462 lines
62 KiB
XML
462 lines
62 KiB
XML
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<mods:title id="4502A2966C23FE13068ADDA29AFA3D08">Designation of a neotype and redescription of Hesionereticulata von Marenzeller, 1879 from Japan (Annelida, Hesionidae)</mods:title>
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<mods:namePart id="BCC4FC871DF9DCCED7645F7E1F2630CC">Jimi, Naoto</mods:namePart>
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<mods:namePart id="BFB9F47B7E6D0EDB682287A04569D3A3">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:namePart id="662129486AE67CD700A9C8A9BAD5C406">Kajihara, Hiroshi</mods:namePart>
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<mods:date id="CF65A02E80F4B20B26F7F4BE5383F9D2">2017</mods:date>
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<mods:number id="3E2FE65F752AC9E8174AAF185F0574D3">657</mods:number>
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<mods:classification id="21E80D516D64A080A27117E1170AA0C1">journal article</mods:classification>
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<mods:identifier id="1F42F4C337A228FE1AB40B24C19AF801" type="DOI">http://dx.doi.org/10.3897/zookeys.657.11064</mods:identifier>
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<treatment id="0EB7AD7CF5F58467AD3946601FE69D74" ID-GBIF-Taxon="127900156" LSID="urn:lsid:plazi:treatment:0EB7AD7CF5F58467AD3946601FE69D74" httpUri="http://treatment.plazi.org/id/0EB7AD7CF5F58467AD3946601FE69D74" lastPageId="8" lastPageNumber="36" pageId="2" pageNumber="31" scope_family="Hesionidae" scope_phylum="Annelida">
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<subSubSection id="D40E3C51AFEC4C680341F71938FC5CBF" pageId="2" pageNumber="31" type="nomenclature">
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<paragraph id="CB2872A04630A2E886871FC0B7CF60E6" pageId="2" pageNumber="31">
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<taxonomicName id="CB9770A3F1DA1045CB49A3BB0CC6B86F" LSID="Japanese name: otohime-gokai" authority="von Marenzeller, 1879" authorityName="von Marenzeller" authorityYear="1879" family="Hesionidae" genus="Hesione" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hesione reticulata" order="Aciculata" pageId="2" pageNumber="31" rank="species" species="reticulata">Hesione reticulata von Marenzeller, 1879</taxonomicName>
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Figs 1, 2, 3
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</paragraph>
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</subSubSection>
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<subSubSection id="9ED02FB875F0610D37DA51CA09348C64" pageId="2" pageNumber="31" type="reference_group">
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<paragraph id="4D2FC3C76AC95BAE40B2AC95FA5D4398" pageId="2" pageNumber="31">
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<taxonomicName id="98C38C9555CC4E0BAFF08A0DA6ED62CF" family="Hesionidae" genus="Hesione" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hesione reticulata" order="Aciculata" pageId="2" pageNumber="31" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
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von Marenzeller, 1879: 129-131, pl. 3, fig. 4;
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<bibRefCitation id="BEA96821DBB49B54097676F408BD9A2D" author="Izuka, A" journalOrPublisher="Journal of the College of Science (Tokyo)" pageId="9" pageNumber="38" pagination="1 - 262" title="The errantiate Polychaeta of Japan." url="http://hdl.handle.net/2261/32884" volume="30" year="1912">Izuka 1912</bibRefCitation>
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: 192-194, pl. 2, fig. 7;
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<bibRefCitation id="1768CE9E69605A9C8F4A8FE73C20214D" author="Imajima, M" journalOrPublisher="Allan Hancock Foundation Publications, Occasional Paper" pageId="9" pageNumber="38" pagination="1 - 166" title="The polychaetous annelids of Japan, 1." url="http://cdm15799.contentdm.oclc.org/cdm/ref/collection/p15799coll82/id/18946" volume="26" year="1964">Imajima and Hartman 1964</bibRefCitation>
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: 80;
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<bibRefCitation id="91523EB518A353E3CA09C2E07C518C3C" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida 2009</bibRefCitation>
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: 36-37, fig. 1.
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</paragraph>
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<paragraph id="600A8CB23EA08E482AD9AA13BD766CC0" pageId="2" pageNumber="31">
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<taxonomicName id="675569BD7C3EBD1E6C85C203E28E978B" family="Hesionidae" genus="Hesione" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hesione splendida" order="Aciculata" pageId="2" pageNumber="31" rank="species" species="splendida">Hesione splendida</taxonomicName>
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<bibRefCitation id="536002239DFA7B7E0A0018CBF6FFCC08" author="Hessle, C" journalOrPublisher="Arkiv foer Zoologi" pageId="9" pageNumber="38" pagination="1 - 36" title="Einiges ueber die Hesioniden und die Stellung der Gattung Ancistrosyllis." volume="17" year="1925">Hessle 1925</bibRefCitation>
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: 13-15 (non Savigny in Lamarck, 1818; partim, smallest specimen with transverse white bands belongs elsewhere).
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</paragraph>
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<paragraph id="6FB1DA2538A407AF893550D47B2039AB" pageId="2" pageNumber="31">
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<taxonomicName id="2DCA909A0B80406FEE3EB26911B5AE85" family="Hesionidae" genus="Hesione" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hesione pantherina" order="Aciculata" pageId="2" pageNumber="31" rank="species" species="pantherina">Hesione pantherina</taxonomicName>
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<bibRefCitation id="F715BA8BDBA11B939B408AE49A4B11B8" author="Fauvel, P" journalOrPublisher="Memoirs of the College of Science, Kyoto Imperial University, Series B" pageId="9" pageNumber="38" pagination="41 - 92" title="Annelides polychetes du Japon." volume="12" year="1937">Fauvel 1937</bibRefCitation>
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: 59-60 (non Risso, 1826).
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</paragraph>
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<paragraph id="8C50B4427808353AC71F657AE3C965E2" pageId="2" pageNumber="31">
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<taxonomicName id="5D514562991EFCA16586EC29A8F8EDE7" family="Hesionidae" genus="Hesione" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hesione intertexta" order="Aciculata" pageId="2" pageNumber="31" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
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<bibRefCitation id="91659F7137BD2A5C50CFE49FDE1BA5B2" author="Wu, B" journalOrPublisher="Studia Marina Sinica" pageId="10" pageNumber="39" pagination="65 - 104" title="[Preliminary report of polychaetous annelids from Xisha Islands, Guangdong Province, China]." volume="10" year="1975">Wu et al. 1975</bibRefCitation>
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: 75, pl. 2, figs 7-8 (non Grube, 1878).
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</paragraph>
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</subSubSection>
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<subSubSection id="CDE2D28D7C0471A315B626D92C07E51E" pageId="2" pageNumber="31" type="type material">
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<paragraph id="C28D01E41370A5F115E01F7F32F5671B" pageId="2" pageNumber="31">Type material.</paragraph>
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<paragraph id="0440CBE3226188ABFDD452772C0A5DCF" pageId="2" pageNumber="31">
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Northwestern Pacific, Japan. Neotype NSMT Pol N-620, and three paraneotypes NSMT Pol 113205, NSMT Pol 113206, NSMT Pol 113207, Zaimokuza (
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<geoCoordinate id="8B03903E4B76A2943ECC124711838837" direction="north" orientation="latitude" precision="1" value="35.300804">35°18'02.9"N</geoCoordinate>
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,
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<geoCoordinate id="E61768A18D85AAC82F112FAB7B1EBB92" direction="east" orientation="longitude" precision="1" value="139.55081">139°33'02.9"E</geoCoordinate>
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), rocky bottom, 1 m depth, 19 Mar. 2016, N. Jimi &
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<taxonomicName id="5D91B777BDB7574916F9CFB7D3915A39" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="2" pageNumber="31" phylum="Annelida" rank="genus">Hesione</taxonomicName>
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Tanaka, coll. Paraneotypes (NSMT Pol 113205 - NSMT Pol 113207) 40-47 mm long, 4 mm wide).
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</paragraph>
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</subSubSection>
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<subSubSection id="C55281C24269BCD8861301696B457475" pageId="2" pageNumber="31" type="neotype locality">
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<paragraph id="C0C735F61BAE6ECE07FE28BC2E3FD61C" pageId="2" pageNumber="31">Neotype locality.</paragraph>
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<paragraph id="7CBB41D7BC343213A06812846ECC79D1" pageId="2" pageNumber="31">
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Zaimokuza (
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<geoCoordinate id="9CF5CF4709734FC183417DB15A4ABEB1" direction="north" orientation="latitude" precision="1" value="35.300804">35°18'02.9"N</geoCoordinate>
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,
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<geoCoordinate id="1255374565C904689460DCB54BA720E3" direction="east" orientation="longitude" precision="1" value="139.55081">139°33'02.9"E</geoCoordinate>
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), rocky bottom, 1 m depth.
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</paragraph>
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</subSubSection>
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<subSubSection id="0C88F5FA5A127CCA9399F2FD72E722C2" lastPageId="4" lastPageNumber="33" pageId="2" pageNumber="31" type="description">
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<paragraph id="0D537AC27E7A298A2677B0223695597A" pageId="2" pageNumber="31">Description.</paragraph>
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<paragraph id="3FB9F12E131B248F36A22592C497ABD7" pageId="2" pageNumber="31">Neotype (NSMT Pol N-620) complete. Body cylindrical, medially swollen (Fig. 1A), damaged, 43 mm long, 4 mm wide in chaetigers 8-9 (not including parapodia), 16 chaetigers (chaetae and parapodia of 2nd left, 8th right, and 9th right chaetigers removed for observation; dorsal cirri of 3rd and 7th chaetigers removed for DNA extraction).</paragraph>
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<paragraph id="00B30D8094E7638205160C44DC6D2FAF" pageId="2" pageNumber="31">
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Dorsal pigmentation pattern consisting of longitudinal, brownish, subcontinuous, irregular lines; no reddish brown longitudinal broken line on median line; single, irregularly-shaped spot (formed by absence of brown pigment, through which basement pale tan to wheat body color seen) on each chaetiger except 2nd, arranged mid-dorsally (larger anteriorly; reduced medially and posteriorly); and additional row of similar but smaller spots on lateral cushion on each side; silvery white spots absent. Cirrophores yellow; cirrostyles yellow to whitish; parapodial lobes whitish (Fig. 1
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<normalizedToken id="82008EAB3BD7C3C66FB5353BD09EFB9D" originalValue="B–E">B-E</normalizedToken>
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). After six months in ethanol, pigmentation limited to dorsal, pale brown, discontinuous longitudinal bands (Fig. 2A).
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</paragraph>
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<paragraph id="A75479C76466286C83DFDEA3413DCFA8" pageId="2" pageNumber="31">Integument smooth, annulated, giving impression of being tuberculated, especially along posterior region; longitudinal ridges absent in lateral cushions.</paragraph>
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<paragraph id="75659E816A0E67F298307CF414B14752" pageId="2" pageNumber="31">Prostomium heart-shaped, wider than long (Fig. 1B, E); anterior margin truncated; lateral margins rounded in anterior body, but expanded posteriorly; posterior margin cleft, as long as 1/6 prostomial length; longitudinal furrow shallow; dark transverse line present on prostomial anterior margin. Antennae digitate, twice longer than wide. Eyes blackish, on center of prostomium; anterior and posterior eyes in trapezoidal arrangement; anterior eyes slightly more separated than posterior eyes; anterior eyes ovoid (appearing longer than wide), posterior eyes rounded.</paragraph>
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<paragraph id="27261901A7ED75953C0DF400E7BDA1CC" pageId="3" pageNumber="32">
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<pageBreakToken id="3332D89AD9C17EF4B0A47EFBC243E3E6" pageId="3" pageNumber="32" start="start">Tentacular</pageBreakToken>
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cirri tapered, longest complete anterior cirri reaching chaetiger 5. Lateral cushions slightly projected, entire, with smooth surface.
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</paragraph>
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<paragraph id="0B861B8C846E9F1632481DBCA29FCACA" pageId="3" pageNumber="32">Parapodia with dorsal cirrophore twice longer than wide, articulated (Fig. 3A, E). Cirrostyle basally cylindrical, medially and distally articulated, as long as body width, including parapodia (Fig. 3A, F). Neuropodia with parallel sides, cylindrical (Fig. 3A). Acicular lobe double; upper tine twice larger than lower one, digitate (Fig. 3B); lower tine of 8th chaetiger of NSMT Pol N-620 and 9th chaetiger of NSMT Pol 113205 adhered or fused to upper tine and difficult to observe (Fig. 3C), it can be clearly confirmed on other four parapodia examined. One acicula present, blackish. Neurochaetae 19-28 per bundle, blade size decreasing ventrally (Fig. 3C); neurochaetal blades bidentate, 3-4 times longer than wide, subdistal tooth shorter and wider than distal one; guard reaching apical tooth (Fig. 3D). Ventral cirrophore three times wider than long; cirrostyle articulated, surpassing chaetal lobe tip.</paragraph>
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<paragraph id="F0B0C4BB15E292B5DF0F582C3618C831" pageId="4" pageNumber="33">
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<pageBreakToken id="2819C321F64E568BC1140800CB32A509" pageId="4" pageNumber="33" start="start">Prepygidial</pageBreakToken>
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segment with two cirri, three times as long as body width of previous chaetiger (chaetiger 16). Pygidium smooth, trapezoidal, as long as wide, cylindrical (Fig. 1D); anus with two anal cirri; anal cirri tapered.
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</paragraph>
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<paragraph id="A680840BABCF38A82D5941B8BEE273B6" pageId="4" pageNumber="33">Venter without pigmentation, with longitudinal midventral depression.</paragraph>
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<paragraph id="7F8EA915717825CBE859B5DBE5AEBAC4" pageId="4" pageNumber="33">Pharynx divided into three rings, with relative lengths 1.5:1.5:1; basal ring with similar pigmentation as anterior end (Fig. 1E); dorsal papilla pale, longer than wide (length: width 1.5:1).</paragraph>
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<paragraph id="AB937AF0E2BC3BBAC12E2380B1A80126" pageId="4" pageNumber="33">Oocytes not visible.</paragraph>
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</subSubSection>
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<subSubSection id="F467D1E79C42E9A29469B514165424FA" pageId="4" pageNumber="33" type="inner anatomy">
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<paragraph id="1A6C6D28F31A277EE78560686B6F25CC" pageId="4" pageNumber="33">Inner anatomy</paragraph>
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<paragraph id="82B7F1D623518D7B8E00A300CF954B62" pageId="4" pageNumber="33">(observed in paraneotypes NSMT Pol 113205-113207). Pharynx and esophagus thick, muscular, yellowish, continuing into darker, shorter stomach; enteric caeca extending anteriorly along two or three chaetigers (Fig. 2B). Stomach contents included gastropod (Fig. 2C) and amphipod remains as prey items.</paragraph>
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<caption id="FF5E0AD896F55466091E33A8CBD8D207" pageId="4" pageNumber="33">
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<paragraph id="82E56271267388199346824D392D855D" pageId="4" pageNumber="33">
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Figure 1.
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<taxonomicName id="CFD46E9D735BF4F0080A1F1CA8C6F2B5" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="4" pageNumber="33" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
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von Marenzeller, 1879, anesthetized living specimens.
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<normalizedToken id="1AA5F9386D695FB14095DD8A58143B77" originalValue="A–D">A-D</normalizedToken>
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(NMST Pol N-620) E (NSMT Pol-113206). A Dorsal view B Anterior end, dorsal view C Medial part D Posterior end, dorsal view E anterior end, dorsal view. Scale bars: 3.2 mm (A); 1.1 mm (
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<normalizedToken id="7CA7E387DC8A694312C08B5CFCFBD7A6" originalValue="B–D">B-D</normalizedToken>
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); 0.7 mm (E).
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</paragraph>
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</caption>
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<caption id="D7A66EE91C6B487939A8D48EC0AF2C88" pageId="4" pageNumber="33">
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<paragraph id="E0E492AF6841F35594A2B44583BFA326" pageId="4" pageNumber="33">
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Figure 2.
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<taxonomicName id="817B4899C0BAA4A84144BA1576D0ED48" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="4" pageNumber="33" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
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Marenzeller, 1879, A (NSMT Pol N-620)
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<normalizedToken id="66C891A1FC703E961F55C18BEA6DF882" originalValue="B–C">B-C</normalizedToken>
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(NSMT Pol-113205) A Anterior end, dorsal view, six months after fixation B Medial part, ventral view after longitudinal dissection (*: enteric caecum, Ph: pharynx, St: stomach) C Same, close up after dissecting stomach, with a gastropod prey (St W: stomach wall). Scale bars: 1.5 mm (A); 1.2 mm (B); 0.6 mm (C).
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</paragraph>
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</caption>
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<caption id="D956057A61AAF421D01E42F1596D2E35" pageId="4" pageNumber="33">
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<paragraph id="C246D20D6CB389E168FAC87A256E1072" pageId="4" pageNumber="33">
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Figure 3.
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<taxonomicName id="0C727C7DC42FB5CD35A6748804291F40" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="4" pageNumber="33" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
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Marenzeller, 1879,
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<normalizedToken id="AE2F94424256709BFF060C83E0F1347A" originalValue="A–F">A-F</normalizedToken>
|
||
(NSMT Pol N-620). A Chaetiger 8, right parapodium, anterior view B Chaetiger 9, right parapodium, anterior view, close-up of acicular lobe C Chaetiger 8, right parapodium, anterior view, close-up of acicular lobe (arrow points to lower tine) and neurochaetal bundle D Same, tip of neurochaetae (inset: blade tip) E Same, close-up of dorsal cirrophore F Same, close-up of cirrostyle distal region. Scale bars: 0.78 mm (A); 0.15 mm (B); 0.17 mm (C), 25
|
||
<normalizedToken id="12D34DCFBCB28BA8DC9E46CCFFA5939E" originalValue="μm">μm</normalizedToken>
|
||
(D); 0.13 mm (E, F).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="AD15521C76CEC80FD4F9494F05F91481" lastPageId="8" lastPageNumber="37" pageId="4" pageNumber="33" type="remarks">
|
||
<paragraph id="B44947870192DB7CA07D39F2EA5BA835" pageId="4" pageNumber="33">Remarks.</paragraph>
|
||
<paragraph id="D80F843484E1512F14BCD0C6CFA71BED" pageId="4" pageNumber="33">
|
||
The nomenclatural status of
|
||
<taxonomicName id="388F9E692FFC83C991F215BB93D08E01" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="4" pageNumber="33" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
von Marenzeller, 1879 has been unclear due to several reasons: there is no type material, some diagnostic features were not clarified in the original description, the species has been recorded from the Red Sea, and some authors have regarded it as a junior synonym of other species within the genus. Consequently, in order to comply with the International Code of Zoological Nomenclature (
|
||
<bibRefCitation id="D82BB95DBBC69F4F5162C2FE325A03D3" author="Imajima, M" journalOrPublisher="International Trust for Zoological Nomenclature, London" pageId="9" pageNumber="38" title="International Code of Zoological Nomenclature (4 th edn)" year="1999">ICZN 1999</bibRefCitation>
|
||
, Art. 75.3) we are herein proposing a neotype.
|
||
</paragraph>
|
||
<paragraph id="C8D1CA3F07DF3959B19A8B590A0C0923" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
|
||
The above description and illustrations will clarify the taxonomic status of
|
||
<taxonomicName id="48AFEB2977D529E9C86B2E0363610854" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="4" pageNumber="33" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
(Art.75.3.1), and its diagnostic and differential features have been included in the description and illustrations (Arts 75.3.2, 75.3.2), and will be contrasted below. Our enquiries on the existence of type material of
|
||
<taxonomicName id="83EDF4B562B8AB3833461200A68A282B" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="4" pageNumber="33" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
to Dr. Helmut Sattmann, Curator of Marine Invertebrates, in the Naturhistorisches Museum, Vienna, where
|
||
<pageBreakToken id="1F349E24681B539FA22500A3AF191442" pageId="5" pageNumber="34" start="start">Emil</pageBreakToken>
|
||
von Marenzeller used to work and produced all of his publications, indicate that type material is absent (Art. 75.3.4), probably destroyed or never deposited. The original collector was Carl Koerbl (
|
||
<bibRefCitation id="D922E4E01D217427EEF5D5A0373F0FE7" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">von Marenzeller 1879</bibRefCitation>
|
||
: 131) and some of his specimens were donated to the Vienna Museum by Richard von Drasche-Wartinberg (
|
||
<bibRefCitation id="3D53E30E963B132F9947848C75F8BB4B" author="Sato, M" journalOrPublisher="Zoological Science" pageId="9" pageNumber="38" pagination="369 - 372" title="Extirpation of Hedistejaponica (Izuka, 1908) (Nereididae, Polychaeta) in Central Japan, evidenced by a museum historical collection." url="https://doi.org/10.2108/zsj.26.369" volume="26" year="2009">Sato and Sattmann 2009</bibRefCitation>
|
||
), but there is no type material available there.
|
||
</paragraph>
|
||
<paragraph id="EC27CC1210A5E0B814398D27F326F8F5" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
|
||
The neotype fits the original description and because it was recently collected, it even matches the general pigmentation pattern which is not long-lasting in ethanol. Further, as happens in some other species of
|
||
<taxonomicName id="6EAE89BF025D36025A90EFA67B2FE539" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="5" pageNumber="34" phylum="Annelida" rank="genus">Hesione</taxonomicName>
|
||
, they are simultaneous hermaphrodites
|
||
<pageBreakToken id="A4312362267602DB938BE34F55072D4C" pageId="6" pageNumber="35" start="start">(</pageBreakToken>
|
||
<bibRefCitation id="0C2E12A57BF785D3E7E74DF7864B77C7" author="Bergmann, W" journalOrPublisher="Zeitschrift fuer wissenschaftliche Zoologie" pageId="8" pageNumber="37" pagination="278 - 301" title="Untersuchungen ueber die Eibildung bei Anneliden und Cephalopoden." url="http://biodiversitylibrary.org/page/43239588" volume="73" year="1902">Bergmann 1902</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="DFBE65B4CA89FB580C560A092279BA72" author="Bergmann, W" journalOrPublisher="Zoologischer Anzeiger" pageId="8" pageNumber="37" pagination="415 - 417" title="Ueber das spaetere Schicksal der Zwitterdruessen von Hesionesicula." url="http://biodiversitylibrary.org/page/30125385" volume="26" year="1903">1903</bibRefCitation>
|
||
), so that differences in pigmentation pattern among different specimens cannot be attributed to sex (Art. 75.3.5). Thus, the morphology of our material does not contradict
|
||
<bibRefCitation id="B66731A773C9E3844D8CB07662071E62" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">
|
||
von
|
||
<normalizedToken id="CAB966945C24435105AEBD16B7B7655A" originalValue="Marenzeller’s">Marenzeller's</normalizedToken>
|
||
(1879)
|
||
</bibRefCitation>
|
||
original description of
|
||
<taxonomicName id="21F2439CCC91ABA1CBA1F048AD569336" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
, nor the general features subsequently described by
|
||
<bibRefCitation id="0D4072282EB72F821A617B3079A77B30" author="Izuka, A" journalOrPublisher="Journal of the College of Science (Tokyo)" pageId="9" pageNumber="38" pagination="1 - 262" title="The errantiate Polychaeta of Japan." url="http://hdl.handle.net/2261/32884" volume="30" year="1912">Izuka (1912</bibRefCitation>
|
||
: 192),
|
||
<bibRefCitation id="07EFB9ED2E753E18E859E2C9D4843E95" author="Imajima, M" journalOrPublisher="Allan Hancock Foundation Publications, Occasional Paper" pageId="9" pageNumber="38" pagination="1 - 166" title="The polychaetous annelids of Japan, 1." url="http://cdm15799.contentdm.oclc.org/cdm/ref/collection/p15799coll82/id/18946" volume="26" year="1964">Imajima and Hartman (1964</bibRefCitation>
|
||
: 80), and
|
||
<bibRefCitation id="90BF825C7F9C99214F07A1FFFA920861" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida (2009</bibRefCitation>
|
||
: 36-37).
|
||
</paragraph>
|
||
<paragraph id="8066F5021CFA488E0E157F15ABCDBDE6" pageId="6" pageNumber="35">
|
||
The original specimen was collected in the east coast of Enoshima (
|
||
<geoCoordinate id="4F7F899094F011326067DD196BCD6B8A" direction="north" orientation="latitude" precision="15" value="35.301945">35°18'07"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="47055FD858D7DBCECCED685DA7FD3A2D" direction="east" orientation="longitude" precision="15" value="139.48334">139°29'00"E</geoCoordinate>
|
||
), and the neotype was found in Zaimokuza (
|
||
<geoCoordinate id="638CE22030F5B9DB1664217BE1BCBA0C" direction="north" orientation="latitude" precision="1" value="35.300804">35°18'02.9"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B3CE801F13B4179FDD0CD58966487A3B" direction="east" orientation="longitude" precision="1" value="139.55081">139°33'02.9"E</geoCoordinate>
|
||
), nearly four kilometers away, such that we are confident these two localities belong to the same ecological unit (Art. 75.3.6). The neotype of
|
||
<taxonomicName id="D54DEBFF28C58246823B77AB61812575" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
has been deposited in the National Science Museum, Tokyo, which holds the most important polychaete collection in Japan, and has a very important tradition in the scientific study of polychaetes from Japan and elsewhere (Art. 75.3.7).
|
||
</paragraph>
|
||
<paragraph id="059D1314CAFF7B50C23B972A29DD0B28" pageId="6" pageNumber="35">
|
||
Another taxonomic relevance of our study lies in the identification of the dorsal color pattern in the living state as a clear distinguishing feature between
|
||
<taxonomicName id="6E30056EFC967A1DAC62285724C0BFC5" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
,
|
||
<taxonomicName id="AC573A91640192DAB508E7E423F4397B" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EA8FF67EF7A6EED2DF3F6B106076251A" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione ehlersi" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="ehlersi">Hesione cf. ehlersi</taxonomicName>
|
||
sensu
|
||
<bibRefCitation id="88F27D4ACC539D1795E9CC17552C22BB" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida (2009)</bibRefCitation>
|
||
. The color pattern agrees with
|
||
<bibRefCitation id="2D7091304AEF92F3DC2EE596B5951BC7" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">
|
||
von
|
||
<normalizedToken id="61073F7663CF2D442904C40C1A55D835" originalValue="Marenzeller’s">Marenzeller's</normalizedToken>
|
||
(1879)
|
||
</bibRefCitation>
|
||
description of the holotype which had, over a reddish-brown background, irregular spots fused into wide bands along some anterior segments continuing to the end of the body.
|
||
<bibRefCitation id="0C52B30FC0C8DA94FEA9C425CAF3D322" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">von Marenzeller (1879)</bibRefCitation>
|
||
mostly relied on this complex reticulated pigmentation pattern for justifying the establishment of
|
||
<taxonomicName id="9574768855ADE8E1D0BD6A7EB1600C0F" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C6E1D155BA6092BB8CDA15F25C952F7" pageId="6" pageNumber="35">
|
||
<bibRefCitation id="977542AA33AFC547EF9B5E17284575F1" author="Ngamniyom, A" journalOrPublisher="Kasetsart Journal, Natural Sciences" pageId="9" pageNumber="38" pagination="719 - 728" title="Morphological and molecular evidence for a new record of Hesionecf. picta (Polychaeta: Hesionidae) from the Western coast of the Gulf of Thailand." url="http://www.thaiscience.info/journals/Article/TKJN/10961404.pdf" volume="48" year="2014">Ngamniyom et al. (2014)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="D77ED2A8D8777BA3C997F7011DCCAE05" author="Lee, Y" journalOrPublisher="Singapore Biodiversity Records" pageId="9" pageNumber="38" pagination="201 - 204" title="New records of two hesionid polychaetes from the Singapore Strait." url="https://lkcnhm.nus.edu.sg/nus/images/pdfs/sbr/2015/sbr2015-201-204.pdf" volume="2015" year="2015">Lee and Ong (2015)</bibRefCitation>
|
||
characterized the two western Pacific species,
|
||
<taxonomicName id="652E60E8657734DC55036C1B20BFDAEC" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione picta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="picta">Hesione cf. picta</taxonomicName>
|
||
and
|
||
<taxonomicName id="12566DDCEBBF8941C8C9E7E21E2832B3" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
. The former has wide dorsal transverse bands, by which
|
||
<taxonomicName id="FE2C7DBF3DA7860C4D891B7BB9F65F53" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione picta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="picta">Hesione cf. picta</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="2E81B20D4651123E358D6D3D9AE4E2E2" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CE9F738026532EE896191CC1CA90597" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
, because the latter two have dorsal, longitudinal, discontinuous dark bands with paler spots mid-dorsally and along dorsal surface of lateral cushions. Furthermore,
|
||
<taxonomicName id="B5B0FF35EA74D7439770986E1C9F4EB4" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
and
|
||
<taxonomicName id="3C6BFCD939D50FF6722F34EF5589F3CA" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
also have tiny antennae and neurochaetal blades with guards approaching distal tooth. Based on these shared characteristics,
|
||
<bibRefCitation id="69454D43FE20FA717058A9B7D038E67D" author="Wu, B" journalOrPublisher="Studia Marina Sinica" pageId="10" pageNumber="39" pagination="65 - 104" title="[Preliminary report of polychaetous annelids from Xisha Islands, Guangdong Province, China]." volume="10" year="1975">Wu et al. (1975)</bibRefCitation>
|
||
viewed
|
||
<taxonomicName id="3D9883387BFFB70CD9CC802A5A8D6B2B" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="723040AEAA101D06CDBD0263BB157C78" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
. Our observation, however, clearly shows that they differ in pigmentation pattern: in
|
||
<taxonomicName id="94111587292CA549A94E6B824CF38A1A" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
the paler spots are smaller, and the mid-dorsal ones tend to be round, whereas in
|
||
<taxonomicName id="A5EFEABB27DB296E061C00051A44F8FD" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
they are longer than wide and markedly larger.
|
||
</paragraph>
|
||
<paragraph id="4182A6582CA3DF2032E173534BFB64A0" pageId="6" pageNumber="35">
|
||
<bibRefCitation id="63DF82957800A05417D8F49E27E3466F" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida (2009)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4650A31411AEB4C9427123391A36ADE4" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione ehlersi" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="ehlersi">Hesione cf. ehlersi</taxonomicName>
|
||
, a species with similar morphological features to
|
||
<taxonomicName id="38C5C26AA0629059BEB4A2B3DD49EDA4" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
. Indeed, von
|
||
<normalizedToken id="E8CE8ACC59D98FDFD37152AB399289E4" originalValue="Marenzeller’s">Marenzeller's</normalizedToken>
|
||
original description of
|
||
<taxonomicName id="C7CB1F49E344ED8D7C93C3426B0D4075" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
could apply to both species. As
|
||
<bibRefCitation id="914974B210EFA82C9E0E40A9D5136FF6" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida (2009)</bibRefCitation>
|
||
stated, dorsal pigmentation in life is useful for discrimination of the two species;
|
||
<taxonomicName id="FC4D018A84CA214C0CF4D0CD5D1650EF" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione ehlersi" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="ehlersi">Hesione cf. ehlersi</taxonomicName>
|
||
has a reddish brown longitudinal broken line on the median line, whereas
|
||
<taxonomicName id="7BD34BC7F1E134A383C8A9E6272BA457" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
lacks this line. Further study is needed to resolve the taxonomic position of
|
||
<taxonomicName id="3C54C42287FAEDBD793B1255D4FFEBCB" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione ehlersi" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="ehlersi">Hesione cf. ehlersi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="82C0635E7952027D6A5CC104945F80B2" pageId="6" pageNumber="35">
|
||
The vivid images of the dorsal color pattern in
|
||
<taxonomicName id="B57181B08B1BAB8BA6E47705AC570668" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
, along with the COI barcoding sequence provided in this paper, will contribute to future taxonomic revision of the genus
|
||
<taxonomicName id="4613264B70B0FF27393A24D636B8D104" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="genus">Hesione</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="00E8430DA59069417B76B6154C917FC5" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35">
|
||
<taxonomicName id="159B9470498DDE5D12BF27965678ABF1" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
was regarded as a distinct species by
|
||
<bibRefCitation id="2452048C1BCB1D9A32BFB2224422270E" author="Hartman, O" journalOrPublisher="Allan Hancock Foundation Publications, Occasional Paper" pageId="9" pageNumber="38" pagination="1 - 628" title="Catalogue of the Polychaetous Annelids of the World." url="http://digitallibrary.usc.edu/cdm/ref/collection/p15799coll82/id/19573" volume="23" year="1959">Hartman (1959</bibRefCitation>
|
||
: 185) and it can be distinguished from its former synonyms
|
||
<taxonomicName id="E8838A3EFAB499E9304E9464DD09E529" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
,
|
||
<taxonomicName id="DA2FB2507CFF54E437BCF71CEC92ADF6" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione splendida" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="splendida">Hesione splendida</taxonomicName>
|
||
as indicated by
|
||
<bibRefCitation id="B5CC3DD89423137B2779EABBEE6C2FC7" author="Augener, H" journalOrPublisher="Die Fauna Suedwest - Australiens" pageId="8" pageNumber="37" pagination="63 - 304" title="Polychaeta I, Errantia." url="http://biodiversitylibrary.org/page/7160888" volume="4" year="1913">Augener (1913)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="F9D2762A770606E3A0CC05AC1243EA2A" author="Hessle, C" journalOrPublisher="Arkiv foer Zoologi" pageId="9" pageNumber="38" pagination="1 - 36" title="Einiges ueber die Hesioniden und die Stellung der Gattung Ancistrosyllis." volume="17" year="1925">Hessle (1925)</bibRefCitation>
|
||
, or from
|
||
<taxonomicName id="C426AB82E30D3F002C9AB5E83B18671C" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione pantherina" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="pantherina">Hesione pantherina</taxonomicName>
|
||
as suggested by
|
||
<bibRefCitation id="190C8A3337DE0FE6BC55978E8BC6832B" author="Fauvel, P" journalOrPublisher="Memoirs of the College of Science, Kyoto Imperial University, Series B" pageId="9" pageNumber="38" pagination="41 - 92" title="Annelides polychetes du Japon." volume="12" year="1937">Fauvel (1937)</bibRefCitation>
|
||
as follows: from
|
||
<taxonomicName id="A6C9762D17ED30484625B53010E11FD4" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione splendida" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="splendida">Hesione splendida</taxonomicName>
|
||
,
|
||
<taxonomicName id="D46E5B0EF9E637092104C9CE0E11808F" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
can be separated by the dorsal pigmentation; it is brownish in
|
||
<taxonomicName id="3DB955AFC8CCD957B4290BD27DDC45AC" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
, but pearly gray in
|
||
<taxonomicName id="8D2364C56831AF34ABD363DE73F5B5AA" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione splendida" order="Phyllodocida" pageId="6" pageNumber="35" phylum="Annelida" rank="species" species="splendida">Hesione splendida</taxonomicName>
|
||
(Savigny,
|
||
<pageBreakToken id="2852048091CDE30A0634A1F33E102760" pageId="7" pageNumber="36" start="start">1822</pageBreakToken>
|
||
), whereas from
|
||
<taxonomicName id="56CEAA5CD62442C1F83F63D7AF89EA0E" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione pantherina" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="pantherina">Hesione pantherina</taxonomicName>
|
||
,
|
||
<taxonomicName id="4F9E527DC02FD5093AF7C8591DCBD27B" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
can be distinguished because the guard tooth in
|
||
<taxonomicName id="D0BDE1F3FCD506E8C442ED56690A3A70" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
reaches the apical tooth, whereas those in
|
||
<taxonomicName id="E31C6D1FDC6DD9C096D2451CBCD7F8B5" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione pantherina" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="pantherina">Hesione pantherina</taxonomicName>
|
||
do not (
|
||
<bibRefCitation id="FEEA1349D6CEF55E5DC67840EF29278E" author="Monro, CCA" journalOrPublisher="Journal of the Linnean Society of London, Zoology" pageId="9" pageNumber="38" pagination="311 - 323" title="Polychaeta of the ' Alert' Expedition. Families Hesionidae and Nereidae." url="https://doi.org/10.1111/j.1096-3642.1926.tb02172.x" volume="36" year="1926">Monro 1926</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="737B88F85F9DC9685E88231C559D1106" pageId="7" pageNumber="36">
|
||
One of the important discoveries in our observation of the specimens of
|
||
<taxonomicName id="380B96E20B60600A8466720351FE26B5" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
is that the acicular lobe in this species is doubled, comprised of the upper and lower tines, a character state that separates
|
||
<taxonomicName id="C6E78B21F8D10080AD649F018EF76435" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="genus">Hesione</taxonomicName>
|
||
species in two groups, each with approximately the same number of species (SISV pers. obs.).
|
||
<bibRefCitation id="F755640D06511C76FD034F9FDCE8E3BB" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">von Marenzeller (1879</bibRefCitation>
|
||
, fig. 4) illustrated a parapodium excised from the middle part of the body in the holotype specimen, indicating that there was a single, thick, finger-shaped acicular lobe, unlike the doubled lobe that we observed in this study.
|
||
<bibRefCitation id="2DE24D40D85F50FD7323EE8997DA91D8" author="Izuka, A" journalOrPublisher="Journal of the College of Science (Tokyo)" pageId="9" pageNumber="38" pagination="1 - 262" title="The errantiate Polychaeta of Japan." url="http://hdl.handle.net/2261/32884" volume="30" year="1912">Izuka (1912)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="506CB5A88E3B023C39ADD48EC6B0661D" author="Imajima, M" journalOrPublisher="Allan Hancock Foundation Publications, Occasional Paper" pageId="9" pageNumber="38" pagination="1 - 166" title="The polychaetous annelids of Japan, 1." url="http://cdm15799.contentdm.oclc.org/cdm/ref/collection/p15799coll82/id/18946" volume="26" year="1964">Imajima and Hartman (1964)</bibRefCitation>
|
||
also described the acicular lobe as a single lobe. In two of the six parapodia examined (left one on the 2nd, right one on the 8th, and right one on the 9th chaetigers from NSMT Pol N-620; right one on the 9th chaetiger from NSMT Pol 113205; and left ones on the 3rd and 9th chaetigers from NSMT Pol 113207), the lower tine adhered to the upper tine. It appeared as if it were a single parapodial lobe, but a careful observation showed that it actually represents a doubled lobe. The reason the acicular lobe was described as
|
||
<normalizedToken id="B80B0DABE12FDA8DB9808A131CCDD567" originalValue="‘single’">'single'</normalizedToken>
|
||
in the previous studies may be that the lower tine in their material was deformed in preservation to lie below the upper tine, or to contact closely to the upper tine. The original illustration (
|
||
<bibRefCitation id="3FA974F46E8402D8A0F341D48C2B07D4" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">von Marenzeller 1879</bibRefCitation>
|
||
, fig. 4) clearly indicates that the acicular lobe was placed under the chaetal bundles on the glass slide. This must have made the acicular lobe difficult to be observed, which would also explain why the adjacent upper and lower tines were hardly detected. This feature further adds to the distinction between
|
||
<taxonomicName id="0363AB22A58D3A6FD83FE4D37A11A562" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
and
|
||
<taxonomicName id="CA128D39692A645AD539A6DFAF749CCD" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
: the acicular lobe in
|
||
<taxonomicName id="A6D8E7573A9CA21E363150F14CA845A3" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
is double whereas it is single in
|
||
<taxonomicName id="F06F08A8AEE90DAD7EACECAA11BF7A33" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="936D3DC86090F2A32E5859CE682657AB" pageId="7" pageNumber="36">
|
||
The record of
|
||
<taxonomicName id="C7FE5BF7718A22B01D168B29AACE1793" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
by
|
||
<bibRefCitation id="88FA6394C2919ADF62B5EF343C996AC1" author="Imajima, M" journalOrPublisher="National Science Museum Monographs (Tokyo)" pageId="9" pageNumber="38" pagination="149 - 228" title="Polychaetous annelids of Suruga Bay, Central Japan." url="http://ci.nii.ac.jp/els/110004312462.pdf?id=ART0006480764&type=pdf&lang=en&host=cinii&order_no=&ppv_type=0&lang_sw=&no=1476474983&cp=" volume="12" year="1997">Imajima (1997</bibRefCitation>
|
||
: 171) might not belong to the same species because he indicated that the acicular lobe was single ("a superior conical papilla"), and because unlike our specimens, his material was collected from 230-250 m depth in Suruga Bay. Other specimens recorded as
|
||
<taxonomicName id="BFA33ABDB51F881122A8183D54C32FA5" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
by
|
||
<bibRefCitation id="F4B942C19DEC4CC1E579B66E705A5556" author="Imajima, M" journalOrPublisher="National Science Museum Monographs" pageId="9" pageNumber="38" pagination="1 - 221" title="Polychaetous annelids from Sagami Bay and Sagami Sea collected by the Emperor Showa of Japan and deposited at the Showa Memorial Institute, National Science Museum, Tokyo, 2. Orders included within the Phyllodocida, Amphinomida, Spintherida and Eunicida." url="http://ci.nii.ac.jp/naid/110004708004/en" volume="23" year="2003">Imajima (2003</bibRefCitation>
|
||
: 132-134), collected in shallow water, were characterized as having acicular lobe single ("a superior conical papilla"), and are regarded as belonging to another species.
|
||
</paragraph>
|
||
<paragraph id="9A6CE003662E966AB83960E5712749C9" pageId="7" pageNumber="36">
|
||
Distribution.
|
||
<taxonomicName id="D9003E45A7E33A39A6791111E85FD726" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
has so far been recorded only from Japan: Kanagawa (
|
||
<bibRefCitation id="91C8C1FA473F83C01289120F9D91F411" author="Von Marenzeller, E" journalOrPublisher="Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften" pageId="10" pageNumber="39" pagination="109 - 154" title="Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea." volume="41" year="1879">von Marenzeller 1879</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="50356A9D3BE39874D6C1FB2F54E2A829" author="Izuka, A" journalOrPublisher="Journal of the College of Science (Tokyo)" pageId="9" pageNumber="38" pagination="1 - 262" title="The errantiate Polychaeta of Japan." url="http://hdl.handle.net/2261/32884" volume="30" year="1912">Izuka 1912</bibRefCitation>
|
||
; this study), Shizuoka and Wakayama (
|
||
<bibRefCitation id="3496601366EBF6D18C853C51DBFD4250" author="Izuka, A" journalOrPublisher="Journal of the College of Science (Tokyo)" pageId="9" pageNumber="38" pagination="1 - 262" title="The errantiate Polychaeta of Japan." url="http://hdl.handle.net/2261/32884" volume="30" year="1912">Izuka 1912</bibRefCitation>
|
||
), and the middle of Honshu to Kyushu (
|
||
<bibRefCitation id="27A1CADAF76CDFE5074715F493B616A3" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<subSection id="7DC5943E5D1C49C5627CC2DC67DF3186" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36" type="key to species of hesione from japan">
|
||
<paragraph id="8C65F49A3B44C571DC4BF61BACC42764" pageId="7" pageNumber="36">
|
||
Key to species of
|
||
<taxonomicName id="463A57643109D2ABF7DD23589A918EEF" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="genus">Hesione</taxonomicName>
|
||
from Japan
|
||
</paragraph>
|
||
<paragraph id="3D4C8A0E9B14CC16055E3EDA427B2862" pageId="7" pageNumber="36">
|
||
(modified from
|
||
<bibRefCitation id="69FD1943435DCCC20A872599AD3C2699" author="Uchida, H" journalOrPublisher="Marine Pavilion, Kushimoto Marine Park" pageId="9" pageNumber="38" pagination="36 - 37" title="[Polychaetologica 55: Key to genus and description of species (40), Hesionidae 4]." volume="38" year="2009">Uchida 2009</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="1B8659F661AC41CDA2031FB6FA31DD3C" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36">
|
||
<table id="20D7E3436FF9BF81DC451D496BD99E45" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36">
|
||
<tr id="78986216015A6DB6C79878B1F35BCE8F" pageId="7" pageNumber="36">
|
||
<td id="0D2A42C1C0DDBA6F115582DB1A0DFAF8" colspan="1" pageId="7" pageNumber="36" rowspan="1">2</td>
|
||
</tr>
|
||
<tr id="8CED855E16819745B6B9D1418B0F6C5E" pageId="7" pageNumber="36">
|
||
<td id="067F792BCE8FCB3278994A5DE27860C6" colspan="1" pageId="7" pageNumber="36" rowspan="1">
|
||
<taxonomicName id="733E774BEE74D198CC9A83E7004ADD53" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="genus">Hesione</taxonomicName>
|
||
? sp.
|
||
</td>
|
||
</tr>
|
||
<tr id="C5F04EAFA5A108D0565DD35FEE08ED37" pageId="7" pageNumber="36">
|
||
<td id="1A972B83E177791513C8937219A03F0B" colspan="1" pageId="7" pageNumber="36" rowspan="1">3</td>
|
||
</tr>
|
||
<tr id="F8A6320F4C3C81F1A5DE9B837D8F2473" pageId="7" pageNumber="36">
|
||
<td id="61F2878380F30F105A4B0BD0874F942C" colspan="1" pageId="7" pageNumber="36" rowspan="1">
|
||
<taxonomicName id="F5866BB8D5A7DE1F553415A120103F7A" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione splendida" order="Phyllodocida" pageId="7" pageNumber="36" phylum="Annelida" rank="species" species="splendida">Hesione splendida?</taxonomicName>
|
||
sensu Monro, 1931
|
||
</td>
|
||
</tr>
|
||
<tr id="F565CD555996AACC18D9E928F32BCAB5" pageId="8" pageNumber="37">
|
||
<td id="8A7771FAED0CC8D0347DC99555DEF5BB" colspan="1" pageId="8" pageNumber="37" rowspan="1">
|
||
<pageBreakToken id="A2841F5DF32554D1E8045D469273B91C" pageId="8" pageNumber="37" start="start">4</pageBreakToken>
|
||
</td>
|
||
</tr>
|
||
<tr id="A471EF8174B951C5AEA07B7BF7AF31B3" pageId="8" pageNumber="37">
|
||
<td id="A78FDAFA7B61624FF2BEB87164565D9A" colspan="1" pageId="8" pageNumber="37" rowspan="1">5</td>
|
||
</tr>
|
||
<tr id="58A3F9D804EB600AE5FFBF962AB9DEDC" pageId="8" pageNumber="37">
|
||
<td id="710E1D5E073F93D58BD8DAFC2D1FFB9A" colspan="1" pageId="8" pageNumber="37" rowspan="1">
|
||
<taxonomicName id="91237B786598D81DE7D08111341D44C1" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione genetta" order="Phyllodocida" pageId="8" pageNumber="37" phylum="Annelida" rank="species" species="genetta">Hesione genetta</taxonomicName>
|
||
Grube, 1867
|
||
</td>
|
||
</tr>
|
||
<tr id="4F073C5DDA7404C73A5200B486BB1CDD" pageId="8" pageNumber="37">
|
||
<td id="F86A851B3DC44887C699DC253F2543F6" colspan="1" pageId="8" pageNumber="37" rowspan="1">
|
||
<taxonomicName id="A5D7D6A01C9584784BBA90A905404E61" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione" order="Phyllodocida" pageId="8" pageNumber="37" phylum="Annelida" rank="genus">Hesione</taxonomicName>
|
||
sp.
|
||
</td>
|
||
</tr>
|
||
<tr id="D94C1C221FB0401DE3942B35D25075CB" pageId="8" pageNumber="37">
|
||
<td id="18AA239ABA778752D1C43FD73B5F7020" colspan="1" pageId="8" pageNumber="37" rowspan="1">
|
||
<taxonomicName id="79C1A8C320E27D98AB23F9B78C1EDD9C" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione intertexta" order="Phyllodocida" pageId="8" pageNumber="37" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
|
||
Grube, 1878
|
||
</td>
|
||
</tr>
|
||
<tr id="4D85DB2FA3CAC6163EE0CD1B784E9CA4" pageId="8" pageNumber="37">
|
||
<td id="801113AEC5516B00E1EE5347E7CED3B5" colspan="1" pageId="8" pageNumber="37" rowspan="1">6</td>
|
||
</tr>
|
||
<tr id="7086B7E12CA6AA92F61E0BFC4AD7C11D" pageId="8" pageNumber="37">
|
||
<td id="285CC7BA41382AB58EC2EE341908DD8D" colspan="1" pageId="8" pageNumber="37" rowspan="1">
|
||
<taxonomicName id="181B7F5A18FB49FAC281FCF206A17DBA" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione ehlersi" order="Phyllodocida" pageId="8" pageNumber="37" phylum="Annelida" rank="species" species="ehlersi">Hesione cf. ehlersi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="674C55241CA771F2A082CAAF8C6D36B0" pageId="8" pageNumber="37">
|
||
<td id="5AD440A700C3CA120484DB4EB0A8B1C2" colspan="1" pageId="8" pageNumber="37" rowspan="1">
|
||
<taxonomicName id="04FED7206C5ED82894B8E6C5410F6EF7" class="Polychaeta" family="Hesionidae" genus="Hesione" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesione reticulata" order="Phyllodocida" pageId="8" pageNumber="37" phylum="Annelida" rank="species" species="reticulata">Hesione reticulata</taxonomicName>
|
||
von Marenzeller, 1879
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |