Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis & Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)
Author
Russell, Peter J. C.
Oakmeadow, Wessex Avenue, East Wittering, West Sussex PO 20 8 NP, U. K.
Author
Lukhtanov, Vladimir A.
Department of Karyosystematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, 199034 Saint Petersburg, Russia.
Author
Tennent, W. John
Scientific Associate, Division of Insects, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. & Honorary Associate, Oxford University Museum of Natural History, Parks Road, Oxford OX 1 3 PW, U. K
text
Zootaxa
2022
2022-05-24
5141
1
25
38
journal article
56757
10.11646/zootaxa.5141.1.2
f0bcd63f-20f7-438b-b72b-77d49b944722
1175-5326
6577627
F1B8B140-4A7C-4AAA-97C6-A0DAA259C8E3
M. phoebe caucasica
Staudinger, 1870
[TL: “Kindermann ganz ähnliche Stücke im Caucasus fing (?- Helenendorf; Kindermann leg.)”]. The name
caucasica
was preoccupied by
M. didyma caucasica
Staudinger, 1861
and the name was replaced first by
M. phoebe ottonis
Fruhstorfer 1917
. A
lectotype
female and a
paralectotype
male were designated by Nekrutenko (
Hesselbarth
et al
. 1995: 2
: 1028) from the Staudinger collection, housed at Zoologisches Museum der Humboldt Universität,
Berlin
(figs 5A, B, C & 6A, B, C). Verity subsequently also proposed a replacement name,
caucasicola
Verity, 1919
, this being a synonym of
ottonis
.
Kemal & Koçak (2011: 44)
used the name ‘
Melitaea
(
Cinclidia
)
(
phoebe
)
sextilis
Jachontov, 1909
’ as a replacement name giving it subspecific(?) status; however,
Jachontov (1909: 285)
used this name for a variety of second generation
M. phoebe
and, so far as the authors are aware, no author since has used the name
sextilis
in favour of
ottonis
Fruhstorfer, 1917
. In fact the
M. phoebe
species group portrayed by
Kemal & Koçak (2011: 44)
, in their article on eastern Mediterranean butterflies, included
M. punica
, a species absent from the eastern Mediterranean. This perpetuates confusion, which the first author with others has been trying to resolve.
Hesselbarth
et al
. (1995: 3
, Tafel 80/81: figs 30–
33 ♂
; Tafel 82/83: figs 1–
4 ♀
) placed
ottonis
as a synonym of
M. phoebe
. Although the
lectotype
female does not show all the characters typical of
M. phoebe
, for instance the underside submarginal black arches do not touch the intervening veins (see
Fig. 5B
), the
paralectotype
underside (
Fig. 6B
) certainly shows all the characters typical of
M. phoebe
. Recent authors, such
Tshikolovets (2011: 497
; 2003: plate 24: figs
16 m
.
and
17 f.
),
Tshikolovets
et al.
(2014: 318–319)
,
van Oorschot & Coutsis (2014: 60)
and
Russell & Tennent (2016: 45
, note 22) have all agreed that this is a subspecies of
M. phoebe
and not
M. ornata
, with which the present authors concur.