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treatments-xml/data/C8/22/F2/C822F213FF4C24FECB88A00674B327F2.xml
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<document id="3D23A438BC243FB2C495C42ADB981970" ID-CLB-Dataset="113069" ID-DOI="10.11646/zootaxa.5111.1.1" ID-GBIF-Dataset="f3992bb2-9962-4f3c-8f5b-131cfe552c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6340729" ID-ZooBank="836B0F69-037C-4D0F-80DB-94FE454F48E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646828425946" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="C822F213FF4C24FECB88A00674B327F2" docLanguage="en" docName="zootaxa.5111.1.1.pdf" docOrigin="Zootaxa 5111 (1)" docSource="http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Sathytes sympatricus Yin 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="138" masterDocId="341B8A6BFFCB2477CB1FA14C7567265F" masterDocTitle="The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="136" updateTime="1699317591893" updateUser="ExternalLinkService">
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<mods:title id="2663342916F19CBC230FFD974865647F">The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="B4C4E444AAD07E6EB744380538768002">Author</mods:roleTerm>
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<mods:namePart id="47240985EEF8E22419263F3E5F981270">Yin, Zi-Wei</mods:namePart>
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<mods:title id="A54BEB30A12D9582A7C7DB2A28AF34EB">Zootaxa</mods:title>
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<mods:date id="03D4F06700ABE220005D31B3940917CB">2022</mods:date>
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<mods:number id="79D41F49F03760BD7CB26D6A293E54C0">2022-03-09</mods:number>
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<mods:number id="B2447137C5A74FCDBC843751C56492F2">5111</mods:number>
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</mods:location>
<mods:classification id="CEFBEAD4E4883FFF0DAFEE5C2E4FDF1A">journal article</mods:classification>
<mods:identifier id="1221234CF8E70C7BE7D3A7DCA49326C8" type="CLB-Dataset">113069</mods:identifier>
<mods:identifier id="241916C57AE10BAE0444E1851FE82258" type="DOI">10.11646/zootaxa.5111.1.1</mods:identifier>
<mods:identifier id="BD28E064767F93F6D800EDE5285E6284" type="GBIF-Dataset">f3992bb2-9962-4f3c-8f5b-131cfe552c4d</mods:identifier>
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<treatment id="C822F213FF4C24FECB88A00674B327F2" ID-DOI="http://doi.org/10.5281/zenodo.6964386" ID-GBIF-Taxon="198579360" ID-Zenodo-Dep="6964386" LSID="urn:lsid:plazi:treatment:C822F213FF4C24FECB88A00674B327F2" httpUri="http://treatment.plazi.org/id/C822F213FF4C24FECB88A00674B327F2" lastPageId="137" lastPageNumber="138" pageId="135" pageNumber="136">
<subSubSection id="0891108EFF4C24F0CB88A006749D273A" box="[151,506,330,357]" pageId="135" pageNumber="136" type="nomenclature">
<paragraph id="40344305FF4C24F0CB88A006749D273A" blockId="135.[151,569,330,429]" box="[151,506,330,357]" pageId="135" pageNumber="136">
<heading id="1B7CF469FF4C24F0CB88A006749D273A" bold="true" box="[151,506,330,357]" fontSize="11" level="2" pageId="135" pageNumber="136" reason="4">
<emphasis id="72FF9F17FF4C24F0CB88A006749D273A" bold="true" box="[151,506,330,357]" pageId="135" pageNumber="136">
<taxonomicName id="878B3886FF4C24F0CB88A00674F2273A" ID-CoL="9GDH7" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[151,405,330,357]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="sympatricus" status="sp. nov.">
<emphasis id="72FF9F17FF4C24F0CB88A00674F2273A" bold="true" box="[151,405,330,357]" italics="true" pageId="135" pageNumber="136">Sathytes sympatricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69CC226CFF4C24F0CA83A007749D273A" box="[412,506,331,357]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4C24F0CB88A023775E27F5" pageId="135" pageNumber="136" type="description">
<paragraph id="40344305FF4C24F0CB88A02374FE27D6" blockId="135.[151,569,330,429]" box="[151,409,366,393]" pageId="135" pageNumber="136">
(
