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<document id="8D0B5F79043DF306B454D25150206728" ID-CLB-Dataset="113069" ID-DOI="10.11646/zootaxa.5111.1.1" ID-GBIF-Dataset="f3992bb2-9962-4f3c-8f5b-131cfe552c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6340729" ID-ZooBank="836B0F69-037C-4D0F-80DB-94FE454F48E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646828425946" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="C822F213FF4E24F0CB88A1DB76C326A6" docLanguage="en" docName="zootaxa.5111.1.1.pdf" docOrigin="Zootaxa 5111 (1)" docSource="http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Sathytes simplex Lobl 1979" docType="treatment" docVersion="8" lastPageNumber="135" masterDocId="341B8A6BFFCB2477CB1FA14C7567265F" masterDocTitle="The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="134" updateTime="1699317591893" updateUser="ExternalLinkService">
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<mods:title id="F58327FAF7F522D49CDD94B56183DE73">The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="F059E3E0FF1E63AC5B466CB8074528B1">Yin, Zi-Wei</mods:namePart>
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<mods:date id="2CDA690E296C5DC2C2CE2108B9D1FF36">2022</mods:date>
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<mods:number id="03869003F7A00B72DF1CB51A03849333">2022-03-09</mods:number>
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<mods:number id="482327133127388994E04C922C8C388F">5111</mods:number>
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<mods:classification id="2159C71FCA669121BC681B008922A198">journal article</mods:classification>
<mods:identifier id="BF7495EC49A6514DC73599DB0466E2A2" type="CLB-Dataset">113069</mods:identifier>
<mods:identifier id="39E5FC95AEECA0E04019F4CFF38D7BFE" type="DOI">10.11646/zootaxa.5111.1.1</mods:identifier>
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<treatment id="C822F213FF4E24F0CB88A1DB76C326A6" ID-DOI="http://doi.org/10.5281/zenodo.6964384" ID-GBIF-Taxon="198579362" ID-Zenodo-Dep="6964384" LSID="urn:lsid:plazi:treatment:C822F213FF4E24F0CB88A1DB76C326A6" httpUri="http://treatment.plazi.org/id/C822F213FF4E24F0CB88A1DB76C326A6" lastPageId="135" lastPageNumber="135" pageId="133" pageNumber="134">
<subSubSection id="0891108EFF4E24F2CB88A1DB748B26EE" box="[151,492,151,178]" pageId="133" pageNumber="134" type="nomenclature">
<paragraph id="40344305FF4E24F2CB88A1DB748B26EE" blockId="133.[151,569,151,250]" box="[151,492,151,178]" pageId="133" pageNumber="134">
<heading id="1B7CF469FF4E24F2CB88A1DB748B26EE" bold="true" box="[151,492,151,178]" fontSize="11" level="2" pageId="133" pageNumber="134" reason="4">
<taxonomicName id="878B3886FF4E24F2CB88A1DB748B26EE" ID-CoL="79QPT" authority="Lobl, 1979" authorityName="Lobl" authorityYear="1979" box="[151,492,151,178]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FF4E24F2CB88A1DB748B26EE" bold="true" box="[151,492,151,178]" pageId="133" pageNumber="134">
<emphasis id="72FF9F17FF4E24F2CB88A1DB740626ED" bold="true" box="[151,353,151,178]" italics="true" pageId="133" pageNumber="134">Sathytes simplex</emphasis>
<bibRefCitation id="241A3EF4FF4E24F2CA77A1DB748B26EE" author="Lobl I." box="[360,492,151,177]" pageId="133" pageNumber="134" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl, 1979</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4E24F2CB88A1F7775E26A8" pageId="133" pageNumber="134" type="description">
<paragraph id="40344305FF4E24F2CB88A1F774FE268A" blockId="133.[151,569,151,250]" box="[151,409,187,214]" pageId="133" pageNumber="134">
(
<figureCitation id="D8B05F80FF4E24F2CB80A1F775912689" box="[159,246,187,214]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="133" pageNumber="134">Figs 58</figureCitation>
,
<figureCitation id="D8B05F80FF4E24F2CA1DA1F77454268A" box="[258,307,187,213]" captionStart="FIGURE 84" captionStartId="186.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-19@186.[151,1437,181,1848]" captionTargetPageId="186" captionText="FIGURE 84. Distribution of Tibetan Batrisini. A. Sathytes magnus Yin &amp; Li (circle), S. panzhaohuii Yin &amp; Shen (triangle) and S. pengzhongi sp. nov. (square). B. S. proclivis sp. nov. (circle), S. pseudograndis sp. nov. (triangle) and S. simplex Löbl (square). C. S. sympatricus sp. nov. (circle), S. transversus Yin &amp; Shen (triangle) and S. virupaksa sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340938" httpUri="https://zenodo.org/record/6340938/files/figure.png" pageId="133" pageNumber="134">84B</figureCitation>
,
<figureCitation id="D8B05F80FF4E24F2CA21A1F774F7268A" box="[318,400,187,213]" captionStart="FIGURE 97" captionStartId="199.[151,250,1355,1379]" captionTargetBox="[151,1437,181,1332]" captionTargetId="figure-15@199.[151,1437,181,1332]" captionTargetPageId="199" captionText="FIGURE 97. Environment of and collecting sites at Nyingchi area. A. Sign of Yigong Tea Farm (ƌŭē厂). B. Forest near Yigong. C. Forest at Pailong Township (ḦË乡). D. Forest near Tongmai Town (ȃ9W)." figureDoi="http://doi.org/10.5281/zenodo.6340967" httpUri="https://zenodo.org/record/6340967/files/figure.png" pageId="133" pageNumber="134">97A–C</figureCitation>
)
</paragraph>
<paragraph id="40344305FF4E24F2CB88A193775E26A8" blockId="133.[151,569,151,250]" box="[151,569,223,250]" pageId="133" pageNumber="134">
Chinese common name:
<emphasis id="72FF9F17FF4E24F2CAB2A1AC775E26A8" bold="true" box="[429,569,224,247]" pageId="133" pageNumber="134">ÊƟḛà甲</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4E24F2CB88A06871CA2764" box="[151,1197,291,316]" pageId="133" pageNumber="134" type="reference_group">
<paragraph id="40344305FF4E24F2CB88A06871CA2764" blockId="133.[151,1197,291,316]" box="[151,1197,291,316]" pageId="133" pageNumber="134">
<treatmentCitationGroup id="609B642BFF4E24F2CB88A06871CA2764" box="[151,1197,291,316]" pageId="133" pageNumber="134">
<treatmentCitation id="C12A6514FF4E24F2CB88A068748C2763" author="Lobl I." box="[151,491,292,316]" page="298" pageId="133" pageNumber="134" year="1979">
<taxonomicName id="878B3886FF4E24F2CB88A068748C2763" ID-CoL="79QPT" authority="Lobl, 1979: 298" authorityName="Lobl" authorityPageNumber="298" authorityYear="1979" box="[151,491,292,316]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FF4E24F2CB88A06874582763" box="[151,319,292,316]" italics="true" pageId="133" pageNumber="134">Sathytes simplex</emphasis>
<bibRefCitation id="241A3EF4FF4E24F2CA59A068748C2763" author="Lobl I." box="[326,491,292,316]" pageId="133" pageNumber="134" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl, 1979: 298</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; figs 19, 35.
<typeStatus id="9F30FDA7FF4E24F2C972A06877F82763" box="[621,671,292,316]" pageId="133" pageNumber="134">Type</typeStatus>
locality:
<collectingCountry id="389C0395FF4E24F2C9E0A06876482763" box="[767,815,292,316]" name="India" pageId="133" pageNumber="134">Inde</collectingCountry>
,
<location id="455415DEFF4E24F2C826A06876872763" LSID="urn:lsid:plazi:treatment:C822F213FF4E24F0CB88A1DB76C326A6:455415DEFF4E24F2C826A06876872763" box="[825,992,292,316]" country="India" municipality="Tiger Hill" name="Darjeeling distr." pageId="133" pageNumber="134">Darjeeling distr.</location>
,
<collectingMunicipality id="A050D97FFF4E24F2C8F6A06871282763" box="[1001,1103,292,316]" pageId="133" pageNumber="134">Tiger Hill</collectingMunicipality>
,
<quantity id="8773EEE0FF4E24F2CF46A06F71CE2764" box="[1113,1193,291,315]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="133" pageNumber="134" unit="m" value="2150.0">
<elevation id="CBA6A436FF4E24F2CF46A06F71CE2764" box="[1113,1193,291,315]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="133" pageNumber="134" unit="m" value="2150.0">2150 m</elevation>
</quantity>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4E24F2CB88A025779A27AF" pageId="133" pageNumber="134" type="materials_examined">
<paragraph id="40344305FF4E24F2CB88A025779A27AF" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
<emphasis id="72FF9F17FF4E24F2CB88A025776E27DC" bold="true" box="[151,521,361,387]" pageId="133" pageNumber="134">Additional material examined</emphasis>
(6 exx.).
<materialsCitation id="F0E34958FF4E24F2C997A026707027F9" ID-GBIF-Occurrence="3879233452" collectingDate="2021-07-11" collectorName="Peng &amp; Yin &amp; Zhang &amp; Affz" country="China" elevation="2300" latitude="30.170555" location="Yigong Tea Farm" longLatPrecision="20" longitude="94.93083" municipality="Nyingchi City" pageId="133" pageNumber="134" specimenCount="5" specimenCount-male="5" stateProvince="Xizang">
<specimenCount id="568D888CFF4E24F2C997A02677A827DB" box="[648,719,362,389]" pageId="133" pageNumber="134" type="male">5 ♂♂</specimenCount>
, ‘
<collectingCountry id="389C0395FF4E24F2C9F9A026765527DB" box="[742,818,362,388]" name="China" pageId="133" pageNumber="134">China</collectingCountry>
:
<collectingRegion id="824F8DE7FF4E24F2C85DA02676F127DB" box="[834,918,362,388]" country="China" name="Xizang" pageId="133" pageNumber="134">Xizang</collectingRegion>
,
<collectingMunicipality id="A050D97FFF4E24F2C8B8A026712927DB" box="[935,1102,362,388]" pageId="133" pageNumber="134">Nyingchi City</collectingMunicipality>
, pass nr.
<location id="455415DEFF4E24F2CFD9A02670FF27DB" LSID="urn:lsid:plazi:treatment:C822F213FF4E24F0CB88A1DB76C326A6:455415DEFF4E24F2CFD9A02670FF27DB" box="[1222,1432,362,388]" country="China" latitude="30.170555" longLatPrecision="20" longitude="94.93083" municipality="Nyingchi City" name="Yigong Tea Farm" pageId="133" pageNumber="134" stateProvince="Xizang">Yigong Tea Farm</location>
,
<geoCoordinate id="25BF25C2FF4E24F2CB88A0C2744427F7" box="[151,291,398,424]" degrees="30" direction="north" minutes="10" orientation="latitude" pageId="133" pageNumber="134" precision="15" seconds="14" value="30.170555">30°10’14”N</geoCoordinate>
,
<geoCoordinate id="25BF25C2FF4E24F2CA30A0C274D027F6" box="[303,439,398,425]" degrees="94" direction="east" minutes="55" orientation="longitude" pageId="133" pageNumber="134" precision="15" seconds="51" value="94.93083">94°55’51”E</geoCoordinate>
,
<quantity id="8773EEE0FF4E24F2CADCA0C2770627F7" box="[451,609,398,424]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="133" pageNumber="134" unit="m" value="2300.0" valueMax="2400.0" valueMin="2200.0">
<elevation id="CBA6A436FF4E24F2CADCA0C2770627F7" box="[451,609,398,424]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="133" pageNumber="134" unit="m" value="2300.0" valueMax="2400.0" valueMin="2200.0">2200–2400 m</elevation>
</quantity>
,
<date id="343565C5FF4E24F2C972A0C2778927F7" box="[621,750,398,424]" pageId="133" pageNumber="134" value="2021-07-11">
<collectingDate id="24719C2DFF4E24F2C972A0C2778927F7" box="[621,750,398,424]" pageId="133" pageNumber="134" value="2021-07-11">11.vii.2021</collectingDate>
</date>
,
<collectorName id="ED7E26D3FF4E24F2C9E5A0C2765227F7" box="[762,821,398,424]" pageId="133" pageNumber="134">Peng</collectorName>
,
<collectorName id="ED7E26D3FF4E24F2C820A0C2760027F7" box="[831,871,398,424]" pageId="133" pageNumber="134">Yin</collectorName>
&amp;
<collectorName id="ED7E26D3FF4E24F2C892A0C276B027F7" box="[909,983,398,424]" pageId="133" pageNumber="134">Zhang</collectorName>
,
<emphasis id="72FF9F17FF4E24F2C8FCA0C3707027F9" bold="true" box="[995,1303,399,422]" pageId="133" pageNumber="134">
Ʊ
<collectorName id="ED7E26D3FF4E24F2CF1EA0C3712927F9" box="[1025,1102,399,422]" pageId="133" pageNumber="134">Ẫffż</collectorName>
波ṁ县ƌŭē厂
</emphasis>
</materialsCitation>
’;
<materialsCitation id="F0E34958FF4E24F2CE30A0C2779E27AF" ID-GBIF-Occurrence="3879233435" collectingDate="2021-07-10" collectionCode="SNUC" collectorName="Z. Peng &amp; Z. Yin &amp; W. Zhang &amp; Ah" country="China" elevation="2180" latitude="30.009722" location="Pailong Township" longLatPrecision="20" longitude="94.96611" municipality="Nyingchi City" pageId="133" pageNumber="134" specimenCount="1" specimenCount-male="1" stateProvince="Xizang">
<specimenCount id="568D888CFF4E24F2CE30A0C2703D27F7" box="[1327,1370,398,424]" pageId="133" pageNumber="134" type="male">1 ♂</specimenCount>
‘China:
<collectingRegion id="824F8DE7FF4E24F2CBDFA0FE74732793" box="[192,276,434,460]" country="China" name="Xizang" pageId="133" pageNumber="134">Xizang</collectingRegion>
,
<collectingMunicipality id="A050D97FFF4E24F2CA00A0FE74D82793" box="[287,447,434,460]" pageId="133" pageNumber="134">Nyingchi City</collectingMunicipality>
,
<location id="455415DEFF4E24F2CAD5A0FE77F02793" LSID="urn:lsid:plazi:treatment:C822F213FF4E24F0CB88A1DB76C326A6:455415DEFF4E24F2CAD5A0FE77F02793" box="[458,663,434,460]" country="China" latitude="30.009722" longLatPrecision="20" longitude="94.96611" municipality="Nyingchi City" name="Pailong Township" pageId="133" pageNumber="134" stateProvince="Xizang">Pailong Township</location>
,
<geoCoordinate id="25BF25C2FF4E24F2C9BDA0FE76782792" box="[674,799,434,461]" degrees="30" direction="north" minutes="0" orientation="latitude" pageId="133" pageNumber="134" precision="15" seconds="35" value="30.009722">30°0’35”N</geoCoordinate>
,
<geoCoordinate id="25BF25C2FF4E24F2C834A0FE76D42792" box="[811,947,434,461]" degrees="94" direction="east" minutes="57" orientation="longitude" pageId="133" pageNumber="134" precision="15" seconds="58" value="94.96611">94°57’58”E</geoCoordinate>
,
<quantity id="8773EEE0FF4E24F2C8A0A0FE71712793" box="[959,1046,434,460]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="133" pageNumber="134" unit="m" value="2180.0">
<elevation id="CBA6A436FF4E24F2C8A0A0FE71712793" box="[959,1046,434,460]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="133" pageNumber="134" unit="m" value="2180.0">2180 m</elevation>
</quantity>
,
<date id="343565C5FF4E24F2CF3DA0FE71C42793" box="[1058,1187,434,460]" pageId="133" pageNumber="134" value="2021-07-10">
<collectingDate id="24719C2DFF4E24F2CF3DA0FE71C42793" box="[1058,1187,434,460]" pageId="133" pageNumber="134" value="2021-07-10">10.vii.2021</collectingDate>
</date>
,
<collectorName id="ED7E26D3FF4E24F2CFB0A0FE706E2793" box="[1199,1289,434,460]" pageId="133" pageNumber="134">Z. Peng</collectorName>
,
<collectorName id="ED7E26D3FF4E24F2CE0BA0FE703D2793" box="[1300,1370,434,460]" pageId="133" pageNumber="134">Z. Yin</collectorName>
&amp;
<collectorName id="ED7E26D3FF4E24F2CE62A0FE75B827AF" pageId="133" pageNumber="134">W. Zhang</collectorName>
leg.
<emphasis id="72FF9F17FF4E24F2CA09A09B772D27B1" bold="true" box="[278,586,471,494]" pageId="133" pageNumber="134">
Ʊ
<collectorName id="ED7E26D3FF4E24F2CA2CA09B740227B1" box="[307,357,471,494]" pageId="133" pageNumber="134">Ẫh</collectorName>
治区ffż市ḦË乡
</emphasis>
’ (all in
<collectionCode id="269ADBC0FF4E24F2C9BEA09A779627AF" box="[673,753,470,496]" pageId="133" pageNumber="134">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4E24F2CBD8A0B5776C24FB" pageId="133" pageNumber="134" type="diagnosis">
<paragraph id="40344305FF4E24F2CBD8A0B5776C24FB" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
<emphasis id="72FF9F17FF4E24F2CBD8A0B57425244C" bold="true" box="[199,322,505,531]" pageId="133" pageNumber="134">Diagnosis.</emphasis>
<emphasis id="72FF9F17FF4E24F2CA57A0B674E5244B" box="[328,386,506,532]" italics="true" pageId="133" pageNumber="134">Male</emphasis>
. Body length
<quantity id="8773EEE0FF4E24F2C93EA0B677DE244B" box="[545,697,506,533]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.6" metricValueMin="2.45" pageId="133" pageNumber="134" unit="mm" value="2.525" valueMax="2.6" valueMin="2.45">2.45–2.6 mm</quantity>
; ratio (length/width) of head 1.04–1.16, pronotum 1.00–1.06, elytra 0.76–0.86, length of eye/tempus 0.86–0.91. Antennae simple, antennomere 9 slightly longer than wide (length/ width 1.06–1.15), lacking hyaline apophysis on mesal margin, antennomere 11 elongate (length/width 1.76–1.95), lacking protuberance at base. Aedeagus broad, slightly constricted at middle, apical margin slightly protuberant at left side.
<emphasis id="72FF9F17FF4E24F2CBE0A3C6743424FB" box="[255,339,650,676]" italics="true" pageId="133" pageNumber="134">Female</emphasis>
. See
<bibRefCitation id="241A3EF4FF4E24F2CA8FA3C6776024FB" author="Lobl I." box="[400,519,650,676]" pageId="133" pageNumber="134" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl 1979</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4E24F1CBD8A3E175B2215C" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134" type="description">
<paragraph id="40344305FF4E24F2CBD8A3E177CA254F" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
<emphasis id="72FF9F17FF4E24F2CBD8A3E1740A2498" bold="true" box="[199,365,685,711]" pageId="133" pageNumber="134">Redescription</emphasis>
(based on populations of
<collectingCountry id="389C0395FF4E24F2C990A3E277B22497" box="[655,725,686,712]" name="China" pageId="133" pageNumber="134">China</collectingCountry>
, measurements of original description included in parentheses).
<emphasis id="72FF9F17FF4E24F2CB88A39E75B624B3" box="[151,209,722,748]" italics="true" pageId="133" pageNumber="134">Male</emphasis>
. Body (
<figureCitation id="D8B05F80FF4E24F2CA36A39E74E924B2" box="[297,398,722,749]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="133" pageNumber="134">Fig. 58A</figureCitation>
) length 2.45–2.56 (2.55–2.60) mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.
</paragraph>
<paragraph id="40344305FF4E24F2CBD8A256716422C3" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
Head (
<figureCitation id="D8B05F80FF4E24F2CA0BA256741E256A" box="[276,377,794,821]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="133" pageNumber="134">Fig. 58B</figureCitation>
) sub-rectangular at base, slightly longer than wide, length
<quantity id="8773EEE0FF4E24F2CF01A25671A2256B" box="[1054,1221,794,821]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8" metricValueMin="5.3" pageId="133" pageNumber="134" unit="mm" value="0.5549999999999999" valueMax="0.58" valueMin="0.53">0.53–0.58 mm</quantity>
, width across eyes
<quantity id="8773EEE0FF4E24F2CB88A27274582507" box="[151,319,830,857]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="133" pageNumber="134" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
, length/width 1.10–1.16 (1.04); vertex weakly raised, with small asetose foveae (dorsal tentorial pits) located slightly posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina thin and short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking median carina or sulcus. Compound eyes prominent, composed of approximately 18 large ommatidia, length of eye/tempus 0.86–0.91. Antenna elongate, length
<quantity id="8773EEE0FF4E24F2C8F8A2BE71EB2253" box="[999,1164,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.28" metricValueMin="1.27" pageId="133" pageNumber="134" unit="mm" value="1.275" valueMax="1.28" valueMin="1.27">1.27–1.28 mm</quantity>
, distinct club (
<figureCitation id="D8B05F80FF4E24F2CE2FA2BE70F32252" box="[1328,1428,1010,1037]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="133" pageNumber="134">Fig. 58C</figureCitation>
) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly broader and 8 slightly narrower than 3–7, 9 enlarged, slightly longer than wide, length/width 1.06–1.15, lacking hyaline apophysis on mesal margin, 10 transverse, length/width 0.82–0.87, 11 largest, slightly longer than 9 and 10 combined (36/33), length/width 1.76–1.95 (1.9), lacking protuberance at base.
</paragraph>
<paragraph id="40344305FF4E24F2CBD8A5EA713F2373" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
Pronotum (
<figureCitation id="D8B05F80FF4E24F2CA59A5EA74CC229E" box="[326,427,1190,1217]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="133" pageNumber="134">Fig. 58B</figureCitation>
) approximately as long as wide, both length and width
<quantity id="8773EEE0FF4E24F2CF3CA5EA71AE229F" box="[1059,1225,1190,1217]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.699999999999999" metricValueMin="5.6000000000000005" pageId="133" pageNumber="134" unit="mm" value="0.565" valueMax="0.57" valueMin="0.56">0.56–0.57 mm</quantity>
, length/width 1.00 (1.06), widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.
</paragraph>
<paragraph id="40344305FF4E24F2CBD8A47A76D423C7" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
Elytra much wider than long, length
<quantity id="8773EEE0FF4E24F2C97CA47A766F230F" box="[611,776,1334,1360]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.199999999999999" pageId="133" pageNumber="134" unit="mm" value="0.73" valueMax="0.74" valueMin="0.72">0.72–0.74 mm</quantity>
, width
<quantity id="8773EEE0FF4E24F2C845A47A7698230F" box="[858,1023,1334,1360]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="9.2" metricValueMin="8.4" pageId="133" pageNumber="134" unit="mm" value="0.88" valueMax="0.92" valueMin="0.84">0.84–0.92 mm</quantity>
, length/width 0.80–0.86 (0.76); each elytron with four large, asetose basal foveae; lacking discal stria; humerus slightly angulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.
</paragraph>
<paragraph id="40344305FF4E24F2CBD8A4EE705F2013" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae contiguous, originating from shared oval opening, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with moderately large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate and shallowly incised at middle.</paragraph>
<paragraph id="40344305FF4E24F2CBD8A71A74A3202F" blockId="133.[151,1437,361,2008]" box="[199,452,1622,1648]" pageId="133" pageNumber="134">Legs elongate, simple.</paragraph>
<paragraph id="40344305FF4E24F2CBD8A73674972187" blockId="133.[151,1437,361,2008]" pageId="133" pageNumber="134">
Abdomen widest at lateral margins of tergite 1 (IV), length
<quantity id="8773EEE0FF4E24F2C842A736716520CB" box="[861,1026,1658,1684]" metricMagnitude="-4" metricUnit="m" metricValue="6.799999999999999" metricValueMax="7.199999999999999" metricValueMin="6.4" pageId="133" pageNumber="134" unit="mm" value="0.6799999999999999" valueMax="0.72" valueMin="0.64">0.64–0.72 mm</quantity>
, width
<quantity id="8773EEE0FF4E24F2CF4CA736719E20CB" box="[1107,1273,1658,1684]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.299999999999999" metricValueMin="7.7" pageId="133" pageNumber="134" unit="mm" value="0.8" valueMax="0.83" valueMin="0.77">0.77–0.83 mm</quantity>
. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites depressed along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (
<figureCitation id="D8B05F80FF4E24F2C8AAA6D6717C21EA" box="[949,1051,1946,1973]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="133" pageNumber="134">Fig. 58D</figureCitation>
) moderately sclerotized, elongate, apex with three elongate setae.
</paragraph>
<paragraph id="40344305FF4D24F1CBD8A1DB714F268A" blockId="134.[151,1436,150,213]" pageId="134" pageNumber="135">
Aedeagus (
<figureCitation id="D8B05F80FF4D24F1CA5AA1DB74CF26EE" box="[325,424,150,177]" captionStart="FIGURE 58" captionStartId="134.[151,250,1590,1614]" captionTargetBox="[153,1436,237,1563]" captionTargetId="figure-48@134.[151,1436,237,1567]" captionTargetPageId="134" captionText="FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D." figureDoi="http://doi.org/10.5281/zenodo.6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="134" pageNumber="135">Fig. 58E</figureCitation>
) 0.29 (0.26) mm long, slightly asymmetric, broad, plate-like; median lobe with large foramen, slightly constricted at middle, apical margin slightly protuberant at left side.
</paragraph>
<caption id="14F4138DFF4D24F1CB88A77A7195202D" ID-DOI="http://doi.org/10.5281/zenodo.6340870" ID-Zenodo-Dep="6340870" httpUri="https://zenodo.org/record/6340870/files/figure.png" pageId="134" pageNumber="135" startId="134.[151,250,1590,1614]" targetBox="[153,1436,237,1563]" targetPageId="134">
<paragraph id="40344305FF4D24F1CB88A77A7195202D" blockId="134.[151,1437,1590,1651]" pageId="134" pageNumber="135">
<emphasis id="72FF9F17FF4D24F1CB88A77A74462011" bold="true" box="[151,289,1590,1614]" pageId="134" pageNumber="135">FIGURE 58.</emphasis>
Morphology of
<taxonomicName id="878B3886FF4D24F1CAD6A77B77CC2010" authority="Lobl" authorityName="Lobl" authorityYear="1979" box="[457,683,1591,1615]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FF4D24F1CAD6A77B77162010" box="[457,625,1591,1615]" italics="true" pageId="134" pageNumber="135">Sathytes simplex</emphasis>
Löbl
</taxonomicName>
, male.
<emphasis id="72FF9F17FF4D24F1C9EAA77A766A2011" bold="true" box="[757,781,1590,1614]" pageId="134" pageNumber="135">A.</emphasis>
Dorsal habitus.
<emphasis id="72FF9F17FF4D24F1C8AAA77B76AB2010" bold="true" box="[949,972,1591,1615]" pageId="134" pageNumber="135">B.</emphasis>
Head and pronotum.
<emphasis id="72FF9F17FF4D24F1CFB4A77A71A32011" bold="true" box="[1195,1220,1590,1614]" pageId="134" pageNumber="135">C.</emphasis>
Antennomeres 9–11.
<emphasis id="72FF9F17FF4D24F1CB88A71775D7202C" bold="true" box="[151,176,1627,1651]" pageId="134" pageNumber="135">D.</emphasis>
Sternite 7 (IX).
<emphasis id="72FF9F17FF4D24F1CA48A7177409202C" bold="true" box="[343,366,1627,1651]" pageId="134" pageNumber="135">E.</emphasis>
Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D.
</paragraph>
</caption>
<paragraph id="40344305FF4D24F1CBD8A7ED712E20E4" blockId="134.[151,1437,1697,2011]" box="[199,1097,1697,1723]" pageId="134" pageNumber="135">
<emphasis id="72FF9F17FF4D24F1CBD8A7ED747C20E4" box="[199,283,1697,1723]" italics="true" pageId="134" pageNumber="135">Female</emphasis>
. No material has been collected in
<collectingCountry id="389C0395FF4D24F1C9B9A7ED778B20E4" box="[678,748,1697,1723]" name="China" pageId="134" pageNumber="135">China</collectingCountry>
, see
<bibRefCitation id="241A3EF4FF4D24F1C83DA7ED76F120E4" author="Lobl I." box="[802,918,1697,1723]" pageId="134" pageNumber="135" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl 1979</bibRefCitation>
for description.
</paragraph>
<paragraph id="40344305FF4D24F1CBD8A78975B2215C" blockId="134.[151,1437,1697,2011]" pageId="134" pageNumber="135">
<emphasis id="72FF9F17FF4D24F1CBD8A78974D72080" bold="true" box="[199,432,1733,1759]" pageId="134" pageNumber="135">Comparative notes.</emphasis>
<taxonomicName id="878B3886FF4D24F1CAA8A78977162080" authorityName="Lobl" authorityYear="1979" box="[439,625,1733,1759]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FF4D24F1CAA8A78977162080" box="[439,625,1733,1759]" italics="true" pageId="134" pageNumber="135">Sathytes simplex</emphasis>
</taxonomicName>
is the only species of the genus known to possess simple antennae in the male.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4D24F1CBD8A64176D82130" pageId="134" pageNumber="135" type="distribution">
<paragraph id="40344305FF4D24F1CBD8A64176D82130" blockId="134.[151,1437,1697,2011]" pageId="134" pageNumber="135">
<emphasis id="72FF9F17FF4D24F1CBD8A64174062178" bold="true" box="[199,353,1805,1831]" pageId="134" pageNumber="135">Distribution.</emphasis>
Darjeeling District, N India; Bomê County and Chagyib District (Nyingchi),
<collectingRegion id="824F8DE7FF4D24F1CFD6A641707A2178" box="[1225,1309,1805,1831]" country="China" name="Xizang" pageId="134" pageNumber="135">Xizang</collectingRegion>
, SW China (
<figureCitation id="D8B05F80FF4D24F1CB80A67D746F2114" box="[159,264,1841,1867]" captionStart="FIGURE 84" captionStartId="186.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-19@186.[151,1437,181,1848]" captionTargetPageId="186" captionText="FIGURE 84. Distribution of Tibetan Batrisini. A. Sathytes magnus Yin &amp; Li (circle), S. panzhaohuii Yin &amp; Shen (triangle) and S. pengzhongi sp. nov. (square). B. S. proclivis sp. nov. (circle), S. pseudograndis sp. nov. (triangle) and S. simplex Löbl (square). C. S. sympatricus sp. nov. (circle), S. transversus Yin &amp; Shen (triangle) and S. virupaksa sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340938" httpUri="https://zenodo.org/record/6340938/files/figure.png" pageId="134" pageNumber="135">Figs 84B</figureCitation>
,
<figureCitation id="D8B05F80FF4D24F1CA0DA67D740F2114" box="[274,360,1841,1867]" captionStart="FIGURE 97" captionStartId="199.[151,250,1355,1379]" captionTargetBox="[151,1437,181,1332]" captionTargetId="figure-15@199.[151,1437,181,1332]" captionTargetPageId="199" captionText="FIGURE 97. Environment of and collecting sites at Nyingchi area. A. Sign of Yigong Tea Farm (ƌŭē厂). B. Forest near Yigong. C. Forest at Pailong Township (ḦË乡). D. Forest near Tongmai Town (ȃ9W)." figureDoi="http://doi.org/10.5281/zenodo.6340967" httpUri="https://zenodo.org/record/6340967/files/figure.png" pageId="134" pageNumber="135">97A–C</figureCitation>
). By discovering the new populations in China, the known range of this species extends considerably for some
<quantity id="8773EEE0FF4D24F1CA27A61974EA2130" box="[312,397,1877,1904]" metricMagnitude="5" metricUnit="m" metricValue="7.5" pageId="134" pageNumber="135" unit="km" value="750.0">750 km</quantity>
to the northeast.
<emphasis id="72FF9F17FF4D24F1C950A61976D82130" bold="true" box="[591,959,1877,1903]" pageId="134" pageNumber="135">
New country record for
<collectingCountry id="389C0395FF4D24F1C870A61976DD2130" box="[879,954,1877,1903]" name="China" pageId="134" pageNumber="135">China</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4D24F0CBD8A63576C326A6" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135" type="discussion">
<paragraph id="40344305FF4D24F0CBD8A63576C326A6" blockId="134.[151,1437,1697,2011]" lastBlockId="135.[151,1437,151,249]" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135">
<emphasis id="72FF9F17FF4D24F1CBD8A635745C21CC" bold="true" box="[199,315,1913,1939]" pageId="134" pageNumber="135">Remarks.</emphasis>
Even when adjusted to a same angle, the aedeagi of the populations from
<collectingCountry id="389C0395FF4D24F1CF8DA63571B221CC" box="[1170,1237,1913,1939]" name="China" pageId="134" pageNumber="135">China</collectingCountry>
appear to be less protuberant at the apical margin than is shown in
<bibRefCitation id="241A3EF4FF4D24F1C9A4A6D1765B21E8" author="Lobl I." box="[699,828,1949,1975]" pageId="134" pageNumber="135" pagination="285 - 307" refId="ref143858" refString="Lobl I. (1979) Revision du genre Sathytes Westwood (Coleoptera, Pselaphidae). Revue suisse de Zoologie, 86, 285 - 307. https: // doi. org / 10.5962 / bhl. part. 82294" type="journal article" year="1979">Löbl (1979</bibRefCitation>
: fig. 35). However, this difference could be reasonably fall into the range of intraspecific variation. The measurements of the body segments show no significant difference from those of specimens from northern
<collectingCountry id="389C0395FF4C24F0C941A1DB77FC26EE" box="[606,667,151,177]" name="India" pageId="135" pageNumber="136">India</collectingCountry>
. The populations of
<collectingCountry id="389C0395FF4C24F0C898A1DB76AD26EE" box="[903,970,151,177]" name="China" pageId="135" pageNumber="136">China</collectingCountry>
are temporarily identified as conspecific with
<taxonomicName id="878B3886FF4C24F0CBCFA1F77424268A" authorityName="Lobl" authorityYear="1979" box="[208,323,187,213]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="72FF9F17FF4C24F0CBCFA1F775B9268A" box="[208,222,187,213]" italics="true" pageId="135" pageNumber="136">S</emphasis>
.
<emphasis id="72FF9F17FF4C24F0CBF2A1F77424268A" box="[237,323,187,213]" italics="true" pageId="135" pageNumber="136">simplex</emphasis>
</taxonomicName>
. A direct comparison with the
<typeStatus id="9F30FDA7FF4C24F0C982A1F77798268A" box="[669,767,187,213]" pageId="135" pageNumber="136" type="holotype">holotype</typeStatus>
in the future is desired, since in many cases this is the only means to distinguish morphologically similar species within
<taxonomicName id="878B3886FF4C24F0C85FA19376FA26A6" authorityName="Westwood" authorityYear="1870" box="[832,925,223,249]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FF4C24F0C85FA19376FA26A6" box="[832,925,223,249]" italics="true" pageId="135" pageNumber="136">Sathytes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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