291 lines
39 KiB
XML
291 lines
39 KiB
XML
<document id="AA72009757BA71CAAC0CE30EEA4DBBD8" ID-CLB-Dataset="113069" ID-DOI="10.11646/zootaxa.5111.1.1" ID-GBIF-Dataset="f3992bb2-9962-4f3c-8f5b-131cfe552c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6340729" ID-ZooBank="836B0F69-037C-4D0F-80DB-94FE454F48E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646828425946" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="C822F213FFD62468CB88A773773D2446" docLanguage="en" docName="zootaxa.5111.1.1.pdf" docOrigin="Zootaxa 5111 (1)" docSource="http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Arthromelodes cylindricus Yin 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="341B8A6BFFCB2477CB1FA14C7567265F" masterDocTitle="The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="30" updateTime="1699317591893" updateUser="ExternalLinkService">
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<mods:title id="A0A7C8D6555D2BF10D2E694E44976366">The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="9BCC4967C631B835CBF9F84C5D87A5BD">Yin, Zi-Wei</mods:namePart>
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<mods:title id="878BB072812B114B37721923C19A1643">Zootaxa</mods:title>
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<mods:part id="9C90B97442624D5C90330C63E2FE446B">
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<mods:date id="4024F49BD2C6109DAF6392155C1078C9">2022</mods:date>
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<mods:number id="F6EAB662A810A80F24D88C8E221317BE">2022-03-09</mods:number>
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<mods:identifier id="00F90322F64D43A3C078E55EE2E60438" type="CLB-Dataset">113069</mods:identifier>
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<mods:identifier id="68FAB4E98E53E0EBE007AA6293605483" type="DOI">10.11646/zootaxa.5111.1.1</mods:identifier>
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<treatment id="C822F213FFD62468CB88A773773D2446" ID-DOI="http://doi.org/10.5281/zenodo.6964218" ID-GBIF-Taxon="198579299" ID-Zenodo-Dep="6964218" LSID="urn:lsid:plazi:treatment:C822F213FFD62468CB88A773773D2446" httpUri="http://treatment.plazi.org/id/C822F213FFD62468CB88A773773D2446" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
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<subSubSection id="0891108EFFD6246ACB88A773775B2005" box="[151,572,1599,1626]" pageId="29" pageNumber="30" type="nomenclature">
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<paragraph id="40344305FFD6246ACB88A773775B2005" blockId="29.[151,572,1599,1698]" box="[151,572,1599,1626]" pageId="29" pageNumber="30">
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<heading id="1B7CF469FFD6246ACB88A773775B2005" bold="true" box="[151,572,1599,1626]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
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<emphasis id="72FF9F17FFD6246ACB88A773775B2005" bold="true" box="[151,572,1599,1626]" pageId="29" pageNumber="30">
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<taxonomicName id="878B3886FFD6246ACB88A77374BF2005" ID-CoL="9GDGV" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[151,472,1599,1626]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cylindricus" status="sp. nov.">
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<emphasis id="72FF9F17FFD6246ACB88A77374BF2005" bold="true" box="[151,472,1599,1626]" italics="true" pageId="29" pageNumber="30">Arthromelodes cylindricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="69CC226CFFD6246ACAC0A70C775B2005" box="[479,572,1600,1626]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0891108EFFD6246ACB88A728775E20FF" pageId="29" pageNumber="30" type="description">
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<paragraph id="40344305FFD6246ACB88A72874F42021" blockId="29.[151,572,1599,1698]" box="[151,403,1636,1662]" pageId="29" pageNumber="30">
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(
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<figureCitation id="D8B05F80FFD6246ACB80A72875922021" box="[159,245,1636,1662]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="29" pageNumber="30">Figs 11</figureCitation>
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,
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<figureCitation id="D8B05F80FFD6246ACA1EA72874532021" box="[257,308,1636,1662]" captionStart="FIGURE 78" captionStartId="180.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-19@180.[151,1437,181,1848]" captionTargetPageId="180" captionText="FIGURE 78. Distribution of Tibetan Batrisini. A. Arthromelodes cylindricus sp. nov. (circle), A. flosculus sp. nov. (triangle) and A. gyamda sp. nov. (square). B. A. intricatus sp. nov. (circle), A. lage sp. nov. (triangle) and A. langjicuo sp. nov. (square). C. A. latithorax sp. nov. (circle), A. lebus sp. nov. (triangle) and A. lotus sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340926" httpUri="https://zenodo.org/record/6340926/files/figure.png" pageId="29" pageNumber="30">78A</figureCitation>
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,
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<figureCitation id="D8B05F80FFD6246ACA20A72874EC2021" box="[319,395,1636,1662]" captionStart="FIGURE 96" captionStartId="198.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-19@198.[151,1436,181,1622]" captionTargetPageId="198" captionText="FIGURE 96. Environment of and collecting sites at Nyingchi area (ffż地区). A. Gongbo'gyamda (工布ȕḭ). B. Zhongzhe Village (ΦĖ村). C. Kading Valley (ϮŤ沟). D. Chongge Cuo (ṗẋś). E, F. A slope near Serjila mountain pass (DzĻŃ山 口). Photo Credit: A, B, E, F by Zhong Peng." figureDoi="http://doi.org/10.5281/zenodo.6340965" httpUri="https://zenodo.org/record/6340965/files/figure.png" pageId="29" pageNumber="30">96E, F</figureCitation>
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)
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</paragraph>
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<paragraph id="40344305FFD6246ACB88A7C4775E20FF" blockId="29.[151,572,1599,1698]" box="[151,569,1672,1698]" pageId="29" pageNumber="30">
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Chinese common name:
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<emphasis id="72FF9F17FFD6246ACAB2A7C5775E20FF" bold="true" box="[429,569,1673,1696]" pageId="29" pageNumber="30">Ñ型njà甲</emphasis>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0891108EFFD6246ACB88A79C74392125" pageId="29" pageNumber="30" type="materials_examined">
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<paragraph id="40344305FFD6246ACB88A79C74392125" blockId="29.[151,1437,1743,2022]" pageId="29" pageNumber="30">
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<emphasis id="72FF9F17FFD6246ACB88A79C742420B6" bold="true" box="[151,323,1743,1770]" pageId="29" pageNumber="30">Type material</emphasis>
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(19 exx.).
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<materialsCitation id="F0E34958FFD6246ACAD1A783766B20B5" ID-GBIF-Occurrence="3879233455" box="[462,780,1743,1770]" country="China" location="CHINA" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="72FF9F17FFD6246ACAD1A783778E20B6" bold="true" box="[462,745,1743,1769]" pageId="29" pageNumber="30">
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||
<typeStatus id="9F30FDA7FFD6246ACAD1A783771620B6" box="[462,625,1743,1769]" pageId="29" pageNumber="30" type="holotype">HOLOTYPE</typeStatus>
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||
:
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<collectingCountry id="389C0395FFD6246AC99DA783778320B6" box="[642,740,1743,1769]" name="China" pageId="29" pageNumber="30">CHINA</collectingCountry>
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||
:
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||
</emphasis>
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<specimenCount id="568D888CFFD6246AC9EAA79C766B20B5" box="[757,780,1744,1770]" pageId="29" pageNumber="30" type="male">♂</specimenCount>
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||
</materialsCitation>
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||
, ‘
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||
<materialsCitation id="F0E34958FFD6246AC83DA79C759F216D" ID-GBIF-Occurrence="3879233304" collectingDate="2018-07-05" collectionCode="SNUC" collectorName="Cheng & Peng & Shen" country="China" county="Nyingchi City" elevation="4340" latitude="29.61389" location="Sheji'ema" longLatPrecision="20" longitude="95.69278" municipality="Shejila" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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<collectingCountry id="389C0395FFD6246AC83DA79C760920B5" box="[802,878,1744,1770]" name="China" pageId="29" pageNumber="30">China</collectingCountry>
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||
:
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<collectingRegion id="824F8DE7FFD6246AC861A79C76B520B5" box="[894,978,1744,1770]" country="China" name="Xizang" pageId="29" pageNumber="30">Xizang</collectingRegion>
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,
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<collectingCounty id="A9553B89FFD6246AC8FCA79C71EF20B5" box="[995,1160,1744,1770]" pageId="29" pageNumber="30">Nyingchi City</collectingCounty>
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||
,
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<collectingMunicipality id="A050D97FFFD6246ACF86A79C718D20B5" box="[1177,1258,1744,1770]" pageId="29" pageNumber="30">Shejila</collectingMunicipality>
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, nr.
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<location id="455415DEFFD6246ACE3BA79C70FF20B5" LSID="urn:lsid:plazi:treatment:C822F213FFD62468CB88A773773D2446:455415DEFFD6246ACE3BA79C70FF20B5" box="[1316,1432,1744,1770]" country="China" county="Nyingchi City" latitude="29.61389" longLatPrecision="20" longitude="95.69278" municipality="Shejila" name="Sheji'ema" pageId="29" pageNumber="30" stateProvince="Xizang">Sheji’ema</location>
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||
,
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<geoCoordinate id="25BF25C2FFD6246ACB88A7B874442150" box="[151,291,1780,1807]" degrees="29" direction="north" minutes="36" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="50" value="29.61389">29°36’50”N</geoCoordinate>
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||
,
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<geoCoordinate id="25BF25C2FFD6246ACA30A7B874DF2150" box="[303,440,1780,1807]" degrees="95" direction="east" minutes="41" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="34" value="95.69278">95°41’34”E</geoCoordinate>
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||
,
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<quantity id="8773EEE0FFD6246ACADAA7B8777A2151" box="[453,541,1780,1806]" metricMagnitude="3" metricUnit="m" metricValue="4.34" pageId="29" pageNumber="30" unit="m" value="4340.0">
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||
<elevation id="CBA6A436FFD6246ACADAA7B8777A2151" box="[453,541,1780,1806]" metricMagnitude="3" metricUnit="m" metricValue="4.34" pageId="29" pageNumber="30" unit="m" value="4340.0">4340 m</elevation>
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</quantity>
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||
,
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||
<date id="343565C5FFD6246AC935A7B877CB2150" box="[554,684,1780,1807]" pageId="29" pageNumber="30" value="2018-07-05">
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||
<collectingDate id="24719C2DFFD6246AC935A7B877CB2150" box="[554,684,1780,1807]" pageId="29" pageNumber="30" value="2018-07-05">05.vii.2018</collectingDate>
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||
</date>
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, leaf litter, sifted,
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<collectorName id="ED7E26D3FFD6246AC864A7B876A02151" box="[891,967,1780,1806]" pageId="29" pageNumber="30">Cheng</collectorName>
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,
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<collectorName id="ED7E26D3FFD6246AC8CCA7B8716C2151" box="[979,1035,1780,1806]" pageId="29" pageNumber="30">Peng</collectorName>
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||
&
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<collectorName id="ED7E26D3FFD6246ACF2EA7B8710E2151" box="[1073,1129,1780,1806]" pageId="29" pageNumber="30">Shen</collectorName>
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||
leg. [
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<emphasis id="72FF9F17FFD6246ACFB3A7B970EB2153" bold="true" box="[1196,1420,1781,1804]" pageId="29" pageNumber="30">ƱẪffż市DzĻŃ</emphasis>
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||
]’ (
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<collectionCode id="269ADBC0FFD6246ACB80A6547597216D" box="[159,240,1816,1842]" pageId="29" pageNumber="30">SNUC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="F0E34958FFD6246ACA1AA65B743D2125" ID-GBIF-Occurrence="3879233444" collectingDate="2021-07-14" collectionCode="SNUC" collectorName="Peng & Yin & Zhang" country="China" county="Nyingchi City" elevation="4300" latitude="29.616945" location="Lulang Tourist Center" longLatPrecision="20" longitude="94.69861" municipality="Shejila" pageId="29" pageNumber="30" specimenCount="18" specimenCount-female="7" specimenCount-male="11" stateProvince="Xizang" typeStatus="paratype">
|
||
<emphasis id="72FF9F17FFD6246ACA1AA65B7745216E" bold="true" box="[261,546,1815,1841]" pageId="29" pageNumber="30">
|
||
<typeStatus id="9F30FDA7FFD6246ACA1AA65B74CA216E" box="[261,429,1815,1841]" pageId="29" pageNumber="30" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<collectingCountry id="389C0395FFD6246ACAA4A65B777A216E" box="[443,541,1815,1841]" name="China" pageId="29" pageNumber="30">CHINA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="568D888CFFD6246AC934A6547709216D" box="[555,622,1816,1842]" pageId="29" pageNumber="30" type="male">9 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="568D888CFFD6246AC963A65477D9216D" box="[636,702,1816,1842]" pageId="29" pageNumber="30" type="female">4 ♀♀</specimenCount>
|
||
, same data as that of
|
||
<typeStatus id="9F30FDA7FFD6246AC8A3A6547145216D" box="[956,1058,1816,1842]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="568D888CFFD6246ACF30A6547115216D" box="[1071,1138,1816,1842]" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="568D888CFFD6246ACF9FA65471A5216D" box="[1152,1218,1816,1842]" pageId="29" pageNumber="30" type="female">3 ♀♀</specimenCount>
|
||
, also from
|
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<collectingMunicipality id="A050D97FFFD6246ACE58A65470FF216D" box="[1351,1432,1816,1842]" pageId="29" pageNumber="30">Shejila</collectingMunicipality>
|
||
, except ‘
|
||
<location id="455415DEFFD6246ACBF4A670748F2109" LSID="urn:lsid:plazi:treatment:C822F213FFD62468CB88A773773D2446:455415DEFFD6246ACBF4A670748F2109" box="[235,488,1852,1878]" country="China" latitude="29.616945" longLatPrecision="20" longitude="94.69861" municipality="Shejila" name="Lulang Tourist Center" pageId="29" pageNumber="30">Lulang Tourist Center</location>
|
||
,
|
||
<geoCoordinate id="25BF25C2FFD6246ACAECA67077192109" box="[499,638,1852,1878]" degrees="29" direction="north" minutes="37" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="01" value="29.616945">29°37’01”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="25BF25C2FFD6246AC995A67076742108" box="[650,787,1852,1879]" degrees="94" direction="east" minutes="41" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="55" value="94.69861">94°41’55”E</geoCoordinate>
|
||
,
|
||
<quantity id="8773EEE0FFD6246AC801A67076122109" box="[798,885,1852,1878]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="29" pageNumber="30" unit="m" value="4300.0">
|
||
<elevation id="CBA6A436FFD6246AC801A67076122109" box="[798,885,1852,1878]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="29" pageNumber="30" unit="m" value="4300.0">4300 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="343565C5FFD6246AC89EA67071652109" box="[897,1026,1852,1878]" pageId="29" pageNumber="30" value="2021-07-14">
|
||
<collectingDate id="24719C2DFFD6246AC89EA67071652109" box="[897,1026,1852,1878]" pageId="29" pageNumber="30" value="2021-07-14">14.vii.2021</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="ED7E26D3FFD6246ACF11A670712E2109" box="[1038,1097,1852,1878]" pageId="29" pageNumber="30">Peng</collectorName>
|
||
,
|
||
<collectorName id="ED7E26D3FFD6246ACF4CA670711C2109" box="[1107,1147,1852,1878]" pageId="29" pageNumber="30">Yin</collectorName>
|
||
&
|
||
<collectorName id="ED7E26D3FFD6246ACF80A670718E2109" box="[1183,1257,1852,1878]" pageId="29" pageNumber="30">Zhang</collectorName>
|
||
,
|
||
<emphasis id="72FF9F17FFD6246ACFEBA671758C2127" bold="true" pageId="29" pageNumber="30">DzĻŃ游Β中 心北坡</emphasis>
|
||
’ (
|
||
<collectionCode id="269ADBC0FFD6246ACA1EA62C74352125" box="[257,338,1888,1914]" pageId="29" pageNumber="30">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFD62469CBD8A6CF779D271E" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
|
||
<paragraph id="40344305FFD62469CBD8A6CF779D271E" blockId="29.[151,1437,1743,2022]" lastBlockId="30.[151,1437,151,2013]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<emphasis id="72FF9F17FFD6246ACBD8A6CF742521C2" bold="true" box="[199,322,1923,1949]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
|
||
<emphasis id="72FF9F17FFD6246ACA54A6C874E221C1" box="[331,389,1924,1950]" italics="true" pageId="29" pageNumber="30">Male</emphasis>
|
||
. Body length over
|
||
<quantity id="8773EEE0FFD6246AC977A6C8767621C1" box="[616,785,1924,1950]" metricMagnitude="-3" metricUnit="m" metricValue="2.635" metricValueMax="2.67" metricValueMin="2.6" pageId="29" pageNumber="30" unit="mm" value="2.635" valueMax="2.67" valueMin="2.6">2.60–2.67 mm</quantity>
|
||
. Head subtruncate at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae moderately large and asetose; antenna long, antennomeres each slightly elongate, lacking modifications. Discal stria of elytron extending to approximately apical 1/4 of elytral length. Fore legs simple, mesotrochanter with small tuft of setae on ventral margin, mesotibia with distinct apical spur, metafemur with dorsal setose patch at apex. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) with deep central cavity. Aedeagus strongly asymmetric, median lobe protruding apically, ventral stalk in dorsal view broad, dorsal lobe laterally strongly curved at middle, parameres reduced and forming single semi-sclerotized structure.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFD52468CBD8A00677F727F2" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
|
||
<paragraph id="40344305FFD52469CBD8A00671D927D6" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
<emphasis id="72FF9F17FFD52469CBD8A006743E273B" bold="true" box="[199,345,330,356]" pageId="30" pageNumber="31">Description.</emphasis>
|
||
<emphasis id="72FF9F17FFD52469CA7FA00774FD273A" box="[352,410,331,357]" italics="true" pageId="30" pageNumber="31">Male</emphasis>
|
||
. Body (
|
||
<figureCitation id="D8B05F80FFD52469CAECA007773F273A" box="[499,600,331,357]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11A</figureCitation>
|
||
) length
|
||
<quantity id="8773EEE0FFD52469C9ABA007763E273A" box="[692,857,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.635" metricValueMax="2.67" metricValueMin="2.6" pageId="30" pageNumber="31" unit="mm" value="2.635" valueMax="2.67" valueMin="2.6">2.60–2.67 mm</quantity>
|
||
, habitus generally cylindrical; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A0DF74A8254A" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
Head (
|
||
<figureCitation id="D8B05F80FFD52469CA0FA0DF741527F2" box="[272,370,403,429]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11B</figureCitation>
|
||
) subtruncate at base, as long as broad, length and width across eyes
|
||
<quantity id="8773EEE0FFD52469CF7EA0DE706127F2" box="[1121,1286,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.1" metricValueMin="5.0" pageId="30" pageNumber="31" unit="mm" value="0.505" valueMax="0.51" valueMin="0.5">0.50–0.51 mm</quantity>
|
||
; vertex, frons and clypeus roughly punctate; vertex with moderately large, asetose vertexal foveae (dorsal tentorial pits), with curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus, with short frontal-clypeal carinae at sides; clypeus with anterior margin carinate and moderately raised; ocular-mandibular carina thick and complete. Venter with gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 19 large ommatidia. Antenna with loosely formed club, lacking modifications, length
|
||
<quantity id="8773EEE0FFD52469CBFAA3FF74EC2492" box="[229,395,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.24" metricValueMin="1.23" pageId="30" pageNumber="31" unit="mm" value="1.2349999999999999" valueMax="1.24" valueMin="1.23">1.23–1.24 mm</quantity>
|
||
; antennomere 1 thick, subcylindrical, anterolateral margin slightly impressed and filled with short setae (but not to form prominent trichome), 2–11 each elongate, 8 smallest, 11 largest, much shorter than 9 and 10 combined, sub-fusiform.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A25370E825B2" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
Pronotum (
|
||
<figureCitation id="D8B05F80FFD52469CA5AA25374C02566" box="[325,423,799,825]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11B</figureCitation>
|
||
) approximately as long as wide, length and width
|
||
<quantity id="8773EEE0FFD52469C8C1A25271E42566" box="[990,1155,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="30" pageNumber="31" unit="mm" value="0.555" valueMax="0.56" valueMin="0.55">0.55–0.56 mm</quantity>
|
||
, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae, with additional pair of marginal antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A2BB76442222" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
Elytra much wider than long, length
|
||
<quantity id="8773EEE0FFD52469C940A2BB7664224E" box="[607,771,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="7.4" metricValueMin="7.3" pageId="30" pageNumber="31" unit="mm" value="0.735" valueMax="0.74" valueMin="0.73">0.73–0.74 mm</quantity>
|
||
, width
|
||
<quantity id="8773EEE0FFD52469C84BA2BB76DE224E" box="[852,953,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="30" pageNumber="31" unit="mm" value="0.82">0.82 mm</quantity>
|
||
, constricted at bases; each elytron with two large, asetose basal foveae, lacking subbasal fovea; humerus lacking prominence or denticle; discal stria extending from outer basal fovea to approximately apical 1/4 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from basal 1/3 to apex. Hind wings absent.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A5CB776E236E" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly and shallowly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A47774DC23C2" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
Legs elongate, fore simple. Mesotrochanter with small setose tuft on ventral margin, mesotibia with distinct apical spur; metatrochanter setose on ventral margin, metafemur (
|
||
<figureCitation id="D8B05F80FFD52469C862A41376872326" box="[893,992,1375,1401]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11C</figureCitation>
|
||
) with dorsal sensory area covered with short, dense setae at apex.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A4EB741A2176" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
Abdomen narrower than elytra, widest at basolateral margins of tergite 1 (IV), length
|
||
<quantity id="8773EEE0FFD52469CFB8A4EA7029239E" box="[1191,1358,1446,1473]" metricMagnitude="-4" metricUnit="m" metricValue="7.65" metricValueMax="7.8" metricValueMin="7.5" pageId="30" pageNumber="31" unit="mm" value="0.765" valueMax="0.78" valueMin="0.75">0.75–0.78 mm</quantity>
|
||
, width
|
||
<quantity id="8773EEE0FFD52469CB88A487745A23BA" box="[151,317,1483,1509]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="7.4" metricValueMin="7.3" pageId="30" pageNumber="31" unit="mm" value="0.735" valueMax="0.74" valueMin="0.73">0.73–0.74 mm</quantity>
|
||
. Tergite 1 (IV) (
|
||
<figureCitation id="D8B05F80FFD52469CAF0A487773423BA" box="[495,595,1483,1509]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11D</figureCitation>
|
||
) strongly modified, much longer than tergites 2–4 (V–VII) combined; with deep central cavity, inside cavity with ‘
|
||
<emphasis id="72FF9F17FFD52469C949A4A377152059" bold="true" box="[598,626,1519,1542]" pageId="30" pageNumber="31">±</emphasis>
|
||
’-shaped thick ridge, with broad, truncate projection posterior to cavity, areas lateral to projection markedly concave; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, lacking discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) slightly longer than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae in broad setose basal impression, with pair of short lateral carinae, with thin marginal carinae much longer than lateral ones; midlength of sternite 2 (IV) approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (
|
||
<figureCitation id="D8B05F80FFD52469C975A7A777AA215A" box="[618,717,1771,1797]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11E</figureCitation>
|
||
) with moderately sclerotized, rounded apex and few long setae along apical margin.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD52469CBD8A67F74662182" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
|
||
Aedeagus (
|
||
<figureCitation id="D8B05F80FFD52469CA5BA67F74A52112" box="[324,450,1843,1869]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="30" pageNumber="31">Fig. 11F, G</figureCitation>
|
||
)
|
||
<quantity id="8773EEE0FFD52469CACEA67E77532112" box="[465,564,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="30" pageNumber="31" unit="mm" value="0.51">0.51 mm</quantity>
|
||
long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk protruding and in dorsal view broad, apex narrowly and shortly split; dorsal lobe in lateral view elongate and bent at middle, in dorsal view strongly curved rightwards (position in figure) at apical 1/5, gradually narrowing apically and with pointed apex; parameres reduced to single semi-membranous structure.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD42468CBD8A1DB7763271E" blockId="31.[151,1437,151,537]" pageId="31" pageNumber="32">
|
||
<emphasis id="72FF9F17FFD42468CBD8A1DB747C26EE" box="[199,283,151,177]" italics="true" pageId="31" pageNumber="32">Female</emphasis>
|
||
. Similar to male in external morphology; antenna slightly shorter; legs and abdomen lacking modifications; each compound eye composed of approximately 19 ommatidia; humeral angle rounded. Measurements (as for male): body length
|
||
<quantity id="8773EEE0FFD42468CA6CA192777D26A6" box="[371,538,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.565" metricValueMax="2.57" metricValueMin="2.56" pageId="31" pageNumber="32" unit="mm" value="2.565" valueMax="2.57" valueMin="2.56">2.56–2.57 mm</quantity>
|
||
; length/width of head 0.51–0.52/
|
||
<quantity id="8773EEE0FFD42468C890A192715326A6" box="[911,1076,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.1" metricValueMin="5.0" pageId="31" pageNumber="32" unit="mm" value="0.505" valueMax="0.51" valueMin="0.5">0.50–0.51 mm</quantity>
|
||
, pronotum 0.55–0.58/
|
||
<quantity id="8773EEE0FFD42468CE32A19275A22742" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="31" pageNumber="32" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
|
||
, elytra 0.70–0.72/
|
||
<quantity id="8773EEE0FFD42468CA8CA04E775E2742" box="[403,569,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="8.7" metricValueMin="8.5" pageId="31" pageNumber="32" unit="mm" value="0.86" valueMax="0.87" valueMin="0.85">0.85–0.87 mm</quantity>
|
||
; abdomen 0.75–0.80/
|
||
<quantity id="8773EEE0FFD42468C833A04E76F32742" box="[812,916,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="mm" value="0.8">0.80 mm</quantity>
|
||
; length of antenna
|
||
<quantity id="8773EEE0FFD42468CF71A04F71B12742" box="[1134,1238,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="31" pageNumber="32" unit="mm" value="1.17">1.17 mm</quantity>
|
||
; maximum width of genitalia (
|
||
<figureCitation id="D8B05F80FFD42468CA3AA06B74EE271E" box="[293,393,295,321]" captionStart="FIGURE 11" captionStartId="31.[151,250,1823,1847]" captionTargetBox="[153,1436,562,1798]" captionTargetId="figure-159@31.[151,1436,561,1800]" captionTargetPageId="31" captionText="FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G." figureDoi="http://doi.org/10.5281/zenodo.6340756" httpUri="https://zenodo.org/record/6340756/files/figure.png" pageId="31" pageNumber="32">Fig. 11H</figureCitation>
|
||
)
|
||
<quantity id="8773EEE0FFD42468CA86A06B7767271E" box="[409,512,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="31" pageNumber="32" unit="mm" value="0.36">0.36 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="40344305FFD42468CBD8A00677F727F2" blockId="31.[151,1437,151,537]" pageId="31" pageNumber="32">
|
||
<emphasis id="72FF9F17FFD42468CBD8A00674C8273A" bold="true" box="[199,431,330,357]" pageId="31" pageNumber="32">Comparative notes.</emphasis>
|
||
The new species can be readily separated from all known congeners by the relatively large body size and a generally cylindrical habitus, combined with the unique sensory patch of the male metafemur and the structure of the abdominal modifications.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFD42468CBD8A0FA7103278E" box="[199,1124,438,466]" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="40344305FFD42468CBD8A0FA7103278E" blockId="31.[151,1437,151,537]" box="[199,1124,438,466]" pageId="31" pageNumber="32">
|
||
<emphasis id="72FF9F17FFD42468CBD8A0FA7406278F" bold="true" box="[199,353,438,464]" pageId="31" pageNumber="32">Distribution.</emphasis>
|
||
Chagyib District (Nyingchi),
|
||
<collectingRegion id="824F8DE7FFD42468C9AEA0FB7662278E" box="[689,773,439,465]" country="China" name="Xizang" pageId="31" pageNumber="32">Xizang</collectingRegion>
|
||
, SW
|
||
<collectingCountry id="389C0395FFD42468C85FA0FB76E4278E" box="[832,899,439,465]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
|
||
(
|
||
<figureCitation id="D8B05F80FFD42468C88DA0FB7699278E" box="[914,1022,439,465]" captionStart="FIGURE 78" captionStartId="180.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-19@180.[151,1437,181,1848]" captionTargetPageId="180" captionText="FIGURE 78. Distribution of Tibetan Batrisini. A. Arthromelodes cylindricus sp. nov. (circle), A. flosculus sp. nov. (triangle) and A. gyamda sp. nov. (square). B. A. intricatus sp. nov. (circle), A. lage sp. nov. (triangle) and A. langjicuo sp. nov. (square). C. A. latithorax sp. nov. (circle), A. lebus sp. nov. (triangle) and A. lotus sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340926" httpUri="https://zenodo.org/record/6340926/files/figure.png" pageId="31" pageNumber="32">Figs 78A</figureCitation>
|
||
,
|
||
<figureCitation id="D8B05F80FFD42468CF16A0FB7130278E" box="[1033,1111,439,465]" captionStart="FIGURE 96" captionStartId="198.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-19@198.[151,1436,181,1622]" captionTargetPageId="198" captionText="FIGURE 96. Environment of and collecting sites at Nyingchi area (ffż地区). A. Gongbo'gyamda (工布ȕḭ). B. Zhongzhe Village (ΦĖ村). C. Kading Valley (ϮŤ沟). D. Chongge Cuo (ṗẋś). E, F. A slope near Serjila mountain pass (DzĻŃ山 口). Photo Credit: A, B, E, F by Zhong Peng." figureDoi="http://doi.org/10.5281/zenodo.6340965" httpUri="https://zenodo.org/record/6340965/files/figure.png" pageId="31" pageNumber="32">96E, F</figureCitation>
|
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).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0891108EFFD42468CBD8A097773D2446" pageId="31" pageNumber="32" type="etymology">
|
||
<paragraph id="40344305FFD42468CBD8A097773D2446" blockId="31.[151,1437,151,537]" pageId="31" pageNumber="32">
|
||
<emphasis id="72FF9F17FFD42468CBD8A097742927AA" bold="true" box="[199,334,475,501]" pageId="31" pageNumber="32">Etymology.</emphasis>
|
||
The specific epithet ‘
|
||
<taxonomicName id="878B3886FFD42468C95DA09777D927AA" authorityName="Yin" authorityYear="2022" box="[578,702,475,501]" class="Insecta" family="Staphylinidae" genus="Arthromelodes" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cylindricus">
|
||
<emphasis id="72FF9F17FFD42468C95DA09777D927AA" box="[578,702,475,501]" italics="true" pageId="31" pageNumber="32">cylindricus</emphasis>
|
||
</taxonomicName>
|
||
(-
|
||
<emphasis id="72FF9F17FFD42468C9C6A097778027AB" box="[729,743,475,500]" italics="true" pageId="31" pageNumber="32">a</emphasis>
|
||
, -
|
||
<emphasis id="72FF9F17FFD42468C9E1A097764727AB" box="[766,800,475,500]" italics="true" pageId="31" pageNumber="32">um</emphasis>
|
||
)’ is a Latin adjective (of Ancient Greek origin) referring to the cylindrical habitus of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |