On the genus Ammonius Thorell, 1899 (Mygalomorphae, Barychelidae): description of the female of A. pupulus, a new species and new distribution records
Author
Gonzalez-Filho, Hector M. O.
8C203170-36D5-4B48-A901-33600BA67BA3
Pós-graduação em Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil.
gonzalezfilho@yahoo.com.br
Author
Guadanucci, José Paulo L.
D4955FF5-FE7F-4E68-AB2F-4A89593F9850
Pós-graduação em Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil. & Departamento de Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil.
jose.guadanucci@unesp.br
Author
Brescovit, Antonio D.
D5B81D79-AFAE-47B1-8A6E-DAB448A24BCC
Pós-graduação em Ecologia, Evolução e Biodiversidade, Instituto de Biociências, Universidade Estadual Paulista, Rio Claro, SP 13506 - 900, Brazil.
antonio.brescovit@butantan.gov.br
text
European Journal of Taxonomy
2023
2023-03-15
861
1
113
131
http://zoobank.org/b55becf4-1716-4439-9bd2-e01b457f092e
journal article
53798
10.5852/ejt.2023.861.2071
e5b1a4ee-623c-40cc-8784-913bf5ad87b8
2118-9773
7737509
B55BECF4-1716-4439-9BD2-E01B457F092E
Genus
Ammonius
Thorell, 1899
Ammonius
Thorell, 1899: 10
.
Ammonius
–
Simon 1903: 915
. —
Benoit 1965: 7
. —
Raven 1985: 113
.
Type
species
Ammonius pupulus
Thorell, 1899
, by monotypy.
Diagnosis
Ammonius
can be distinguished from the other barychelids by the apical segment of the PLS digitiform (
Fig. 1F
); compact eye group rectangular in shape (
Figs 1D
,
4B
,
5D
,
9B
), eye tubercle flat;
A. pupulus
with eight eyes (
Figs 1D–E
,
4B
,
5D
); absence of AME in
A. benoiti
sp. nov.
(
Fig. 9B
); male cymbium unequal, with prolateral lobe inconspicuous and retrolateral lobe strongly projected (
Figs 2
,
3C–D
,
8C–E
). Males resemble
Pisenor notius
Simon, 1889
(see
Benoit 1966
: figs 24, 26) by the strongly pronounced retrolateral lobe of the cymbium but differ by the lack of a spur on tibia I, and the modified palpal tibia, with a tibial concavity in the ventro-prolateral side of the palp on
A. pupulus
(
Fig. 3A–B
), and by the presence of a tibial thorn on the retrolateral side of the palp on
A. benoiti
(
Figs 8D–E
,
9D
). Females of the genus can be distinguished from the other genera by the bilobed spermathecae, with the external stalk small and emerging from the basal region of the receptacle (
Fig. 5E
).
Description
Very small spiders, total length between 3.9–7.3. General coloration yellowish light brown (
Figs 1A–B
,
5A–B
,
8A–B
). Abdomen dorsally pale without pattern and spinnerets pale with some brown light setae on dorsal side (
Figs 1F
,
8A
). Carapace ovate (
Figs 1A
,
5A
,
8A
,
9A
). Thoracic fovea small and straight (
Figs 1A
,
5A
,
8A
). Clypeus narrow (
Figs 1D
,
4B
,
5D
,
9B
). Eye tubercle black and flat (
Figs 1D
,
5D
). Eye group in rectangular shape, anterior and posterior eye rows straight (
Figs 1D
,
4B
,
5D
,
9D
).
Ammonius pupulus
with eight eyes (
Figs 1D–E
,
4B
,
5D
);
A. benoiti
sp. nov.
with six eyes, AME absent (
Fig. 9A– B
). Labium rectangular, 2–6 cuspules rounded. Maxillae rectangular with 3–20 cuspules on inner corner, maxillary hell reduced. Anterior lobe short, not projected (
Figs 1C
,
4C
,
5C
). Maxillary lyra absent. Rastellum absent, distal edge of maxilla with thick setae (
Figs 1C
,
5C
). Intercheliceral tumescence in males absent. Labiosternal groove shallow, flat, with pair of sigilla (
Fig. 4C
). Book lungs semi-circular, opening elliptical, and book lung combs absent. Plumose clavate trichobothria in two rows on distal half on all tarsi (
Fig. 4D
). Claws: ITC absent. STC with parallel row of teeth present on lateral edge on males and absent in females (
Figs 4E
,
6A
,
7A
). Frictional setae present in all tarsi, very evident, longer and inserted below the claw tuft plate (
Figs 4E–F
,
6A–B
,
7A
). Scopulae: on metatarsi I–II scopulate and absent on metatarsi III–IV; on tarsi I fully scopulate and tarsi II–IV slightly scopulate. Four spinnerets, PMS very small, PLS long with apical segment digitiform (
Fig. 1F
), with several spigots on apical half (
Fig. 4A
). Male palpal tibia with a ventral concavity in
A. pupulus
(
Figs 2E
,
3A–B
), absent in
A. benoiti
.
Tibial thorns on the retrolateral side of the palp in
A. benoiti
(
Figs 8D–E
,
9D
). Tibial spur absent. Megaspine on the tibia I in
A. benoiti
(
Figs 8J
,
9E
). Male palpal bulb with globose subtegulum, with thin and sinuous embolus in
A. pupulus
(
Fig. 2A–G
), and slightly curved embolus in
A. benoiti
(
Fig. 8F–I
). Spermathecae with two straight receptacles, external receptacle with globose lobe, emerging from the internal receptacle (
Fig. 5E
).
Included species
Ammonius pupulus
Thorell, 1899
;
A. benoiti
sp. nov.
Distribution
Cameroon
and
Ivory Coast
(
Fig. 10
).