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<mods:title id="193824E07AF6FC1BF3EEBF94AA4E5335">On the genus Ammonius Thorell, 1899 (Mygalomorphae, Barychelidae): description of the female of A. pupulus, a new species and new distribution records</mods:title>
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<mods:namePart id="8571FCD92F0B1127DBE8FD7628F65C07">Gonzalez-Filho, Hector M. O.</mods:namePart>
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<mods:affiliation id="57BABE63A993F4378ECB77B81396047F">Pós-graduação em Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil.</mods:affiliation>
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<mods:affiliation id="7F3F932514F5DBC1FFA4304EF744FEAA">Pós-graduação em Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil. &amp; Departamento de Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil.</mods:affiliation>
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<mods:affiliation id="B5855C24E6D701A457968E1E506A7752">Pós-graduação em Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil.</mods:affiliation>
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Genus 
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<bibRefCitation id="EFA34B5BFFFAFFD5FC8FF9CF799DE6D6" author="Thorell T." box="[824,984,1650,1676]" pageId="2" pageNumber="115" pagination="1 - 105" refId="ref8554" refString="Thorell T. 1899. Araneae Camerunenses (Africae occidentalis) quas anno 1891 collegerunt Cel. Dr. Y. Sjostedt aliique. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 25 (4, 1): 1 - 105. Available from https: // www. biodiversitylibrary. org / page / 13682607 [accessed 8 Feb. 2023]." type="journal article" year="1899">Thorell, 1899</bibRefCitation>
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<taxonomicName id="4C324D29FFFAFFD5FF0AF9097855E697" ID-CoL="TXW" authority="Thorell, 1899: 10" authorityName="Thorell" authorityPageNumber="10" authorityYear="1899" box="[189,528,1715,1742]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAB8FFFAFFD5FF0AF9097B7CE694" box="[189,313,1716,1742]" italics="true" pageId="2" pageNumber="115">Ammonius</emphasis>
<bibRefCitation id="EFA34B5BFFFAFFD5FEF7F90E7855E697" author="Thorell T." box="[320,528,1715,1741]" pageId="2" pageNumber="115" pagination="1 - 105" refId="ref8554" refString="Thorell T. 1899. Araneae Camerunenses (Africae occidentalis) quas anno 1891 collegerunt Cel. Dr. Y. Sjostedt aliique. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 25 (4, 1): 1 - 105. Available from https: // www. biodiversitylibrary. org / page / 13682607 [accessed 8 Feb. 2023]." type="journal article" year="1899">Thorell, 1899: 10</bibRefCitation>
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.
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<taxonomicName id="4C324D29FFFAFFD5FF0AF9487B7CE755" ID-CoL="TXW" authorityName="Thorell" authorityYear="1899" box="[189,313,1781,1807]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAB8FFFAFFD5FF0AF9487B7CE755" box="[189,313,1781,1807]" italics="true" pageId="2" pageNumber="115">Ammonius</emphasis>
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– 
<treatmentCitation id="0A9310BBFFFAFFD5FEE1F9497861E755" author="Simon E." box="[342,548,1780,1807]" page="915" pageId="2" pageNumber="115" year="1903">
<bibRefCitation id="EFA34B5BFFFAFFD5FEE1F9497861E755" author="Simon E." box="[342,548,1780,1807]" pageId="2" pageNumber="115" refId="ref8520" refString="Simon E. 1903. Histoire naturelle des Araignees. Deuxieme edition, tome second. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1903">Simon 1903: 915</bibRefCitation>
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. — 
<treatmentCitation id="0A9310BBFFFAFFD5FDE3F9497941E755" author="Benoit P. L. G." box="[596,772,1780,1807]" page="7" pageId="2" pageNumber="115" year="1965">
<bibRefCitation id="EFA34B5BFFFAFFD5FDE3F9497941E755" author="Benoit P. L. G." box="[596,772,1780,1807]" pageId="2" pageNumber="115" pagination="72 - 78" refId="ref7608" refString="Benoit P. L. G. 1965. Les genres des Barychelidae-Leptopelmatinae africains (Araneae - Orthognatha). Revue de Zoologie et de Botanique africaines 72: 72 - 78." type="journal article" year="1965">Benoit 1965: 7</bibRefCitation>
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. — 
<treatmentCitation id="0A9310BBFFFAFFD5FC81F9487E45E755" author="Raven R. J." box="[822,1024,1780,1807]" page="113" pageId="2" pageNumber="115" year="1985">
<bibRefCitation id="EFA34B5BFFFAFFD5FC81F9487E45E755" author="Raven R. J." box="[822,1024,1780,1807]" pageId="2" pageNumber="115" pagination="1 - 180" refId="ref8328" refString="Raven R. J. 1985. The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History 182: 1 - 180. Available from http: // hdl. handle. net / 2246 / 955 [accessed 8 Feb. 2023]." type="journal article" year="1985">Raven 1985: 113</bibRefCitation>
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.
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species
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<taxonomicName id="4C324D29FFFAFFD5FF0AF8D87803E725" ID-CoL="66CNM" authority="Thorell, 1899" authorityName="Thorell" authorityYear="1899" box="[189,582,1892,1919]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFAFFD5FF0AF8D87BD8E724" box="[189,413,1892,1919]" italics="true" pageId="2" pageNumber="115">Ammonius pupulus</emphasis>
<bibRefCitation id="EFA34B5BFFFAFFD5FE13F8D97803E725" author="Thorell T." box="[420,582,1892,1919]" pageId="2" pageNumber="115" pagination="1 - 105" refId="ref8554" refString="Thorell T. 1899. Araneae Camerunenses (Africae occidentalis) quas anno 1891 collegerunt Cel. Dr. Y. Sjostedt aliique. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 25 (4, 1): 1 - 105. Available from https: // www. biodiversitylibrary. org / page / 13682607 [accessed 8 Feb. 2023]." type="journal article" year="1899">Thorell, 1899</bibRefCitation>
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, by monotypy.
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<emphasis id="B946EAB8FFFBFFD4FF0AFEB57B73E178" bold="true" box="[189,310,264,290]" pageId="3" pageNumber="116">Diagnosis</emphasis>
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<taxonomicName id="4C324D29FFFBFFD4FF0AFE8A7B7CE10B" authorityName="Thorell" authorityYear="1899" box="[189,313,311,337]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B946EAB8FFFBFFD4FF0AFE8A7B7CE10B" box="[189,313,311,337]" italics="true" pageId="3" pageNumber="116">Ammonius</emphasis>
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can be distinguished from the other barychelids by the apical segment of the PLS digitiform (
<figureCitation id="13092A2FFFFBFFD4FF71FEE77B5AE12E" box="[198,287,346,372]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Fig. 1F</figureCitation>
); compact eye group rectangular in shape (
<figureCitation id="13092A2FFFFBFFD4FCABFEE77938E12E" box="[796,893,346,372]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC3CFEE779F5E12E" box="[907,944,346,372]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">4B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC0AFEE779A6E12F" box="[957,995,346,373]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC47FEE77E50E12E" box="[1008,1045,346,372]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9B</figureCitation>
), eye tubercle flat; 
<taxonomicName id="4C324D29FFFBFFD4FB4FFEE67F32E12E" authorityName="Thorell" authorityYear="1899" box="[1272,1399,346,373]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFBFFD4FB4FFEE67F32E12E" box="[1272,1399,346,373]" italics="true" pageId="3" pageNumber="116">A. pupulus</emphasis>
</taxonomicName>
with eight eyes (
<figureCitation id="13092A2FFFFBFFD4FE37FEC07846E1C2" box="[384,515,381,408]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1D–E</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FDA7FEC37870E1C2" box="[528,565,382,408]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">4B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FDF5FEC0782CE1C2" box="[578,617,381,408]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5D</figureCitation>
); absence of AME in 
<taxonomicName id="4C324D29FFFBFFD4FCDAFEC3799AE1CD" authority="Guadanucci &amp; Brescovit, 2023" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[877,991,381,408]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti" status="sp. nov.">
<emphasis id="B946EAB8FFFBFFD4FCDAFEC3799AE1CD" box="[877,991,381,408]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557C3FFFBFFD4FC50FEC37E07E1C2" box="[999,1090,382,408]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13092A2FFFFBFFD4FBE4FEC07EEAE1C2" box="[1107,1199,381,408]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">Fig. 9B</figureCitation>
); male cymbium unequal, with prolateral lobe inconspicuous and retrolateral lobe strongly projected (
<figureCitation id="13092A2FFFFBFFD4FB7FFE1C7F5CE1E1" box="[1224,1305,417,443]" captionStart="Fig" captionStartId="5.[189,232,1856,1882]" captionTargetBox="[219,1367,692,1816]" captionTargetId="figure-108@5.[213,1374,689,1820]" captionTargetPageId="5" captionText="Fig. 2. Male palp of Ammonius pupulus Thorell, 1899. A–D. Holotype, ♂ (NHRS-GULI37848). A. Dorsal view. B. Ventral view. C. Prolateral view. D. Retrolateral view. E–F. SEM micrographs of the male palp of Ammonius pupulus (RMCA 227221). E. Prolateral view. F. Retrolateral view. G. Drawing from Benoit (1965: fig. 1), ventral view. Abbreviation: Tc = Tibial concavity. Scale bars: A–D = 1 mm; E–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737513" httpUri="https://zenodo.org/record/7737513/files/figure.png" pageId="3" pageNumber="116">Figs 2</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FA9DFE1C7F37E1E1" box="[1322,1394,417,443]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[195,1392,640,1843]" captionTargetId="figure-135@6.[189,1398,634,1849]" captionTargetPageId="6" captionText="Fig. 3. SEM micrographs of the male palp of Ammonius pupulus Thorell, 1899 (RMCA 227221). A–B. Tibial concavity. A. Prolateral view. B. Retrolateral view. C–D. Cymbium. C. Prolateral view. D. Retrolateral view. Abbreviations: Em = embolus; PL = prolateral lobe; So = slit organ; Tc = tibial concavity. Scale bars: A = 400 µm; B = 200 µm; C–D = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.7737515" httpUri="https://zenodo.org/record/7737515/files/figure.png" pageId="3" pageNumber="116">3C–D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FF0AFE797B41E184" box="[189,260,452,478]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">8C–E</figureCitation>
). Males resemble 
<taxonomicName id="4C324D29FFFBFFD4FE52FE78797CE184" authority="Simon, 1889" authorityName="Simon" authorityYear="1889" box="[485,825,452,478]" class="Arachnida" family="Barychelidae" genus="Pisenor" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="notius">
<emphasis id="B946EAB8FFFBFFD4FE52FE7878D6E184" box="[485,659,453,478]" italics="true" pageId="3" pageNumber="116">Pisenor notius</emphasis>
Simon, 1889
</taxonomicName>
(see 
<bibRefCitation id="EFA34B5BFFFBFFD4FCC9FE797E52E184" author="Benoit P. L. G." box="[894,1047,452,478]" pageId="3" pageNumber="116" pagination="209 - 241" refId="ref7641" refString="Benoit P. L. G. 1966. Les Barychelidae-Barychelinae africains et malgaches (Aran. - Orthogn.). Revue de Zoologie et de Botanique africaines 74 (3 - 4): 209 - 241." type="journal article" year="1966">Benoit 1966</bibRefCitation>
: figs 24, 26) by the strongly pronounced retrolateral lobe of the cymbium but differ by the lack of a spur on tibia I, and the modified palpal tibia, with a tibial concavity in the ventro-prolateral side of the palp on 
<taxonomicName id="4C324D29FFFBFFD4FBEDFDB67E9DE27E" authorityName="Thorell" authorityYear="1899" box="[1114,1240,522,549]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFBFFD4FBEDFDB67E9DE27E" box="[1114,1240,522,549]" italics="true" pageId="3" pageNumber="116">A. pupulus</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FB5FFDB77F2CE27E" box="[1256,1385,522,548]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[195,1392,640,1843]" captionTargetId="figure-135@6.[189,1398,634,1849]" captionTargetPageId="6" captionText="Fig. 3. SEM micrographs of the male palp of Ammonius pupulus Thorell, 1899 (RMCA 227221). A–B. Tibial concavity. A. Prolateral view. B. Retrolateral view. C–D. Cymbium. C. Prolateral view. D. Retrolateral view. Abbreviations: Em = embolus; PL = prolateral lobe; So = slit organ; Tc = tibial concavity. Scale bars: A = 400 µm; B = 200 µm; C–D = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.7737515" httpUri="https://zenodo.org/record/7737515/files/figure.png" pageId="3" pageNumber="116">Fig. 3A–B</figureCitation>
), and by the presence of a tibial thorn on the retrolateral side of the palp on 
<taxonomicName id="4C324D29FFFBFFD4FB98FD937EE4E21D" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[1071,1185,557,584]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti">
<emphasis id="B946EAB8FFFBFFD4FB98FD937EE4E21D" box="[1071,1185,557,584]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FB06FD907F71E212" box="[1201,1332,557,584]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">Figs 8D–E</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FAF6FD937F2CE212" box="[1345,1385,558,584]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9D</figureCitation>
). Females of the genus can be distinguished from the other genera by the bilobed spermathecae, with the external stalk small and emerging from the basal region of the receptacle (
<figureCitation id="13092A2FFFFBFFD4FB94FDC97E3BE2D5" box="[1059,1150,628,655]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">Fig. 5E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3286521FFFBFFD4FF0AFD087962E608" pageId="3" pageNumber="116" type="description">
<paragraph id="8B8D36AAFFFBFFD4FF0AFD087B0BE295" blockId="3.[189,334,693,719]" box="[189,334,693,719]" pageId="3" pageNumber="116">
<heading id="D0C581C6FFFBFFD4FF0AFD087B0BE295" bold="true" box="[189,334,693,719]" fontSize="11" level="3" pageId="3" pageNumber="116" reason="3">
<emphasis id="B946EAB8FFFBFFD4FF0AFD087B0BE295" bold="true" box="[189,334,693,719]" pageId="3" pageNumber="116">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36AAFFFBFFD4FF0AFD59784AE5B8" blockId="3.[188,1400,740,1506]" pageId="3" pageNumber="116">