<figureCitation id="D8B05F80FF4C24F0CB80A023759127D6" box="[159,246,366,393]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Figs 59</figureCitation>
,
<figureCitation id="D8B05F80FF4C24F0CA1DA023745427D6" box="[258,307,367,393]" captionStart="FIGURE 84" captionStartId="186.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-19@186.[151,1437,181,1848]" captionTargetPageId="186" captionText="FIGURE 84. Distribution of Tibetan Batrisini. A. Sathytes magnus Yin &amp; Li (circle), S. panzhaohuii Yin &amp; Shen (triangle) and S. pengzhongi sp. nov. (square). B. S. proclivis sp. nov. (circle), S. pseudograndis sp. nov. (triangle) and S. simplex Löbl (square). C. S. sympatricus sp. nov. (circle), S. transversus Yin &amp; Shen (triangle) and S. virupaksa sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340938" httpUri="https://zenodo.org/record/6340938/files/figure.png" pageId="135" pageNumber="136">84C</figureCitation>
,
<figureCitation id="D8B05F80FF4C24F0CA21A02274F727D6" box="[318,400,366,393]" captionStart="FIGURE 95" captionStartId="197.[151,250,1166,1190]" captionTargetBox="[151,1436,181,1142]" captionTargetId="figure-15@197.[151,1436,181,1142]" captionTargetPageId="197" captionText="FIGURE 95. Environment of and collecting sites at Lebu Valley (勒布沟). Forests at 2400 m (A, B), and 3650 m (C, D)." figureDoi="http://doi.org/10.5281/zenodo.6340963" httpUri="https://zenodo.org/record/6340963/files/figure.png" pageId="135" pageNumber="136">95A, B</figureCitation>
)
</paragraph>
<paragraph id="40344305FF4C24F0CB88A0DF775E27F5" blockId="135.[151,569,330,429]" box="[151,569,403,429]" pageId="135" pageNumber="136">
Chinese common name:
<emphasis id="72FF9F17FF4C24F0CAB2A0DF775E27F5" bold="true" box="[429,569,403,426]" pageId="135" pageNumber="136">ḲRḛà甲</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4C24F0CB88A09770E42462" pageId="135" pageNumber="136" type="materials_examined">
<paragraph id="40344305FF4C24F0CB88A09770E42462" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">
<emphasis id="72FF9F17FF4C24F0CB88A097742727AB" bold="true" box="[151,320,474,501]" pageId="135" pageNumber="136">Type material</emphasis>
(16 exx.).
<materialsCitation id="F0E34958FF4C24F0CAA0A096779127AA" ID-GBIF-Occurrence="3879233319" box="[447,758,474,501]" country="China" location="CHINA" pageId="135" pageNumber="136" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="72FF9F17FF4C24F0CAA0A09677B127AB" bold="true" box="[447,726,474,500]" pageId="135" pageNumber="136">
<typeStatus id="9F30FDA7FF4C24F0CAA0A096770527AB" box="[447,610,474,500]" pageId="135" pageNumber="136" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="389C0395FF4C24F0C96FA09677B527AB" box="[624,722,474,500]" name="China" pageId="135" pageNumber="136">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="568D888CFF4C24F0C9C0A097779127AA" box="[735,758,475,501]" pageId="135" pageNumber="136" type="male">♂</specimenCount>
</materialsCitation>
, ‘
<materialsCitation id="F0E34958FF4C24F0C816A097743D2462" ID-GBIF-Occurrence="3879233370" collectingDate="2021-07-04" collectionCode="SNUC" collectorName="Z. Peng &amp; Z. Yin &amp; W. Zhang" country="China" county="Cona County" elevation="2550" latitude="27.837778" location="Lebu Valley" longLatPrecision="20" longitude="91.763336" pageId="135" pageNumber="136" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<collectingCountry id="389C0395FF4C24F0C816A097763227AA" box="[777,853,475,501]" name="China" pageId="135" pageNumber="136">China</collectingCountry>
:
<collectingRegion id="824F8DE7FF4C24F0C87EA09776D227AA" box="[865,949,475,501]" country="China" name="Xizang" pageId="135" pageNumber="136">Xizang</collectingRegion>
,
<collectingCounty id="A9553B89FF4C24F0C8DEA097713E27AA" box="[961,1113,475,501]" pageId="135" pageNumber="136">Cona County</collectingCounty>
,
<location id="455415DEFF4C24F0CF79A097719627AA" LSID="urn:lsid:plazi:treatment:C822F213FF4C24FECB88A00674B327F2:455415DEFF4C24F0CF79A097719627AA" box="[1126,1265,475,501]" country="China" county="Cona County" latitude="27.837778" longLatPrecision="20" longitude="91.763336" name="Lebu Valley" pageId="135" pageNumber="136" stateProvince="Xizang">Lebu Valley</location>
, pass to Mama Vill.,
<geoCoordinate id="25BF25C2FF4C24F0CBC8A0B274052446" box="[215,354,510,537]" degrees="27" direction="north" minutes="50" orientation="latitude" pageId="135" pageNumber="136" precision="15" seconds="16" value="27.837778">27°50’16”N</geoCoordinate>
,
<geoCoordinate id="25BF25C2FF4C24F0CA70A0B274902446" box="[367,503,510,537]" degrees="91" direction="east" minutes="45" orientation="longitude" pageId="135" pageNumber="136" precision="15" seconds="48" value="91.763336">91°45’48”E</geoCoordinate>
,
<quantity id="8773EEE0FF4C24F0C91CA0B377C62446" box="[515,673,511,537]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="135" pageNumber="136" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">
<elevation id="CBA6A436FF4C24F0C91CA0B377C62446" box="[515,673,511,537]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="135" pageNumber="136" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">2400–2700 m</elevation>
</quantity>
,
<date id="343565C5FF4C24F0C9B1A0B376572446" box="[686,816,511,537]" pageId="135" pageNumber="136" value="2021-07-04">
<collectingDate id="24719C2DFF4C24F0C9B1A0B376572446" box="[686,816,511,537]" pageId="135" pageNumber="136" value="2021-07-04">04.vii.2021</collectingDate>
</date>
,
<collectorName id="ED7E26D3FF4C24F0C823A0B376F02446" box="[828,919,511,537]" pageId="135" pageNumber="136">Z. Peng</collectorName>
,
<collectorName id="ED7E26D3FF4C24F0C8BCA0B3768E2446" box="[931,1001,511,537]" pageId="135" pageNumber="136">Z. Yin</collectorName>
&amp;
<collectorName id="ED7E26D3FF4C24F0CF11A0B3711A2446" box="[1038,1149,511,537]" pageId="135" pageNumber="136">W. Zhang</collectorName>
leg.,
<emphasis id="72FF9F17FF4C24F0CFA3A0B3758B2465" bold="true" pageId="135" pageNumber="136">ƱẪDZḄ县勒布沟 Þ玛乡</emphasis>
’ (
<collectionCode id="269ADBC0FF4C24F0CA1EA36F74352462" box="[257,338,547,573]" pageId="135" pageNumber="136">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F0E34958FF4C24F0CA7AA36E70182462" ID-GBIF-Occurrence="3879233376" box="[357,1407,546,573]" collectingDate="2021-07-04" collectionCode="SNUC" collectorName="Z. Peng &amp; Z. Yin &amp; W. Zhang" country="China" county="Cona County" elevation="2550" latitude="27.837778" location="Lebu Valley" longLatPrecision="20" longitude="91.763336" pageId="135" pageNumber="136" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="72FF9F17FF4C24F0CA7AA36E77172463" bold="true" box="[357,624,546,572]" pageId="135" pageNumber="136">
<typeStatus id="9F30FDA7FF4C24F0CA7AA36E74992463" box="[357,510,546,572]" pageId="135" pageNumber="136" type="paratype">PARATYPE</typeStatus>
:
<collectingCountry id="389C0395FF4C24F0C916A36E770C2463" box="[521,619,546,572]" name="China" pageId="135" pageNumber="136">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="568D888CFF4C24F0C968A36F77A12462" box="[631,710,547,573]" pageId="135" pageNumber="136" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="568D888CFF4C24F0C9CDA36E76752462" box="[722,786,546,573]" pageId="135" pageNumber="136" type="female">5 ♀♀</specimenCount>
, same data as that of
<typeStatus id="9F30FDA7FF4C24F0CF1CA36F71022462" box="[1027,1125,547,573]" pageId="135" pageNumber="136" type="holotype">holotype</typeStatus>
(all
<typeStatus id="9F30FDA7FF4C24F0CF87A36F70642462" box="[1176,1283,547,573]" pageId="135" pageNumber="136" type="paratype">paratypes</typeStatus>
in
<collectionCode id="269ADBC0FF4C24F0CE38A36F70102462" box="[1319,1399,547,573]" pageId="135" pageNumber="136">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4C24F0CBD8A30A77C1254A" pageId="135" pageNumber="136" type="diagnosis">
<paragraph id="40344305FF4C24F0CBD8A30A77C1254A" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">
<emphasis id="72FF9F17FF4C24F0CBD8A30A7425243F" bold="true" box="[199,322,582,608]" pageId="135" pageNumber="136">Diagnosis.</emphasis>
<emphasis id="72FF9F17FF4C24F0CA56A30B74E4243E" box="[329,387,583,609]" italics="true" pageId="135" pageNumber="136">Male</emphasis>
. Body length approximately
<quantity id="8773EEE0FF4C24F0C9D2A30B7641243E" box="[717,806,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="135" pageNumber="136" unit="mm" value="2.7">2.7 mm</quantity>
; ratio (length/width) of head 1.00–1.08, pronotum 1.05, elytra 0.83–0.84, length of eye/tempus 0.95. Antennomere 9 as long as wide (length/width excluding apophysis 1.0), with hyaline apophysis at middle of mesal margin, antennomere 11 elongate (length/width 1.64), with short protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad, slightly constricted at middle, apical margin slightly emarginate and convex on right side.
<emphasis id="72FF9F17FF4C24F0C820A39B76F424AE" box="[831,915,727,753]" italics="true" pageId="135" pageNumber="136">Female</emphasis>
. Body length approximately
<quantity id="8773EEE0FF4C24F0CFC0A39B705E24AE" box="[1247,1337,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="135" pageNumber="136" unit="mm" value="2.6">2.6 mm</quantity>
; antenna lacking modifications, genitalia as in
<figureCitation id="D8B05F80FF4C24F0C924A3B777C5254A" box="[571,674,762,789]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Fig. 59G</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4C24FECBD8A252776B271F" lastPageId="137" lastPageNumber="138" pageId="135" pageNumber="136" type="description">
<paragraph id="40344305FF4C24F0CBD8A25277992502" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">
<emphasis id="72FF9F17FF4C24F0CBD8A252743E2567" bold="true" box="[199,345,798,824]" pageId="135" pageNumber="136">Description.</emphasis>
<emphasis id="72FF9F17FF4C24F0CA7FA25374FD2566" box="[352,410,799,825]" italics="true" pageId="135" pageNumber="136">Male</emphasis>
. Body (
<figureCitation id="D8B05F80FF4C24F0CAEAA253773B2566" box="[501,604,798,825]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Fig. 59A</figureCitation>
) length
<quantity id="8773EEE0FF4C24F0C9A5A25376072566" box="[698,864,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.68" metricValueMin="2.66" pageId="135" pageNumber="136" unit="mm" value="2.67" valueMax="2.68" valueMin="2.66">2.66–2.68 mm</quantity>
; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.
</paragraph>
<paragraph id="40344305FF4C24F0CBD8A22B74AE236E" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">
Head (
<figureCitation id="D8B05F80FF4C24F0CA0CA22B741E25DE" box="[275,377,870,897]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Fig. 59B</figureCitation>
) sub-rectangular at base, approximately as long as wide, length
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, width across eyes
<quantity id="8773EEE0FF4C24F0CBCEA2C6741025FA" box="[209,375,906,933]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="135" pageNumber="136" unit="mm" value="0.545" valueMax="0.56" valueMin="0.53">0.53–0.56 mm</quantity>
, length/width 1.00–1.08; vertex moderately raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, shallowly impressed at middle; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking distinct median carina or sulcus. Compound eyes prominent, composed of approximately 17 large ommatidia, length of eye/tempus 0.95. Antenna elongate, length
<quantity id="8773EEE0FF4C24F0CF76A57371A92206" box="[1129,1230,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="135" pageNumber="136" unit="mm" value="1.33">1.33 mm</quantity>
, distinct club (
<figureCitation id="D8B05F80FF4C24F0CE70A57375A02222" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Fig. 59C</figureCitation>
) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each transverse, subequal in width, 8 as wide as and much shorter than 7, 9 (
<figureCitation id="D8B05F80FF4C24F0C810A5CB761222FE" box="[783,885,1158,1185]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Fig. 59D</figureCitation>
) strongly enlarged, as long as wide, length/width excluding apophysis 1.0, mesal margin moderately expanded at middle and strongly oblique anteriorly and posteriorly, hyaline apophysis located at middle of mesal margin, 10 transverse, length/width 0.71, 11 largest, as long as 9 and 10 combined, length/width 1.64, with short, apically truncate protuberance at base, apex of protuberance with curved tuft of golden setae.
</paragraph>
<paragraph id="40344305FF4C24F0CBD8A4777688239E" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">
Pronotum (
<figureCitation id="D8B05F80FF4C24F0CA5AA47774CF230A" box="[325,424,1338,1365]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="135" pageNumber="136">Fig. 59B</figureCitation>
) slightly longer than wide, length
<quantity id="8773EEE0FF4C24F0C836A47676A9230A" box="[809,974,1338,1365]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.1" metricValueMin="5.8" pageId="135" pageNumber="136" unit="mm" value="0.595" valueMax="0.61" valueMin="0.58">0.58–0.61 mm</quantity>
, width
<quantity id="8773EEE0FF4C24F0CF3FA47671A2230A" box="[1056,1221,1338,1365]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="135" pageNumber="136" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
, length/width 1.05, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; large asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.
</paragraph>
<paragraph id="40344305FF4C24F0CBD8A48776852072" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">
Elytra much wider than long, length
<quantity id="8773EEE0FF4C24F0C941A487766523BA" box="[606,770,1483,1509]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" metricValueMax="8.6" metricValueMin="8.2" pageId="135" pageNumber="136" unit="mm" value="0.84" valueMax="0.86" valueMin="0.82">0.82–0.86 mm</quantity>
, width
<quantity id="8773EEE0FF4C24F0C84CA487769023BA" box="[851,1015,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.005" metricValueMax="1.02" metricValueMin="0.99" pageId="135" pageNumber="136" unit="mm" value="1.005" valueMax="1.02" valueMin="0.99">0.99–1.02 mm</quantity>
, length/width 0.83–0.84; each elytron with four large, asetose basal foveae; discal stria shallow and short; humerus weakly angulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.
</paragraph>
<paragraph id="40344305FF4C24F0CBD8A77B765820BE" blockId="135.[151,1437,474,2013]" pageId="135" pageNumber="136">Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, in oval opening, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate at middle, lacking split.</paragraph>
<paragraph id="40344305FF4C24F0CBD8A7A774A3215A" blockId="135.[151,1437,474,2013]" box="[199,452,1771,1797]" pageId="135" pageNumber="136">Legs elongate, simple.</paragraph>
<paragraph id="40344305FF4C24FFCBD8A64377422742" blockId="135.[151,1437,474,2013]" lastBlockId="136.[151,1437,150,537]" lastPageId="136" lastPageNumber="137" pageId="135" pageNumber="136">
Abdomen widest at lateral margins of tergite 1 (IV), length
<quantity id="8773EEE0FF4C24F0C842A64371652176" box="[861,1026,1807,1833]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="7.8" metricValueMin="7.7" pageId="135" pageNumber="136" unit="mm" value="0.775" valueMax="0.78" valueMin="0.77">0.77–0.78 mm</quantity>
, width
<quantity id="8773EEE0FF4C24F0CF4CA643719E2176" box="[1107,1273,1807,1833]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="8.8" metricValueMin="8.7" pageId="135" pageNumber="136" unit="mm" value="0.875" valueMax="0.88" valueMin="0.87">0.87–0.88 mm</quantity>
. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of large mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites flat along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (
<figureCitation id="D8B05F80FF4324FFC867A19376BB26A6" box="[888,988,222,249]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="136" pageNumber="137">Fig. 59E</figureCitation>
) moderately sclerotized, elongate, with few long setae along apical margin.
</paragraph>
<paragraph id="40344305FF4324FFCBD8A06B70E2273A" blockId="136.[151,1437,150,537]" pageId="136" pageNumber="137">
Aedeagus (
<figureCitation id="D8B05F80FF4324FFCA59A06B74C0271E" box="[326,423,294,321]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="136" pageNumber="137">Fig. 59F</figureCitation>
)
<quantity id="8773EEE0FF4324FFCAA8A06B777B271E" box="[439,540,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="136" pageNumber="137" unit="mm" value="0.29">0.29 mm</quantity>
long, slightly asymmetric, plate-like; median lobe with large foramen, broad at base and slightly constricted at middle, apical margin moderately emarginate at middle and convex on right side.
</paragraph>
<paragraph id="40344305FF4324FFCBD8A02377622446" blockId="136.[151,1437,150,537]" pageId="136" pageNumber="137">
<emphasis id="72FF9F17FF4324FFCBD8A023747C27D6" box="[199,283,367,393]" italics="true" pageId="136" pageNumber="137">Female</emphasis>
. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 14 ommatidia; humerus weakly prominent. Measurements (as for male): body length
<quantity id="8773EEE0FF4324FFCA39A0FA74A9278E" box="[294,462,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="2.575" metricValueMax="2.59" metricValueMin="2.56" pageId="136" pageNumber="137" unit="mm" value="2.575" valueMax="2.59" valueMin="2.56">2.56–2.59 mm</quantity>
; length/width of head 0.56/
<quantity id="8773EEE0FF4324FFC817A0FA76C9278E" box="[776,942,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.6" metricValueMin="5.2" pageId="136" pageNumber="137" unit="mm" value="0.54" valueMax="0.56" valueMin="0.52">0.52–0.56 mm</quantity>
, pronotum 0.58–0.60/
<quantity id="8773EEE0FF4324FFCFB6A0FA7028278E" box="[1193,1359,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="5.8999999999999995" metricValueMin="5.6" pageId="136" pageNumber="137" unit="mm" value="0.575" valueMax="0.59" valueMin="0.56">0.56–0.59 mm</quantity>
, elytra 0.76–0.77/
<quantity id="8773EEE0FF4324FFCA10A09774D027AA" box="[271,439,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="9.9" metricValueMin="9.7" pageId="136" pageNumber="137" unit="mm" value="0.98" valueMax="0.99" valueMin="0.97">0.97–0.99 mm</quantity>
; abdomen 0.77–0.80/
<quantity id="8773EEE0FF4324FFC9B4A097763527AA" box="[683,850,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="9.35" metricValueMax="9.399999999999999" metricValueMin="9.3" pageId="136" pageNumber="137" unit="mm" value="0.935" valueMax="0.94" valueMin="0.93">0.93–0.94 mm</quantity>
; length of antenna
<quantity id="8773EEE0FF4324FFCF31A09771B227AA" box="[1070,1237,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" metricValueMax="1.23" metricValueMin="1.19" pageId="136" pageNumber="137" unit="mm" value="1.21" valueMax="1.23" valueMin="1.19">1.19–1.23 mm</quantity>
; maximum width of genitalia (
<figureCitation id="D8B05F80FF4324FFCA3AA0B374EC2446" box="[293,395,510,537]" captionStart="FIGURE 59" captionStartId="136.[151,250,1836,1860]" captionTargetBox="[155,1436,562,1813]" captionTargetId="figure-169@136.[151,1436,561,1813]" captionTargetPageId="136" captionText="FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G." figureDoi="http://doi.org/10.5281/zenodo.6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="136" pageNumber="137">Fig. 59G</figureCitation>
)
<quantity id="8773EEE0FF4324FFCA85A0B377662446" box="[410,513,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="136" pageNumber="137" unit="mm" value="0.26">0.26 mm</quantity>
.
</paragraph>
<caption id="14F4138DFF4324FFCB88A66074A621D3" ID-DOI="http://doi.org/10.5281/zenodo.6340872" ID-Zenodo-Dep="6340872" httpUri="https://zenodo.org/record/6340872/files/figure.png" pageId="136" pageNumber="137" startId="136.[151,250,1836,1860]" targetBox="[155,1436,562,1813]" targetPageId="136">
<paragraph id="40344305FF4324FFCB88A66074A621D3" blockId="136.[151,1437,1836,1932]" pageId="136" pageNumber="137">
<emphasis id="72FF9F17FF4324FFCB88A6607478211B" bold="true" box="[151,287,1836,1860]" pageId="136" pageNumber="137">FIGURE 59.</emphasis>
Morphology of
<taxonomicName id="878B3886FF4324FFCADAA66077FE211B" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[453,665,1836,1860]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="sympatricus" status="sp. nov.">
<emphasis id="72FF9F17FF4324FFCADAA66077FE211B" box="[453,665,1836,1860]" italics="true" pageId="136" pageNumber="137">Sathytes sympatricus</emphasis>
</taxonomicName>
<emphasis id="72FF9F17FF4324FFC980A6607788211B" bold="true" box="[671,751,1836,1860]" pageId="136" pageNumber="137">
<taxonomicNameLabel id="69CC226CFF4324FFC980A6607788211B" box="[671,751,1836,1860]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A–F. Male. G. Female).
<emphasis id="72FF9F17FF4324FFC8ECA660716B211B" bold="true" box="[1011,1036,1836,1860]" pageId="136" pageNumber="137">A.</emphasis>
Dorsal habitus.
<emphasis id="72FF9F17FF4324FFCFAEA66071AF211B" bold="true" box="[1201,1224,1836,1860]" pageId="136" pageNumber="137">B.</emphasis>
Head and pronotum.
<emphasis id="72FF9F17FF4324FFCB88A61C75D72137" bold="true" box="[151,176,1872,1896]" pageId="136" pageNumber="137">C.</emphasis>
Antennomeres 9–11.
<emphasis id="72FF9F17FF4324FFCA91A61C74C12137" bold="true" box="[398,422,1872,1896]" pageId="136" pageNumber="137">D.</emphasis>
Antennomere 9.
<emphasis id="72FF9F17FF4324FFC949A61C77092137" bold="true" box="[598,622,1872,1896]" pageId="136" pageNumber="137">E.</emphasis>
Sternite 7 (IX).
<emphasis id="72FF9F17FF4324FFC80AA61C764F2137" bold="true" box="[789,808,1872,1896]" pageId="136" pageNumber="137">F.</emphasis>
Aedeagus, ventral.
<emphasis id="72FF9F17FF4324FFC8EEA61C716C2137" bold="true" box="[1009,1035,1872,1896]" pageId="136" pageNumber="137">G.</emphasis>
Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G.
</paragraph>
</caption>
<paragraph id="40344305FF4224FECBD8A1DA776B271F" blockId="137.[151,1437,150,429]" pageId="137" pageNumber="138">
<emphasis id="72FF9F17FF4224FECBD8A1DA74D726EF" bold="true" box="[199,432,150,176]" pageId="137" pageNumber="138">Comparative notes.</emphasis>
The new species is related to a number of species centered on
<taxonomicName id="878B3886FF4224FECF70A1DB718626EF" authorityName="Lobl" authorityYear="1979" box="[1135,1249,150,177]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="72FF9F17FF4224FECF70A1DB711A26EE" box="[1135,1149,151,177]" italics="true" pageId="137" pageNumber="138">S</emphasis>
.
<emphasis id="72FF9F17FF4224FECF94A1DA718626EF" box="[1163,1249,150,176]" italics="true" pageId="137" pageNumber="138">grandis</emphasis>
</taxonomicName>
, with all sharing a hyaline apophysis at the oblique mesal margin of antennomeres 9, and with a single basal protuberance consisting of a curved apical setose tuft on antennomere 11.
<taxonomicName id="878B3886FF4224FEC9CDA19276D826A7" authorityName="Yin" authorityYear="2022" box="[722,959,222,248]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="sympatricus">
<emphasis id="72FF9F17FF4224FEC9CDA19276D826A7" box="[722,959,222,248]" italics="true" pageId="137" pageNumber="138">Sathytes sympatricus</emphasis>
</taxonomicName>
differs particularly in the relatively small body size (
<quantity id="8773EEE0FF4224FECA0EA04E740A2742" box="[273,365,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="137" pageNumber="138" unit="mm" value="2.7">2.7 mm</quantity>
), the long hyaline apophyses of antennomeres 9, the short basal protuberances of antennomeres 11, and the form of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4224FECBD8A00676C5273A" box="[199,930,330,357]" pageId="137" pageNumber="138" type="distribution">
<paragraph id="40344305FF4224FECBD8A00676C5273A" blockId="137.[151,1437,150,429]" box="[199,930,330,357]" pageId="137" pageNumber="138">
<emphasis id="72FF9F17FF4224FECBD8A0067406273B" bold="true" box="[199,353,330,356]" pageId="137" pageNumber="138">Distribution.</emphasis>
Cona County,
<collectingRegion id="824F8DE7FF4224FEC917A0067722273B" box="[520,581,330,356]" country="China" name="Xizang" pageId="137" pageNumber="138">Tibet</collectingRegion>
, SW
<collectingCountry id="389C0395FF4224FEC99FA00677A4273B" box="[640,707,330,356]" name="China" pageId="137" pageNumber="138">China</collectingCountry>
(
<figureCitation id="D8B05F80FF4224FEC9CDA006765B273B" box="[722,828,330,356]" captionStart="FIGURE 84" captionStartId="186.[151,250,1872,1896]" captionTargetBox="[151,1437,181,1848]" captionTargetId="figure-19@186.[151,1437,181,1848]" captionTargetPageId="186" captionText="FIGURE 84. Distribution of Tibetan Batrisini. A. Sathytes magnus Yin &amp; Li (circle), S. panzhaohuii Yin &amp; Shen (triangle) and S. pengzhongi sp. nov. (square). B. S. proclivis sp. nov. (circle), S. pseudograndis sp. nov. (triangle) and S. simplex Löbl (square). C. S. sympatricus sp. nov. (circle), S. transversus Yin &amp; Shen (triangle) and S. virupaksa sp. nov. (square)." figureDoi="http://doi.org/10.5281/zenodo.6340938" httpUri="https://zenodo.org/record/6340938/files/figure.png" pageId="137" pageNumber="138">Figs 84C</figureCitation>
,
<figureCitation id="D8B05F80FF4224FEC858A00676FE273A" box="[839,921,330,357]" captionStart="FIGURE 95" captionStartId="197.[151,250,1166,1190]" captionTargetBox="[151,1436,181,1142]" captionTargetId="figure-15@197.[151,1436,181,1142]" captionTargetPageId="197" captionText="FIGURE 95. Environment of and collecting sites at Lebu Valley (勒布沟). Forests at 2400 m (A, B), and 3650 m (C, D)." figureDoi="http://doi.org/10.5281/zenodo.6340963" httpUri="https://zenodo.org/record/6340963/files/figure.png" pageId="137" pageNumber="138">95A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF4224FECBD8A02274B327F2" pageId="137" pageNumber="138" type="etymology">
<paragraph id="40344305FF4224FECBD8A02274B327F2" blockId="137.[151,1437,150,429]" pageId="137" pageNumber="138">
<emphasis id="72FF9F17FF4224FECBD8A022742927D7" bold="true" box="[199,334,366,392]" pageId="137" pageNumber="138">Etymology.</emphasis>
The specific epithet ‘
<taxonomicName id="878B3886FF4224FEC95BA02377AC27D7" authorityName="Yin" authorityYear="2022" box="[580,715,367,392]" class="Insecta" family="Staphylinidae" genus="Sathytes" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="sympatricus">
<emphasis id="72FF9F17FF4224FEC95BA02377AC27D7" box="[580,715,367,392]" italics="true" pageId="137" pageNumber="138">sympatricus</emphasis>
</taxonomicName>
, -
<emphasis id="72FF9F17FF4224FEC9FCA023779627D7" box="[739,753,367,392]" italics="true" pageId="137" pageNumber="138">a</emphasis>
, -
<emphasis id="72FF9F17FF4224FEC816A023764C27D7" box="[777,811,367,392]" italics="true" pageId="137" pageNumber="138">um</emphasis>
’ is a Latin adjective indicating that this species is sympatric with a few congeners.
</paragraph>
</subSubSection>
</treatment>
</document>
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