis id="61B6EAB4FFF9FFE7E0F80601FD9BF9B6" box="[450,568,1627,1653]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
|
||
</taxonomicName>
|
||
and not
|
||
<taxonomicName id="94C24D25FFF9FFE7E3A30601FCA9F9B7" authorityName="Christoph" authorityYear="1893" box="[665,778,1627,1652]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="61B6EAB4FFF9FFE7E3A30601FCA9F9B7" box="[665,778,1627,1652]" italics="true" pageId="1" pageNumber="26">M. ornata</emphasis>
|
||
</taxonomicName>
|
||
, with which the present authors concur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |