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<document id="152E8D7693B54753E2F0CF03316A959E" ID-DOI="http://dx.doi.org/10.3897/zookeys.349.6164" ID-GBIF-Dataset="cd871e7e-7e75-4f2e-8c8b-7af9cb4803d4" ID-PMC="PMC3837407" ID-Pensoft-Pub="1313-2970-349-81" ID-PubMed="24294080" ID-ZooBank="45895E0B79E74794A0155CD74FEDDF22" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-349-81" ModsDocOrigin="ZooKeys 349" ModsDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" checkinTime="1451246687136" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2013" docId="7042A3ACA274291EC08F0DC89473D832" docLanguage="en" docName="ZooKeys 349: 81-100" docOrigin="ZooKeys 349" docSource="http://dx.doi.org/10.3897/zookeys.349.6164" docTitle="Orthogonius lancangjiang Tian &amp; Deuve 2006" docType="treatment" docVersion="5" lastPageNumber="86" masterDocId="B203D41F1D41C83E8603FFE5D1499035" masterDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" masterLastPageNumber="100" masterPageNumber="81" pageNumber="83" updateTime="1701317668806" updateUser="plazi">
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<mods:title id="6E495FBC275FCDD4B4A65EA929DE7BD9">Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)</mods:title>
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<mods:name id="6A3DB95F07FB6B9B2EB1766C2E6F033A" type="personal">
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<mods:namePart id="06144ADC7AD781D7F39181B3BA07DE46">Tian, Mingyi</mods:namePart>
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<mods:namePart id="24A77E5E3EEC2B29308C3263E4C6ADF6">Deuve, Thierry</mods:namePart>
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<mods:date id="F51662E6632C3F47AA4754FDD923110F">2013</mods:date>
<mods:detail id="D8F11D359A05AA98108D9BA0B50EAEEF" type="volume">
<mods:number id="58FCF1882F62C42A43ABAE3CA496B6FD">349</mods:number>
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<mods:url id="388C38035CE77704BC17EF111315F6B4">http://dx.doi.org/10.3897/zookeys.349.6164</mods:url>
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<mods:classification id="31B64EB3FF5A98A47F1EC0ACD061CDCF">journal article</mods:classification>
<mods:identifier id="28070A711723EDE242C12E1141716BA7" type="DOI">http://dx.doi.org/10.3897/zookeys.349.6164</mods:identifier>
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<treatment id="7042A3ACA274291EC08F0DC89473D832" ID-GBIF-Taxon="152049510" LSID="urn:lsid:plazi:treatment:7042A3ACA274291EC08F0DC89473D832" httpUri="http://treatment.plazi.org/id/7042A3ACA274291EC08F0DC89473D832" lastPageId="5" lastPageNumber="86" pageId="2" pageNumber="83">
<subSubSection id="58AF420EF2CF5A8979C0A16F67156215" pageId="2" pageNumber="83" type="nomenclature">
<paragraph id="B7F73CA25000C4CC6D111F9261D2A375" pageId="2" pageNumber="83">
<taxonomicName id="E4A3E4C422A8FB4C10DB904C8CEC12BC" ID-CoL="6SX9X" LSID="http://species-id.net/wiki/Orthogonius_lancangjiang" authority="Tian &amp; Deuve, 2006" authorityName="Tian &amp; Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang Tian &amp; Deuve, 2006</taxonomicName>
Figs 1-3, 12-13
</paragraph>
</subSubSection>
<subSubSection id="CC04C12A05918FF057F851306F8B07BC" pageId="2" pageNumber="83" type="reference_group">
<paragraph id="7EB08A704ECCE7E14DD9E4B2805D852F" pageId="2" pageNumber="83">
<taxonomicName id="E1B24A2A19411C23FA253821AEB38C0C" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
Tian &amp; Deuve, 2006: 134
</paragraph>
</subSubSection>
