381 lines
51 KiB
XML
381 lines
51 KiB
XML
<document id="3A67F0C1CF9765165732055A7745CC6F" ID-CLB-Dataset="113069" ID-DOI="10.11646/zootaxa.5111.1.1" ID-GBIF-Dataset="f3992bb2-9962-4f3c-8f5b-131cfe552c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6340729" ID-ZooBank="836B0F69-037C-4D0F-80DB-94FE454F48E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646828425946" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="C822F213FFF62448CB88A0B274082206" docLanguage="en" docName="zootaxa.5111.1.1.pdf" docOrigin="Zootaxa 5111 (1)" docSource="http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Arthromelodes torus Yin 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="341B8A6BFFCB2477CB1FA14C7567265F" masterDocTitle="The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="62" updateTime="1699317591893" updateUser="ExternalLinkService">
|
||
<mods:mods id="0DD431FCEB9B6476F0E060D3986F372C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="09DCB9363F11F39E778F69B5158DC350">
|
||
<mods:title id="4871C47926D910B6E7E729A776CA0F8F">The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A1065490C7327C06C676361A1146FC27" type="personal">
|
||
<mods:role id="43F550FC1DF10A4E442C1B490F0198BC">
|
||
<mods:roleTerm id="9ADCACD4DA56A9CBDAC2413CABDFCC74">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5D6C27715B22E53F246C4D13E1FA778B">Yin, Zi-Wei</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DCFDEB1F540705CA933D0842C55D4FB7">text</mods:typeOfResource>
|
||
<mods:relatedItem id="CF4EB50FEBB49F5A16E194CEE3ABD7DB" type="host">
|
||
<mods:titleInfo id="AF4B83403C2026F7318CDAAD585C9AE1">
|
||
<mods:title id="20CFFE1CFED62D47773EA7D6EA8440B8">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="A2A5F504D762FAF9F4C4F1AC75582BC2">
|
||
<mods:date id="404F4E021E2B9800861203CE10DF0C45">2022</mods:date>
|
||
<mods:detail id="E34DBA584649B9C6B5DC2D73EC833E9D" type="pubDate">
|
||
<mods:number id="59EF5AB9F92CAFC35A8DABE29175D846">2022-03-09</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="27EA3F3FCB2A2927C70125AD4F17FEF9" type="volume">
|
||
<mods:number id="BB43588AFC3F99720FA507ADE7D58471">5111</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AE3500961095153996D57A063130C8A3" type="issue">
|
||
<mods:number id="5537BB1E2CA3BB6BCDAA709D3DDC9365">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="F6462EB608FA22DBD88B8FC1D52CEDE3" unit="page">
|
||
<mods:start id="AD09A0E4020ECED691303B006F3C035A">1</mods:start>
|
||
<mods:end id="2D8570E2408E611631EC2967A98BB7EA">211</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="4D6D25D1371C1A5FD06947992BF8F38A">
|
||
<mods:url id="7A43A68EBA7B3E77B03C858A4253966B">http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="08DA1A69C97D1B6F5FDD6AC902F0AB46">journal article</mods:classification>
|
||
<mods:identifier id="98F1C374B24EC7578508F199B6FCB373" type="CLB-Dataset">113069</mods:identifier>
|
||
<mods:identifier id="9DF0DC26A2E522D1378536535D6CD7C1" type="DOI">10.11646/zootaxa.5111.1.1</mods:identifier>
|
||
<mods:identifier id="7C012386DAAE5224CBD3FDCA8DDC6069" type="GBIF-Dataset">f3992bb2-9962-4f3c-8f5b-131cfe552c4d</mods:identifier>
|
||
<mods:identifier id="8F0D4C2AA3C7A9CAF94EF2ACB4AA1594" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="DF63F3A9F254FE8E59AABF199E1B1150" type="Zenodo-Dep">6340729</mods:identifier>
|
||
<mods:identifier id="3DFD215208CD64B059FDD4E88A721AF0" type="ZooBank">836B0F69-037C-4D0F-80DB-94FE454F48E3</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="C822F213FFF62448CB88A0B274082206" ID-DOI="http://doi.org/10.5281/zenodo.6964265" ID-GBIF-Taxon="198579317" ID-Zenodo-Dep="6964265" LSID="urn:lsid:plazi:treatment:C822F213FFF62448CB88A0B274082206" httpUri="http://treatment.plazi.org/id/C822F213FFF62448CB88A0B274082206" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62">
|
||
<subSubSection id="0891108EFFF6244ACB88A0B274922446" box="[151,501,510,537]" pageId="61" pageNumber="62" type="nomenclature">
|
||
<paragraph id="40344305FFF6244ACB88A0B274922446" blockId="61.[151,569,510,609]" box="[151,501,510,537]" pageId="61" pageNumber="62">
|
||
<heading id="1B7CF469FFF6244ACB88A0B274922446" bold="true" box="[151,501,510,537]" fontSize="11" level="2" pageId="61" pageNumber="62" reason="4">
|
||
<emphasis id="72FF9F17FFF6244ACB88A0B274922446" bold="true" box="[151,501,510,537]" pageId="61" pageNumber="62">
|
||
<taxonomicName id="878B3886FFF6244ACB88A0B274F72446" ID-CoL="9GDJQ" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[151,400,510,537]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="torus" status="sp. nov.">
|
||
<emphasis id="72FF9F17FFF6244ACB88A0B274F72446" bold="true" box="[151,400,510,537]" italics="true" pageId="61" pageNumber="62">Arthromelodes torus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69CC226CFFF6244ACA88A0B374922446" box="[407,501,511,537]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFF6244ACB88A36F775E2401" pageId="61" pageNumber="62" type="description">
|
||
<paragraph id="40344305FFF6244ACB88A36F74F22462" blockId="61.[151,569,510,609]" box="[151,405,547,573]" pageId="61" pageNumber="62">
|
||
(
|
||
<figureCitation id="D8B05F80FFF6244ACB80A36F75912462" box="[159,246,547,573]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="61" pageNumber="62">Figs 26</figureCitation>
|
||
,
|
||
<figureCitation id="D8B05F80FFF6244ACA1DA36F74542462" box="[258,307,547,573]" captionStart="FIGURE 79" captionStartId="181.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-15@181.[151,1437,181,1848]" captionTargetPageId="181" captionText="FIGURE 79. Distribution of Tibetan Batrisini. A. Arthromelodes markam sp. nov. (circle), A. monba sp. nov. (triangle) and A. nepaeformis sp. nov. (square). B. A. nepaeformis sp. nov. (circle), A. songxiaobini sp. nov. (triangle) and A. speciosus sp. nov. (square). C. A. torus sp. nov. (circle), A. zhangmu sp. nov. (triangle) and A. zhentangensis sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340928" httpUri="https://zenodo.org/record/6340928/files/figure.png" pageId="61" pageNumber="62">79C</figureCitation>
|
||
,
|
||
<figureCitation id="D8B05F80FFF6244ACA21A36F74EA2462" box="[318,397,547,573]" captionStart="FIGURE 93" captionStartId="195.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-15@195.[151,1436,181,1623]" captionTargetPageId="195" captionText="FIGURE 93. Environment of and collecting sites at Zhêntang Valley (ĿƋ沟). Slopes at altitudes of 3700 m (A), 4000 m (B), and 3050 m (C). D. Nadang Village (Ḅ当村). E. Road to Xiuxiongma Village (šŏ玛村). F, G. Environment of Ganma Zangbo Valley (HḼẪ布河ě). Photo Credit: F, G by Zhong Peng." figureDoi="http://doi.org/10.5281/zenodo.6340959" httpUri="https://zenodo.org/record/6340959/files/figure.png" pageId="61" pageNumber="62">93C, E</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="40344305FFF6244ACB88A30B775E2401" blockId="61.[151,569,510,609]" box="[151,569,583,609]" pageId="61" pageNumber="62">
|
||
Chinese common name:
|
||
<emphasis id="72FF9F17FFF6244ACAB2A30B775E2401" bold="true" box="[429,569,583,606]" pageId="61" pageNumber="62">瘤únjà甲</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFF6244ACB88A3C377032566" pageId="61" pageNumber="62" type="materials_examined">
|
||
<paragraph id="40344305FFF6244ACB88A3C377032566" blockId="61.[151,1437,654,2013]" pageId="61" pageNumber="62">
|
||
<emphasis id="72FF9F17FFF6244ACB88A3C3742724F7" bold="true" box="[151,320,654,681]" pageId="61" pageNumber="62">Type material</emphasis>
|
||
(
|
||
<materialsCitation id="F0E34958FFF6244ACA4EA3C370242492" ID-GBIF-Occurrence="3879233475" collectingDate="2021-06-25" collectingDateMax="2021-06-28" collectingDateMin="2021-06-25" collectionCode="SNUC" collectorName="Yin & Zhang " country="China" county="Dinggye County" elevation="3060" latitude="27.903055" location="Zhentang Town" longLatPrecision="20" longitude="87.378334" pageId="61" pageNumber="62" specimenCount="20" specimenCount-female="8" specimenCount-male="12" stateProvince="Xizang" typeStatus="holotype">
|
||
20 exx.).
|
||
<emphasis id="72FF9F17FFF6244ACAA0A3C277B024F7" bold="true" box="[447,727,654,680]" pageId="61" pageNumber="62">
|
||
<typeStatus id="9F30FDA7FFF6244ACAA0A3C2770524F7" box="[447,610,654,680]" pageId="61" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<collectingCountry id="389C0395FFF6244AC96FA3C277B524F7" box="[624,722,654,680]" name="China" pageId="61" pageNumber="62">CHINA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="568D888CFFF6244AC9FFA3C3779024F6" box="[736,759,655,681]" pageId="61" pageNumber="62" type="male">♂</specimenCount>
|
||
, ‘China:
|
||
<collectingRegion id="824F8DE7FFF6244AC87DA3C376D124F6" box="[866,950,655,681]" country="China" name="Xizang" pageId="61" pageNumber="62">Xizang</collectingRegion>
|
||
,
|
||
<collectingCounty id="A9553B89FFF6244AC8DDA3C371E724F6" box="[962,1152,655,681]" pageId="61" pageNumber="62">Dinggyê County</collectingCounty>
|
||
, pass to
|
||
<location id="455415DEFFF6244ACFFBA3C370FF24F6" LSID="urn:lsid:plazi:treatment:C822F213FFF62448CB88A0B274082206:455415DEFFF6244ACFFBA3C370FF24F6" box="[1252,1432,655,681]" country="China" county="Dinggye County" latitude="27.903055" longLatPrecision="20" longitude="87.378334" name="Zhentang Town" pageId="61" pageNumber="62" stateProvince="Xizang">Zhêntang Town</location>
|
||
,
|
||
<geoCoordinate id="25BF25C2FFF6244ACB88A3FE74452492" box="[151,290,690,717]" degrees="27" direction="north" minutes="53" orientation="latitude" pageId="61" pageNumber="62" precision="15" seconds="58" value="27.899445">27°53’58”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="25BF25C2FFF6244ACA33A3FF74D32492" box="[300,436,691,717]" degrees="87" direction="east" minutes="27" orientation="longitude" pageId="61" pageNumber="62" precision="15" seconds="21" value="87.45583">87°27’21”E</geoCoordinate>
|
||
,
|
||
<quantity id="8773EEE0FFF6244ACAA1A3FF77752492" box="[446,530,691,717]" metricMagnitude="3" metricUnit="m" metricValue="3.06" pageId="61" pageNumber="62" unit="m" value="3060.0">
|
||
<elevation id="CBA6A436FFF6244ACAA1A3FF77752492" box="[446,530,691,717]" metricMagnitude="3" metricUnit="m" metricValue="3.06" pageId="61" pageNumber="62" unit="m" value="3060.0">3060 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="343565C5FFF6244AC903A3FF77F12492" box="[540,662,691,717]" pageId="61" pageNumber="62" value="2021-06-28">
|
||
<collectingDate id="24719C2DFFF6244AC903A3FF77F12492" box="[540,662,691,717]" pageId="61" pageNumber="62" value="2021-06-28">28.vi.2021</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="ED7E26D3FFF6244AC981A3FF77A12492" box="[670,710,691,717]" pageId="61" pageNumber="62">Yin</collectorName>
|
||
&
|
||
<collectorName id="ED7E26D3FFF6244AC9FAA3FF764A2492" box="[741,813,691,717]" pageId="61" pageNumber="62">Zhang</collectorName>
|
||
leg.,
|
||
<emphasis id="72FF9F17FFF6244AC879A3FF71B42495" bold="true" box="[870,1235,691,714]" pageId="61" pageNumber="62">ƱẪŤĠ县ĿƋ沟公ḡṅðff</emphasis>
|
||
’ (
|
||
<collectionCode id="269ADBC0FFF6244ACFF9A3FF70502492" box="[1254,1335,691,717]" pageId="61" pageNumber="62">SNUC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="F0E34958FFF6244ACE58A3FE77032566" ID-GBIF-Occurrence="3879233454" collectingDate="2021-06-25" collectingDateMax="2021-06-28" collectingDateMin="2021-06-25" collectionCode="SNUC" collectorName="Yin & Zhang & Zhentang & Xiuxiongma Vill. & Z. Peng & Z. Yin & W. Zhang" country="China" county="Dinggye County" elevation="3060" latitude="27.903055" location="Zhentang Town" longLatPrecision="20" longitude="87.378334" pageId="61" pageNumber="62" specimenCount="20" specimenCount-female="8" specimenCount-male="12" stateProvince="Xizang" typeStatus="holotype">
|
||
<emphasis id="72FF9F17FFF6244ACE58A3FE740D24AF" bold="true" pageId="61" pageNumber="62">
|
||
PARATYPES:
|
||
<collectingCountry id="389C0395FFF6244ACA1BA39A740124AF" box="[260,358,726,752]" name="China" pageId="61" pageNumber="62">CHINA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="568D888CFFF6244ACA6CA39B74D124AE" box="[371,438,727,753]" pageId="61" pageNumber="62" type="male">6 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="568D888CFFF6244ACADBA39B776124AE" box="[452,518,727,753]" pageId="61" pageNumber="62" type="female">8 ♀♀</specimenCount>
|
||
, same data as that of
|
||
<typeStatus id="9F30FDA7FFF6244AC81DA39B760F24AE" box="[770,872,727,753]" pageId="61" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="568D888CFFF6244AC86AA39A76DF24AE" box="[885,952,726,753]" pageId="61" pageNumber="62" type="male">5 ♂♂</specimenCount>
|
||
, also from
|
||
<collectorName id="ED7E26D3FFF6244ACF24A39B71CF24AE" box="[1083,1192,727,753]" pageId="61" pageNumber="62">Zhêntang</collectorName>
|
||
, except ‘
|
||
<collectorName id="ED7E26D3FFF6244ACE14A39B75AC254A" pageId="61" pageNumber="62">Xiuxiongma Vill.</collectorName>
|
||
,
|
||
<geoCoordinate id="25BF25C2FFF6244ACBC9A3B67406254A" box="[214,353,762,789]" degrees="27" direction="north" minutes="54" orientation="latitude" pageId="61" pageNumber="62" precision="15" seconds="11" value="27.903055">27°54’11”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="25BF25C2FFF6244ACA73A3B77493254A" box="[364,500,763,789]" degrees="87" direction="east" minutes="22" orientation="longitude" pageId="61" pageNumber="62" precision="15" seconds="42" value="87.378334">87°22’42”E</geoCoordinate>
|
||
,
|
||
<quantity id="8773EEE0FFF6244ACAE0A3B777F0254A" box="[511,663,763,789]" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="61" pageNumber="62" unit="m" value="2850.0" valueMax="3000.0" valueMin="2700.0">
|
||
<elevation id="CBA6A436FFF6244ACAE0A3B777F0254A" box="[511,663,763,789]" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="61" pageNumber="62" unit="m" value="2850.0" valueMax="3000.0" valueMin="2700.0">2700-3000 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="343565C5FFF6244AC9BCA3B6767A254A" box="[675,797,762,789]" pageId="61" pageNumber="62" value="2021-06-25">
|
||
<collectingDate id="24719C2DFFF6244AC9BCA3B6767A254A" box="[675,797,762,789]" pageId="61" pageNumber="62" value="2021-06-25">25.vi.2021</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="ED7E26D3FFF6244AC837A3B776E5254A" box="[808,898,763,789]" pageId="61" pageNumber="62">Z. Peng</collectorName>
|
||
,
|
||
<collectorName id="ED7E26D3FFF6244AC892A3B776B5254A" box="[909,978,763,789]" pageId="61" pageNumber="62">Z. Yin</collectorName>
|
||
&
|
||
<collectorName id="ED7E26D3FFF6244AC8E9A3B77104254A" box="[1014,1123,763,789]" pageId="61" pageNumber="62">W. Zhang</collectorName>
|
||
leg.,
|
||
<emphasis id="72FF9F17FFF6244ACFBFA3B774582569" bold="true" pageId="61" pageNumber="62">ƱẪU喀NJŤĠ县Ŀ ƋWšŏ玛村</emphasis>
|
||
’ (all
|
||
<typeStatus id="9F30FDA7FFF6244ACA65A25374822566" box="[378,485,799,825]" pageId="61" pageNumber="62" type="paratype">paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="269ADBC0FFF6244AC916A253773E2566" box="[521,601,799,825]" pageId="61" pageNumber="62">SNUC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFF6244ACBD8A20E740D22FE" pageId="61" pageNumber="62" type="diagnosis">
|
||
<paragraph id="40344305FFF6244ACBD8A20E740D22FE" blockId="61.[151,1437,654,2013]" pageId="61" pageNumber="62">
|
||
<emphasis id="72FF9F17FFF6244ACBD8A20E74252503" bold="true" box="[199,322,834,860]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
|
||
<emphasis id="72FF9F17FFF6244ACA56A20F74E42502" box="[329,387,835,861]" italics="true" pageId="61" pageNumber="62">Male</emphasis>
|
||
. Body length over
|
||
<quantity id="8773EEE0FFF6244AC940A20F77DE2502" box="[607,697,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="61" pageNumber="62" unit="mm" value="2.6">2.6 mm</quantity>
|
||
. Head subtruncate at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 8 smallest, 11 slightly shorter than 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to approximately apical 2/5 of elytral length. Fore and hind legs simple, mesotrochanter with setose tuft on ventral margin, mesotibia with large apical spine. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 with strongly raised protuberance at middle. Aedeagus strongly asymmetric, median lobe with large basal capsule and oval foramen, ventral stalk narrowed and protruding, dilated at apex, dorsal lobe elongate, forked at apex, parameres reduced and forming single membranous structure.
|
||
<emphasis id="72FF9F17FFF6244ACABFA52F749C2222" box="[416,507,1123,1149]" italics="true" pageId="61" pageNumber="62">Female.</emphasis>
|
||
Body length slightly over
|
||
<quantity id="8773EEE0FFF6244AC836A52E76E62222" box="[809,897,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="61" pageNumber="62" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, legs and abdomen lacking modifications, genitalia as in
|
||
<figureCitation id="D8B05F80FFF6244ACA15A5CB740122FE" box="[266,358,1159,1185]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="61" pageNumber="62">Fig. 26I</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFF62448CBD8A5E6779425B2" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="description">
|
||
<paragraph id="40344305FFF6244ACBD8A5E6779922B6" blockId="61.[151,1437,654,2013]" pageId="61" pageNumber="62">
|
||
<emphasis id="72FF9F17FFF6244ACBD8A5E6743E229B" bold="true" box="[199,345,1194,1220]" pageId="61" pageNumber="62">Description.</emphasis>
|
||
<emphasis id="72FF9F17FFF6244ACA41A5E774FF229A" box="[350,408,1195,1221]" italics="true" pageId="61" pageNumber="62">Male</emphasis>
|
||
. Body (
|
||
<figureCitation id="D8B05F80FFF6244ACAF0A5E777332299" box="[495,596,1195,1222]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="61" pageNumber="62">Fig. 26A</figureCitation>
|
||
) length
|
||
<quantity id="8773EEE0FFF6244AC9B2A5E77674229A" box="[685,787,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" pageId="61" pageNumber="62" unit="mm" value="2.64">2.64 mm</quantity>
|
||
; color reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with short pubescence.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF6244ACBD8A5BF7053200E" blockId="61.[151,1437,654,2013]" pageId="61" pageNumber="62">
|
||
Head (
|
||
<figureCitation id="D8B05F80FFF6244ACA0DA5BF74102351" box="[274,375,1267,1294]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="61" pageNumber="62">Fig. 26B</figureCitation>
|
||
) subtruncate at base, approximately as long as wide, length
|
||
<quantity id="8773EEE0FFF6244ACF3DA5BE71AF2352" box="[1058,1224,1266,1293]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="61" pageNumber="62" unit="mm" value="0.535" valueMax="0.55" valueMin="0.52">0.52–0.55 mm</quantity>
|
||
, width across eyes
|
||
<quantity id="8773EEE0FFF6244ACB88A45A7427236E" box="[151,320,1302,1329]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.5" metricValueMin="5.4" pageId="61" pageNumber="62" unit="mm" value="0.545" valueMax="0.55" valueMin="0.54">0.54–0.55 mm</quantity>
|
||
; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with distinct transverse sulcus between antennal tubercles, mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina present, thin carina present below posteroventral articulation of mandible. Venter with tiny gular foveae (posterior tentorial pits) in single small pit, median carina extending from fovea anteriorly to mouthpart, with thin basal half and thick apical half. Eyes moderately prominent, composed of approximately 25 ommatidia. Antenna
|
||
<quantity id="8773EEE0FFF6244ACFA7A4A3703B2056" box="[1208,1372,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="1.38" metricValueMin="1.34" pageId="61" pageNumber="62" unit="mm" value="1.3599999999999999" valueMax="1.38" valueMin="1.34">1.34–1.38 mm</quantity>
|
||
long, lacking modifications; antennomere 1 thick, subcylindrical, 2–8 each elongate, 8 smallest, much shorter and slightly narrower than 7, 9 longer but narrower than 10, 11 largest, shorter than 9 and 10 combined, sub-fusiform.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF6244ACBD8A717715A2176" blockId="61.[151,1437,654,2013]" pageId="61" pageNumber="62">
|
||
Pronotum (
|
||
<figureCitation id="D8B05F80FFF6244ACA59A71774CD2029" box="[326,426,1627,1654]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="61" pageNumber="62">Fig. 26B</figureCitation>
|
||
) slightly longer than wide, length
|
||
<quantity id="8773EEE0FFF6244AC830A71776B2202A" box="[815,981,1627,1653]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="6.2" metricValueMin="6.0" pageId="61" pageNumber="62" unit="mm" value="0.61" valueMax="0.62" valueMin="0.6">0.60–0.62 mm</quantity>
|
||
, width
|
||
<quantity id="8773EEE0FFF6244ACF37A71671A9202A" box="[1064,1230,1626,1653]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="5.8999999999999995" metricValueMin="5.8" pageId="61" pageNumber="62" unit="mm" value="0.585" valueMax="0.59" valueMin="0.58">0.58–0.59 mm</quantity>
|
||
, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF6244ACBD8A67F710821E6" blockId="61.[151,1437,654,2013]" pageId="61" pageNumber="62">
|
||
Elytra much wider than long, length
|
||
<quantity id="8773EEE0FFF6244AC970A67F76712112" box="[623,790,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.2" metricValueMin="8.1" pageId="61" pageNumber="62" unit="mm" value="0.815" valueMax="0.82" valueMin="0.81">0.81–0.82 mm</quantity>
|
||
, width
|
||
<quantity id="8773EEE0FFF6244AC872A67F71732112" box="[877,1044,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.399999999999999" metricValueMin="9.1" pageId="61" pageNumber="62" unit="mm" value="0.925" valueMax="0.94" valueMin="0.91">0.91–0.94 mm</quantity>
|
||
, constricted at bases; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea to approximately apical 2/5 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from posterior to middle to posterior margin of elytron.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF62449CBD8A68F758A2742" blockId="61.[151,1437,654,2013]" lastBlockId="62.[151,1437,151,358]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</paragraph>
|
||
<paragraph id="40344305FFF52449CBD8A06B7797273A" blockId="62.[151,1437,151,358]" pageId="62" pageNumber="63">
|
||
Legs elongate, fore and hind legs simple. Mesotrochanter (
|
||
<figureCitation id="D8B05F80FFF52449C84EA06B76D3271D" box="[849,948,295,322]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="62" pageNumber="63">Fig. 26C</figureCitation>
|
||
) with moderately long setose tuft on ventral margin, mesotibia (
|
||
<figureCitation id="D8B05F80FFF52449CA6FA00774B02739" box="[368,471,331,358]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="62" pageNumber="63">Fig. 26D</figureCitation>
|
||
) with long spine at apex.
|
||
</paragraph>
|
||
<caption id="14F4138DFFF52449CB88A7D0760320A3" ID-DOI="http://doi.org/10.5281/zenodo.6340791" ID-Zenodo-Dep="6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,250,1692,1716]" targetBox="[154,1437,382,1667]" targetPageId="62">
|
||
<paragraph id="40344305FFF52449CB88A7D0760320A3" blockId="62.[151,1437,1692,1789]" pageId="62" pageNumber="63">
|
||
<emphasis id="72FF9F17FFF52449CB88A7D0744720EA" bold="true" box="[151,288,1692,1717]" pageId="62" pageNumber="63">FIGURE 26.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="878B3886FFF52449CAD8A7D177F020EB" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[455,663,1693,1717]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="torus" status="sp. nov.">
|
||
<emphasis id="72FF9F17FFF52449CAD8A7D177F020EB" box="[455,663,1693,1717]" italics="true" pageId="62" pageNumber="63">Arthromelodes torus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72FF9F17FFF52449C982A7D1778920EA" bold="true" box="[669,750,1693,1717]" pageId="62" pageNumber="63">
|
||
<taxonomicNameLabel id="69CC226CFFF52449C982A7D1778920EA" box="[669,750,1693,1717]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A–H. Male. I. Female).
|
||
<emphasis id="72FF9F17FFF52449C8EFA7D0716E20EB" bold="true" box="[1008,1033,1692,1716]" pageId="62" pageNumber="63">A.</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="72FF9F17FFF52449CFB0A7D171A120EA" bold="true" box="[1199,1222,1693,1717]" pageId="62" pageNumber="63">B.</emphasis>
|
||
Head and pronotum.
|
||
<emphasis id="72FF9F17FFF52449CB88A78C75D72087" bold="true" box="[151,176,1728,1752]" pageId="62" pageNumber="63">C.</emphasis>
|
||
Mesotrochanter.
|
||
<emphasis id="72FF9F17FFF52449CA7FA78D741E2086" bold="true" box="[352,377,1729,1753]" pageId="62" pageNumber="63">D.</emphasis>
|
||
Mesotibia.
|
||
<emphasis id="72FF9F17FFF52449CAEFA78D776F2086" bold="true" box="[496,520,1729,1753]" pageId="62" pageNumber="63">E.</emphasis>
|
||
Tergite 1 (IV).
|
||
<emphasis id="72FF9F17FFF52449C9BAA78D77DF2086" bold="true" box="[677,696,1729,1753]" pageId="62" pageNumber="63">F.</emphasis>
|
||
Sternite 7 (IX).
|
||
<emphasis id="72FF9F17FFF52449C841A78C76FE2086" bold="true" box="[862,921,1728,1753]" pageId="62" pageNumber="63">G, H.</emphasis>
|
||
Aedeagus, lateral (G) and ventral (H).
|
||
<emphasis id="72FF9F17FFF52449CE3CA78D70542086" bold="true" box="[1315,1331,1729,1753]" pageId="62" pageNumber="63">I.</emphasis>
|
||
Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="40344305FFF52448CBD8A66774DB271E" blockId="62.[151,1437,1835,2042]" lastBlockId="63.[151,1437,151,1113]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length
|
||
<quantity id="8773EEE0FFF52449CFB3A6677036211A" box="[1196,1361,1835,1861]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="7.9" metricValueMin="7.7" pageId="62" pageNumber="63" unit="mm" value="0.78" valueMax="0.79" valueMin="0.77">0.77–0.79 mm</quantity>
|
||
, width
|
||
<quantity id="8773EEE0FFF52449CB88A60374592136" box="[151,318,1871,1898]" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="8.6" metricValueMin="8.5" pageId="62" pageNumber="63" unit="mm" value="0.855" valueMax="0.86" valueMin="0.85">0.85–0.86 mm</quantity>
|
||
; with modified tergite 1 (IV). Tergite 1 (
|
||
<figureCitation id="D8B05F80FFF52449C819A603760E2135" box="[774,873,1871,1898]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="62" pageNumber="63">Fig. 26E</figureCitation>
|
||
) longer than tergites 2–4 (V–VII) combined, with single, strongly raised projection at middle, degree of development of projection slightly varying among individuals, basal impression broken by mediobasal and basolateral foveae, with pair of short discal carinae, with incomplete outer marginal carina; tergites 2 and 3 (V and VI) subequal in length along midline, 4 (VII) slightly longer than 2 and 3 combined along middle, 2–4 (V–VII) each with one pair of small basolateral foveae, 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with two mediobasal foveae and large basolateral sockets, lacking basal impression or sulcus, with pair of short lateral carinae; midlength of sternite 2 (IV) as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (
|
||
<figureCitation id="D8B05F80FFF42448C91BA04F77002741" box="[516,615,259,286]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="63" pageNumber="64">Fig. 26F</figureCitation>
|
||
) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF42448CBD8A007711327F2" blockId="63.[151,1437,151,1113]" pageId="63" pageNumber="64">
|
||
Aedeagus (
|
||
<figureCitation id="D8B05F80FFF42448CA58A00774B5273A" box="[327,466,331,357]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="63" pageNumber="64">Fig. 26G, H</figureCitation>
|
||
)
|
||
<quantity id="8773EEE0FFF42448CAFBA007772D273A" box="[484,586,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="63" pageNumber="64" unit="mm" value="0.42">0.42 mm</quantity>
|
||
long, strongly asymmetric; median lobe with large basal capsule and oval foramen, basoventral projection short, ventral stalk protruding and in lateral view dilated at apex; dorsal lobe recumbent, apically forked into two lobes; parameres reduced to single membranous structure.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF42448CBD8A0FB77E5243E" blockId="63.[151,1437,151,1113]" pageId="63" pageNumber="64">
|
||
<emphasis id="72FF9F17FFF42448CBD8A0FB747C278E" box="[199,283,439,465]" italics="true" pageId="63" pageNumber="64">Female</emphasis>
|
||
. Similar to male in external morphology; antenna shorter; legs and abdomen lacking spines/projections or modifications; each compound eye composed of approximately 23 ommatidia; humeral angle rounded. Measurements (as for male): body length
|
||
<quantity id="8773EEE0FFF42448C91DA0B277C12446" box="[514,678,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.61" metricValueMin="2.53" pageId="63" pageNumber="64" unit="mm" value="2.57" valueMax="2.61" valueMin="2.53">2.53–2.61 mm</quantity>
|
||
; length/width of head 0.53–0.54/
|
||
<quantity id="8773EEE0FFF42448CF0CA0B271102446" box="[1043,1143,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="63" pageNumber="64" unit="mm" value="0.53">0.53 mm</quantity>
|
||
, pronotum 0.56–0.57/
|
||
<quantity id="8773EEE0FFF42448CE74A0B275A22462" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="63" pageNumber="64" unit="mm" value="0.56">0.56 mm</quantity>
|
||
, elytra 0.72–0.76/
|
||
<quantity id="8773EEE0FFF42448CA8EA36F77502462" box="[401,567,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.0" metricValueMin="8.9" pageId="63" pageNumber="64" unit="mm" value="0.895" valueMax="0.9" valueMin="0.89">0.89–0.90 mm</quantity>
|
||
; abdomen 0.77–0.80/
|
||
<quantity id="8773EEE0FFF42448C834A36E76B72462" box="[811,976,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="8.7" metricValueMin="8.5" pageId="63" pageNumber="64" unit="mm" value="0.86" valueMax="0.87" valueMin="0.85">0.85–0.87 mm</quantity>
|
||
; length of antenna
|
||
<quantity id="8773EEE0FFF42448CFB7A36F70282462" box="[1192,1359,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.265" metricValueMax="1.27" metricValueMin="1.26" pageId="63" pageNumber="64" unit="mm" value="1.265" valueMax="1.27" valueMin="1.26">1.26–1.27 mm</quantity>
|
||
; maximum width of genitalia (
|
||
<figureCitation id="D8B05F80FFF42448CAB2A30B7760243D" box="[429,519,583,610]" captionStart="FIGURE 26" captionStartId="62.[151,250,1692,1716]" captionTargetBox="[154,1437,382,1667]" captionTargetId="figure-94@62.[151,1437,381,1669]" captionTargetPageId="62" captionText="FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I." figureDoi="http://doi.org/10.5281/zenodo.6340791" httpUri="https://zenodo.org/record/6340791/files/figure.png" pageId="63" pageNumber="64">Fig. 26I</figureCitation>
|
||
)
|
||
<quantity id="8773EEE0FFF42448C908A30B7719243E" box="[535,638,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="mm" value="0.3">0.30 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="40344305FFF42448CBD8A326779425B2" blockId="63.[151,1437,151,1113]" pageId="63" pageNumber="64">
|
||
<emphasis id="72FF9F17FFF42448CBD8A32674D624DA" bold="true" box="[199,433,618,645]" pageId="63" pageNumber="64">Comparative notes.</emphasis>
|
||
<taxonomicName id="878B3886FFF42448CAA7A32777F824DB" authorityName="Yin" authorityYear="2022" box="[440,671,619,645]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="torus">
|
||
<emphasis id="72FF9F17FFF42448CAA7A32777F824DB" box="[440,671,619,645]" italics="true" pageId="63" pageNumber="64">Arthromelodes torus</emphasis>
|
||
</taxonomicName>
|
||
is morphologically most similar to
|
||
<taxonomicName id="878B3886FFF42448CF2EA32771D124DA" authorityName="Yin" authorityYear="2022" box="[1073,1206,619,645]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="alpitorus">
|
||
<emphasis id="72FF9F17FFF42448CF2EA327712524DA" box="[1073,1090,619,645]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448CF4EA32771D124DA" box="[1105,1206,619,645]" italics="true" pageId="63" pageNumber="64">alpitorus</emphasis>
|
||
</taxonomicName>
|
||
by sharing most external characters, as well as a similar male tergal modification and the form of the aedeagus. The new species can be separated by the smaller body size (
|
||
<quantity id="8773EEE0FFF42448C94EA3FF77D02492" box="[593,695,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" pageId="63" pageNumber="64" unit="mm" value="2.64">2.64 mm</quantity>
|
||
vs.
|
||
<quantity id="8773EEE0FFF42448C9F8A3FF76EB2492" box="[743,908,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.84" metricValueMax="2.88" metricValueMin="2.8" pageId="63" pageNumber="64" unit="mm" value="2.84" valueMax="2.88" valueMin="2.8">2.80–2.88 mm</quantity>
|
||
for male;
|
||
<quantity id="8773EEE0FFF42448CF1CA3FE71C02492" box="[1027,1191,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.61" metricValueMin="2.53" pageId="63" pageNumber="64" unit="mm" value="2.57" valueMax="2.61" valueMin="2.53">2.53–2.61 mm</quantity>
|
||
vs
|
||
<quantity id="8773EEE0FFF42448CFCFA3FF70132492" box="[1232,1396,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.73" metricValueMax="2.8" metricValueMin="2.66" pageId="63" pageNumber="64" unit="mm" value="2.73" valueMax="2.8" valueMin="2.66">2.66–2.80 mm</quantity>
|
||
for female), a much longer median longitudinal sulcus of the pronotum, the male having a thinner and slightly curved spine at the apex of the mesotibia, the median projection of tergite 1 (IV) with single posteromedial corner (with acute posterolateral angles in
|
||
<taxonomicName id="878B3886FFF42448CAF6A25377092566" authorityName="Yin" authorityYear="2022" box="[489,622,799,825]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="alpitorus">
|
||
<emphasis id="72FF9F17FFF42448CAF6A253749D2566" box="[489,506,799,825]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448C916A25377092566" box="[521,622,799,825]" italics="true" pageId="63" pageNumber="64">alpitorus</emphasis>
|
||
</taxonomicName>
|
||
), the area posterior to the projection flat along the midline (raised in
|
||
<taxonomicName id="878B3886FFF42448CE9BA25375B62503" authorityName="Yin" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="torus">
|
||
<emphasis id="72FF9F17FFF42448CE9BA25370F22566" box="[1412,1429,799,825]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448CB88A20F75B62503" box="[151,209,835,860]" italics="true" pageId="63" pageNumber="64">torus</emphasis>
|
||
</taxonomicName>
|
||
), and a less expanded apex of the median lobe of the aedeagus. These two species appear to be well isolated by altitudinal barriers.
|
||
<taxonomicName id="878B3886FFF42448CA88A22B771925DF" authorityName="Yin" authorityYear="2022" box="[407,638,871,897]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="torus">
|
||
<emphasis id="72FF9F17FFF42448CA88A22B771925DF" box="[407,638,871,897]" italics="true" pageId="63" pageNumber="64">Arthromelodes torus</emphasis>
|
||
</taxonomicName>
|
||
is also close to
|
||
<taxonomicName id="878B3886FFF42448C82DA22B76A725DE" authorityName="Yin" authorityYear="2022" box="[818,960,871,897]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="angulatus">
|
||
<emphasis id="72FF9F17FFF42448C82DA22B762425DE" box="[818,835,871,897]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448C84DA22B76A725DE" box="[850,960,871,897]" italics="true" pageId="63" pageNumber="64">angulatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="878B3886FFF42448C8D1A22B712325DF" authorityName="Yin" authorityYear="2022" box="[974,1092,871,897]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="aniqiao">
|
||
<emphasis id="72FF9F17FFF42448C8D1A22B76B825DE" box="[974,991,871,897]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448C8F1A22B712325DF" box="[1006,1092,871,896]" italics="true" pageId="63" pageNumber="64">aniqiao</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="878B3886FFF42448CF4DA22B718F25DE" authorityName="Yin" authorityYear="2022" box="[1106,1256,871,897]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="complexus">
|
||
<emphasis id="72FF9F17FFF42448CF4DA22B710425DE" box="[1106,1123,871,897]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448CF6DA22B718F25DE" box="[1138,1256,871,897]" italics="true" pageId="63" pageNumber="64">complexus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="878B3886FFF42448CFE9A22B702125DE" authorityName="Yin" authorityYear="2022" box="[1270,1350,871,897]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="lage">
|
||
<emphasis id="72FF9F17FFF42448CFE9A22B706025DE" box="[1270,1287,871,897]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448CE09A22B702125DE" box="[1302,1350,871,897]" italics="true" pageId="63" pageNumber="64">lage</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="878B3886FFF42448CE9BA22B746625FA" authorityName="Yin" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="langjicuo">
|
||
<emphasis id="72FF9F17FFF42448CE9BA22B70F225DE" box="[1412,1429,871,897]" italics="true" pageId="63" pageNumber="64">A</emphasis>
|
||
.
|
||
<emphasis id="72FF9F17FFF42448CB88A2C7746625FA" box="[151,257,907,933]" italics="true" pageId="63" pageNumber="64">langjicuo</emphasis>
|
||
</taxonomicName>
|
||
, sharing with them a setose male mesotrochanter, as well as a similar general form of the aedeagus. The male of the new species can be readily separated by the unique presence of a large central projection on tergite 1 (IV), as well as the long apical spine of the mesotibia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFF42448CBD8A2BA7153224E" box="[199,1076,1014,1041]" pageId="63" pageNumber="64" type="distribution">
|
||
<paragraph id="40344305FFF42448CBD8A2BA7153224E" blockId="63.[151,1437,151,1113]" box="[199,1076,1014,1041]" pageId="63" pageNumber="64">
|
||
<emphasis id="72FF9F17FFF42448CBD8A2BA743E224F" bold="true" box="[199,345,1014,1040]" pageId="63" pageNumber="64">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="D8B05F80FFF42448CA76A2BB74DD224E" box="[361,442,1015,1041]" captionStart="FIGURE 92" captionStartId="194.[151,250,1169,1193]" captionTargetBox="[151,1437,181,1145]" captionTargetId="figure-19@194.[151,1437,181,1145]" captionTargetPageId="194" captionText="FIGURE 92. Environment of Zhangmu Valley (樟木沟). Environment of Youyiqiao (友谊桥) (A, B), Lixin Village (ĒẾ村) (C) and Qu Township (DZ乡) (D). Photo Credit: A, B by Jian-Qing Zhu; C, D by Wen-Xuan Bi." figureDoi="http://doi.org/10.5281/zenodo.6340957" httpUri="https://zenodo.org/record/6340957/files/figure.png" pageId="63" pageNumber="64">Fig. 92</figureCitation>
|
||
). Dinggyê County,
|
||
<collectingRegion id="824F8DE7FFF42448C989A2BB77B4224E" box="[662,723,1015,1041]" country="China" name="Xizang" pageId="63" pageNumber="64">Tibet</collectingRegion>
|
||
, SW
|
||
<collectingCountry id="389C0395FFF42448C810A2BB7635224E" box="[783,850,1015,1041]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
|
||
(
|
||
<figureCitation id="D8B05F80FFF42448C87FA2BB76AD224E" box="[864,970,1015,1041]" captionStart="FIGURE 79" captionStartId="181.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-15@181.[151,1437,181,1848]" captionTargetPageId="181" captionText="FIGURE 79. Distribution of Tibetan Batrisini. A. Arthromelodes markam sp. nov. (circle), A. monba sp. nov. (triangle) and A. nepaeformis sp. nov. (square). B. A. nepaeformis sp. nov. (circle), A. songxiaobini sp. nov. (triangle) and A. speciosus sp. nov. (square). C. A. torus sp. nov. (circle), A. zhangmu sp. nov. (triangle) and A. zhentangensis sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340928" httpUri="https://zenodo.org/record/6340928/files/figure.png" pageId="63" pageNumber="64">Figs 79C</figureCitation>
|
||
,
|
||
<figureCitation id="D8B05F80FFF42448C8C9A2BB7140224E" box="[982,1063,1015,1041]" captionStart="FIGURE 93" captionStartId="195.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-15@195.[151,1436,181,1623]" captionTargetPageId="195" captionText="FIGURE 93. Environment of and collecting sites at Zhêntang Valley (ĿƋ沟). Slopes at altitudes of 3700 m (A), 4000 m (B), and 3050 m (C). D. Nadang Village (Ḅ当村). E. Road to Xiuxiongma Village (šŏ玛村). F, G. Environment of Ganma Zangbo Valley (HḼẪ布河ě). Photo Credit: F, G by Zhong Peng." figureDoi="http://doi.org/10.5281/zenodo.6340959" httpUri="https://zenodo.org/record/6340959/files/figure.png" pageId="63" pageNumber="64">93C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFF42448CBD8A55774082206" pageId="63" pageNumber="64" type="etymology">
|
||
<paragraph id="40344305FFF42448CBD8A55774082206" blockId="63.[151,1437,151,1113]" pageId="63" pageNumber="64">
|
||
<emphasis id="72FF9F17FFF42448CBD8A5577429226A" bold="true" box="[199,334,1051,1077]" pageId="63" pageNumber="64">Etymology.</emphasis>
|
||
The new species name ‘
|
||
<taxonomicName id="878B3886FFF42448C97FA55777FD226B" authorityName="Yin" authorityYear="2022" box="[608,666,1051,1076]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="torus">
|
||
<emphasis id="72FF9F17FFF42448C97FA55777FD226B" box="[608,666,1051,1076]" italics="true" pageId="63" pageNumber="64">torus</emphasis>
|
||
</taxonomicName>
|
||
(swelling, protuberance)’ is a Latin noun referring to the protuberant male tergite 1 (IV).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |