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<document id="14F9E8C7FF936245B05C24F3E0BAA5E7" ID-CLB-Dataset="113069" ID-DOI="10.11646/zootaxa.5111.1.1" ID-GBIF-Dataset="f3992bb2-9962-4f3c-8f5b-131cfe552c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6340729" ID-ZooBank="836B0F69-037C-4D0F-80DB-94FE454F48E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646828425946" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="C822F213FFA42406CB88A76A70FB271F" docLanguage="en" docName="zootaxa.5111.1.1.pdf" docOrigin="Zootaxa 5111 (1)" docSource="http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Sathytes Westwood 1870" docType="key" docVersion="9" lastPageNumber="114" masterDocId="341B8A6BFFCB2477CB1FA14C7567265F" masterDocTitle="The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earths “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="112" updateTime="1699317591893" updateUser="ExternalLinkService">
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<mods:title id="ED90291B387445715C40165CFDD9A7CC">The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earths “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<treatment id="C822F213FFA42406CB88A76A70FB271F" ID-DOI="http://doi.org/10.5281/zenodo.6964335" ID-GBIF-Taxon="198579332" ID-Zenodo-Dep="6964335" LSID="urn:lsid:plazi:treatment:C822F213FFA42406CB88A76A70FB271F" httpUri="http://treatment.plazi.org/id/C822F213FFA42406CB88A76A70FB271F" lastPageId="113" lastPageNumber="114" pageId="111" pageNumber="112">
<subSubSection id="0891108EFFA42418CB88A76A777B203D" pageId="111" pageNumber="112" type="nomenclature">
<paragraph id="40344305FFA42418CB88A76A74AA201F" blockId="111.[151,540,1574,1637]" box="[151,461,1574,1601]" pageId="111" pageNumber="112">
<heading id="1B7CF469FFA42418CB88A76A74AA201F" bold="true" box="[151,461,1574,1601]" fontSize="11" level="2" pageId="111" pageNumber="112" reason="4">
<taxonomicName id="878B3886FFA42418CB88A76A74AA201F" ID-CoL="8W3BW" authority="Westwood, 1870" authorityName="Westwood" authorityYear="1870" box="[151,461,1574,1601]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FFA42418CB88A76A74AA201F" bold="true" box="[151,461,1574,1601]" pageId="111" pageNumber="112">
<emphasis id="72FF9F17FFA42418CB88A76A759B201E" bold="true" box="[151,252,1574,1601]" italics="true" pageId="111" pageNumber="112">Sathytes</emphasis>
<bibRefCitation id="241A3EF4FFA42418CA1CA76A74AA201F" author="Westwood, J. O." box="[259,461,1574,1600]" pageId="111" pageNumber="112" pagination="125 - 132" refId="ref145492" refString="Westwood, J. O. (1870) Descriptions of twelve new exotic species of the Coleopterous family Pselaphidae. Transactions of the Royal Entomological Society of London, 1870, 125 - 132. https: // doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01868. x" type="journal article" year="1870">Westwood, 1870</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="40344305FFA42418CB88A706777B203D" blockId="111.[151,540,1574,1637]" box="[151,540,1610,1637]" pageId="111" pageNumber="112">
Chinese common name:
<emphasis id="72FF9F17FFA42418CAB2A707777B203D" bold="true" box="[429,540,1611,1634]" pageId="111" pageNumber="112">ḛà甲¤</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFFA42418CB88A7C37453215D" pageId="111" pageNumber="112" type="reference_group">
<paragraph id="40344305FFA42418CB88A7C3713D209A" blockId="111.[151,1436,1679,1795]" pageId="111" pageNumber="112">
<treatmentCitationGroup id="609B642BFFA42418CB88A7C3713D209A" pageId="111" pageNumber="112">
<treatmentCitation id="C12A6514FFA42418CB88A7C374A920F8" author="Westwood, J. O." box="[151,462,1679,1703]" page="128" pageId="111" pageNumber="112" year="1870">
<taxonomicName id="878B3886FFA42418CB88A7C374A920F8" ID-CoL="8W3BW" authority="Westwood, 1870: 128" authorityName="Westwood" authorityPageNumber="128" authorityYear="1870" box="[151,462,1679,1703]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FFA42418CB88A7C3758C20F8" box="[151,235,1679,1703]" italics="true" pageId="111" pageNumber="112">Sathytes</emphasis>
<bibRefCitation id="241A3EF4FFA42418CBEFA7C374A920F8" author="Westwood, J. O." box="[240,462,1679,1703]" pageId="111" pageNumber="112" pagination="125 - 132" refId="ref145492" refString="Westwood, J. O. (1870) Descriptions of twelve new exotic species of the Coleopterous family Pselaphidae. Transactions of the Royal Entomological Society of London, 1870, 125 - 132. https: // doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01868. x" type="journal article" year="1870">Westwood, 1870: 128</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<bibRefCitation id="241A3EF4FFA42418CAC8A7C3775820F8" author="Lobl I." box="[471,575,1679,1703]" pageId="111" pageNumber="112" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl 1979</bibRefCitation>
(revision of world species);
<bibRefCitation id="241A3EF4FFA42418C845A7C3769E20F8" author="Yin, Z. - W. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[858,1017,1679,1703]" pageId="111" pageNumber="112" pagination="831 - 857" refId="ref146753" refString="Yin, Z. - W., Li, L. - Z. &amp; Zhao, M. - J. (2012 a) Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History, 46 (13 - 14), 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651" type="journal article" year="2012">
Yin
<emphasis id="72FF9F17FFA42418C89BA7C376D720F8" box="[900,944,1679,1703]" italics="true" pageId="111" pageNumber="112">et al</emphasis>
. 2012a
</bibRefCitation>
(redescription);
<bibRefCitation id="241A3EF4FFA42418CF81A7C3702B20F8" author="Shen, Q. &amp; Yin, Z. - W." box="[1182,1356,1679,1703]" pageId="111" pageNumber="112" pagination="386 - 394" refId="ref145156" refString="Shen, Q. &amp; Yin, Z. - W. (2020) Five new species and a new record of Sathytes Westwood from eastern China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4751 (2), 386 - 394. https: // doi. org / 10.11646 / zootaxa. 4751.2.13" type="journal article" year="2020">Yin &amp; Shen 2020</bibRefCitation>
(checklist of world species).
<typeStatus id="9F30FDA7FFA42418CAB8A7E274BE2099" box="[423,473,1710,1734]" pageId="111" pageNumber="112">Type</typeStatus>
species:
<treatmentCitation id="C12A6514FFA42418C929A7E176B2209A" author="Westwood, J. O." box="[566,981,1709,1733]" page="128" pageId="111" pageNumber="112" year="1870">
<taxonomicName id="878B3886FFA42418C929A7E176B2209A" ID-CoL="6XQPT" authority="Westwood, 1870: 128" authorityName="Westwood" authorityPageNumber="128" authorityYear="1870" box="[566,981,1709,1733]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="punctiger">
<emphasis id="72FF9F17FFA42418C929A7E17795209A" box="[566,754,1709,1733]" italics="true" pageId="111" pageNumber="112">Sathytes punctiger</emphasis>
<bibRefCitation id="241A3EF4FFA42418C9E7A7E176B2209A" author="Westwood, J. O." box="[760,981,1709,1733]" pageId="111" pageNumber="112" pagination="125 - 132" refId="ref145492" refString="Westwood, J. O. (1870) Descriptions of twelve new exotic species of the Coleopterous family Pselaphidae. Transactions of the Royal Entomological Society of London, 1870, 125 - 132. https: // doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01868. x" type="journal article" year="1870">Westwood, 1870: 128</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(monotypy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="40344305FFA42418CB88A7807453215D" blockId="111.[151,1436,1679,1795]" pageId="111" pageNumber="112">
=
<emphasis id="72FF9F17FFA42418CBB3A780747D20BB" box="[172,282,1740,1764]" italics="true" pageId="111" pageNumber="112">Batoxylina</emphasis>
<bibRefCitation id="241A3EF4FFA42418CA3FA78074A020BB" author="Jeannel, R." box="[288,455,1740,1764]" pageId="111" pageNumber="112" pagination="5 - 32" refId="ref143142" refString="Jeannel, R. (1957) Sur quelques Pselaphides du Tonkin recueillis par le Pere A. de Cooman. Revue Francaise d'Entomologie, 24 (1), 5 - 32." type="journal article" year="1957">Jeannel, 1957: 8</bibRefCitation>
.
<typeStatus id="9F30FDA7FFA42418CACEA780776420BB" box="[465,515,1740,1764]" pageId="111" pageNumber="112">Type</typeStatus>
species:
<taxonomicName id="878B3886FFA42418C97FA78076A920BB" ID-CoL="KZ5R" authority="Jeannel, 1957: 9" authorityName="Jeannel" authorityPageNumber="9" authorityYear="1957" box="[608,974,1740,1764]" class="Insecta" family="Staphylinidae" genus="Batoxylina" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="clavalis">
<emphasis id="72FF9F17FFA42418C97FA780764420BB" box="[608,803,1740,1764]" italics="true" pageId="111" pageNumber="112">Batoxylina clavalis</emphasis>
<bibRefCitation id="241A3EF4FFA42418C836A78076A920BB" author="Jeannel, R." box="[809,974,1740,1764]" pageId="111" pageNumber="112" pagination="5 - 32" refId="ref143142" refString="Jeannel, R. (1957) Sur quelques Pselaphides du Tonkin recueillis par le Pere A. de Cooman. Revue Francaise d'Entomologie, 24 (1), 5 - 32." type="journal article" year="1957">Jeannel, 1957: 9</bibRefCitation>
</taxonomicName>
(original designation). Synonymized by
<bibRefCitation id="241A3EF4FFA42418CE73A7807457215D" author="Lobl I." pageId="111" pageNumber="112" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl 1979: 285</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFFA42407CB88A67D76CA273A" lastPageId="112" lastPageNumber="113" pageId="111" pageNumber="112" type="description">
<paragraph id="40344305FFA42407CB88A67D76DF26A6" blockId="111.[151,1437,1841,2011]" lastBlockId="112.[151,1436,151,357]" lastPageId="112" lastPageNumber="113" pageId="111" pageNumber="112">
This is a rather easily recognizable genus with all included members possessing a coarsely punctate dorsal surface of the body, and a great number of foveae at the elytral bases and the bases of the abdominal segments. The males of all species, except
<taxonomicName id="878B3886FFA42418CA5AA635749221CC" authority="Lobl" authorityName="Lobl" authorityYear="1979" box="[325,501,1913,1939]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FFA42418CA5AA635743421CC" box="[325,339,1913,1939]" italics="true" pageId="111" pageNumber="112">S</emphasis>
.
<emphasis id="72FF9F17FFA42418CA7EA63574D021CC" box="[353,439,1913,1939]" italics="true" pageId="111" pageNumber="112">simplex</emphasis>
Löbl
</taxonomicName>
, possess strongly modified and enlarged antennal clubs of characteristic forms, thus making identifications less difficult in most cases with enough careful comparisons. In
<collectingRegion id="824F8DE7FFA42418CF71A6D171CC21E8" box="[1134,1195,1949,1975]" country="China" name="Xizang" pageId="111" pageNumber="112">Tibet</collectingRegion>
, it is common to find several species at the same location, making it sometimes impossible to associate females with males of species that have a similar body size with 100% assurance.
<taxonomicName id="878B3886FFBB2407C9BCA1DB766726EE" authorityName="Westwood" authorityYear="1870" box="[675,768,151,177]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FFBB2407C9BCA1DB766726EE" box="[675,768,151,177]" italics="true" pageId="112" pageNumber="113">Sathytes</emphasis>
</taxonomicName>
is diverse in
<collectingCountry id="389C0395FFBB2407C88CA1DB76BE26EE" box="[915,985,151,177]" name="China" pageId="112" pageNumber="113">China</collectingCountry>
, and seven species have been previously reported from
<collectingRegion id="824F8DE7FFBB2407CA28A1F77416268A" box="[311,369,187,213]" country="China" name="Xizang" pageId="112" pageNumber="113">Tibet</collectingRegion>
(
<bibRefCitation id="241A3EF4FFBB2407CA61A1F77754268A" author="Yin, Z. - W. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[382,563,187,213]" pageId="112" pageNumber="113" pagination="831 - 857" refId="ref146753" refString="Yin, Z. - W., Li, L. - Z. &amp; Zhao, M. - J. (2012 a) Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History, 46 (13 - 14), 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651" type="journal article" year="2012">
Yin
<emphasis id="72FF9F17FFBB2407CAB0A1F774B8268A" box="[431,479,187,213]" italics="true" pageId="112" pageNumber="113">et al</emphasis>
. 2012a
</bibRefCitation>
;
<bibRefCitation id="241A3EF4FFBB2407C921A1F77665268A" author="Shen, Q. &amp; Yin, Z. - W." box="[574,770,187,213]" pageId="112" pageNumber="113" pagination="386 - 394" refId="ref145156" refString="Shen, Q. &amp; Yin, Z. - W. (2020) Five new species and a new record of Sathytes Westwood from eastern China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4751 (2), 386 - 394. https: // doi. org / 10.11646 / zootaxa. 4751.2.13" type="journal article" year="2020">Shen &amp; Yin 2020</bibRefCitation>
;
<bibRefCitation id="241A3EF4FFBB2407C814A1F776A8268A" author="Shen, Q. &amp; Yin, Z. - W." box="[779,975,187,213]" pageId="112" pageNumber="113" pagination="386 - 394" refId="ref145156" refString="Shen, Q. &amp; Yin, Z. - W. (2020) Five new species and a new record of Sathytes Westwood from eastern China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4751 (2), 386 - 394. https: // doi. org / 10.11646 / zootaxa. 4751.2.13" type="journal article" year="2020">Yin &amp; Shen 2020</bibRefCitation>
), with another 12 species being described here. Additionally,
<taxonomicName id="878B3886FFBB2407CA71A193774F26A6" authorityName="Lobl" authorityYear="1979" box="[366,552,223,249]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FFBB2407CA71A193774F26A6" box="[366,552,223,249]" italics="true" pageId="112" pageNumber="113">Sathytes simplex</emphasis>
</taxonomicName>
is newly reported to occur in
<collectingRegion id="824F8DE7FFBB2407C869A19376D326A6" box="[886,948,223,249]" country="China" name="Xizang" pageId="112" pageNumber="113">Tibet</collectingRegion>
.
</paragraph>
<paragraph id="40344305FFBB2407CBD8A04F76CA273A" blockId="112.[151,1436,151,357]" pageId="112" pageNumber="113">The 20 Tibetan species are keyed below. It should be kept in mind that in some cases a direct and careful comparison between the specimens is the only means for separating morphologically similar species due to often different angles showing the antennal modifications in published literatures.</paragraph>
</subSubSection>
<subSubSection id="0891108EFFBB2406CB88A0FB70FB271F" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113" type="key">
<paragraph id="40344305FFBB2407CB88A0FB77F5278F" blockId="112.[151,658,438,465]" box="[151,658,438,465]" pageId="112" pageNumber="113">
<emphasis id="72FF9F17FFBB2407CB88A0FB77F5278F" bold="true" box="[151,658,438,465]" pageId="112" pageNumber="113">
Key to Tibetan species of
<taxonomicName id="878B3886FFBB2407CAC9A0FA775C278E" ID-CoL="8W3BW" authorityName="Westwood" authorityYear="1870" box="[470,571,438,465]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FFBB2407CAC9A0FA775C278E" bold="true" box="[470,571,438,465]" italics="true" pageId="112" pageNumber="113">Sathytes</emphasis>
</taxonomicName>
(male)
</emphasis>
</paragraph>
<key id="341A1434FFBB2406CB88A0B070FB271F" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113">
<keyStep id="FB7F5BA0FFBB2407CB88A0B070FB2471" pageId="112" pageNumber="113">
<paragraph id="40344305FFBB2407CB88A0B070FB244D" blockId="112.[151,1437,507,2026]" box="[151,1436,507,530]" pageId="112" pageNumber="113">
<keyLead id="FB7AE030FFBB2407CB88A0B070FB244D" box="[151,1436,507,530]" pageId="112" pageNumber="113">
1 Antenna simple, lacking modifications (
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)...............................................
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- Antenna with antennomeres 9 and 11 modified (
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Morphology of Sathytes aequalis sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." captionText-1="FIGURE 50. Morphology of Sathytes asura sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in DG." captionText-2="FIGURE 51. Morphology of Sathytes caichenyangi sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." captionText-3="FIGURE 52. Morphology of Sathytes dawai sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." captionText-4="FIGURE 53. Morphology of Sathytes duojii sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DF." captionText-5="FIGURE 54. Morphology of Sathytes laticornis sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in DG." captionText-6="FIGURE 55. Morphology of Sathytes pengzhongi sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.05 mm in D, 0.1 mm in EG." captionText-7="FIGURE 56. Morphology of Sathytes proclivis sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." captionText-8="FIGURE 57. Morphology of Sathytes pseudograndis sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." figureDoi-0="http://doi.org/10.5281/zenodo.6343319" figureDoi-1="http://doi.org/10.5281/zenodo.6343325" figureDoi-2="http://doi.org/10.5281/zenodo.6343333" figureDoi-3="http://doi.org/10.5281/zenodo.6343339" figureDoi-4="http://doi.org/10.5281/zenodo.6343343" figureDoi-5="http://doi.org/10.5281/zenodo.6343350" figureDoi-6="http://doi.org/10.5281/zenodo.6343358" figureDoi-7="http://doi.org/10.5281/zenodo.6343362" figureDoi-8="http://doi.org/10.5281/zenodo.6340853" httpUri-0="https://zenodo.org/record/6343319/files/figure.png" httpUri-1="https://zenodo.org/record/6343325/files/figure.png" httpUri-2="https://zenodo.org/record/6343333/files/figure.png" httpUri-3="https://zenodo.org/record/6343339/files/figure.png" httpUri-4="https://zenodo.org/record/6343343/files/figure.png" httpUri-5="https://zenodo.org/record/6343350/files/figure.png" httpUri-6="https://zenodo.org/record/6343358/files/figure.png" httpUri-7="https://zenodo.org/record/6343362/files/figure.png" httpUri-8="https://zenodo.org/record/6340853/files/figure.png" pageId="112" pageNumber="113">Figs 4957</figureCitation>
,
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)............................................. 2
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2 Antennomere 11 markedly elongate, ratio (length/width) greater than 1.85 (
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;
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: fig. 3B)..... 3
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3 Body length less than
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); ratio (length/width) of antennomere 11 less than 1.9 (1.89) (
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); aedeagus expanded apically and with broadly emarginate apical margin (
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)..........................
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or more; ratio (length/width) of antennomere 11 greater than 2.0; aedeagus apically as broad as basal half................................................................................................ 4
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4 Setae of head relatively much longer and sparser (
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); distributed in Cona.....................................................................
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- Setae of head relatively much shorter and denser (
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: fig. 3A); pronotum less elongate, ratio (length/width) 80/78 (
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: fig. 3A); distributed in Zayü..........................................
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5 Antennomere 11 with two basal protuberances (
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: fig. 6B, D)..................
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<emphasis id="72FF9F17FFBB2407CFA4A2177043252E" bold="true" box="[1211,1316,859,881]" italics="true" pageId="112" pageNumber="113">linzhiensis</emphasis>
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Yin &amp; Shen
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<keyLead id="FB7AE030FFBB2407CB88A23570FB25D0" box="[151,1436,889,911]" pageId="112" pageNumber="113">- Antennomere 11 with single basal protuberance............................................................. 6</keyLead>
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6 Hyaline apophysis located on apical 1/2 of antennomere 9 (
<figureCitation id="D8B05F80FFBB2407C803A2DA761025F3" box="[796,887,918,940]" captionStart="FIGURE 55" captionStartId="127.[151,250,1612,1636]" captionTargetBox="[153,1434,417,1588]" captionTargetId="figure-103@127.[151,1436,416,1588]" captionTargetPageId="127" captionText="FIGURE 55. Morphology of Sathytes pengzhongi sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.05 mm in D, 0.1 mm in EG." figureDoi="http://doi.org/10.5281/zenodo.6343358" httpUri="https://zenodo.org/record/6343358/files/figure.png" pageId="112" pageNumber="113">Figs 55D</figureCitation>
,
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;
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: figs 10C, 13C)........ 7
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- Hyaline apophysis located at middle or on basal 1/2 of antennomere 9 (e.g., D in
<figureCitation id="D8B05F80FFBB2407C8F1A2F8713B2596" box="[1006,1116,947,970]" captionStart-0="FIGURE 50" captionStart-1="FIGURE 51" captionStart-2="FIGURE 52" captionStart-3="FIGURE 53" captionStart-4="FIGURE 54" captionStartId-0="116.[151,250,1851,1875]" captionStartId-1="118.[151,250,1891,1915]" captionStartId-2="121.[151,250,1730,1754]" captionStartId-3="123.[151,250,1544,1568]" captionStartId-4="125.[151,250,1525,1549]" captionTargetBox-0="[153,1435,597,1827]" captionTargetBox-1="[153,1436,561,1867]" captionTargetBox-2="[154,1435,417,1705]" captionTargetBox-3="[152,1435,309,1521]" captionTargetBox-4="[153,1435,310,1502]" captionTargetId-0="figure-192@116.[151,1436,597,1828]" captionTargetId-1="figure-184@118.[151,1436,561,1868]" captionTargetId-2="figure-100@121.[151,1436,417,1706]" captionTargetId-3="figure-75@123.[151,1436,309,1521]" captionTargetId-4="figure-60@125.[151,1435,309,1502]" captionTargetPageId-0="116" captionTargetPageId-1="118" captionTargetPageId-2="121" captionTargetPageId-3="123" captionTargetPageId-4="125" captionText-0="FIGURE 50. Morphology of Sathytes asura sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in DG." captionText-1="FIGURE 51. Morphology of Sathytes caichenyangi sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." captionText-2="FIGURE 52. Morphology of Sathytes dawai sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." captionText-3="FIGURE 53. Morphology of Sathytes duojii sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DF." captionText-4="FIGURE 54. Morphology of Sathytes laticornis sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in DG." figureDoi-0="http://doi.org/10.5281/zenodo.6343325" figureDoi-1="http://doi.org/10.5281/zenodo.6343333" figureDoi-2="http://doi.org/10.5281/zenodo.6343339" figureDoi-3="http://doi.org/10.5281/zenodo.6343343" figureDoi-4="http://doi.org/10.5281/zenodo.6343350" httpUri-0="https://zenodo.org/record/6343325/files/figure.png" httpUri-1="https://zenodo.org/record/6343333/files/figure.png" httpUri-2="https://zenodo.org/record/6343339/files/figure.png" httpUri-3="https://zenodo.org/record/6343343/files/figure.png" httpUri-4="https://zenodo.org/record/6343350/files/figure.png" pageId="112" pageNumber="113">Figs 5054</figureCitation>
,
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,
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).................. 11
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7 Body length less than 2.0 mm (
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); antennomere 10 of unique form (
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)..........
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<emphasis id="72FF9F17FFBB2407CFC8A29C702D25B9" bold="true" box="[1239,1354,976,998]" italics="true" pageId="112" pageNumber="113">pengzhongi</emphasis>
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8 Body length greater than
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(
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); antennomere 9 strongly expanded mesally (
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: fig. 13C); basal protuberance of antennomere 11 conspicuously extended posteriorly to reach antennomere 10 (
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: fig. 13B)............................................................................
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<emphasis id="72FF9F17FFBB2407CFABA50B70422203" bold="true" box="[1204,1317,1095,1116]" italics="true" pageId="112" pageNumber="113">transversus</emphasis>
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Yin &amp; Shen
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- Body length less than
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; antennomere 9 moderately expanded mesally or obliquely expanded at anteromesal portion; basal protuberance of antennomere 11 not short and not reach antennomere 10… 9
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9 Anterior margin of antennomere 9 distinctly oblique (
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); antennomere 11 with relatively shorter basal protuberance, apical margin subtruncate (
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).......................................................
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<emphasis id="72FF9F17FFBB2407CFE9A5F7702D228E" bold="true" box="[1270,1354,1211,1233]" italics="true" pageId="112" pageNumber="113">proclivis</emphasis>
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<keyLead id="FB7AE030FFBB2407CB88A59570FB2354" pageId="112" pageNumber="113">- Anterior margin of antennomere 9 even; antennomere 11 with relatively longer basal protuberance and angulate apical margin .................................................................................................. 10</keyLead>
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10 Antennomere 11 relatively longer, ratio (length/width) 1.72 (
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); hyaline apophysis of antennomere 9 more anteriorly positioned (
<figureCitation id="D8B05F80FFBB2407CA4CA47D74CC2318" box="[339,427,1329,1351]" captionStart="FIGURE 61" captionStartId="141.[151,250,1504,1528]" captionTargetBox="[154,1436,345,1480]" captionTargetId="figure-87@141.[151,1436,345,1481]" captionTargetPageId="141" captionText="FIGURE 61. Morphology of Sathytes yigongensis sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in DG." figureDoi="http://doi.org/10.5281/zenodo.6340876" httpUri="https://zenodo.org/record/6340876/files/figure.png" pageId="112" pageNumber="113">Fig. 61D</figureCitation>
).................................................................
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<emphasis id="72FF9F17FFBB2407CFCAA47D702E2319" bold="true" box="[1237,1353,1329,1350]" italics="true" pageId="112" pageNumber="113">yigongensis</emphasis>
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- Antennomere 11 relatively shorter, ratio (length/width) 1.6 (
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: fig. 10B); hyaline apophysis of antennomere 9 more posteriorly positioned (
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: fig. 10C).................................
<taxonomicName id="878B3886FFBB2407CF90A42770FB23DD" ID-CoL="8Y99K" authority="Yin &amp; Shen" authorityName="Yin &amp; Shen" authorityYear="2020" box="[1167,1436,1387,1410]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="panzhaohuii">
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<emphasis id="72FF9F17FFBB2407CF90A42771FB23DF" bold="true" box="[1167,1180,1387,1408]" italics="true" pageId="112" pageNumber="113">S</emphasis>
.
<emphasis id="72FF9F17FFBB2407CFB6A427704223DE" bold="true" box="[1193,1317,1387,1409]" italics="true" pageId="112" pageNumber="113">panzhaohuii</emphasis>
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Yin &amp; Shen
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<keyLead id="FB7AE030FFBB2407CB88A4C570FB23C1" box="[151,1436,1417,1439]" pageId="112" pageNumber="113">11 Body length greater than 3.0 mm........................................................................ 12</keyLead>
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<keyLead id="FB7AE030FFBB2407CB88A4EB70FB23E4" box="[151,1436,1446,1468]" pageId="112" pageNumber="113">- Body length less than 3.0 mm.......................................................................... 14</keyLead>
</paragraph>
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<paragraph id="40344305FFBB2407CB88A48870FB23A8" blockId="112.[151,1437,507,2026]" pageId="112" pageNumber="113">
<keyLead id="FB7AE030FFBB2407CB88A48870FB23A8" pageId="112" pageNumber="113">
12 Antennomere 9 strongly inclined at mesal margin (
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Yin
<emphasis id="72FF9F17FFBB2407C9EBA48876792386" box="[756,798,1476,1497]" italics="true" pageId="112" pageNumber="113">et al</emphasis>
. 2012a
</bibRefCitation>
: fig. 7F); aedeagus (
<bibRefCitation id="241A3EF4FFBB2407CF35A48871AF2386" author="Yin, Z. - W. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[1066,1224,1476,1498]" pageId="112" pageNumber="113" pagination="831 - 857" refId="ref146753" refString="Yin, Z. - W., Li, L. - Z. &amp; Zhao, M. - J. (2012 a) Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History, 46 (13 - 14), 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651" type="journal article" year="2012">Yin et al. 2012a</bibRefCitation>
: fig. 8F)...............................................................................................
<taxonomicName id="878B3886FFBB2407CFC8A4AD70FB23A8" ID-CoL="6Y2Q4" authority="Yin &amp; Li" authorityName="Yin &amp; Li" authorityYear="2012" box="[1239,1436,1505,1527]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="72FF9F17FFBB2407CFC8A4AD702723A9" bold="true" box="[1239,1344,1505,1526]" pageId="112" pageNumber="113">
<emphasis id="72FF9F17FFBB2407CFC8A4AD718323A9" bold="true" box="[1239,1252,1505,1526]" italics="true" pageId="112" pageNumber="113">S</emphasis>
.
<emphasis id="72FF9F17FFBB2407CFEEA4AD702723A9" bold="true" box="[1265,1344,1505,1526]" italics="true" pageId="112" pageNumber="113">magnus</emphasis>
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Yin &amp; Li
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<keyLead id="FB7AE030FFBB2407CB88A4B370FB204C" box="[151,1436,1534,1556]" pageId="112" pageNumber="113">- Antennomere 9 oblique, but not as inclined as above; aedeagus of a different form................................. 13</keyLead>
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<keyStep id="FB7F5BA0FFBB2407CB88A75070FB20D6" pageId="112" pageNumber="113">
<paragraph id="40344305FFBB2407CB88A75070FB2011" blockId="112.[151,1437,507,2026]" pageId="112" pageNumber="113">
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13 Body length approximately
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; tempus relatively longer, length of eye/tempus 0.78 (
<figureCitation id="D8B05F80FFBB2407CF2BA75071EC206E" box="[1076,1163,1563,1586]" captionStart="FIGURE 51" captionStartId="118.[151,250,1891,1915]" captionTargetBox="[153,1436,561,1867]" captionTargetId="figure-184@118.[151,1436,561,1868]" captionTargetPageId="118" captionText="FIGURE 51. Morphology of Sathytes caichenyangi sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." figureDoi="http://doi.org/10.5281/zenodo.6343333" httpUri="https://zenodo.org/record/6343333/files/figure.png" pageId="112" pageNumber="113">Fig. 51B</figureCitation>
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).................................................................
<emphasis id="72FF9F17FFBB2407CFB3A77570FB2011" bold="true" box="[1196,1436,1592,1614]" pageId="112" pageNumber="113">
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<emphasis id="72FF9F17FFBB2407CFD9A774702D2011" bold="true" box="[1222,1354,1592,1614]" italics="true" pageId="112" pageNumber="113">caichenyangi</emphasis>
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<taxonomicNameLabel id="69CC226CFFBB2407CE4FA77570FB2011" box="[1360,1436,1593,1614]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="40344305FFBB2407CB88A71B70FB20D6" blockId="112.[151,1437,507,2026]" pageId="112" pageNumber="113">
<keyLead id="FB7AE030FFBB2407CB88A71B70FB20D6" pageId="112" pageNumber="113">
- Body length approximately
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; tempus relatively shorter, length of eye/tempus 0.81 (
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)................................................................
<emphasis id="72FF9F17FFBB2407CFE8A73F70FB20D6" bold="true" box="[1271,1436,1651,1673]" pageId="112" pageNumber="113">
<taxonomicName id="878B3886FFBB2407CFE8A73F702E20D6" ID-CoL="9GDGK" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[1271,1353,1651,1673]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="duojii" status="sp. nov.">
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<emphasis id="72FF9F17FFBB2407CE0FA73F702E20D6" bold="true" box="[1296,1353,1651,1673]" italics="true" pageId="112" pageNumber="113">duojii</emphasis>
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14 Body length
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or slightly less (
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)..............................................
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.
<emphasis id="72FF9F17FFBB2407CFF3A7DC702E20F9" bold="true" box="[1260,1353,1680,1702]" italics="true" pageId="112" pageNumber="113">laticornis</emphasis>
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- Body length greater than
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........................................................................ 15
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<keyStep id="FB7F5BA0FFBB2407CB88A78770FB20A0" pageId="112" pageNumber="113">
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15 Antennomere 9 relatively more elongate (
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), ratio (length/width) 1.181.23....................
<emphasis id="72FF9F17FFBB2407CFEAA78770FB20BE" bold="true" box="[1269,1436,1739,1761]" pageId="112" pageNumber="113">
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.
<emphasis id="72FF9F17FFBB2407CE10A787702D20BE" bold="true" box="[1295,1354,1739,1761]" italics="true" pageId="112" pageNumber="113">dawai</emphasis>
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16 Hyaline apophysis located at approximately middle of antennomere 9 (
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: fig. 16C)....................................................................................................
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.
<emphasis id="72FF9F17FFBB2407CFADA66F70422167" bold="true" box="[1202,1317,1827,1848]" italics="true" pageId="112" pageNumber="113">xizangensis</emphasis>
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Yin &amp; Shen
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<keyLead id="FB7AE030FFBB2407CB88A60D70FB2109" box="[151,1436,1857,1879]" pageId="112" pageNumber="113">- Hyaline apophysis located on basal 1/2 of antennomere 9..................................................... 17</keyLead>
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<paragraph id="40344305FFBB2407CB88A61270FB212C" blockId="112.[151,1437,507,2026]" box="[151,1436,1886,1908]" pageId="112" pageNumber="113">
<keyLead id="FB7AE030FFBB2407CB88A61270FB212C" box="[151,1436,1886,1908]" pageId="112" pageNumber="113">
17 Aedeagus broadened apically (
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)...............................................
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<emphasis id="72FF9F17FFBB2407CFBFA61271CA212C" bold="true" box="[1184,1197,1886,1907]" italics="true" pageId="112" pageNumber="113">S</emphasis>
.
<emphasis id="72FF9F17FFBB2407CFA6A612702E212B" bold="true" box="[1209,1353,1886,1908]" italics="true" pageId="112" pageNumber="113">pseudograndis</emphasis>
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<taxonomicNameLabel id="69CC226CFFBB2407CE4FA61270FB212C" box="[1360,1436,1886,1907]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="40344305FFBB2407CB88A63070FB21CE" blockId="112.[151,1437,507,2026]" box="[151,1436,1916,1938]" pageId="112" pageNumber="113">
<keyLead id="FB7AE030FFBB2407CB88A63070FB21CE" box="[151,1436,1916,1938]" pageId="112" pageNumber="113">- Apical part of aedeagus as broad as basal half.............................................................. 18</keyLead>
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<paragraph id="40344305FFBB2407CB88A6D570FB2194" blockId="112.[151,1437,507,2026]" pageId="112" pageNumber="113">
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18 Hyaline apophysis of antennomere 9 relatively longer (
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); basal protuberance of antennomere 11 relatively short (
<figureCitation id="D8B05F80FFBB2407CE6AA6D5746A2193" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in DG." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="112" pageNumber="113">Fig. 59C</figureCitation>
)..............................................................................
<emphasis id="72FF9F17FFBB2407CFA6A6FA70FB2194" bold="true" box="[1209,1436,1974,1995]" pageId="112" pageNumber="113">
<taxonomicName id="878B3886FFBB2407CFA6A6FA702D2194" ID-CoL="9GDH7" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[1209,1354,1974,1995]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="sympatricus" status="sp. nov.">
<emphasis id="72FF9F17FFBB2407CFA6A6FA71A12194" bold="true" box="[1209,1222,1974,1995]" italics="true" pageId="112" pageNumber="113">S</emphasis>
.
<emphasis id="72FF9F17FFBB2407CFCCA6FA702D2194" bold="true" box="[1235,1354,1974,1995]" italics="true" pageId="112" pageNumber="113">sympatricus</emphasis>
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<paragraph id="40344305FFBB2407CB88A69870FB21B6" blockId="112.[151,1437,507,2026]" box="[151,1436,2004,2026]" pageId="112" pageNumber="113">
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19 Body smaller in size (length
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or slightly more), antenna shorter (
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), antennomeres 9 slightly more elongate (
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) and antennomere 11 stouter (
<figureCitation id="D8B05F80FFBA2406C97CA1F977DC2694" box="[611,699,181,203]" captionStart="FIGURE 50" captionStartId="116.[151,250,1851,1875]" captionTargetBox="[153,1435,597,1827]" captionTargetId="figure-192@116.[151,1436,597,1828]" captionTargetPageId="116" captionText="FIGURE 50. Morphology of Sathytes asura sp. nov. (AF. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 911. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in DG." figureDoi="http://doi.org/10.5281/zenodo.6343325" httpUri="https://zenodo.org/record/6343325/files/figure.png" pageId="113" pageNumber="114">Fig. 50C</figureCitation>
), apical margin of aedeagus not expanded on left side (
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)................................................................................................
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- Body larger in size (length
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or slightly more), antenna longer (
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), antennomeres 9 slightly stouter (
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: fig. 1C) and antennomere 11 more elongate (
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: fig. 1B), apical margin of aedeagus expanded on left side (
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: fig. 1E).............................................................
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.
<emphasis id="72FF9F17FFBA2406CFC8A0667041271F" bold="true" box="[1239,1318,298,320]" italics="true" pageId="113" pageNumber="114">alpicola</emphasis>
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Yin &amp; Shen
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