<subSubSection id="D9C21E8E2FB88575D875468B4C8CD74A" pageId="2" pageNumber="83" type="diagnosis">
<paragraph id="C2E6E8B4CB74BB6776E565CFB602A09D" pageId="2" pageNumber="83">Diagnosis.</paragraph>
<paragraph id="2C62C551FCD4F55181BAB6A00B4FE34A" pageId="2" pageNumber="83">Medium to large sized, elytra slender or elongate ovate; head longer than wide; upper surface covered with dense punctures; antennae long; eyes moderate for orthogoniines; labrum round at front; pronotum quite narrow, but transverse in form; whole lateral expanded margins slightly reflexed; elytra not carinate on interval 7 in both sexes.</paragraph>
<paragraph id="B1625332C7F1C24D4E43C8CDA9374493" pageId="2" pageNumber="83">Length: 12.5-16.0 mm; width (=EW): 5.2-6.5 mm.</paragraph>
</subSubSection>
<subSubSection id="834AE49DDA26055019EE30E6E4FA2EB8" lastPageId="4" lastPageNumber="85" pageId="2" pageNumber="83" type="description">
<paragraph id="7ACBF877FD96625372880C33D0A8EB87" pageId="2" pageNumber="83">Description.</paragraph>
<paragraph id="C49D984BAF9F89957CBDB0020CF0CB8B" pageId="2" pageNumber="83">Body elongate, strongly shiny. Habitus as in Figs 2-3.</paragraph>
<paragraph id="7EF5781A7F410DE40DC2013FED6BA6C0" pageId="3" pageNumber="84">
<pageBreakToken id="DCAEF5061882B30BCD3B316C744943EA" pageId="3" pageNumber="84" start="start">Head</pageBreakToken>
and disc of pronotum black; elytra black to dark brown; lateral expanded margins of pronotum, antennomere 1 and 3, mandibles, and tibiae dark brown; other parts brown.
</paragraph>
<paragraph id="A3274FD021B96149D450DDE7EA867370" pageId="3" pageNumber="84">Macrosculpture: Surface of head, pronotum and elytra (including all intervals) with dense punctures, head obscurely striate, pronotum impunctate in most specimens, or with a few faint punctures in one or two individuals, moderately striate or not; underside surface smooth and glabrous, impunctate.</paragraph>
<paragraph id="17CEC0254FD52ADD7EF6CE841980EADF" pageId="4" pageNumber="85">
<pageBreakToken id="B26A3DC5569810AC646DD07B6B122DEB" pageId="4" pageNumber="85" start="start">Microsculpture</pageBreakToken>
: Engraved meshes densely isodiametric or slightly transverse on head, pronotum and elytra.
</paragraph>
<paragraph id="29101C7F99C372F0ABF76AAD75181217" pageId="4" pageNumber="85">Head longer than wide, HL/HW=1.15-1.20, eyes small but prominent, frons and vertex convex, frontal impressions large and deep; clypeus bisetose, basal portion unevenly convex; labrum broad at apical margin, sexsetose; palps rather stout, subcylindrical, maxillary palpomere 3 as long as 4, palpomere 4 glabrous, palpomere 3 with two short setae at apex; labial palpomere 2 slightly longer than 3, bisetose on inner margin, with several additional setae at subapex and apex, palpomere 3 with a few tiny setae; ligula small, bisetose at apex; mentum without tooth, each of mentum and submentum bisetose, palpiger asetose. Antennae very long, extending to middle of elytra; pubescent and slightly expanded from basal 1/3 of antennomere 4; antennomere 3 almost as long as 4, and 1.40 times longer than 2; antennomere 1 unisetose on subapex.</paragraph>
<paragraph id="AAD1D7F7C4375B0B32B96838B56AA9E3" pageId="4" pageNumber="85">Pronotum strongly transverse, PW/PL=1.56-1.60, disc moderately convex, fore and basal margins well beaded, sides evenly expanded, widest at about middle; lateral expanded margin well defined, flat, very smooth and hardly reflexed; both transversal impressions distinct, basal foveae small and deep.</paragraph>
<paragraph id="5E19AC4EAA0C0B5C37D67666834DA498" pageId="4" pageNumber="85">Elytra elongate ovate, EL/EW=1.45-1.77, widest at about middle, sides slightly expanded at middle, base completely bordered, apex broadly sinuate, inner angle nearly rectangular, obtuse; striae very deep, intervals strongly convex, subequal in width in middle, interval 3 with basal and middle setiferous pores, the subapical one absent.</paragraph>
<paragraph id="77C5D8901A97823683C152E87B546FA2" pageId="4" pageNumber="85">Legs slender, fore tibia distinctly expanded at apex, apex obliquely truncate, outer margin slightly subserrate; middle and hind coxae smooth and glabrous; middle tibia slender and slightly expanded at apex, strongly curved in male, normal in female; hind femora with two setae posteriorly; hind tibia slightly expanded at apex, apical spurs long and sharp; fore tarsi slightly wider than the middle, and distinctly wider than the hind ones; hind tarsomeres 1 and 3 much longer than 2 and 4 respectively, length ratio of tarsomeres1 to 4 as 3.2: 1.9: 1.5: 1.0; tarsomere 4 asymmetrically emarginated at apex, longer lobe as long as 1/3 of the whole joint; all tarsal claws strongly pectinate.</paragraph>
<paragraph id="7FE7A803876BDCE63FBC4768EAB9A67D" pageId="4" pageNumber="85">Prosternal process well bordered at apex, abdominal ventrite VII of male distinctly and deeply emarginate at apical margin.</paragraph>
<paragraph id="994A9F3EA548C7C9F28EEF76FF279E41" pageId="4" pageNumber="85">Male genitalia (Figs 12-13): Slender and quite straight, slightly sinuate on ventral surface, apex blunt, the apical lamella as long as wide, broad at tip.</paragraph>
<caption id="E09C6A206C03687D577085BABFFD19AF" pageId="4" pageNumber="85">
<paragraph id="385167901A5A6F230429358D15FC72E6" pageId="4" pageNumber="85">
Figures 12-15. Male genitalia of the lancangjiang species group 12
<taxonomicName id="1645F01722C83BB113622DD06671C79E" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
, lateral view 13 idem, dorsal view of apical portion 14
<taxonomicName id="27FBB44E26404462482A722F86D1F4FD" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius euthyphallus" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="euthyphallus">Orthogonius euthyphallus</taxonomicName>
sp. n., lateral view 15 idem, dorsal view of apical portion.
</paragraph>
</caption>
<paragraph id="E2A8D01E61E72A2D5FB3A7A20DC76FFF" pageId="4" pageNumber="85">Female. Similar to male, except the abdominal ventrite VII complete at apical margin, fore tarsomeres 1-3 without spongy setae ventrally, middle tibiae not curved in middle portion, pronotum impunctate, and antennae slightly shorter.</paragraph>
</subSubSection>
<subSubSection id="53E46FD434BC285D863F595146B84771" pageId="4" pageNumber="85" type="variability">
<paragraph id="C80E240CA233CC32244272C74CBADC82" pageId="4" pageNumber="85">Variability.</paragraph>
<paragraph id="1B70416BF6CB219E78B8EC7511FF1EE5" pageId="4" pageNumber="85">Body slender or a little stouter, in general pronotum impunctate in female, but one or two individuals with faint and sparse punctures.</paragraph>
</subSubSection>
<subSubSection id="CB0AE932F38EC212B5B1E79058B5598C" lastPageId="5" lastPageNumber="86" pageId="4" pageNumber="85" type="materials examined">
<paragraph id="1A1597A4258E725D940EB640E259DCF0" pageId="4" pageNumber="85">Materials examined.</paragraph>
<paragraph id="E14355384E513CA893748D215A7F13F4" lastPageId="5" lastPageNumber="86" pageId="4" pageNumber="85">
1 male, the holotype, &quot;Haut Mekong, Vien Poukha, 3. V. 1918, R. V. de Salvaza&quot;, &quot;
<taxonomicName id="DCA2E7B1023333785B71DB1AC20CD056" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius" order="Coleoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">
<pageBreakToken id="8D429FF3E1BA4AD2F2D8848A5FE5DBBB" pageId="5" pageNumber="86" start="start">Orthogonius</pageBreakToken>
</taxonomicName>
sp.&quot; (by Andrewes) and &quot;Brit. Mus. 1921
<normalizedToken id="A44267FE9308BCB6340DDD8A6B6A3481" originalValue="89”">-89&quot;</normalizedToken>
, from Laos, deposited in NHML; 8 males and 7 females, &quot;Laos-NE: Xieng Khouang Province, Phonsaven (30 km NE), Phou Sane Mt., 19.37-38N/ 103.20E, 1400-1500 m, 10.-30.V. 2009, Z. Kraus Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 6 males and 4 females, &quot;Laos-NE: Xieng Khouang Province, Phonsaven (30 km NE), Phou Sane Mt., 19.37-38N/ 103.20-21E, 1400-1700 m, 10.-30.V. 2009, D. Hauck Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 7 males and 8 females,
<normalizedToken id="F2BF9AA41180FA61285B68B4F0E3ED94" originalValue="“LaosNE">&quot;Laos-NE</normalizedToken>
: Xieng Khouang Province, 30 km NE Phonsaven, Phou Sane Mt., 19.37-8N/ 103.20-21E, 1420 m, 30 km NE Phonsaven, Ban Na Lam to Phou Sane Mt., 1300-1700 m, 10.-30.V. 2009, M. Geiser Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 1 male,
<normalizedToken id="764F92F73E7E473B70723363B9A930E9" originalValue="“LaosNE">&quot;Laos-NE</normalizedToken>
: Xieng Khouang Province, 30 km NE Phonsaven, Phou Sane Mt., 19.3820N/ 103.2020E, 1420 m, 10.-30.V. 2009, D. Hauck Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 7 male 4 females,
<normalizedToken id="015AA9734E1E9E0169D88B91AC779CC1" originalValue="“LaosNE">&quot;Laos-NE</normalizedToken>
: Xieng Khouang Province, 30 km NE Phonsaven: Ban Na Lam to Phou Sane Mt., 19.37-38N/ 103.20E, 1300-1500 m, 10.-30.V. 2009, M. Brancucci Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 1 male,
<normalizedToken id="8D1379D6AE16AC3FD1F74121A40CDCB2" originalValue="“LaosNE">&quot;Laos-NE</normalizedToken>
: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.11-13N/ 103.59-104.01E, 1300-1900 m, 9-17.VI. 2009, Michael Geiser Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 2 males and 1 female,
<normalizedToken id="8AAEB061D718B0D32A0A4938E37C1B72" originalValue="“LaosNE">&quot;Laos-NE</normalizedToken>
: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.12-13N/ 103.595-104.01E, 1340-1870 m, 10. V.-16. VI. 2009, M. Brancucci and local coll. Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 2 males, &quot;Laos-NE: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.13N/ 104.00E, 1350-1500 m, 1-16. VI. 2009, M. Brancucci Leg.&quot;, and &quot;NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban&quot;; 8 females,
<normalizedToken id="1387C4937CF63A5DBCC3F0A17D37AA65" originalValue="“LaosNE">&quot;Laos-NE</normalizedToken>
: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.12N/ 104.01E, 1500-1900 m, 17.V.-3.VI. 2007, M. Brancucci Leg.&quot;, and &quot;NHMB Basel Expeditionto Laos, 2007&quot;. All in NHMB, except for eight males and eight females in MNHN and SCAU, respectively.
</paragraph>
</subSubSection>
<subSubSection id="0388A1C012E0F46BC24E7EC47C151A45" pageId="5" pageNumber="86" type="distribution">
<paragraph id="2738C23FDF516BEE970E88E36A61CBC7" pageId="5" pageNumber="86">Distribution.</paragraph>
<paragraph id="522C5A569C9CE0521ADDFEF2A956B888" pageId="5" pageNumber="86">Laos (Fig. 1).</paragraph>
</subSubSection>
</treatment>
</document>