Very small spiders, total length between 3.9–7.3. General coloration yellowish light brown (
<figureCitation id="13092A2FFFFBFFD4FB59FD597F34E2A4" box="[1262,1393,740,766]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1A–B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FF0AFCBA7B40E378" box="[189,261,775,802]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5A–B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FEA6FCBA7B19E37B" box="[273,348,775,801]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">8A–B</figureCitation>
). Abdomen dorsally pale without pattern and spinnerets pale with some brown light setae on dorsal side (
<figureCitation id="13092A2FFFFBFFD4FEC6FC977B89E31E" box="[369,460,810,836]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1F</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FE6EFC977845E31E" box="[473,512,810,836]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">8A</figureCitation>
). Carapace ovate (
<figureCitation id="13092A2FFFFBFFD4FD6DFC977979E31E" box="[730,828,810,836]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1A</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FCFEFC977935E31F" box="[841,880,810,837]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5A</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FCCBFC9779E7E31E" box="[892,930,810,836]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">8A</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC18FC97799DE31E" box="[943,984,810,836]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9A</figureCitation>
). Thoracic fovea small and straight (
<figureCitation id="13092A2FFFFBFFD4FF71FCF07B60E332" box="[198,293,845,872]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1A</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FE87FCF07B13E332" box="[304,342,845,872]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5A</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FED7FCF37BCDE332" box="[352,392,846,872]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">8A</figureCitation>
). Clypeus narrow (
<figureCitation id="13092A2FFFFBFFD4FDD6FCF07885E332" box="[609,704,845,872]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FD7BFCF378B5E332" box="[716,752,846,872]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">4B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FD4CFCF07964E332" box="[763,801,845,872]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC9BFCF37917E332" box="[812,850,846,872]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9B</figureCitation>
). Eye tubercle black and flat (
<figureCitation id="13092A2FFFFBFFD4FB13FCF07F46E332" box="[1188,1283,845,872]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FAB9FCF07F73E332" box="[1294,1334,845,872]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5D</figureCitation>
). Eye group in rectangular shape, anterior and posterior eye rows straight (
<figureCitation id="13092A2FFFFBFFD4FC5FFCCC7E0FE3D1" box="[1000,1098,881,907]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FBEFFCCC7E39E3D1" box="[1112,1148,881,907]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">4B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FB3DFCCD7EF5E3D1" box="[1162,1200,880,907]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5D</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FB0AFCCC7EA0E3D1" box="[1213,1253,881,907]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9D</figureCitation>
). 
<taxonomicName id="4C324D29FFFBFFD4FB4CFCCC7B5FE3F4" authorityName="Thorell" authorityYear="1899" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFBFFD4FB4CFCCC7B5FE3F4" italics="true" pageId="3" pageNumber="116">Ammonius pupulus</emphasis>
</taxonomicName>
with eight eyes (
<figureCitation id="13092A2FFFFBFFD4FE53FC297822E3F4" box="[484,615,916,942]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1D–E</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FDC3FC2978DDE3F4" box="[628,664,916,942]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">4B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FD12FC297889E3F5" box="[677,716,916,943]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5D</figureCitation>
); 
<taxonomicName id="4C324D29FFFBFFD4FD53FC297913E3F4" authority="Guadanucci &amp; Brescovit, 2023" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[740,854,916,942]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti" status="sp. nov.">
<emphasis id="B946EAB8FFFBFFD4FD53FC297913E3F4" box="[740,854,916,942]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557C3FFFBFFD4FCE9FC2979FCE3F4" box="[862,953,916,942]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
with six eyes, AME absent (
<figureCitation id="13092A2FFFFBFFD4FABAFC297A96E388" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">Fig. 9A– B</figureCitation>
). Labium rectangular, 2–6 cuspules rounded. Maxillae rectangular with 3–20 cuspules on inner corner, maxillary hell reduced. Anterior lobe short, not projected (
<figureCitation id="13092A2FFFFBFFD4FC32FC6779A2E3AE" box="[901,999,986,1012]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1C</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC4EFC677E58E3AE" box="[1017,1053,986,1012]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">4C</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FB9AFC677E16E3AF" box="[1069,1107,986,1013]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5C</figureCitation>
). Maxillary lyra absent. Rastellum absent, distal edge of maxilla with thick setae (
<figureCitation id="13092A2FFFFBFFD4FCC7FC407995E442" box="[880,976,1021,1048]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Figs 1C</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FC68FC407E40E442" box="[991,1029,1021,1048]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">5C</figureCitation>
). Intercheliceral tumescence in males absent. Labiosternal groove shallow, flat, with pair of sigilla (
<figureCitation id="13092A2FFFFBFFD4FC6EFB9C7E71E461" box="[985,1076,1057,1083]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">Fig. 4C</figureCitation>
). Book lungs semi-circular, opening elliptical, and book lung combs absent. Plumose clavate trichobothria in two rows on distal half on all tarsi (
<figureCitation id="13092A2FFFFBFFD4FEC9FBDA7B9BE4DB" box="[382,478,1127,1153]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">Fig. 4D</figureCitation>
). Claws: ITC absent. STC with parallel row of teeth present on lateral edge on males and absent in females (
<figureCitation id="13092A2FFFFBFFD4FDA1FB377830E4FE" box="[534,629,1162,1188]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">Figs 4E</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FD36FB3778E2E4FE" box="[641,679,1162,1188]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[196,1391,862,1842]" captionTargetId="figure-174@9.[189,1398,855,1849]" captionTargetPageId="9" captionText="Fig. 6. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 169417), tarsus I. A. Lateral view. B. Detail of frictional setae, lateral view. C–D. Detail of the chemosensory setae. Abbreviations: Ch = chemosensory setae; ChM = chemosensory setae male; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A = 100 µm; B–C = 50 µm; D = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.7737522" httpUri="https://zenodo.org/record/7737522/files/figure.png" pageId="3" pageNumber="116">6A</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FD04FB37789EE4FE" box="[691,731,1162,1188]" captionStart="Fig" captionStartId="10.[189,232,1795,1821]" captionTargetBox="[195,1391,274,1747]" captionTargetId="figure-20@10.[189,1398,268,1753]" captionTargetPageId="10" captionText="Fig. 7. SEM micrographs of Ammonius pupulus Thorell, 1899, ♀ (RMCA 169416), tarsus IV. A. Lateral view. B. Detail of macrosetae, lateral view.C. Chemosensory setae.D. Apical detail of the chemosensory setae. E. Scale setae. F. Chemosensory setae. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A = 100 µm; B–C = 50 µm; D = 20 µm; E = 50 µm; F = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.7737524" httpUri="https://zenodo.org/record/7737524/files/figure.png" pageId="3" pageNumber="116">7A</figureCitation>
). Frictional setae present in all tarsi, very evident, longer and inserted below the claw tuft plate (
<figureCitation id="13092A2FFFFBFFD4FDC8FB1078BCE492" box="[639,761,1197,1224]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">Figs 4E–F</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FCB2FB107908E49D" box="[773,845,1197,1223]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[196,1391,862,1842]" captionTargetId="figure-174@9.[189,1398,855,1849]" captionTargetPageId="9" captionText="Fig. 6. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 169417), tarsus I. A. Lateral view. B. Detail of frictional setae, lateral view. C–D. Detail of the chemosensory setae. Abbreviations: Ch = chemosensory setae; ChM = chemosensory setae male; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A = 100 µm; B–C = 50 µm; D = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.7737522" httpUri="https://zenodo.org/record/7737522/files/figure.png" pageId="3" pageNumber="116">6A–B</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FCEEFB1379C4E492" box="[857,897,1198,1224]" captionStart="Fig" captionStartId="10.[189,232,1795,1821]" captionTargetBox="[195,1391,274,1747]" captionTargetId="figure-20@10.[189,1398,268,1753]" captionTargetPageId="10" captionText="Fig. 7. SEM micrographs of Ammonius pupulus Thorell, 1899, ♀ (RMCA 169416), tarsus IV. A. Lateral view. B. Detail of macrosetae, lateral view.C. Chemosensory setae.D. Apical detail of the chemosensory setae. E. Scale setae. F. Chemosensory setae. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A = 100 µm; B–C = 50 µm; D = 20 µm; E = 50 µm; F = 30 µm." figureDoi="http://doi.org/10.5281/zenodo.7737524" httpUri="https://zenodo.org/record/7737524/files/figure.png" pageId="3" pageNumber="116">7A</figureCitation>
). Scopulae: on metatarsi I–II scopulate and absent on metatarsi III–IV; on tarsi I fully scopulate and tarsi II–IV slightly scopulate. Four spinnerets, PMS very small, PLS long with apical segment digitiform (
<figureCitation id="13092A2FFFFBFFD4FC2FFB4979B6E554" box="[920,1011,1268,1294]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[196,1390,533,1878]" captionTargetId="figure-70@4.[189,1398,526,1885]" captionTargetPageId="4" captionText="Fig. 1. Ammonius pupulus Thorell, 1899, holotype, ♂ (NHRS-GULI37848). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D–E. Eye group. D. Dorsal view. E. Detail of the eye arrangement. F. Abdomen, lateral view. Scale bars: A–B, F = 1 mm; C–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737511" httpUri="https://zenodo.org/record/7737511/files/figure.png" pageId="3" pageNumber="116">Fig. 1F</figureCitation>
), with several spigots on apical half (
<figureCitation id="13092A2FFFFBFFD4FF4BFAAA7B19E56B" box="[252,348,1303,1329]" captionStart="Fig" captionStartId="7.[189,232,1771,1797]" captionTargetBox="[195,1391,270,1722]" captionTargetId="figure-12@7.[189,1398,263,1728]" captionTargetPageId="7" captionText="Fig. 4. SEM micrographs of Ammonius pupulus Thorell, 1899, ♂ (RMCA 227221). A. Apical segment of PLS. B. Eye group, lateral view. C. Maxillae and labium, ventral view. D. Clavate trichobothria. E. Apical of the tarsus I, lateral view. F. Detail of frictional setae, lateral view. Abbreviations: Ch = chemosensory setae; CT = claw tufts; FS = frictional setae; STC = superior tarsal claws. Scale bars: A, F = 50 µm; B = 300 µm; C = 500 µm; D = 5 µm; E = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7737518" httpUri="https://zenodo.org/record/7737518/files/figure.png" pageId="3" pageNumber="116">Fig. 4A</figureCitation>
). Male palpal tibia with a ventral concavity in 
<taxonomicName id="4C324D29FFFBFFD4FC39FAAA7E48E56B" authorityName="Thorell" authorityYear="1899" box="[910,1037,1303,1329]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFBFFD4FC39FAAA7E48E56B" box="[910,1037,1303,1329]" italics="true" pageId="3" pageNumber="116">A. pupulus</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FBABFAAA7E39E56B" box="[1052,1148,1303,1329]" captionStart="Fig" captionStartId="5.[189,232,1856,1882]" captionTargetBox="[219,1367,692,1816]" captionTargetId="figure-108@5.[213,1374,689,1820]" captionTargetPageId="5" captionText="Fig. 2. Male palp of Ammonius pupulus Thorell, 1899. A–D. Holotype, ♂ (NHRS-GULI37848). A. Dorsal view. B. Ventral view. C. Prolateral view. D. Retrolateral view. E–F. SEM micrographs of the male palp of Ammonius pupulus (RMCA 227221). E. Prolateral view. F. Retrolateral view. G. Drawing from Benoit (1965: fig. 1), ventral view. Abbreviation: Tc = Tibial concavity. Scale bars: A–D = 1 mm; E–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737513" httpUri="https://zenodo.org/record/7737513/files/figure.png" pageId="3" pageNumber="116">Figs 2E</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FB3DFAAA7E91E56B" box="[1162,1236,1303,1329]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[195,1392,640,1843]" captionTargetId="figure-135@6.[189,1398,634,1849]" captionTargetPageId="6" captionText="Fig. 3. SEM micrographs of the male palp of Ammonius pupulus Thorell, 1899 (RMCA 227221). A–B. Tibial concavity. A. Prolateral view. B. Retrolateral view. C–D. Cymbium. C. Prolateral view. D. Retrolateral view. Abbreviations: Em = embolus; PL = prolateral lobe; So = slit organ; Tc = tibial concavity. Scale bars: A = 400 µm; B = 200 µm; C–D = 400 µm." figureDoi="http://doi.org/10.5281/zenodo.7737515" httpUri="https://zenodo.org/record/7737515/files/figure.png" pageId="3" pageNumber="116">3A–B</figureCitation>
), absent in 
<emphasis id="B946EAB8FFFBFFD4FAE8FAAA7B53E50E" italics="true" pageId="3" pageNumber="116">
<taxonomicName id="4C324D29FFFBFFD4FAE8FAAA7B57E50E" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti">A. benoiti</taxonomicName>
.
</emphasis>
Tibial thorns on the retrolateral side of the palp in 
<taxonomicName id="4C324D29FFFBFFD4FCEDFA86798EE50E" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[858,971,1338,1365]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti">
<emphasis id="B946EAB8FFFBFFD4FCEDFA86798EE50E" box="[858,971,1338,1365]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FC6CFA877E1EE50E" box="[987,1115,1338,1364]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">Figs 8D–E</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FBD1FA877ECBE50E" box="[1126,1166,1338,1364]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9D</figureCitation>
). Tibial spur absent. Megaspine on the tibia I in 
<taxonomicName id="4C324D29FFFBFFD4FE4FFAE3782DE52D" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[504,616,1373,1400]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti">
<emphasis id="B946EAB8FFFBFFD4FE4FFAE3782DE52D" box="[504,616,1373,1400]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FDCFFAE0788AE522" box="[632,719,1373,1400]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">Figs 8J</figureCitation>
, 
<figureCitation id="13092A2FFFFBFFD4FD6DFAE378BBE522" box="[730,766,1374,1400]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[193,1388,567,1843]" captionTargetId="figure-126@13.[189,1398,560,1851]" captionTargetPageId="13" captionText="Fig. 9. SEM micrographs of Ammonius benoiti sp. nov. (RMCA 202354). A. Carapace, dorsal view. B. Eye tubercle, dorsal view. C. Tibia I, prolateral view. D. Detail of tibial thorns, retrolateral view. Abbreviations: ALE = anterior lateral eyes; PLE = posterior lateral eyes; PME = posterior median eyes. Scale bars: A = 0.5 mm; B–C = 0.3 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737528" httpUri="https://zenodo.org/record/7737528/files/figure.png" pageId="3" pageNumber="116">9E</figureCitation>
). Male palpal bulb with globose subtegulum, with thin and sinuous embolus in 
<taxonomicName id="4C324D29FFFBFFD4FE6CFA3C781CE5C1" authorityName="Thorell" authorityYear="1899" box="[475,601,1409,1435]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFBFFD4FE6CFA3C781CE5C1" box="[475,601,1409,1435]" italics="true" pageId="3" pageNumber="116">A. pupulus</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FDDEFA3C78A9E5C1" box="[617,748,1409,1435]" captionStart="Fig" captionStartId="5.[189,232,1856,1882]" captionTargetBox="[219,1367,692,1816]" captionTargetId="figure-108@5.[213,1374,689,1820]" captionTargetPageId="5" captionText="Fig. 2. Male palp of Ammonius pupulus Thorell, 1899. A–D. Holotype, ♂ (NHRS-GULI37848). A. Dorsal view. B. Ventral view. C. Prolateral view. D. Retrolateral view. E–F. SEM micrographs of the male palp of Ammonius pupulus (RMCA 227221). E. Prolateral view. F. Retrolateral view. G. Drawing from Benoit (1965: fig. 1), ventral view. Abbreviation: Tc = Tibial concavity. Scale bars: A–D = 1 mm; E–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737513" httpUri="https://zenodo.org/record/7737513/files/figure.png" pageId="3" pageNumber="116">Fig. 2A–G</figureCitation>
), and slightly curved embolus in 
<taxonomicName id="4C324D29FFFBFFD4FBC3FA3C7EA3E5C1" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[1140,1254,1409,1435]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti">
<emphasis id="B946EAB8FFFBFFD4FBC3FA3C7EA3E5C1" box="[1140,1254,1409,1435]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
(
<figureCitation id="13092A2FFFFBFFD4FB40FA3C7F2CE5C1" box="[1271,1385,1409,1435]" captionStart="Fig" captionStartId="12.[191,234,1857,1883]" captionTargetBox="[196,1391,547,1807]" captionTargetId="figure-86@12.[189,1398,540,1814]" captionTargetPageId="12" captionText="Fig. 8. Ammonius benoiti sp. nov. A–E. Holotype, ♂ (RMCA 202482). A. Habitus dorsal. B. Habitus ventral. C–E. Male palp. C. Prolateral view. D. Ventral view. E. Retrolateral view. F–J. Paratype, ♂ (RMCA 202354), male palpal bulb. F. Ventral view. G. Dorsal view. H. Prolateral view. I. Retrolateral view. J. Tibia I, prolateral view. Abbreviations: Ms = Megaspine; Tt = Tibial thorns. Scale bars: A–E, J = 0.5 mm; F–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7737526" httpUri="https://zenodo.org/record/7737526/files/figure.png" pageId="3" pageNumber="116">Fig. 8F–I</figureCitation>
). Spermathecae with two straight receptacles, external receptacle with globose lobe, emerging from the internal receptacle (
<figureCitation id="13092A2FFFFBFFD4FE11FA7A7844E5B8" box="[422,513,1479,1506]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[196,1391,539,1877]" captionTargetId="figure-113@8.[189,1398,532,1884]" captionTargetPageId="8" captionText="Fig. 5. Ammonius pupulus Thorell, 1899, ♀ (RMCA 227221). A. Habitus dorsal. B. Habitus ventral. C. Maxillae and labium, ventral view. D. Eye group, dorsal view. E. Spermathecae. F. Sternum, ventral view. Scale bars: A–B = 1 mm; C–D, F = 0.5 mm; E = 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.7737520" httpUri="https://zenodo.org/record/7737520/files/figure.png" pageId="3" pageNumber="116">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="8B8D36AAFFFBFFD4FF0AF9B47BCCE678" blockId="3.[189,393,1544,1571]" box="[189,393,1544,1571]" pageId="3" pageNumber="116">
<heading id="D0C581C6FFFBFFD4FF0AF9B47BCCE678" bold="true" box="[189,393,1544,1571]" fontSize="11" level="3" pageId="3" pageNumber="116" reason="3">
<emphasis id="B946EAB8FFFBFFD4FF0AF9B47BCCE678" bold="true" box="[189,393,1544,1571]" pageId="3" pageNumber="116">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36AAFFFBFFD4FF0AF98A7962E608" blockId="3.[189,807,1591,1618]" box="[189,807,1591,1618]" pageId="3" pageNumber="116">
<taxonomicName id="4C324D29FFFBFFD4FF0AF98A7802E60B" authority="Thorell, 1899" authorityName="Thorell" authorityYear="1899" box="[189,583,1591,1617]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="pupulus">
<emphasis id="B946EAB8FFFBFFD4FF0AF98A7BD8E60B" box="[189,413,1591,1617]" italics="true" pageId="3" pageNumber="116">Ammonius pupulus</emphasis>
<bibRefCitation id="EFA34B5BFFFBFFD4FE13F98A7802E60B" author="Thorell T." box="[420,583,1591,1617]" pageId="3" pageNumber="116" pagination="1 - 105" refId="ref8554" refString="Thorell T. 1899. Araneae Camerunenses (Africae occidentalis) quas anno 1891 collegerunt Cel. Dr. Y. Sjostedt aliique. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 25 (4, 1): 1 - 105. Available from https: // www. biodiversitylibrary. org / page / 13682607 [accessed 8 Feb. 2023]." type="journal article" year="1899">Thorell, 1899</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C324D29FFFBFFD4FDE5F98A7881E60B" authority="Guadanucci &amp; Brescovit, 2023" authorityName="Guadanucci &amp; Brescovit" authorityYear="2023" box="[594,708,1591,1617]" class="Arachnida" family="Barychelidae" genus="Ammonius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="benoiti" status="sp. nov.">
<emphasis id="B946EAB8FFFBFFD4FDE5F98A7881E60B" box="[594,708,1591,1617]" italics="true" pageId="3" pageNumber="116">A. benoiti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557C3FFFBFFD4FD7BF9857962E608" box="[716,807,1592,1618]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3286521FFFBFFD4FF0AF9C5782AE69B" box="[189,623,1656,1729]" pageId="3" pageNumber="116" type="distribution">
<paragraph id="8B8D36AAFFFBFFD4FF0AF9C57B13E6C8" blockId="3.[189,342,1656,1682]" box="[189,342,1656,1682]" pageId="3" pageNumber="116">
<heading id="D0C581C6FFFBFFD4FF0AF9C57B13E6C8" bold="true" box="[189,342,1656,1682]" fontSize="11" level="3" pageId="3" pageNumber="116" reason="3">
<emphasis id="B946EAB8FFFBFFD4FF0AF9C57B13E6C8" bold="true" box="[189,342,1656,1682]" pageId="3" pageNumber="116">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36AAFFFBFFD4FF0AF91A782AE69B" blockId="3.[189,623,1703,1729]" box="[189,623,1703,1729]" pageId="3" pageNumber="116">
<collectingCountry id="F325763AFFFBFFD4FF0AF91A7B72E69B" box="[189,311,1703,1729]" name="Cameroon" pageId="3" pageNumber="116">Cameroon</collectingCountry>
and 
<collectingCountry id="F325763AFFFBFFD4FEC7F91A7BBFE69B" box="[368,506,1703,1729]" name="Ivory Coast" pageId="3" pageNumber="116">Ivory Coast</collectingCountry>
(
<figureCitation id="13092A2FFFFBFFD4FDBDF91A7827E69B" box="[522,610,1703,1729]" captionStart="Fig" captionStartId="14.[187,230,1998,2024]" captionTargetBox="[189,1398,1383,1950]" captionTargetId="figure-293@14.[189,1398,1383,1950]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of the presently known species of the genus Ammonius Thorell, 1899." figureDoi="http://doi.org/10.5281/zenodo.7737530" httpUri="https://zenodo.org/record/7737530/files/figure.png" pageId="3" pageNumber="116">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>